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<document id="2FC4633C58E9A4AF99AC5633A034387E" ID-CLB-Dataset="295775" ID-DOI="10.3906/zoo-1811-29" ID-GBIF-Dataset="b077f4af-500b-4fd8-85f4-31d00f8b77e5" ID-ISSN="1303-6114" ID-Zenodo-Dep="10973876" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="diego" checkinTime="1713182503328" checkinUser="felipe" docAuthor="Francis, Sanu Vengasseril &amp; Nandanb, Sivasankaran Bijoy" docDate="2019" docId="1142FC51FFFF9940FCC2248FFD25F8DB" docLanguage="en" docName="TurkJZool.43.5.425-436.pdf" docOrigin="Turkish Journal of Zoology 43 (5)" docSource="http://dx.doi.org/10.3906/zoo-1811-29" docStyle="DocumentStyle:8169CCCB0FD041ECF72354F5C265AA0F.2:TurkJZool.2013-.journal_article.1cover" docStyleId="8169CCCB0FD041ECF72354F5C265AA0F" docStyleName="TurkJZool.2013-.journal_article.1cover" docStyleVersion="2" docTitle="Atortus (Atortus) Ohtsuka 1992" docType="key" docVersion="5" lastPageNumber="434" masterDocId="ED7B8429FFF6994AFFE32431FFE3FF9C" masterDocTitle="A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea" masterLastPageNumber="436" masterPageNumber="425" pageNumber="433" updateTime="1727471465737" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A46E8166DA0B6EAB828AC1ED7624DBA1">A new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from Minicoy Island, southeastern Arabian Sea</mods:title>
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<mods:namePart id="F9D05B3A73CD840266D28EA3D22B3D6F">Francis, Sanu Vengasseril</mods:namePart>
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<mods:namePart id="2548A42710840694C22E03BD99FB27CD">Nandanb, Sivasankaran Bijoy</mods:namePart>
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<mods:title id="A89C9636285CB92588DF6C0DA8C503DD">Turkish Journal of Zoology</mods:title>
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<mods:date id="0F12C6348F2B9B9FB5258CED1206DCBC">2019</mods:date>
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<treatment id="1142FC51FFFF9940FCC2248FFD25F8DB" ID-GBIF-Taxon="229726372" LSID="urn:lsid:plazi:treatment:1142FC51FFFF9940FCC2248FFD25F8DB" httpUri="http://treatment.plazi.org/id/1142FC51FFFF9940FCC2248FFD25F8DB" lastPageId="10" lastPageNumber="434" pageId="9" pageNumber="433">
<subSubSection id="D1F11ECCFFFF9943FCC2248FFC79FF6B" pageId="9" pageNumber="433" type="nomenclature">
<paragraph id="99544D47FFFF9943FCC2248FFC79FF6B" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
<heading id="C21CFA2BFFFF9943FCC2248FFC79FF6B" fontSize="10" level="2" pageId="9" pageNumber="433" reason="7">
<emphasis id="AB9F9155FFFF9943FCC2248FFC79FF6B" bold="true" pageId="9" pageNumber="433">
Key to the species of subgenus
<taxonomicName id="5EEB36C4FFFF9943FB7E248FFB0CFF4A" authority="Ohtsuka, 1992" authorityName="Ohtsuka" authorityYear="1992" box="[1181,1263,190,214]" class="Copepoda" family="Tortanidae" genus="Atortus" higherTaxonomySource="CoL" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="AB9F9155FFFF9943FB7E248FFB0CFF4A" bold="true" box="[1181,1263,190,214]" italics="true" pageId="9" pageNumber="433">Atortus</emphasis>
</taxonomicName>
from the Indian Ocean:
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D1F11ECCFFFF9940FCC22531FD25F8DB" lastPageId="10" lastPageNumber="434" pageId="9" pageNumber="433" type="key">
<paragraph id="99544D47FFFF9943FCC22531FA87FEA5" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">Adult females: 1. Genital compound somite asymmetrical ................. 2</paragraph>
<paragraph id="99544D47FFFF9943FCC22573FA82FEE7" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
2. Genital compound somite right anterolateral margin slightly swollen ...........................................
<taxonomicName id="5EEB36C4FFFF9943FB522555FA82FEE7" ID-CoL="57GV2" box="[1201,1377,355,379]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="andamanensis">
<emphasis id="AB9F9155FFFF9943FB522555FA82FEE7" box="[1201,1377,355,379]" italics="true" pageId="9" pageNumber="433">T. andamanensis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC225B5FA80FE21" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
3. Caudal rami slightly asymmetrical with left ramus stouter and shorter than the right one ...............
<taxonomicName id="5EEB36C4FFFF9943FB132597FA80FE21" ID-CoL="57GVC" authorityName="Ohtsuka &amp; Conway" authorityYear="2003" box="[1264,1379,421,445]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="AB9F9155FFFF9943FB132597FA80FE21" box="[1264,1379,421,445]" italics="true" pageId="9" pageNumber="433">T. insularis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC225F7FA80FE63" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">4. Genital compound somite with a bulge on the left or right side ................................................................................. 5</paragraph>
<paragraph id="99544D47FFFF9943FCC22639FA80FDFE" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
- Genital compound somite strongly asymmetrical, with the pointed process on the right ventrolateral side .... ..............................................................................
<taxonomicName id="5EEB36C4FFFF9943FB3F267AFA80FDFE" ID-CoL="57GVG" authorityName="Ohtsuka &amp; Conway" authorityYear="2005" box="[1244,1379,587,610]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="magnonyx">
<emphasis id="AB9F9155FFFF9943FB3F267AFA80FDFE" box="[1244,1379,587,610]" italics="true" pageId="9" pageNumber="433">T. magnonyx</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC2265AFA81FD7A" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
5. Genital compound somite expanded anterolaterally on both sides ending in small papilla with apical spinule, right expansion strongly convex to straight laterally .......... ..............................................................................
<taxonomicName id="5EEB36C4FFFF9943FB3F26FFFA81FD7A" authorityName="Giesbrecht" authorityYear="1889" box="[1244,1378,718,742]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="recticauda">
<emphasis id="AB9F9155FFFF9943FB3F26FFFA81FD7A" box="[1244,1378,718,742]" italics="true" pageId="9" pageNumber="433">T. recticauda</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC226DEFA82FCF5" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
6. Genital compound somite with a small bulge on the left side, caudal rami asymmetrical with left ramus stouter and right ramus slightly curved, fifth leg with spine at distal end ................................................................
<taxonomicName id="5EEB36C4FFFF9943FB172763FA82FCF5" ID-CoL="57GVW" authorityName="Sewell" authorityYear="1932" box="[1268,1377,850,873]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="tropicus">
<emphasis id="AB9F9155FFFF9943FB172763FA82FCF5" box="[1268,1377,850,873]" italics="true" pageId="9" pageNumber="433">T. tropicus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC22742FA83FB93" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
- Genital compound somite with a prominent bulge on the right side with 2 dorsoventral spines, caudal rami asymmetrical with left ramus slightly larger and right ramus not curved, fifth legs without spines at distal end .... ...........................................................
<taxonomicName id="5EEB36C4FFFF9943FB8B27C6FAE5FB92" ID-CoL="7ZGRV" authority="Francis &amp; Nandanb, 2019" authorityName="Francis &amp; Nandanb" authorityYear="2019" box="[1128,1286,1015,1038]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="minicoyensis" status="sp. nov.">
<emphasis id="AB9F9155FFFF9943FB8B27C6FAE5FB92" box="[1128,1286,1015,1038]" italics="true" pageId="9" pageNumber="433">T. minicoyensis</emphasis>
</taxonomicName>
<emphasis id="AB9F9155FFFF9943FAEE27C6FA83FB93" bold="true" box="[1293,1376,1015,1039]" pageId="9" pageNumber="433">
<taxonomicNameLabel id="B0AC2C2EFFFF9943FAEE27C6FA83FB93" box="[1293,1376,1015,1039]" pageId="9" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="99544D47FFFF9943FCC22026FC4BFBB3" blockId="9.[762,1382,190,1896]" box="[801,936,1047,1071]" pageId="9" pageNumber="433">Adult males:</paragraph>
<paragraph id="99544D47FFFF9943FCC22009FA81FB48" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
1. Left leg 5 subdistal seta strongly curved along hemispherical tip with the granular surface, and right coxa of leg 5 truncate with an uncinated corner, basis bearing medial bilobed process near midlength ................................ ..............................................................................
<taxonomicName id="5EEB36C4FFFF9943FB3F208CFA81FB48" authorityName="Giesbrecht" authorityYear="1889" box="[1244,1378,1212,1236]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="recticauda">
<emphasis id="AB9F9155FFFF9943FB3F208CFA81FB48" box="[1244,1378,1212,1236]" italics="true" pageId="9" pageNumber="433">T. recticauda</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC220ECFA80FAC4" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
2. Left leg 5 with subdistal spiniform terminal seta, and right leg coxa produced into the rectangular process, basis bearing rounded, flattened process located at inner distal corner ......................................................................
<taxonomicName id="5EEB36C4FFFF9943FB152170FA80FAC4" ID-CoL="57GVK" authorityName="Ohtsuka, El Sherbiny &amp; Ueda" authorityYear="2000" box="[1270,1379,1344,1368]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="nishidai">
<emphasis id="AB9F9155FFFF9943FB152170FA80FAC4" box="[1270,1379,1344,1368]" italics="true" pageId="9" pageNumber="433">T. nishidai</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC22150FA87FAE5" blockId="9.[762,1382,190,1896]" box="[801,1380,1377,1402]" pageId="9" pageNumber="433">3. Right leg 5 coxa without any process ........................ 4</paragraph>
<paragraph id="99544D47FFFF9943FCC221B2FA81FA06" blockId="9.[762,1382,190,1896]" box="[801,1378,1410,1434]" pageId="9" pageNumber="433">
- Basis bearing small rounded process ...........
<taxonomicName id="5EEB36C4FFFF9943FB1421B2FA81FA06" ID-CoL="57GVW" authorityName="Sewell" authorityYear="1932" box="[1271,1378,1411,1434]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="tropicus">
<emphasis id="AB9F9155FFFF9943FB1421B2FA81FA06" box="[1271,1378,1411,1434]" italics="true" pageId="9" pageNumber="433">T. tropicus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC22192FA80FA61" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
4. Left leg 5 subdistal seta serrated along inner margin, and right leg 5 basis tapering proximally with triangular process ................................................................
<taxonomicName id="5EEB36C4FFFF9943FB3F21D7FA80FA61" ID-CoL="57GVG" authorityName="Ohtsuka &amp; Conway" authorityYear="2005" box="[1244,1379,1510,1533]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="magnonyx">
<emphasis id="AB9F9155FFFF9943FB3F21D7FA80FA61" box="[1244,1379,1510,1533]" italics="true" pageId="9" pageNumber="433">T. magnonyx</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC22237FA81F93E" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
5. Left leg 5 terminal portion of second exopodal segment acutely pointed at tip, with serrated subdistal seta, and right leg 5 coxa bearing 3 acutely pointed prominences of unequal size, basis bearing large bilobed process at midlength ..............................................................
<taxonomicName id="5EEB36C4FFFF9943FB0C22BAFA81F93E" ID-CoL="57GVC" authorityName="Ohtsuka &amp; Conway" authorityYear="2003" box="[1263,1378,1674,1698]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="AB9F9155FFFF9943FB0C22BAFA81F93E" box="[1263,1378,1674,1698]" italics="true" pageId="9" pageNumber="433">T. insularis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFF9943FCC2229AFA82F8F4" blockId="9.[762,1382,190,1896]" pageId="9" pageNumber="433">
6. Left leg 5 with subdistal blunt seta, and right leg 5 coxa semispherical with semispherical medial process, basis semicircular with the medial process being ocarinashaped with depression on distomedial margin and with small rounded process at distal side of its base ................. .......................................................................
<taxonomicName id="5EEB36C4FFFF9943FB522361FA82F8F4" ID-CoL="57GV2" box="[1201,1377,1872,1896]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="433" phylum="Arthropoda" rank="species" species="andamanensis">
<emphasis id="AB9F9155FFFF9943FB522361FA82F8F4" box="[1201,1377,1872,1896]" italics="true" pageId="9" pageNumber="433">T. andamanensis</emphasis>
</taxonomicName>
</paragraph>
<caption id="CD941DCFFFFC9940FFB8248FFF46FF6D" ID-Table-UUID="CD941DCFFFFC9940FFB8248FFF46FF6D" httpUri="http://table.plazi.org/id/CD941DCFFFFC9940FFB8248FFF46FF6D" pageId="10" pageNumber="434" startId="10.[91,145,190,211]" targetBox="[103,1374,290,1540]" targetIsTable="true" targetPageId="10" targetType="table">
<paragraph id="99544D47FFFC9940FFB8248FFF46FF6D" blockId="10.[91,1379,190,241]" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FFB8248FFF48FF4F" bold="true" box="[91,171,190,211]" pageId="10" pageNumber="434">Table 4.</emphasis>
Differential diagnosis of the female
<taxonomicName id="5EEB36C4FFFC9940FDF6248EFD46FF48" authority="Francis &amp; Nandanb, 2019" authorityName="Francis &amp; Nandanb" authorityYear="2019" box="[533,677,191,212]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="minicoyensis" status="sp. nov.">
<emphasis id="AB9F9155FFFC9940FDF6248EFD46FF48" box="[533,677,191,212]" italics="true" pageId="10" pageNumber="434">T. minicoyensis</emphasis>
</taxonomicName>
<emphasis id="AB9F9155FFFC9940FD48248FFD14FF4F" bold="true" box="[683,759,190,211]" pageId="10" pageNumber="434">
<taxonomicNameLabel id="B0AC2C2EFFFC9940FD48248FFD14FF4F" box="[683,759,190,211]" pageId="10" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from all other species of the subgenus
<taxonomicName id="5EEB36C4FFFC9940FB98248FFB21FF4F" authorityName="Ohtsuka" authorityYear="1992" box="[1147,1218,190,211]" pageId="10" pageNumber="434" rank="subGenus" subGenus="Atortus">
<emphasis id="AB9F9155FFFC9940FB98248FFB21FF4F" box="[1147,1218,190,211]" italics="true" pageId="10" pageNumber="434">Atortus</emphasis>
</taxonomicName>
from the Indian Ocean.
</paragraph>
</caption>
<paragraph id="99544D47FFFC9940FF842501FC7EF998" pageId="10" pageNumber="434">
<table id="EBEBBFE7FFFC66B5FF842513FABDF998" box="[103,1374,290,1540]" gridcols="5" gridrows="15" pageId="10" pageNumber="434">
<tr id="27DB4F05FFFC66B5FF842513FABDFEC9" box="[103,1374,290,341]" gridrow="0" pageId="10" pageNumber="434">
<th id="640A2679FFFC66B5FF842513FEFDFEC9" box="[103,286,290,341]" gridcol="0" gridrow="0" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FF842501FF53FEDA" bold="true" box="[103,176,304,326]" pageId="10" pageNumber="434">Species</emphasis>
</th>
<th id="640A2679FFFC66B5FEDA2513FDD8FEC9" box="[313,571,290,341]" gridcol="1" gridrow="0" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FED92501FDC4FEDA" bold="true" box="[314,551,304,326]" pageId="10" pageNumber="434">Fifth pedigerous somite</emphasis>
</th>
<th id="640A2679FFFC66B5FDA52513FCA7FEC9" box="[582,836,290,341]" gridcol="2" gridrow="0" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FDA52501FD09FEDA" bold="true" box="[582,746,304,326]" pageId="10" pageNumber="434">Coupling device</emphasis>
</th>
<th id="640A2679FFFC66B5FCB12513FBB2FEC9" box="[850,1105,290,341]" gridcol="3" gridrow="0" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FCB12513FC76FEC9" bold="true" pageId="10" pageNumber="434">Genital compound somite</emphasis>
</th>
<th id="640A2679FFFC66B5FBBC2513FABDFEC9" box="[1119,1374,290,341]" gridcol="4" gridrow="0" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FBBC2501FB3FFEDA" bold="true" box="[1119,1244,304,326]" pageId="10" pageNumber="434">Caudal rami</emphasis>
</th>
</tr>
<tr id="27DB4F05FFFC66B5FF84255BFABDFDAC" box="[103,1374,362,560]" gridrow="1" pageId="10" pageNumber="434">
<th id="640A2679FFFC66B5FF84255BFEFDFDAC" box="[103,286,362,560]" gridcol="0" gridrow="1" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FF842582FF73FE7A" bold="true" pageId="10" pageNumber="434">
<taxonomicName id="5EEB36C4FFFC9940FF842582FF1EFE54" authority="Francis &amp; Nandanb, 2019" authorityName="Francis &amp; Nandanb" authorityYear="2019" box="[103,253,435,456]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="minicoyensis" status="sp. nov.">
<emphasis id="AB9F9155FFFC9940FF842582FF1EFE54" bold="true" box="[103,253,435,456]" italics="true" pageId="10" pageNumber="434">T. minicoyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0AC2C2EFFFC9940FEE02582FF73FE7A" pageId="10" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="640A2679FFFC66B5FEDA255BFDD8FDAC" box="[313,571,362,560]" gridcol="1" gridrow="1" pageId="10" pageNumber="434">Asymmetrical, left margin with a notch and right margin with a single downward directed triangular lobe (Figures 2A, 2C, 2D)</td>
<td id="640A2679FFFC66B5FDA5255BFBB2FDAC" box="[582,1105,362,560]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="10" pageNumber="434">Left process larger than right covering the Asymmetrical, right dorsolateral process of fifth margin with a bulge pedigerous somite and at midpoint with 2 lateral surface of genital ventrolateral spines compound somite (Figure 2D) (Figures 2C, 2D)</td>
<td id="640A2679FFFC66B5FBBC255BFABDFDAC" box="[1119,1374,362,560]" gridcol="4" gridrow="1" pageId="10" pageNumber="434">Asymmetrical with 2 rounded medial processes, left ramus slightly larger than the right (Figures 2C, 2D)</td>
</tr>
<tr id="27DB4F05FFFC66B5FF842675FABDFC96" box="[103,1374,580,778]" gridrow="2" pageId="10" pageNumber="434">
<th id="640A2679FFFC66B5FF842675FEFDFC96" box="[103,286,580,778]" gridcol="0" gridrow="2" pageId="10" pageNumber="434">
<taxonomicName id="5EEB36C4FFFC9940FF8426ADFEEDFD2E" box="[103,270,668,690]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="andamanensis">
<emphasis id="AB9F9155FFFC9940FF8426ADFEEDFD2E" bold="true" box="[103,270,668,690]" italics="true" pageId="10" pageNumber="434">T. andamanensis</emphasis>
</taxonomicName>
</th>
<td id="640A2679FFFC66B5FEDA2675FDD8FC96" box="[313,571,580,778]" gridcol="1" gridrow="2" pageId="10" pageNumber="434">
The right process produced more posteriorly than the left (
<bibRefCitation id="FD7A30B6FFFC9940FEA32688FE19FD53" author="Nishida S &amp; Anandavelu I &amp; Padmavati G" box="[320,506,697,719]" pageId="10" pageNumber="434" pagination="216 - 230" refId="ref7452" refString="Nishida S, Anandavelu I, Padmavati G (2015). Two new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from the Andaman Islands. Crustaceana 88: 216 - 230." type="journal article" year="2015">Nishida et al., 2015</bibRefCitation>
, fig. 1A)
</td>
<td id="640A2679FFFC66B5FDA52675FCA7FC96" box="[582,836,580,778]" gridcol="2" gridrow="2" pageId="10" pageNumber="434">
Right process larger, covering the dorsolateral process of the fifth pediger and right lateral surface of genital compound somite (
<bibRefCitation id="FD7A30B6FFFC9940FDAF26E6FCE4FD70" author="Nishida S &amp; Anandavelu I &amp; Padmavati G" box="[588,775,726,749]" pageId="10" pageNumber="434" pagination="216 - 230" refId="ref7452" refString="Nishida S, Anandavelu I, Padmavati G (2015). Two new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from the Andaman Islands. Crustaceana 88: 216 - 230." type="journal article" year="2015">Nishida et al., 2015</bibRefCitation>
, figs. 1A, 1B)
</td>
<td id="640A2679FFFC66B5FCB12675FBB2FC96" box="[850,1105,580,778]" gridcol="3" gridrow="2" pageId="10" pageNumber="434">
Right anterolateral margin slightly swollen (
<bibRefCitation id="FD7A30B6FFFC9940FCBA269AFBF0FD5C" author="Nishida S &amp; Anandavelu I &amp; Padmavati G" box="[857,1043,682,705]" pageId="10" pageNumber="434" pagination="216 - 230" refId="ref7452" refString="Nishida S, Anandavelu I, Padmavati G (2015). Two new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from the Andaman Islands. Crustaceana 88: 216 - 230." type="journal article" year="2015">Nishida et al., 2015</bibRefCitation>
, fig. 1D)
</td>
<td id="640A2679FFFC66B5FBBC2675FABDFC96" box="[1119,1374,580,778]" gridcol="4" gridrow="2" pageId="10" pageNumber="434">
Symmetrical with left ramus slightly produced medially while the right ramus concave (
<bibRefCitation id="FD7A30B6FFFC9940FB8626F9FAFCFD42" author="Nishida S &amp; Anandavelu I &amp; Padmavati G" box="[1125,1311,712,734]" pageId="10" pageNumber="434" pagination="216 - 230" refId="ref7452" refString="Nishida S, Anandavelu I, Padmavati G (2015). Two new species of Tortanus (Atortus) (Copepoda, Calanoida, Tortanidae) from the Andaman Islands. Crustaceana 88: 216 - 230." type="journal article" year="2015">Nishida et al., 2015</bibRefCitation>
, figs. 1A, 1D)
</td>
</tr>
<tr id="27DB4F05FFFC66B5FF842711FABDFC54" box="[103,1374,800,968]" gridrow="3" pageId="10" pageNumber="434">
<th id="640A2679FFFC66B5FF842711FEFDFC54" box="[103,286,800,968]" gridcol="0" gridrow="3" pageId="10" pageNumber="434">
<taxonomicName id="5EEB36C4FFFC9940FF842758FF05FCE2" authority="Asymmetrical" authorityName="Asymmetrical" box="[103,230,873,894]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="magnonyx">
<emphasis id="AB9F9155FFFC9940FF842758FF05FCE2" bold="true" box="[103,230,873,894]" italics="true" pageId="10" pageNumber="434">T. magnonyx</emphasis>
</taxonomicName>
</th>
<td id="640A2679FFFC66B5FEDA2711FDD8FC54" box="[313,571,800,968]" gridcol="1" gridrow="3" pageId="10" pageNumber="434">Asymmetrical, with right side more swollen. (Ohtsuka and Conway, 2005, fig. 2A)</td>
<td id="640A2679FFFC66B5FDA52711FCA7FC54" box="[582,836,800,968]" gridcol="2" gridrow="3" pageId="10" pageNumber="434">Not described</td>
<td id="640A2679FFFC66B5FCB12711FBB2FC54" box="[850,1105,800,968]" gridcol="3" gridrow="3" pageId="10" pageNumber="434">Asymmetrical, expanded anterolaterally, with a pointed process on the right ventrolateral side (Ohtsuka and Conway 2005, figs. 2A–2H)</td>
<td id="640A2679FFFC66B5FBBC2711FABDFC54" box="[1119,1374,800,968]" gridcol="4" gridrow="3" pageId="10" pageNumber="434">Asymmetrical, left ramus larger and wider than right, constricted at base (Ohtsuka and Conway, 2005, figs. 2A, 2C, 2D)</td>
</tr>
<tr id="27DB4F05FFFC66B5FF8427D2FABDFBF2" box="[103,1374,995,1134]" gridrow="4" pageId="10" pageNumber="434">
<th id="640A2679FFFC66B5FF8427D2FEFDFBF2" box="[103,286,995,1134]" gridcol="0" gridrow="4" pageId="10" pageNumber="434">
<taxonomicName id="5EEB36C4FFFC9940FF84202FFF36FBAF" authority="Symmetrical" authorityName="Symmetrical" box="[103,213,1053,1075]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="AB9F9155FFFC9940FF84202FFF36FBAF" bold="true" box="[103,213,1053,1075]" italics="true" pageId="10" pageNumber="434">T. insularis</emphasis>
</taxonomicName>
</th>
<td id="640A2679FFFC66B5FEDA27D2FCA7FBF2" box="[313,836,995,1134]" colspan="2" colspanRight="1" gridcol="1" gridrow="4" pageId="10" pageNumber="434">Symmetrical in dorsal view (Ohstuka and Conway, Not described 2003, figs. A, F)</td>
<td id="640A2679FFFC66B5FCB127D2FBB2FBF2" box="[850,1105,995,1134]" gridcol="3" gridrow="4" pageId="10" pageNumber="434">Asymmetrical, expanded anterolaterally on both sides with peg sensillum at tip (Ohstuka and Conway, 2003, figs. 2C–2E)</td>
<td id="640A2679FFFC66B5FBBC27D2FABDFBF2" box="[1119,1374,995,1134]" gridcol="4" gridrow="4" pageId="10" pageNumber="434">Asymmetrical, left ramus larger than right (Ohstuka and Conway, 2003, figs. 2A, 2C, 2D)</td>
</tr>
<tr id="27DB4F05FFFC66B5FF8420B4FABDFB07" box="[103,1374,1157,1179]" gridrow="5" pageId="10" pageNumber="434" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="640A2679FFFC66B5FEDA20B4FDD8FB07" box="[313,571,1157,1179]" gridcol="1" gridrow="5" pageId="10" pageNumber="434">Nearly symmetrical, the</td>
<td id="640A2679FFFC66B5FBBC20B4FABDFB07" box="[1119,1374,1157,1179]" gridcol="4" gridrow="5" pageId="10" pageNumber="434">Asymmetrical with</td>
</tr>
<tr id="27DB4F05FFFC66B5FF842092FABDFB6F" box="[103,1374,1187,1267]" gridrow="6" pageId="10" pageNumber="434">
<th id="640A2679FFFC66B5FF842092FEFDFB6F" box="[103,286,1187,1267]" gridcol="0" gridrow="6" pageId="10" pageNumber="434">
<taxonomicName id="5EEB36C4FFFC9940FF8420F1FF2DFB49" authorityName="Sewell" authorityYear="1932" box="[103,206,1216,1237]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="tropicus">
<emphasis id="AB9F9155FFFC9940FF8420F1FF2DFB49" bold="true" box="[103,206,1216,1237]" italics="true" pageId="10" pageNumber="434">T. tropicus</emphasis>
</taxonomicName>
</th>
<td id="640A2679FFFC66B5FEDA2092FDD8FB6F" box="[313,571,1187,1267]" gridcol="1" gridrow="6" pageId="10" pageNumber="434">right side being produced backwards in a wing-like process</td>
<td id="640A2679FFFC66B5FDA52092FCA7FB6F" box="[582,836,1187,1267]" gridcol="2" gridrow="6" pageId="10" pageNumber="434">
Not described (
<bibRefCitation id="FD7A30B6FFFC9940FDAF20FEFD25FB78" author="Sewell RBS" box="[588,710,1231,1253]" pageId="10" pageNumber="434" pagination="223 - 407" refId="ref7711" refString="Sewell RBS (1932). The Copepoda of Indian seas. Calanoida. Memoirs of the Indian Museum 10: 223 - 407." type="journal article" year="1932">Sewell, 1932</bibRefCitation>
, fig. 131a)
</td>
<td id="640A2679FFFC66B5FCB12092FBB2FB6F" box="[850,1105,1187,1267]" gridcol="3" gridrow="6" pageId="10" pageNumber="434">
Asymmetrical, small bulge on the left side (
<bibRefCitation id="FD7A30B6FFFC9940FCBA20ECFC31FB6E" author="Sewell RBS" box="[857,978,1245,1267]" pageId="10" pageNumber="434" pagination="223 - 407" refId="ref7711" refString="Sewell RBS (1932). The Copepoda of Indian seas. Calanoida. Memoirs of the Indian Museum 10: 223 - 407." type="journal article" year="1932">Sewell, 1932</bibRefCitation>
, fig. 131a)
</td>
<td id="640A2679FFFC66B5FBBC2092FABDFB6F" box="[1119,1374,1187,1267]" gridcol="4" gridrow="6" pageId="10" pageNumber="434">
distinctly stouter left ramus at its base than that of right (
<bibRefCitation id="FD7A30B6FFFC9940FB5120ECFAC8FB6E" author="Sewell RBS" box="[1202,1323,1245,1267]" pageId="10" pageNumber="434" pagination="223 - 407" refId="ref7711" refString="Sewell RBS (1932). The Copepoda of Indian seas. Calanoida. Memoirs of the Indian Museum 10: 223 - 407." type="journal article" year="1932">Sewell, 1932</bibRefCitation>
, fig.
</td>
</tr>
<tr id="27DB4F05FFFC66B5FF8420CAFABDFA8D" box="[103,1374,1275,1297]" gridrow="7" pageId="10" pageNumber="434" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="640A2679FFFC66B5FEDA20CAFDD8FA8D" box="[313,571,1275,1297]" gridcol="1" gridrow="7" pageId="10" pageNumber="434">
(
<bibRefCitation id="FD7A30B6FFFC9940FEA320CAFE5AFA8C" author="Sewell RBS" box="[320,441,1275,1297]" pageId="10" pageNumber="434" pagination="223 - 407" refId="ref7711" refString="Sewell RBS (1932). The Copepoda of Indian seas. Calanoida. Memoirs of the Indian Museum 10: 223 - 407." type="journal article" year="1932">Sewell, 1932</bibRefCitation>
, fig. 131a)
</td>
<td id="640A2679FFFC66B5FBBC20CAFABDFA8D" box="[1119,1374,1275,1297]" gridcol="4" gridrow="7" pageId="10" pageNumber="434">131a)</td>
</tr>
<tr id="27DB4F05FFFC66B5FF842111FABDFAAA" box="[103,1374,1312,1334]" gridrow="8" pageId="10" pageNumber="434" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="640A2679FFFC66B5FCB12111FBB2FAAA" box="[850,1105,1312,1334]" gridcol="3" gridrow="8" pageId="10" pageNumber="434">Asymmetrical, expanded</td>
</tr>
<tr id="27DB4F05FFFC66B5FF84210FFABDFAC8" box="[103,1374,1342,1364]" gridrow="9" pageId="10" pageNumber="434" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="640A2679FFFC66B5FCB1210FFBB2FAC8" box="[850,1105,1342,1364]" gridcol="3" gridrow="9" pageId="10" pageNumber="434">anterolaterally on both</td>
</tr>
<tr id="27DB4F05FFFC66B5FF84216AFABDFAED" box="[103,1374,1371,1393]" gridrow="10" pageId="10" pageNumber="434" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="640A2679FFFC66B5FCB1216AFBB2FAED" box="[850,1105,1371,1393]" gridcol="3" gridrow="10" pageId="10" pageNumber="434">sides ending in small</td>
<td id="640A2679FFFC66B5FBBC216AFABDFAED" box="[1119,1374,1371,1393]" gridcol="4" gridrow="10" pageId="10" pageNumber="434">Asymmetrical, left ramus</td>
</tr>
<tr id="27DB4F05FFFC66B5FF842149FABDFA30" box="[103,1374,1400,1452]" gridrow="11" pageId="10" pageNumber="434">
<th id="640A2679FFFC66B5FF842149FEFDFA30" box="[103,286,1400,1452]" gridcol="0" gridrow="11" pageId="10" pageNumber="434">
<taxonomicName id="5EEB36C4FFFC9940FF8421B6FF04FA01" authority="Symmetrical" authorityName="Symmetrical" box="[103,231,1415,1437]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="recticauda">
<emphasis id="AB9F9155FFFC9940FF8421B6FF04FA01" bold="true" box="[103,231,1415,1437]" italics="true" pageId="10" pageNumber="434">T. recticauda</emphasis>
</taxonomicName>
</th>
<td id="640A2679FFFC66B5FEDA2149FDD8FA30" box="[313,571,1400,1452]" gridcol="1" gridrow="11" pageId="10" pageNumber="434">Symmetrical (Bowman, 1987, fig. 1A)</td>
<td id="640A2679FFFC66B5FDA52149FCA7FA30" box="[582,836,1400,1452]" gridcol="2" gridrow="11" pageId="10" pageNumber="434">Not described</td>
<td id="640A2679FFFC66B5FCB12149FBB2FA30" box="[850,1105,1400,1452]" gridcol="3" gridrow="11" pageId="10" pageNumber="434">papilla with apical spinule; right expansion strongly</td>
<td id="640A2679FFFC66B5FBBC2149FABDFA30" box="[1119,1374,1400,1452]" gridcol="4" gridrow="11" pageId="10" pageNumber="434">slightly wider and longer than right (Bowman,</td>
</tr>
<tr id="27DB4F05FFFC66B5FF842182FABDFA55" box="[103,1374,1459,1481]" gridrow="12" pageId="10" pageNumber="434" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="640A2679FFFC66B5FCB12182FBB2FA55" box="[850,1105,1459,1481]" gridcol="3" gridrow="12" pageId="10" pageNumber="434">convex to straight laterally</td>
<td id="640A2679FFFC66B5FBBC2182FABDFA55" box="[1119,1374,1459,1481]" gridcol="4" gridrow="12" pageId="10" pageNumber="434">1987, figs. 1A, 1C)</td>
</tr>
<tr id="27DB4F05FFFC66B5FF8421E1FABDFA7A" box="[103,1374,1488,1510]" gridrow="13" pageId="10" pageNumber="434" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="640A2679FFFC66B5FCB121E1FBB2FA7A" box="[850,1105,1488,1510]" gridcol="3" gridrow="13" pageId="10" pageNumber="434">(Bowman, 1987, figs.</td>
</tr>
<tr id="27DB4F05FFFC66B5FF8421DFFABDF998" box="[103,1374,1518,1540]" gridrow="14" pageId="10" pageNumber="434" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="640A2679FFFC66B5FCB121DFFBB2F998" box="[850,1105,1518,1540]" gridcol="3" gridrow="14" pageId="10" pageNumber="434">1A–1C)</td>
</tr>
</table>
</paragraph>
<paragraph id="99544D47FFFC9940FF67227EFD26F93B" blockId="10.[94,712,1615,1863]" pageId="10" pageNumber="434">
7. Left leg 5 with subdistal short seta, and right leg 5 coxa semicircular with small bilobed medial process, basis semicircular with fingerlike medial process .....
<taxonomicName id="5EEB36C4FFFC9940FDAB22A1FD26F93B" ID-CoL="57GVS" authorityName="Nishida, Anandavelu &amp; Padmavati" authorityYear="2015" box="[584,709,1679,1703]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="sigmoides">
<emphasis id="AB9F9155FFFC9940FDAB22A1FD26F93B" box="[584,709,1679,1703]" italics="true" pageId="10" pageNumber="434">T. sigmoides</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="99544D47FFFC9940FF67229EFD25F8DB" blockId="10.[94,712,1615,1863]" pageId="10" pageNumber="434">
8. Left leg 5 subdistal seta strongly curved along hemispherical tip with the granular surface, and right leg 5 coxa semitrapezoid with a beak-shaped medial process, basis semicircular and bearing triangular medial process .. ............................................................
<taxonomicName id="5EEB36C4FFFC9940FE322301FD8DF8DB" ID-CoL="7ZGRV" authority="Francis &amp; Nandanb, 2019" authorityName="Francis &amp; Nandanb" authorityYear="2019" box="[465,622,1840,1863]" class="Copepoda" family="Tortanidae" genus="Tortanus" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="434" phylum="Arthropoda" rank="species" species="minicoyensis" status="sp. nov.">
<emphasis id="AB9F9155FFFC9940FE322301FD8DF8DB" box="[465,622,1840,1863]" italics="true" pageId="10" pageNumber="434">T. minicoyensis</emphasis>
</taxonomicName>
<emphasis id="AB9F9155FFFC9940FD97231EFBE3F9FB" bold="true" box="[628,1024,1615,1863]" pageId="10" pageNumber="434">
<taxonomicNameLabel id="B0AC2C2EFFFC9940FD97231EFD25F8DB" box="[628,710,1839,1863]" pageId="10" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
<subSection id="F06454ACFFFC9941FCC2227EFBDCF8FA" lastPageId="11" lastPageNumber="435" pageId="10" pageNumber="434" type="multiple">
<paragraph id="99544D47FFFC9940FCC2227EFC4AF97B" blockId="10.[762,1380,1615,1767]" pageId="10" pageNumber="434">
Nomenclatural acts: This work and the nomenclatural acts it contains have been registered in ZooBank. The ZooBank Life Science Identifier (
<collectionCode id="FFFAD582FFFC9940FBB722BEFB73F93B" box="[1108,1168,1679,1703]" pageId="10" pageNumber="434">LSID</collectionCode>
) for this publication is:
<uri id="ED7A4145FFFC9940FCC2229EFC4AF97B" pageId="10" pageNumber="434">
urn:lsid:zoobank.org:pub:
<uuid id="ED4D7792FFFC9940FBD2229EFC4AF97B" pageId="10" pageNumber="434">3F2596B8-E1F9-4321-9157- BED7BC773638</uuid>
</uri>
</paragraph>
<paragraph id="99544D47FFFC9940FD18233EFC2AF8BB" blockId="10.[762,1380,1807,1895]" box="[763,969,1807,1831]" pageId="10" pageNumber="434">
<emphasis id="AB9F9155FFFC9940FD18233EFC2AF8BB" bold="true" box="[763,969,1807,1831]" pageId="10" pageNumber="434">Acknowledgments</emphasis>
</paragraph>
<paragraph id="99544D47FFFC9941FD18231EFBDCF8FA" blockId="10.[762,1380,1807,1895]" lastBlockId="11.[764,1381,1710,1894]" lastPageId="11" lastPageNumber="435" pageId="10" pageNumber="434">
This study was funded by the Department of Biotechnology of the Government of
<collectingCountry id="E1FC0DD7FFFC9940FC35237EFBEDF8FB" box="[982,1038,1871,1895]" name="India" pageId="10" pageNumber="434">India</collectingCountry>
[BT/PR4258/AAQ/3/575/2011]. The authors are thankful to the Head of the Department of Marine Biology, Microbiology, and Biochemistry, School of Marine Sciences,
<collectingCountry id="E1FC0DD7FFFD9941FEAA22DFFE74F89A" box="[329,407,1774,1798]" name="India" pageId="11" pageNumber="435">Cochin</collectingCountry>
University of Science and Technology for facilities given, and gratefully acknowledge the assistance rendered by the Department of Science and Technology,
<collectingCountry id="E1FC0DD7FFFD9941FF0E237FFE9DF8FA" box="[237,382,1870,1894]" name="India" pageId="11" pageNumber="435">Lakshadweep</collectingCountry>
. Thanks also to the research fellowship provided to SVF under the University Grants Commission Basic Science Research Program of the Government of
<collectingCountry id="E1FC0DD7FFFD9941FC4622DFFC3CF89A" box="[933,991,1774,1798]" name="India" pageId="11" pageNumber="435">India</collectingCountry>
. Special thanks are due to Dr Shuhei Nishida, Atmosphere and Ocean Research Institute, University of Tokyo, for specimen identification and initial preparation of the manuscript.
</paragraph>
</subSection>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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