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<document id="F5DF9445A824323C14CB700FE173C15B" ID-CLB-Dataset="25155" ID-DOI="10.5852/ejt.2019.569" ID-GBIF-Dataset="ee4333ce-4fde-4a61-85e9-effa3d1b5999" ID-Zenodo-Dep="3517522" ID-ZooBank="79BE13FC-B840-4C39-8D25-3328BDCC44D2" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571893727309" checkinUser="plazi" docAuthor="Cole, Mary L." docDate="2019" docId="101687E3D576FFE1FDC0ACAAFDF1D12C" docLanguage="en" docName="ejt-569_cole.pdf" docOrigin="European Journal of Taxonomy 569" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chondrocyclus pulcherrimus Cole 2019, sp. nov." docType="treatment" docUuid="F19B8F07-075C-4A7F-BD19-BB875003E7D0" docUuidSource="ZooBank" docVersion="11" lastPageNumber="42" masterDocId="EC2FFF9BD550FFC8FFF9A93CFFA3D317" masterDocTitle="Revision of Chondrocyclus s. l. (Mollusca: Cyclophoridae), with description of a new genus and twelve new species" masterLastPageNumber="92" masterPageNumber="1" pageNumber="39" updateTime="1698760991196" updateUser="ExternalLinkService">
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<mods:title id="5A84E025E4121DAC31FDD165E87023DF">Revision of Chondrocyclus s. l. (Mollusca: Cyclophoridae), with description of a new genus and twelve new species</mods:title>
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<mods:namePart id="5E64411EA8BC5F3228D5AACD92E09873">Cole, Mary L.</mods:namePart>
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<mods:date id="CAD6B04BED024D279749CACE2CAF886A">2019</mods:date>
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<treatment id="101687E3D576FFE1FDC0ACAAFDF1D12C" ID-DOI="http://doi.org/10.5281/zenodo.5586681" ID-GBIF-Taxon="159834416" ID-Zenodo-Dep="5586681" LSID="urn:lsid:zoobank.org:act:F19B8F07-075C-4A7F-BD19-BB875003E7D0" httpUri="http://treatment.plazi.org/id/101687E3D576FFE1FDC0ACAAFDF1D12C" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39">
<subSubSection id="D0A5657ED576FFEEFDC0ACAAFB66D6C0" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="980036F5D576FFEEFDC0ACAAFC58D6A6" blockId="38.[367,1221,1430,1532]" box="[569,1019,1430,1457]" pageId="38" pageNumber="39">
<heading id="C3488199D576FFEEFDC0ACAAFC58D6A6" box="[569,1019,1430,1457]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<taxonomicName id="5FBF4D76D576FFEEFDC0ACAAFC3AD6A6" ID-CoL="7QNT8" authority="Cole, 2019" authorityName="Cole" authorityYear="2019" box="[569,921,1430,1457]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="pulcherrimus" status="sp. nov.">
<emphasis id="AACBEAE7D576FFEEFDC0ACAAFC3AD6A6" bold="true" box="[569,921,1430,1457]" italics="true" pageId="38" pageNumber="39">Chondrocyclus pulcherrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1F8579CD576FFEEFC58ACABFC58D6A6" box="[929,1019,1431,1457]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<uri id="EC2E3AF7D576FFEEFE96AC80FB66D6C0" box="[367,1221,1468,1495]" pageId="38" pageNumber="39">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="D0A5657ED576FFEEFD59ACDDFC30D6EB" box="[672,915,1505,1532]" pageId="38" pageNumber="39" type="description">
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<figureCitation id="00842A70D576FFEEFD59ACDDFD55D6EB" box="[672,758,1505,1532]" captionStart="Fig" captionStartId="19.[189,232,1789,1815]" captionTargetBox="[278,1299,265,1766]" captionTargetId="figure@19.[278,1299,265,1766]" captionTargetPageId="19" captionText="Fig. 10. Holotypes of species of Chondrocyclus Ancey, 1898. AB. C. herberti sp. nov., diameter 3.70 mm (NMSA P1043/T4277). C, F. C. silvicolus sp. nov., diameter 5.82 mm (NMSA P0639/T4156). DE. C. amathole sp. nov., diameter 6.01 mm (NMSA P0640/T4157). GH. C. pondoensis sp. nov., diameter 5.92 mm (NMSA P0643/T4160). I, L. C. devilliersi sp. nov., diameter 5.23 mm (NMSA P1141/ T4312). JK. C. pulcherrimus sp. nov., diameter 4.34 mm (NMSA W9272/T3075). MN. C. cooperae sp. nov., diameter 4.93 mm (NMSA W9271/T3074). O, R. C. langebergensis sp. nov., diameter 5.13 mm (NMSA P0642/T4159). PQ. C. kevincolei sp. nov., diameter 4.24 mm (NMSA P9270/T3073)." figureDoi="http://doi.org/10.5281/zenodo.3517542" httpUri="https://zenodo.org/record/3517542/files/figure.png" pageId="38" pageNumber="39">Figs 10</figureCitation>
JK, 16A, 19
</paragraph>
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<subSubSection id="D0A5657ED576FFEEFF44AF19FD34D43C" pageId="38" pageNumber="39" type="diagnosis">
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<heading id="C3488199D576FFEEFF44AF19FE95D528" bold="true" box="[189,310,1573,1599]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="AACBEAE7D576FFEEFF44AF19FE95D528" bold="true" box="[189,310,1573,1599]" pageId="38" pageNumber="39">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="980036F5D576FFEEFF44AF6AFD34D43C" blockId="38.[189,1399,1622,1835]" pageId="38" pageNumber="39">Shell very small, very depressed, discoidal; periostracum with very dense axial costae developing at the periphery a row of quadrangularly-shaped flanges and several spiral rows of semicircular flanges above and below this; operculum duplex, exterior saucer-shaped portion with step-shaped multispiral lamella terminating in a long solid fringe reflexed over peristome; radula with three large cusps on second lateral tooth, cusps on rachidian uniform in size; penis flattened dorsoventrally and laterally expanded on left side from about midway down the shaft.</paragraph>
</subSubSection>
<subSubSection id="D0A5657ED576FFEEFF44AE69FE6AD4FC" pageId="38" pageNumber="39" type="etymology">
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<emphasis id="AACBEAE7D576FFEEFF44AE69FEE7D478" bold="true" box="[189,324,1877,1903]" pageId="38" pageNumber="39">Etymology</emphasis>
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The specific name is derived from the Latin
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<emphasis id="AACBEAE7D576FFEEFD23AEBAFCD5D4B7" box="[730,886,1926,1952]" italics="true" pageId="38" pageNumber="39">pulcherrimus</emphasis>
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, meaning most beautiful, with reference to the ornate periostracum with very dense axial lamellae and a larger number of spiral rows of flanges than any other species.
</paragraph>
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<emphasis id="AACBEAE7D577FFEFFF44A837FE4FD232" bold="true" box="[189,492,267,293]" pageId="39" pageNumber="40">Type material examined</emphasis>
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<emphasis id="AACBEAE7D577FFEFFF13A800FEFFD241" bold="true" box="[234,348,316,342]" pageId="39" pageNumber="40">
<typeStatus id="47048857D577FFEFFF13A800FEFFD241" box="[234,348,316,342]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="980036F5D577FFEFFF44A85DFE0DD2D2" blockId="39.[189,1399,316,454]" pageId="39" pageNumber="40">
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, west of
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,
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;
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27.8245°
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,
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31.4191°
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;
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<date id="EC011035D577FFEFFE25A8B9FDB5D288" box="[476,534,389,415]" pageId="39" pageNumber="40" value="1136">1136</date>
m
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a.s.l.
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;
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19
<date id="EC011035D577FFEFFD51A8BAFD74D2B7" box="[680,727,390,416]" month="01" monthName="Jan" pageId="39" pageNumber="40" year="2010">Jan.</date>
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<date id="EC011035D577FFEFFD27A8BAFCBFD2B7" box="[734,796,390,416]" pageId="39" pageNumber="40" value="2010" year="2010">2010</date>
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;
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. and
<collectionCode id="FEAEAE30D577FFEFFC7CA8BAFC3DD2B7" box="[901,926,390,416]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="39" pageNumber="40">K</collectionCode>
. Cole
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leg.; in leaf litter;
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9272
</specimenCode>
/
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3075. (
<figureCitation id="00842A70D577FFEFFED8A897FED7D2D2" box="[289,372,427,453]" captionStart="Fig" captionStartId="19.[189,232,1789,1815]" captionTargetBox="[278,1299,265,1766]" captionTargetId="figure@19.[278,1299,265,1766]" captionTargetPageId="19" captionText="Fig. 10. Holotypes of species of Chondrocyclus Ancey, 1898. AB. C. herberti sp. nov., diameter 3.70 mm (NMSA P1043/T4277). C, F. C. silvicolus sp. nov., diameter 5.82 mm (NMSA P0639/T4156). DE. C. amathole sp. nov., diameter 6.01 mm (NMSA P0640/T4157). GH. C. pondoensis sp. nov., diameter 5.92 mm (NMSA P0643/T4160). I, L. C. devilliersi sp. nov., diameter 5.23 mm (NMSA P1141/ T4312). JK. C. pulcherrimus sp. nov., diameter 4.34 mm (NMSA W9272/T3075). MN. C. cooperae sp. nov., diameter 4.93 mm (NMSA W9271/T3074). O, R. C. langebergensis sp. nov., diameter 5.13 mm (NMSA P0642/T4159). PQ. C. kevincolei sp. nov., diameter 4.24 mm (NMSA P9270/T3073)." figureDoi="http://doi.org/10.5281/zenodo.3517542" httpUri="https://zenodo.org/record/3517542/files/figure.png" pageId="39" pageNumber="40">Fig. 10</figureCitation>
JK)
</materialsCitation>
</paragraph>
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<typeStatus id="47048857D577FFEFFF13A8D2FEC4D11F" box="[234,359,494,520]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
</emphasis>
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<materialsCitation id="28D73CA8D577FFEFFF44AB2FFEC2D145" ID-GBIF-Occurrence="2429182869" collectingDate="2010-01-19" collectionCode="ELM" collectorName="M. and K. Cole" country="South Africa" county="Ngome Forest" elevation="1136" latitude="-27.8245" location="Southern Mistbelt forest" longLatPrecision="6" longitude="31.4191" municipality="Nongoma" pageId="39" pageNumber="40" specimenCount="66" stateProvince="KwaZulu-Natal" typeStatus="paratype">
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AFRICA
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<collectingRegion id="5A7BF817D577FFEFFE44AB2FFD21D13A" box="[445,642,531,557]" country="South Africa" name="KwaZulu-Natal" pageId="39" pageNumber="40">KwaZulu-Natal</collectingRegion>
</emphasis>
<emphasis id="AACBEAE7D577FFEFFD76AB28FD3AD13A" box="[655,665,532,557]" italics="true" pageId="39" pageNumber="40"></emphasis>
<specimenCount id="8EB9FD7CD577FFEFFD5FAB2FFCECD13A" box="[678,847,531,557]" pageId="39" pageNumber="40" type="generic">66 specimens</specimenCount>
; same collection data as for holotype;
<collectionCode id="FEAEAE30D577FFEFFAC1AB28FAD5D139" box="[1336,1398,532,558]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="39" pageNumber="40">ELM</collectionCode>
D16939/
<collectionCode id="FEAEAE30D577FFEFFEDCAB04FE94D145" box="[293,311,568,594]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="40">T</collectionCode>
118
</materialsCitation>
<materialsCitation id="28D73CA8D577FFEFFE87AB05FB1DD145" ID-GBIF-Occurrence="2429182931" box="[382,1214,568,595]" collectingDate="2010-01-19" collectionCode="ELM" collectorName="M. and K. Cole" country="South Africa" county="Ngome Forest" elevation="1136" latitude="-27.8245" location="Southern Mistbelt forest" longLatPrecision="6" longitude="31.4191" municipality="Nongoma" pageId="39" pageNumber="40" specimenCode="W3626" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="8EB9FD7CD577FFEFFE87AB05FDA9D145" box="[382,522,568,595]" pageId="39" pageNumber="40" type="generic">1 specimen</specimenCount>
; same collection data as for holotype;
<collectionCode id="FEAEAE30D577FFEFFC25AB05FBB9D144" box="[988,1050,569,595]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="39" pageNumber="40">ELM</collectionCode>
<specimenCode id="C8199E8ED577FFEFFBDAAB04FBD9D145" box="[1059,1146,568,594]" collectionCode="ELM" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="39" pageNumber="40">
<collectionCode id="FEAEAE30D577FFEFFBDAAB04FB9CD145" box="[1059,1087,568,594]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="39" pageNumber="40" type="Herbarium">W</collectionCode>
3626
</specimenCode>
/
<collectionCode id="FEAEAE30D577FFEFFB7AAB04FB36D145" box="[1155,1173,568,594]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="40">T</collectionCode>
119
</materialsCitation>
<materialsCitation id="28D73CA8D577FFEFFB25AB04FCD7D16F" ID-GBIF-Occurrence="2429182926" collectingDate="2010-01-19" collectionCode="NHMUK" collectorName="M. and K. Cole" country="South Africa" county="Ngome Forest" elevation="1136" latitude="-27.8245" location="Southern Mistbelt forest" longLatPrecision="6" longitude="31.4191" municipality="Nongoma" pageId="39" pageNumber="40" specimenCode="NHMUK 20120281" specimenCount="5" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="8EB9FD7CD577FFEFFB25AB04FAD1D145" box="[1244,1394,568,595]" pageId="39" pageNumber="40" type="generic">5 specimens</specimenCount>
; same collection data as for holotype;
<specimenCode id="C8199E8ED577FFEFFD7FAB62FCD7D16F" box="[646,884,606,632]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="39" pageNumber="40" type="Museum">NHMUK 20120281</specimenCode>
</materialsCitation>
<materialsCitation id="28D73CA8D577FFEFFC6DAB62FD7ED18A" ID-GBIF-Occurrence="2429182815" collectingDate="2010-01-19" collectionCode="NMW" collectorName="M. and K. Cole" country="South Africa" county="Ngome Forest" elevation="1136" latitude="-27.8245" location="Southern Mistbelt forest" longLatPrecision="6" longitude="31.4191" municipality="Nongoma" pageId="39" pageNumber="40" specimenCount="3" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="8EB9FD7CD577FFEFFC6DAB62FB84D160" box="[916,1063,605,632]" pageId="39" pageNumber="40" type="generic">3 specimens</specimenCount>
in ethanol; same collection data as for holotype;
<collectionCode id="FEAEAE30D577FFEFFE4FABBFFDA1D18A" box="[438,514,643,669]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="39" pageNumber="40">NMW</collectionCode>
.
<collectionCode id="FEAEAE30D577FFEFFDFFABBFFDBBD18A" box="[518,536,643,669]" country="Switzerland" httpUri="http://grbio.org/cool/e6hc-1y8v" name="Universit<69>t Z<>rich" pageId="39" pageNumber="40" type="Herbarium">Z</collectionCode>
.2019.004.00009
</materialsCitation>
<materialsCitation id="28D73CA8D577FFEFFD0EABBFFE47D1D5" ID-GBIF-Occurrence="2429182916" collectingDate="2010-01-19" collectionCode="NMW" collectorName="M. and K. Cole" country="South Africa" county="Ngome Forest" elevation="1136" latitude="-27.8245" location="Southern Mistbelt forest" longLatPrecision="6" longitude="31.4191" municipality="Nongoma" pageId="39" pageNumber="40" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
2 dry specimens; same collection data as for holotype;
<collectionCode id="FEAEAE30D577FFEFFF44AB94FEA9D1D5" box="[189,266,680,706]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="39" pageNumber="40">NMW</collectionCode>
.
<collectionCode id="FEAEAE30D577FFEFFEF4AB94FEBCD1D5" box="[269,287,680,706]" country="Switzerland" httpUri="http://grbio.org/cool/e6hc-1y8v" name="Universit<69>t Z<>rich" pageId="39" pageNumber="40" type="Herbarium">Z</collectionCode>
.2019.004.00008
</materialsCitation>
<materialsCitation id="28D73CA8D577FFEFFDF9AB94FAC0D1D4" ID-GBIF-Occurrence="2429182809" box="[512,1379,680,707]" collectingDate="2010-01-19" collectionCode="RMNH" collectorName="M. and K. Cole" country="South Africa" county="Ngome Forest" elevation="1136" latitude="-27.8245" location="Southern Mistbelt forest" longLatPrecision="6" longitude="31.4191" municipality="Nongoma" pageId="39" pageNumber="40" specimenCount="4" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="8EB9FD7CD577FFEFFDF9AB94FD35D1D5" box="[512,662,680,706]" pageId="39" pageNumber="40" type="generic">4 specimens</specimenCount>
; same collection data as for holotype;
<collectionCode id="FEAEAE30D577FFEFFB9BAB94FB19D1D5" box="[1122,1210,680,706]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="39" pageNumber="40" type="Museum">RMNH</collectionCode>
MOL.330504
</materialsCitation>
<materialsCitation id="28D73CA8D577FFEFFF44ABF1FE67D026" ID-GBIF-Occurrence="2429182841" collectingDate="1998-12-02" collectionCode="NMSA" collectorName="Herbert &amp; Seddon &amp; Tattersfield" country="South Africa" elevation="1200" latitude="-27.825" location="Ngome Forest" longLatPrecision="6" longitude="31.4183" pageId="39" pageNumber="40" specimenCode="V7103" specimenCount="11" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="8EB9FD7CD577FFEFFF44ABF1FEC6D1F0" box="[189,357,717,743]" pageId="39" pageNumber="40" type="generic">11 specimens</specimenCount>
;
<location id="9D60602ED577FFEFFE8FABF2FD85D1FF" LSID="urn:lsid:plazi:treatment:101687E3D576FFE1FDC0ACAAFDF1D12C:9D60602ED577FFEFFE8FABF2FD85D1FF" box="[374,550,718,744]" country="South Africa" latitude="-27.825" longLatPrecision="6" longitude="31.4183" name="Ngome Forest" pageId="39" pageNumber="40" stateProvince="KwaZulu-Natal">Ngome Forest</location>
, mist-belt
<taxonomicName id="5FBF4D76D577FFEFFD57ABF1FC9FD1F0" box="[686,828,717,743]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="AACBEAE7D577FFEFFD57ABF1FC9FD1F0" box="[686,828,717,743]" italics="true" pageId="39" pageNumber="40">Podocarpus</emphasis>
</taxonomicName>
forest;
<geoCoordinate id="FD8B5032D577FFEFFC58ABF1FB81D1F0" box="[929,1058,717,744]" degrees="27.8250" direction="south" orientation="latitude" pageId="39" pageNumber="40" precision="5" value="-27.825">
27.8250°
<collectionCode id="FEAEAE30D577FFEFFBE9ABF1FB81D1F0" box="[1040,1058,717,743]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="39" pageNumber="40" type="Herbarium">S</collectionCode>
</geoCoordinate>
,
<geoCoordinate id="FD8B5032D577FFEFFBCCABF1FB1AD1FF" box="[1077,1209,717,744]" degrees="31.4183" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="5" value="31.4183">
31.4183°
<collectionCode id="FEAEAE30D577FFEFFB5DABF2FB1AD1FF" box="[1188,1209,718,744]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="39" pageNumber="40">E</collectionCode>
</geoCoordinate>
;
<elevation id="1392D1C6D577FFEFFB32ABF1FAD3D1F0" box="[1227,1392,717,744]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="39" pageNumber="40" unit="m" value="1200.0">
<quantity id="5F479B10D577FFEFFB32ABF1FA89D1FF" box="[1227,1322,717,744]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="39" pageNumber="40" unit="m" value="1200.0">
<date id="EC011035D577FFEFFB32ABF1FAA5D1F0" box="[1227,1286,717,743]" pageId="39" pageNumber="40" value="1200">1200</date>
m
</quantity>
a.s.l.
</elevation>
;
<date id="EC011035D577FFEFFF44ABCEFEF6D01B" box="[189,341,754,781]" pageId="39" pageNumber="40" value="1998-12-02">
<collectingDate id="FC45E9DDD577FFEFFF44ABCEFEF6D01B" box="[189,341,754,781]" pageId="39" pageNumber="40" value="1998-12-02">
<date id="EC011035D577FFEFFF44ABCEFEAED01A" box="[189,269,754,781]" day="02" month="12" monthName="Dec" pageId="39" pageNumber="40">
2
<date id="EC011035D577FFEFFF2FABCFFEAED01A" box="[214,269,755,781]" month="12" monthName="Dec" pageId="39" pageNumber="40" year="1998">Dec.</date>
</date>
<date id="EC011035D577FFEFFEEEABCEFEF6D01B" box="[279,341,754,780]" pageId="39" pageNumber="40" value="1998" year="1998">1998</date>
</collectingDate>
</date>
;
<collectorName id="354A5323D577FFEFFE9AABCEFE63D01B" box="[355,448,754,780]" pageId="39" pageNumber="40">Herbert</collectorName>
,
<collectorName id="354A5323D577FFEFFE36ABCEFD84D01B" box="[463,551,754,780]" pageId="39" pageNumber="40">Seddon</collectorName>
and
<collectorName id="354A5323D577FFEFFD9CABCEFD4AD01B" box="[613,745,754,780]" pageId="39" pageNumber="40">Tattersfield</collectorName>
leg.; in leaf-litter beneath
<collectionCode id="FEAEAE30D577FFEFFBD4ABCFFB9CD01A" box="[1069,1087,755,781]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="40">T</collectionCode>
<emphasis id="AACBEAE7D577FFEFFBC6ABCEFAD5D01B" box="[1087,1398,754,780]" italics="true" pageId="39" pageNumber="40">richocladus grandilflorus;</emphasis>
<collectionCode id="FEAEAE30D577FFEFFF44AA2BFEB3D026" box="[189,272,791,817]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="39" pageNumber="40" type="Museum">NMSA</collectionCode>
<specimenCode id="C8199E8ED577FFEFFED8AA2BFECCD026" box="[289,367,791,817]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="39" pageNumber="40" type="Museum">
<collectionCode id="FEAEAE30D577FFEFFED8AA2BFE95D026" box="[289,310,791,817]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="39" pageNumber="40" type="Museum">V</collectionCode>
7103
</specimenCode>
/
<collectionCode id="FEAEAE30D577FFEFFE81AA2BFE2AD026" box="[376,393,791,817]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="40">T</collectionCode>
4112
</materialsCitation>
<materialsCitation id="28D73CA8D577FFEFFE08AA2BFEC6D06B" ID-GBIF-Occurrence="2429182741" collectingDate="1998-12-02" collectionCode="NMSA" collectorName="Herbert &amp; Seddon &amp; Tattersfield" country="South Africa" elevation="1100" latitude="-27.825" location="Ngome Forest" longLatPrecision="6" longitude="31.4833" pageId="39" pageNumber="40" specimenCode="V8530" specimenCount="24" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount id="8EB9FD7CD577FFEFFE08AA2BFD3ED026" box="[497,669,791,817]" pageId="39" pageNumber="40" type="generic">24 specimens</specimenCount>
;
<location id="9D60602ED577FFEFFD4AAA24FCC5D025" LSID="urn:lsid:plazi:treatment:101687E3D576FFE1FDC0ACAAFDF1D12C:9D60602ED577FFEFFD4AAA24FCC5D025" box="[691,870,792,818]" country="South Africa" latitude="-27.825" longLatPrecision="6" longitude="31.4833" name="Ngome Forest" pageId="39" pageNumber="40" stateProvince="KwaZulu-Natal">Ngome Forest</location>
, mist-belt
<taxonomicName id="5FBF4D76D577FFEFFC0CAA2BFB20D026" box="[1013,1155,791,817]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="AACBEAE7D577FFEFFC0CAA2BFB20D026" box="[1013,1155,791,817]" italics="true" pageId="39" pageNumber="40">Podocarpus</emphasis>
</taxonomicName>
forest;
<geoCoordinate id="FD8B5032D577FFEFFB09AA2BFAD2D026" box="[1264,1393,791,818]" degrees="27.8250" direction="south" orientation="latitude" pageId="39" pageNumber="40" precision="5" value="-27.825">
27.8250°
<collectionCode id="FEAEAE30D577FFEFFAA6AA2BFAD2D026" box="[1375,1393,791,817]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="39" pageNumber="40" type="Herbarium">S</collectionCode>
</geoCoordinate>
,
<geoCoordinate id="FD8B5032D577FFEFFF44AA01FEE2D040" box="[189,321,829,855]" degrees="31.4833" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="5" value="31.4833">
31.4833°
<collectionCode id="FEAEAE30D577FFEFFED5AA01FEE2D040" box="[300,321,829,855]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="39" pageNumber="40">E</collectionCode>
</geoCoordinate>
;
<elevation id="1392D1C6D577FFEFFEA8AA01FE52D041" box="[337,497,828,855]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="39" pageNumber="40" unit="m" value="1100.0">
<quantity id="5F479B10D577FFEFFEA8AA01FE0ED040" box="[337,429,829,855]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="39" pageNumber="40" unit="m" value="1100.0">
<date id="EC011035D577FFEFFEA8AA01FE28D040" box="[337,395,829,855]" pageId="39" pageNumber="40" value="1100">1100</date>
m
</quantity>
a.s.l.
</elevation>
;
<date id="EC011035D577FFEFFDFAAA01FD3ED040" box="[515,669,829,855]" pageId="39" pageNumber="40" value="1998-12-02">
<collectingDate id="FC45E9DDD577FFEFFDFAAA01FD3ED040" box="[515,669,829,855]" pageId="39" pageNumber="40" value="1998-12-02">
<date id="EC011035D577FFEFFDFAAA01FDF7D040" box="[515,596,829,855]" day="02" month="12" monthName="Dec" pageId="39" pageNumber="40">
2
<date id="EC011035D577FFEFFDE4AA01FDF7D040" box="[541,596,829,855]" month="12" monthName="Dec" pageId="39" pageNumber="40" year="1998">Dec.</date>
</date>
<date id="EC011035D577FFEFFDA6AA01FD3ED040" box="[607,669,829,855]" pageId="39" pageNumber="40" value="1998" year="1998">1998</date>
</collectingDate>
</date>
;
<collectorName id="354A5323D577FFEFFD54AA00FCA9D041" box="[685,778,828,854]" pageId="39" pageNumber="40">Herbert</collectorName>
,
<collectorName id="354A5323D577FFEFFCE3AA00FCD1D041" box="[794,882,828,854]" pageId="39" pageNumber="40">Seddon</collectorName>
and
<collectorName id="354A5323D577FFEFFC4BAA00FB95D041" box="[946,1078,828,854]" pageId="39" pageNumber="40">Tattersfield</collectorName>
leg.; in leaf-litter;
<collectionCode id="FEAEAE30D577FFEFFADAAA01FAD5D040" box="[1315,1398,829,855]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="39" pageNumber="40" type="Museum">NMSA</collectionCode>
<specimenCode id="C8199E8ED577FFEFFF44AA5DFEADD06B" box="[189,270,865,892]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="39" pageNumber="40" type="Museum">
<collectionCode id="FEAEAE30D577FFEFFF44AA5DFF71D06B" box="[189,210,865,892]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="39" pageNumber="40" type="Museum">V</collectionCode>
8530
</specimenCode>
/
<collectionCode id="FEAEAE30D577FFEFFEEEAA5DFE8AD06B" box="[279,297,865,892]" country="0" httpUri="http://grbio.org/cool/ey0i-ifzt" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="40">T</collectionCode>
4113
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D0A5657ED577FFEFFF44AA99FAF4D4FC" pageId="39" pageNumber="40" type="description">
<paragraph id="980036F5D577FFEFFF44AA99FEEDD0A8" blockId="39.[189,334,933,959]" box="[189,334,933,959]" pageId="39" pageNumber="40">
<heading id="C3488199D577FFEFFF44AA99FEEDD0A8" bold="true" box="[189,334,933,959]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="AACBEAE7D577FFEFFF44AA99FEEDD0A8" bold="true" box="[189,334,933,959]" pageId="39" pageNumber="40">Description</emphasis>
</heading>
</paragraph>
<paragraph id="980036F5D577FFEFFF44AAEAFC09D6BA" blockId="39.[189,1399,981,1454]" pageId="39" pageNumber="40">
SHELL (
<figureCitation id="00842A70D577FFEFFEE1AAE9FECCD0E7" box="[280,367,981,1008]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
AC). Small, very depressed, discoidal, adult diameter
<quantity id="5F479B10D577FFEFFBE5AAE9FB6CD0E7" box="[1052,1231,981,1008]" metricMagnitude="-3" metricUnit="m" metricValue="4.175" metricValueMax="4.85" metricValueMin="3.5" pageId="39" pageNumber="40" unit="mm" value="4.175" valueMax="4.85" valueMin="3.5">3.504.85 mm</quantity>
, height
<quantity id="5F479B10D577FFEFFACDAAEAFE86D702" metricMagnitude="-3" metricUnit="m" metricValue="1.97" metricValueMax="2.56" metricValueMin="1.38" pageId="39" pageNumber="40" unit="mm" value="1.97" valueMax="2.56" valueMin="1.38">1.38 2.56 mm</quantity>
, diameter:height 1.872.54(n = 22). Spire almost flat with sometimes only protoconch projecting (
<figureCitation id="00842A70D577FFEFFF3FAD1CFE90D72D" box="[198,307,1056,1082]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19A</figureCitation>
), suture deeply impressed. Embryonic shell (
<figureCitation id="00842A70D577FFEFFCBCAD1CFC13D72D" box="[837,944,1056,1082]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19D</figureCitation>
) just over two whorls, sculptured with pock marks, junction between embryonic shell and teleoconch not particularly distinct, with weak costae at first, becoming stronger. Teleoconch comprising two whorls, rapidly increasing, suture deeply impressed. Aperture circular, last whorl descending near aperture, peristome simple, continuous and free. Umbilicus very wide, exposing all the whorls (
<figureCitation id="00842A70D577FFEFFCCCAD88FC07D7D9" box="[821,932,1204,1230]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19C</figureCitation>
). Periostracum glossy, light yellowishbrown and lacquer-like, with very dense lamellate axial costae at regular intervals, approx. 300 on last whorl (300 and
<quantity id="5F479B10D577FFEFFE8EADC3FE62D60F" box="[375,449,1278,1305]" metricMagnitude="0" metricUnit="m" metricValue="8.636" pageId="39" pageNumber="40" unit="in" value="340.0">340 in</quantity>
<figureCitation id="00842A70D577FFEFFE31ADC2FDDDD60E" box="[456,638,1278,1305]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19B and A</figureCitation>
, respectively), developing at the periphery quarangularly-shaped flanges, their expanded distal ends fused to form a solid fringe (
<figureCitation id="00842A70D577FFEFFC66AC18FBAAD629" box="[927,1033,1316,1342]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19E</figureCitation>
); above periphery are fourfive spiral rows of flanges and sixseven rows below it spiraling into umbilicus, height of flanges forming spiral rows varies; row of flanges below suture forms a channel (see
<figureCitation id="00842A70D577FFEFFBFAAC52FB2ED69F" box="[1027,1165,1390,1416]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19B,D</figureCitation>
); intervals between costae smooth. Shell pale, corneous and translucent when fresh.
</paragraph>
<paragraph id="980036F5D577FFEFFF44ACEBFC17D5E3" blockId="39.[189,1399,1494,1781]" pageId="39" pageNumber="40">
OPERCULUM (
<figureCitation id="00842A70D577FFEFFEACACEAFE0AD6E6" box="[341,425,1494,1521]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
GI). Duplex, outer portion consists of multispiral lamella with about 4.5 whorls, each step-shaped; at the growing edge the top of lamellar blade slopes down at an angle then curves under the step formed by the previous whorl and is thinner and more or less perpendicular where it is attached to the disc; very long fringe of fused bristles emanates from near base of vertical portion of step and curves upwards and then outwards forming a deep furrow between fringe and lamellar blade, spanned by a few bristles; a very short solid horizontal fringe emanates just below main fringe; top edge of lamella projects above level of fringe; fringe of each whorl fused to lamella of following whorl; outermost lamellar fringe very long and reflexed over peristome.
</paragraph>
<paragraph id="980036F5D577FFEFFF44AE22FB6AD495" blockId="39.[189,1399,1822,1923]" pageId="39" pageNumber="40">
RADULA (
<figureCitation id="00842A70D577FFEFFED3AE22FE32D42F" box="[298,401,1822,1848]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19F</figureCitation>
). Rachidian with five cusps of approx. equivalent length; first lateral tooth with three cusps of approx. equivalent length and a very small fourth; second lateral tooth with three cusps of approx. equivalent length and two vestigial outer cusps; teeth do not differ much in size.
</paragraph>
<paragraph id="980036F5D577FFEFFF44AE90FAF4D4FC" blockId="39.[189,1399,1963,2027]" pageId="39" pageNumber="40">
PENIS (
<figureCitation id="00842A70D577FFEFFEE9AE97FEC6D4D1" box="[272,357,1963,1990]" captionStart="Fig" captionStartId="40.[189,232,1894,1920]" captionTargetBox="[189,1398,580,1870]" captionTargetId="figure@40.[189,1398,580,1870]" captionTargetPageId="40" captionText="Fig. 19. Chondrocyclus pulcherrimus sp. nov. AK. ELM W3626/T119. A. Aperture view. B. Dorsal view. C. Ventral view. D Protoconch. E. Detail of axial lamellae of periostracum. F. Portion of radula. GI. Operculum. J. Penis, lateral view. K. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
JK). flattened dorsoventrally and laterally expanded on left side from about midway down the shaft, with numerous annular rugae, smooth distal end occupies about one quarter of shaft.
</paragraph>
</subSubSection>
<subSubSection id="D0A5657ED578FFE0FF44A837FC65D260" pageId="40" pageNumber="41" type="distribution">
<paragraph id="980036F5D578FFE0FF44A837FE4DD232" blockId="40.[189,494,267,293]" box="[189,494,267,293]" pageId="40" pageNumber="41">
<heading id="C3488199D578FFE0FF44A837FE4DD232" bold="true" box="[189,494,267,293]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="AACBEAE7D578FFE0FF44A837FE4DD232" bold="true" box="[189,494,267,293]" pageId="40" pageNumber="41">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="980036F5D578FFE0FF44A806FC65D260" blockId="40.[189,1399,313,375]" pageId="40" pageNumber="41">
Known only from Ngome Forest,
<collectingRegion id="5A7BF817D578FFE0FD98A805FCBED244" box="[609,797,313,339]" country="South Africa" name="KwaZulu-Natal" pageId="40" pageNumber="41">KwaZulu-Natal</collectingRegion>
. Indigenous Eastern Mistbelt Forest (
<bibRefCitation id="FC2E4B04D578FFE0FB11A806FE98D260" author="von Maltitz G. &amp; Mucina L. &amp; Geldenhuys C. &amp; Lawes M. &amp; Eeley H. &amp; Adie H. &amp; Vink D. &amp; Fleming G. &amp; Bailey C." pageId="40" pageNumber="41" refId="ref52802" refString="von Maltitz G., Mucina L., Geldenhuys C., Lawes M., Eeley H., Adie H., Vink D., Fleming G. &amp; Bailey C. 2003. Classification System for South African Indigenous Forests. CSIR, Pretoria." type="book" year="2003">
von Maltitz
<emphasis id="AACBEAE7D578FFE0FF44A861FF5BD260" box="[189,248,349,375]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
2003
</bibRefCitation>
) with large trees, south-facing; in leaf-litter (
<figureCitation id="00842A70D578FFE0FCB2A860FC1BD260" box="[843,952,348,375]" captionStart="Fig" captionStartId="32.[189,232,740,766]" captionTargetBox="[194,1391,261,694]" captionTargetId="figure@32.[189,1398,255,699]" captionTargetPageId="32" captionText="Fig. 16. Distribution map of Chondrocyclus species in the Eastern clade. A. Species in KwaZulu-Natal and the eastern half of the Eastern Cape, on the coast and extending inland. B. Species on the Eastern Cape coast. Note that the range of C. pondoensis sp. nov. overlaps with that of C. putealis Connolly, 1939 and C. cooperae sp. nov. T indicates type localities. Major cities and rivers are labelled." figureDoi="http://doi.org/10.5281/zenodo.3517554" httpUri="https://zenodo.org/record/3517554/files/figure.png" pageId="40" pageNumber="41">Fig. 16A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D0A5657ED578FFE1FF44A8A2FDF1D12C" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="discussion">
<paragraph id="980036F5D578FFE0FF44A8A2FE8CD2AF" blockId="40.[189,303,414,440]" box="[189,303,414,440]" pageId="40" pageNumber="41">
<heading id="C3488199D578FFE0FF44A8A2FE8CD2AF" bold="true" box="[189,303,414,440]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="AACBEAE7D578FFE0FF44A8A2FE8CD2AF" bold="true" box="[189,303,414,440]" pageId="40" pageNumber="41">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="980036F5D578FFE0FF44A8F0FED7D13A" blockId="40.[189,1398,460,557]" pageId="40" pageNumber="41">
The very dense costae of the periostracum of
<taxonomicName id="5FBF4D76D578FFE0FD0EA8F1FC1AD2F0" authority="Cole, 2019" authorityName="Cole" authorityYear="2019" box="[759,953,461,487]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="pulcherrimus" status="sp. nov.">
<emphasis id="AACBEAE7D578FFE0FD0EA8F1FC1AD2F0" box="[759,953,461,487]" italics="true" pageId="40" pageNumber="41">C. pulcherrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1F8579CD578FFE0FC3CA8F1FB87D2F0" box="[965,1060,461,487]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
and large number of spiral rows of flanges, as well as the flat spire, immediately distinguish this species from all other species of
<taxonomicName id="5FBF4D76D578FFE0FF44AB2FFECDD13A" authorityName="Ancey" authorityYear="1898" box="[189,366,531,557]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="40" pageNumber="41" phylum="Mollusca" rank="genus">
<emphasis id="AACBEAE7D578FFE0FF44AB2FFECDD13A" box="[189,366,531,557]" italics="true" pageId="40" pageNumber="41">Chondrocyclus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="CCC0667DD578FFE0FF44AE5AFDA0D4FC" ID-DOI="http://doi.org/10.5281/zenodo.3517560" ID-Zenodo-Dep="3517560" httpUri="https://zenodo.org/record/3517560/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,232,1894,1920]" targetBox="[189,1398,580,1870]" targetPageId="40">
<paragraph id="980036F5D578FFE0FF44AE5AFDA0D4FC" blockId="40.[189,1399,1894,2027]" pageId="40" pageNumber="41">
<emphasis id="AACBEAE7D578FFE0FF44AE5AFEB8D497" bold="true" box="[189,283,1894,1920]" pageId="40" pageNumber="41">Fig. 19.</emphasis>
<taxonomicName id="5FBF4D76D578FFE0FEDDAE5AFDD9D497" authority="Cole, 2019" authorityName="Cole" authorityYear="2019" box="[292,634,1894,1920]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="pulcherrimus" status="sp. nov.">
<emphasis id="AACBEAE7D578FFE0FEDDAE5AFDD9D497" box="[292,634,1894,1920]" italics="true" pageId="40" pageNumber="41">Chondrocyclus pulcherrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1F8579CD578FFE0FD7DAE5BFD43D496" box="[644,736,1895,1921]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="AACBEAE7D578FFE0FD10AE5AFC87D497" bold="true" box="[745,804,1894,1920]" pageId="40" pageNumber="41">AK</emphasis>
. ELM W3626/T119.
<emphasis id="AACBEAE7D578FFE0FBDEAE5AFB9FD497" bold="true" box="[1063,1084,1894,1920]" pageId="40" pageNumber="41">A</emphasis>
. Aperture view.
<emphasis id="AACBEAE7D578FFE0FAFDAE5BFABBD496" bold="true" box="[1284,1304,1895,1921]" pageId="40" pageNumber="41">B</emphasis>
. Dorsal view.
<emphasis id="AACBEAE7D578FFE0FEFAAEB5FEBBD4B4" bold="true" box="[259,280,1929,1955]" pageId="40" pageNumber="41">C</emphasis>
. Ventral view.
<emphasis id="AACBEAE7D578FFE0FE32AEB6FE43D4B3" bold="true" box="[459,480,1930,1956]" pageId="40" pageNumber="41">D</emphasis>
Protoconch.
<emphasis id="AACBEAE7D578FFE0FD84AEB6FD32D4B3" bold="true" box="[637,657,1930,1956]" pageId="40" pageNumber="41">E</emphasis>
. Detail of axial lamellae of periostracum.
<emphasis id="AACBEAE7D578FFE0FB7FAEB6FB3BD4B3" bold="true" box="[1158,1176,1930,1956]" pageId="40" pageNumber="41">F</emphasis>
. Portion of radula.
<emphasis id="AACBEAE7D578FFE0FF44AE91FF4DD4D0" bold="true" box="[189,238,1965,1991]" pageId="40" pageNumber="41">GI</emphasis>
. Operculum.
<emphasis id="AACBEAE7D578FFE0FE77AE91FE3ED4D0" bold="true" box="[398,413,1965,1991]" pageId="40" pageNumber="41">J</emphasis>
. Penis, lateral view.
<emphasis id="AACBEAE7D578FFE0FD77AE91FD06D4D0" bold="true" box="[654,677,1965,1991]" pageId="40" pageNumber="41">K</emphasis>
. Penis, dorsal view. Scale bars: AC = 2 mm; DE = 200 µm; F = 20 µm; GK = 500 µm.
</paragraph>
</caption>
<paragraph id="980036F5D579FFE1FF44A837FD49D2A4" blockId="41.[189,1399,267,435]" pageId="41" pageNumber="42">
<taxonomicName id="5FBF4D76D579FFE1FF44A837FDB0D232" authority="Cole, 2019" authorityName="Cole" authorityYear="2019" box="[189,531,267,293]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="pulcherrimus" status="sp. nov.">
<emphasis id="AACBEAE7D579FFE1FF44A837FDB0D232" box="[189,531,267,293]" italics="true" pageId="41" pageNumber="42">Chondrocyclus pulcherrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1F8579CD579FFE1FDE4A830FDD9D231" box="[541,634,268,294]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="5FBF4D76D579FFE1FCF8A830FC1AD232" authorityName="Connolly" authorityYear="1929" box="[769,953,267,294]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="trifimbriatus">
<emphasis id="AACBEAE7D579FFE1FCF8A830FC1AD232" box="[769,953,267,294]" italics="true" pageId="41" pageNumber="42">C. trifimbriatus</emphasis>
</taxonomicName>
in features of the operculum, radula, penis and the rough texture of the protoconch. The operculum is similar to that of
<taxonomicName id="5FBF4D76D579FFE1FB75A813FAE1D25E" authorityName="Connolly" authorityYear="1929" box="[1164,1346,303,329]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="trifimbriatus">
<emphasis id="AACBEAE7D579FFE1FB75A813FAE1D25E" box="[1164,1346,303,329]" italics="true" pageId="41" pageNumber="42">C. trifimbriatus</emphasis>
</taxonomicName>
, but the fringe emanates from near the bottom of the vertical portion of the stepped lamellar blade forming a deep groove while in
<taxonomicName id="5FBF4D76D579FFE1FE21A84AFD2CD298" authorityName="Connolly" authorityYear="1929" box="[472,655,373,400]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="trifimbriatus">
<emphasis id="AACBEAE7D579FFE1FE21A84AFD2CD298" box="[472,655,373,400]" italics="true" pageId="41" pageNumber="42">C. trifimbriatus</emphasis>
</taxonomicName>
it emanates from near the top of the blade and there is a very shallow groove between the fringe and lamella.
</paragraph>
<paragraph id="980036F5D579FFE1FF44A8E6FDF1D12C" blockId="41.[189,1399,474,571]" pageId="41" pageNumber="42">
The Ngome Forest is 3360 ha and much of the forest is protected in the Ntendeka Wilderness Area.
<taxonomicName id="5FBF4D76D579FFE1FF44A8C2FDB1D10F" authority="Cole, 2019" authorityName="Cole" authorityYear="2019" box="[189,530,510,536]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="pulcherrimus" status="sp. nov.">
<emphasis id="AACBEAE7D579FFE1FF44A8C2FDB1D10F" box="[189,530,510,536]" italics="true" pageId="41" pageNumber="42">Chondrocyclus pulcherrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1F8579CD579FFE1FDE2A8C2FDD5D10F" box="[539,630,510,536]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
is locally very common unlike
<taxonomicName id="5FBF4D76D579FFE1FC16A8C2FB05D10F" authorityName="Connolly" authorityYear="1929" box="[1007,1190,510,536]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="trifimbriatus">
<emphasis id="AACBEAE7D579FFE1FC16A8C2FB05D10F" box="[1007,1190,510,536]" italics="true" pageId="41" pageNumber="42">C. trifimbriatus</emphasis>
</taxonomicName>
which appears to be rare in all populations sampled.
</paragraph>
</subSubSection>
</treatment>
</document>