349 lines
32 KiB
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349 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.714.13646" ID-GBIF-Dataset="eaf90d26-0c01-4a36-b2e0-dc53101ac2cf" ID-Pensoft-Pub="1313-2970-714-33" ID-ZBK="4339F0211886401B85B5357761BD24E6" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-714-33" ModsDocOrigin="ZooKeys 714" ModsDocTitle="A new species of the genus Teloganopsis with setaceous mouthparts and forelegs from southern China (Ephemeroptera, Ephemerellidae)" checkinTime="1509980586151" checkinUser="pensoft" docAuthor="Zhang, Wei, Ma, Zhen-Xing, Hu, Ze, Luo, Juan-Yan & Zhou, Chang-Fa" docDate="2017" docId="11DB7B9F72DD22866F1EC19D728DE7C0" docLanguage="en" docName="ZooKeys 714: 33-46" docOrigin="ZooKeys 714" docSource="http://dx.doi.org/10.3897/zookeys.714.13646" docTitle="Teloganopsis setosa Zhou, sp. n." docType="treatment" docUuid="507C1DC5-0D06-41BF-9320-74B75695901D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="43" masterDocId="5225FFDDB679A957FFF7274FFF806248" masterDocTitle="A new species of the genus Teloganopsis with setaceous mouthparts and forelegs from southern China (Ephemeroptera, Ephemerellidae)" masterLastPageNumber="46" masterPageNumber="33" pageNumber="35" updateTime="1643555747617" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of the genus Teloganopsis with setaceous mouthparts and forelegs from southern China (Ephemeroptera, Ephemerellidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Wei</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ma, Zhen-Xing</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hu, Ze</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Luo, Juan-Yan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhou, Chang-Fa</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>714</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.714.13646</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.714.13646</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-714-33</mods:identifier>
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<mods:identifier type="ZBK">4339F0211886401B85B5357761BD24E6</mods:identifier>
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<mods:identifier type="ZooBank">4339F0211886401B85B5357761BD24E6</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137239409" LSID="urn:lsid:zoobank.org:act:507C1DC5-0D06-41BF-9320-74B75695901D" httpUri="http://treatment.plazi.org/id/11DB7B9F72DD22866F1EC19D728DE7C0" lastPageId="10" lastPageNumber="43" pageId="2" pageNumber="35">
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<subSubSection pageId="2" pageNumber="35" type="nomenclature">
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<paragraph pageId="2" pageNumber="35">
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<taxonomicName LSID="http://zoobank.org/507C1DC5-0D06-41BF-9320-74B75695901D" authority="Zhou" class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa Zhou</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="35">sp. n.</taxonomicNameLabel>
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Figs 1, 2, 3, 4, 5, 6, 7
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="35" type="holotype">
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<paragraph pageId="2" pageNumber="35">Holotype.</paragraph>
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<paragraph pageId="2" pageNumber="35">>, Ba-Wang-Ling National Forest Park (19°12.12?N, 109°09.35?E, alt. 300 m), Chang-Jiang county, Hainan Province, China, collected by Qin SI, Jun-Zhi SUN, Juan-Yan LUO, 16 Nov 2015; Paratypes: 2>>1>subimago 2++30L, same as the holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="35" type="diagnosis">
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<paragraph pageId="2" pageNumber="35">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="35">Nymph: Inner margins of forelegs and mouthparts (labrum, mandibles, and labium) have dense and long setae. Paraglossae and palpi of labium are enlarged. Abdominal tergum IV has two dark spots. Male adults: The dorsolateral projections of the penes are visible in ventral view. The caudal filaments have wide reddish-brown bands. Two pigmented spots or dots are on tergum IV.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="40" pageId="2" pageNumber="35" type="description">
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<paragraph pageId="2" pageNumber="35">Description.</paragraph>
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<paragraph pageId="2" pageNumber="35">Nymph (in alcohol, Figs 1 4): Body length 3.0 5.0 mm, caudal filaments 1.2 1.6 mm, cerci subequal to terminal filament (Figs 1, 2). Body reddish to dark brown. Head hypognathous, antennae located near front margin of head, with very tiny setae on articulations of flagellae; antennal length approximately 1.5X head width. Dark base of ocelli and male compound eyes clear.</paragraph>
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<caption pageId="2" pageNumber="35">
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<paragraph pageId="2" pageNumber="35">
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Figure 1. Digital photograph of male nymph of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="35">
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<paragraph pageId="2" pageNumber="35">
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Figure 2. Male nymphal habitus of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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<paragraph pageId="2" pageNumber="35">
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Mouthparts: labrum with relatively long setae on dorsal surface and free margin, ventral surface with shorter but stouter setae (Figs 3A, 4A). Both mandibles with long setae on outer surface; outer and inner incisors of left mandible divided into three teeth apically, but inner incisor of right mandible serrated into two teeth only; prosthecae of mandibles with a tuft of spines on common base (Figs 3B C, 4C D).
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<taxonomicName genus="Maxillae" lsidName="Maxillae" pageId="2" pageNumber="35" rank="genus">Maxillae</taxonomicName>
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: dense long setae and bristles on apex and dorsal surface (Fig. 4F); two rows of bristles on apical half of inner margin; basal half of outer margin and cardo also with shorter setae (Figs 3D, 4E). Labium: paraglossae enlarged, its posterolateral angle projected significantly, this makes paraglossae triangular; ventral surface of glossae with denser and longer setae than dorsal surface; labial palpi elongated remarkably into long broad filamentous process-like structure, 3-segmented, basal and apical one smooth; basal one slightly longer than half of segment II; apical one very short, less than half of basal one; segment II with very long and distinct setae on lateral margins and dorsal surface (Figs 3E, 4G); hypopharynx with denser and longer setae on superlinguae surface, other parts of lingua and superlinguae with shorter setae (Figs 3F, 4B).
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</paragraph>
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<caption pageId="2" pageNumber="35">
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<paragraph pageId="2" pageNumber="35">
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Figure 3. Structures of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n. nymph (digital photographs) A labrum (dorsal view) B right mandible C left mandible D maxilla E labium (ventral view) F hypopharynx (dorsal view) G foreleg.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="35">
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<paragraph pageId="2" pageNumber="35">
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Figure 4. Nymphal characters of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n. A labrum (dorsal view on left; ventral view on right) B hypopharynx (dorsal view on left; ventral view on right) C right mandible D left mandible E maxilla F apex of maxilla G labium (dorsal view on left; ventral view on right) H gill I (ventral view) I gill V (dorsal view) J claw K foreleg L midleg M hindleg N articulation of caudal filaments.
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</paragraph>
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</caption>
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<paragraph lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
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Legs (Figs 3G, 4K M): femora of all legs slightly shorter than tibiae, tarsi slightly longer than half tibiae; basal half of femora, tibiae and tarsi darker or more reddish brown than other parts, an additional reddish to dark ring on median tibiae; foreleg with blunt but relatively long bristles on outer margin of femora, 4 5 more bristles lined up a row sub-apically on femoral surface; inner surface of fore-tibiae and tarsi with rows of setae, those setae of tibiae longer than others; outer margin of them with shorter setae (Figs 1, 2, 3G, 4K); mid- and hind legs similar in structure: outer margin of femora with 3 5 bristles, tibiae with 2 4 bristles on dorsal surface, rows of spines
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<pageBreakToken pageId="3" pageNumber="36" start="start">on</pageBreakToken>
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inner margin while outer margin with tiny setae; setae and spine pattern of tarsi similar to tibiae but without bristle; inner margin of all femora with tiny setae too (Fig. 4L, M); claws of all legs similar, with six denticles from middle to apex, apical one larger than others (Figs 3G, 4J).
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</paragraph>
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<paragraph pageId="3" pageNumber="36">Abdomen: reddish brown to brown, tergum IV with a pair of clear reddish to dark dots, terga V VI distinctly paler than others; terga washed with brown to dark pigments without regular markings (Figs 1, 2). Sterna generally brown to dark brown but with pale median line, especially on posterior half; an additional pair of oblique dark stripes present laterally. Gills on terga III VII, anterior four pairs of gills similar in structure: dorsal lamellae plate-like, with tri-lobed marking dorsally; ventral lamellae bifurcated into two parts, each one with 4 6 leaf-like lobes (Fig. 4H); gills on tergum VII much smaller than anterior ones, ventral lamellae divided into three lobes (Fig. 4I). All posterolateral corners of terga IV IX extended into small but sharp angles, progressively larger posteriorly (Figs 1, 2). Posterior and lateral margins of each tergum with tiny spines. Caudal filaments with reddish to dark median band (Figs 1, 2); articulations with distinct spines (Fig. 4N).</paragraph>
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<paragraph lastPageId="5" lastPageNumber="38" pageId="3" pageNumber="36">
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Male imago (in alcohol, Figs 5 6). Body length 5.0 6.0 mm, caudal filaments 7.0 8.0 mm, forewing 6.5 7.0 mm, hindwing only approx. 1/5 of forewing in length. Upper portion of compound eyes reddish, basal portion dark, two eyes separated with a distance less than width of median ocellus (Fig. 5A).
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<taxonomicName genus="Forewings" lsidName="Forewings" pageId="3" pageNumber="36" rank="genus">Forewings</taxonomicName>
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hyaline, but subcostal brace and bulla of Sc pigmented with reddish brown dots; cross veins in stigma area
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<pageBreakToken pageId="4" pageNumber="37" start="start">oblique</pageBreakToken>
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, those between C and Sc separated into two parts by a short vein (Figs 5B, 6A).
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<taxonomicName genus="Hindwings" lsidName="Hindwings" pageId="4" pageNumber="37" rank="genus">Hindwings</taxonomicName>
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transparent, with a projection at median leading margin (Figs 5C, 6B). Femora subequal to tarsi in length of foreleg, each of them approx. 2/3 of tibiae. Tarsi 5-segmented, basal one the shortest, with hook-like structure near tibiae (Figs 5D, 6C); other segments of fore-tarsi progressively shorter apically. Mid- and hind-leg similar: femora distinctly shorter than tibiae, tarsi less than half of tibiae; tarsi 4-segmented,
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<pageBreakToken pageId="5" pageNumber="38" start="start">length</pageBreakToken>
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arrangement in decreasing order as 4, 1, 2, 3. All claws of legs similar, one blunt and one hooked. Abdominal terga pale to brown, tergum IV with a pair of reddish to dark dots dorsally (Fig. 5A). Sterna pale but with reddishly pigmented lateral margins. Basal half of each segment of caudal filaments reddish, apical half pale (Fig. 5A).
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</paragraph>
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<caption pageId="5" pageNumber="38">
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<paragraph pageId="5" pageNumber="38">
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Figure 5. Male structures of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n. (digital photographs) A male adult B forewing C hindwing D proximal hook of foreleg E genitalia (ventral view).
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="38">
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<paragraph pageId="5" pageNumber="38">
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Figure 6. Male structures of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n. A forewing B hindwing C joint of tarsi and tibiae of foreleg D genitalia (ventral view).
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</paragraph>
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</caption>
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<paragraph pageId="5" pageNumber="38">Genitalia (Figs 5E, 6D): basal segment of forceps very short, less than 1/6 of segment II; the latter straight, narrowed progressively from base to apex, with tiny projections on inner margin; apical segment approx. 1/6 to 1/5 of segment II, mesal margin emarginated at base, making segment III appear somewhat pointed or hooked; penes fused at 2/3 base and separated at apical 1/3; each penis with broad lobe-like projection dorsally; posterior margin of subgenital plate convex.</paragraph>
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<paragraph lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39">
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<pageBreakToken pageId="6" pageNumber="39" start="start">Female</pageBreakToken>
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imago (in alcohol). Body length 6.5 7.0 mm, forewing 7.5 8.0 mm, caudal filaments 6.5 7.0 mm. Body pale to pale reddish, washed with reddish brown dots and markings laterally.
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<taxonomicName genus="Forewings" lsidName="Forewings" pageId="6" pageNumber="39" rank="genus">Forewings</taxonomicName>
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transparent but subcostal brace and bulla of Sc with dark pigments; Tergum IV also with a pair of brown dots but much smaller and indistinct
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<pageBreakToken pageId="7" pageNumber="40" start="start">than</pageBreakToken>
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those of male. Sterna dark brown with pairs of pale dots. Tibiae longer than femora, the latter much longer than tarsi. One of claws blunt, the other hooked and sharp.
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</paragraph>
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<paragraph pageId="7" pageNumber="40">Male Subimago (in alcohol). Body length 5.0 mm, forewing 6.5 7.0 mm, caudal filaments 5.0 mm; resembles male imago except dull; veins and cross veins of wings much clearer than male imago.</paragraph>
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<paragraph pageId="7" pageNumber="40">Egg (Fig. 7). Egg scanned with length 0.133 mm, width 0.084 mm. Egg oval with one polar helmet-shaped cap (Fig. 7A). Egg surface sculptured with hexagonal structures and decorated with sparse tubercle-like projections (Fig. 7B).</paragraph>
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<caption pageId="7" pageNumber="40">
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<paragraph pageId="7" pageNumber="40">
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Figure 7.
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n. Egg morphology (SEM photos) A whole egg B egg surface enlarged (arrow indicates the projection).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="40" type="distribution">
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<paragraph pageId="7" pageNumber="40">Distribution.</paragraph>
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<paragraph pageId="7" pageNumber="40">China (Hainan Province).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="40" type="etymology">
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<paragraph pageId="7" pageNumber="40">Etymology.</paragraph>
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<paragraph pageId="7" pageNumber="40">
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The name
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<taxonomicName lsidName="setosa" pageId="7" pageNumber="40" rank="species" species="setosa">setosa</taxonomicName>
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(from Latin epithet setosus) means having numerous setae. It indicates here that the nymphs of the new species have long and dense setae on the mouthparts and forelegs.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="42" pageId="8" pageNumber="41" type="remarks">
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<paragraph pageId="8" pageNumber="41">
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<pageBreakToken pageId="8" pageNumber="41" start="start">Remarks</pageBreakToken>
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</paragraph>
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<paragraph pageId="8" pageNumber="41">
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Based on the following characters (gills on abdominal segments III VII, ventral lamellae of gills IV (on segment VI) bifurcated, body without any tubercles or projections, maxillae without palpi and shape of claw), the nymph of this new species belongs to the genus
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis" order="Ephemeroptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">Teloganopsis</taxonomicName>
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based on the nymphal keys provided by
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<bibRefCitation author="Jacobus, LM" journalOrPublisher="Annotationes Zoologicae Japonenses" pageId="11" pageNumber="44" url="https://doi.org/10.3157/0002-8320(2008)134[185:ROEGE]2.0.CO;2" year="2008">Jacobus and McCafferty (2008)</bibRefCitation>
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and
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<bibRefCitation author="Ubero-Pascal, N" journalOrPublisher="Aquatic Insects" pageId="11" pageNumber="44" url="https://doi.org/10.1080/01650420902819276" year="2009">Ubero-Pascal and Sartori (2009)</bibRefCitation>
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. The proximal hook of the forelegs, the shape of the penis, and forceps also imply that the male imagoes of this new species belong to the genus
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis" order="Ephemeroptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">Teloganopsis</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="8" pageNumber="41">
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In the nymphal key of all known species of the genus
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis" order="Ephemeroptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">Teloganopsis</taxonomicName>
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prepared by
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<bibRefCitation author="Ubero-Pascal, N" journalOrPublisher="Aquatic Insects" pageId="11" pageNumber="44" url="https://doi.org/10.1080/01650420902819276" year="2009">Ubero-Pascal and Sartori (2009)</bibRefCitation>
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,
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<taxonomicName genus="T." lsidName="T. setosa" pageId="8" pageNumber="41" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. is most similar to the species
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<taxonomicName genus="T." lsidName="T. puigae" pageId="8" pageNumber="41" rank="species" species="puigae">T. puigae</taxonomicName>
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because of their uniform body color pattern, missing maxillary palpi, and without any tubercles on body. However, the setal pattern on the forelegs and the shapes of maxillae and labia are different. The nymphs of
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<taxonomicName genus="T." lsidName="T. setosa" pageId="8" pageNumber="41" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. have longer setae on forelegs and longer labial palpi. In the imaginal stage of these two species, the shape of their hindwings and abdominal colour patterns are dissimilar. The hindwing of
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<taxonomicName genus="T." lsidName="T. setosa" pageId="8" pageNumber="41" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. has a shallow marginal projection while the counterpart in
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<taxonomicName genus="T." lsidName="T. puigae" pageId="8" pageNumber="41" rank="species" species="puigae">T. puigae</taxonomicName>
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has a larger and sharper projection. Furthermore, the imaginal abdomen of the latter species is uniformly dark while that of
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<taxonomicName genus="T." lsidName="T. setosa" pageId="8" pageNumber="41" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. is pale with a pair of reddish brown dots on tergum IV.
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</paragraph>
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<paragraph pageId="8" pageNumber="41">
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Among Chinese ephemerelliids belonging to the genus
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis" order="Ephemeroptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">Teloganopsis</taxonomicName>
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, the nymphs of
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<taxonomicName genus="T." lsidName="T. setosa" pageId="8" pageNumber="41" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. is closest to
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="8" pageNumber="41" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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(Matsumura, 1931) and
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="8" pageNumber="41" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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(
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<bibRefCitation author="Xu, JZ" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="11" pageNumber="44" pagination="413 - 415" title="A new species of Ephemerella (Ephemeroptera: Ephemerellidae)." volume="9" year="1984">Xu et al., 1984</bibRefCitation>
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) as they have no maxillary palpi nor any tubercles on the body. As imagoes, they are also similar because of their longer tibiae (distinctly longer than the femora and tarsi) on the forelegs and the morphology of the genitalia (penis with dorsal projection).
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</paragraph>
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<paragraph pageId="8" pageNumber="41">
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However, the nymph of
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<taxonomicName genus="T." lsidName="T. setosa" pageId="8" pageNumber="41" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. is unique because of its setaceous mouthparts and forelegs. Four diagnostic characters are remarkable: 1) the outer margin of mandible has long setae (Figs 3B C, 4C D); 2) the paraglossae of the labium are expanded and bear dense setae ventrally (Figs 3E, 4G); 3) segment II of the labial palpi are greatly elongated and have setae dorsally (Figs 3E, 4G); 4) forelegs possess long setae (Figs 1, 2, 3G, 4K). The absence of projections, spines, or tubercles on nymphal body also contributes to its identification (Figs 1, 2). Besides these characters, when compared to the similar species
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="8" pageNumber="41" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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and
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="8" pageNumber="41" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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, the nymph of this new species has more and longer setae on its mouthparts (the setae on the labrum, maxillae, hypopharynx, and labium are more numerous and denser). The nymphal femora of the new species are broader, with fewer spines compared to those of
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="8" pageNumber="41" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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and
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="8" pageNumber="41" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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, and their color pattern is different. Tergum IV of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n. is more colorful than that of these two species (Fig. 1), while the latter two species have longitudinal pale stripes on body (Fig. 8A, B), which are not present in the new species.
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</paragraph>
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<caption pageId="8" pageNumber="41">
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<paragraph pageId="8" pageNumber="41">
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Figure 8. Structures of
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis punctisetae" order="Ephemeroptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="punctisetae">Teloganopsis punctisetae</taxonomicName>
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and
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="8" pageNumber="41" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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(digital photographs) A
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="8" pageNumber="41" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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nymph (dorsal view) B
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="8" pageNumber="41" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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nymph (dorsal view) C
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="8" pageNumber="41" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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genitalia (ventral view) D
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="8" pageNumber="41" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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genitalia (ventral view).
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</paragraph>
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</caption>
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<paragraph pageId="9" pageNumber="42">
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<pageBreakToken pageId="9" pageNumber="42" start="start">In</pageBreakToken>
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male imagoes, however, the distinguishing characters of the new species mostly rely on color; 1) the penes and dorsolateral projections of the penes are larger and more distinctive than those of
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="9" pageNumber="42" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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and
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="9" pageNumber="42" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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(Figs 5E, 6D, 8C, 8D); 2) the caudal filaments of
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<taxonomicName genus="T." lsidName="T. setosa" pageId="9" pageNumber="42" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. have broad reddish brown bands on the basal half of each segment (Fig. 5A) while those of the
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="9" pageNumber="42" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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and
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="9" pageNumber="42" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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have rings on the articulations, while broad bands are rare; 3) both the male and female of
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<taxonomicName genus="T." lsidName="T. setosa" pageId="9" pageNumber="42" rank="species" species="setosa">T. setosa</taxonomicName>
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sp. n. have relatively obvious and big spots on tergum IV (Fig. 5A) but in
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<taxonomicName genus="T." lsidName="T. punctisetae" pageId="9" pageNumber="42" rank="species" species="punctisetae">T. punctisetae</taxonomicName>
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and
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<taxonomicName genus="T." lsidName="T. jinghongensis" pageId="9" pageNumber="42" rank="species" species="jinghongensis">T. jinghongensis</taxonomicName>
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there are no recognizable markings on the terga.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="key to the three close chinese teloganopsis species (adult)">
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<paragraph pageId="10" pageNumber="43">
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<pageBreakToken pageId="10" pageNumber="43" start="start">Key</pageBreakToken>
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to the three close Chinese
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="genus">Teloganopsis</taxonomicName>
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species (adult)
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</paragraph>
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<paragraph pageId="10" pageNumber="43">
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<table pageId="10" pageNumber="43">
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">5E8C</td>
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">2</td>
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</tr>
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">8D</td>
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis jinghongensis" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="jinghongensis">Teloganopsis jinghongensis</taxonomicName>
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</td>
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</tr>
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName genus="Tergum" lsidName="Tergum" pageId="10" pageNumber="43" rank="genus">Tergum</taxonomicName>
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5A5A
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</td>
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n.
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</td>
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</tr>
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName genus="Tergum" lsidName="Tergum" pageId="10" pageNumber="43" rank="genus">Tergum</taxonomicName>
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</td>
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis punctisetae" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="punctisetae">Teloganopsis punctisetae</taxonomicName>
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</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="key to the three close chinese teloganopsis species (nymph)">
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<paragraph pageId="10" pageNumber="43">
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Key to the three close Chinese
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="genus">Teloganopsis</taxonomicName>
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species (nymph)
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</paragraph>
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<paragraph pageId="10" pageNumber="43">
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<table pageId="10" pageNumber="43">
|
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">123G4K3E4G</td>
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis setosa" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="setosa">Teloganopsis setosa</taxonomicName>
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sp. n.
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</td>
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</tr>
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">2</td>
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</tr>
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">8A</td>
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis punctisetae" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="punctisetae">Teloganopsis punctisetae</taxonomicName>
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</td>
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</tr>
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<tr pageId="10" pageNumber="43">
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">8B</td>
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<td colspan="1" pageId="10" pageNumber="43" rowspan="1">
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<taxonomicName class="Insecta" family="Ephemerellidae" genus="Teloganopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teloganopsis jinghongensis" order="Ephemeroptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="jinghongensis">Teloganopsis jinghongensis</taxonomicName>
|
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</td>
|
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</tr>
|
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</table>
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |