162 lines
19 KiB
XML
162 lines
19 KiB
XML
<document id="FCA8DA7405DF358F41F538CF4670AACF" ID-CLB-Dataset="34496" ID-DOI="http://dx.doi.org/10.3897/zookeys.527.9575" ID-GBIF-Dataset="05e9fc25-2559-4e72-ae9c-76c9b6c569ed" ID-PMC="PMC4668888" ID-Pensoft-Pub="1313-2970-527-57" ID-PubMed="26692788" ID-ZooBank="05826BC127464BAE97EF5BC06BD63D5C" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-527-57" ModsDocOrigin="ZooKeys 527" ModsDocTitle="Revision of the genus Aseptis McDunnough (Lepidoptera, Noctuidae, Noctuinae, Xylenini) with a description of two new genera, Paraseptis and Viridiseptis" checkinTime="1451243894865" checkinUser="pensoft" docAuthor="Mustelin, Tomas & Crabo, Lars G." docDate="2015" docId="0CEEE5403168800CAD5343258B36D5BB" docLanguage="en" docName="ZooKeys 527: 57-102" docOrigin="ZooKeys 527" docSource="http://dx.doi.org/10.3897/zookeys.527.9575" docTitle="Paraseptis Mustelin & Crabo, 2015, gen. n." docType="treatment" docUuid="E3418CA4-70AB-4F26-8EE6-ED770970B171" docUuidSource="ZooBank" docVersion="7" lastPageNumber="81" masterDocId="AA6DFFF1FFC1FFF1FFB9FFD0FFC7B325" masterDocTitle="Revision of the genus Aseptis McDunnough (Lepidoptera, Noctuidae, Noctuinae, Xylenini) with a description of two new genera, Paraseptis and Viridiseptis" masterLastPageNumber="102" masterPageNumber="57" pageNumber="79" updateTime="1732603197740" updateUser="ExternalLinkService">
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<mods:title id="B0829AF97BD6ECB27BCDB7F758854AE1">Revision of the genus Aseptis McDunnough (Lepidoptera, Noctuidae, Noctuinae, Xylenini) with a description of two new genera, Paraseptis and Viridiseptis</mods:title>
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<mods:namePart id="55525AD91CBE9A8167C0E423F71A5D6C">Mustelin, Tomas</mods:namePart>
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<mods:namePart id="17B6D277491720BEA0C383FF63DD7977">Crabo, Lars G.</mods:namePart>
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<mods:title id="7E6B75D64C7B4BBFA9E0C7C3CF7971CE">ZooKeys</mods:title>
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<mods:date id="85C3DAEF52D871696417F59250041A73">2015</mods:date>
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<mods:detail id="BB378AE794D9A16C078071E52B377EE4" type="volume">
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<mods:number id="562682645DDB2B1672515EF53FE9B7F3">527</mods:number>
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<mods:start id="F644786AEE215B4B5093E087C8BD3006">57</mods:start>
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<mods:end id="94B70423BDF290E34CF0942004142828">102</mods:end>
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<mods:url id="84BA814BB9D366021A322BA670E1752C">http://dx.doi.org/10.3897/zookeys.527.9575</mods:url>
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<mods:classification id="89103D7E3F2FF2112C494C10553961EA">journal article</mods:classification>
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<mods:identifier id="A590EB981751939BF573308ED6B32236" type="DOI">http://dx.doi.org/10.3897/zookeys.527.9575</mods:identifier>
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<mods:identifier id="143FA37E4B3F52D197B4F893A5CE979F" type="Pensoft-Pub">1313-2970-527-57</mods:identifier>
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</mods:mods>
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<treatment id="0CEEE5403168800CAD5343258B36D5BB" ID-GBIF-Taxon="152062765" LSID="urn:lsid:zoobank.org:act:E3418CA4-70AB-4F26-8EE6-ED770970B171" httpUri="http://treatment.plazi.org/id/0CEEE5403168800CAD5343258B36D5BB" lastPageId="24" lastPageNumber="81" pageId="22" pageNumber="79" scope_family="Noctuidae" scope_order="Lepidoptera" scope_subFamily="Noctuinae" scope_tribe="Xylenini">
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<subSubSection id="3EF4C81F49ABFAEE1CA4FAC3AAF8CC71" pageId="22" pageNumber="79" type="multiple">
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<paragraph id="CF110348C29C450B6BEA0F886E2B7764" pageId="22" pageNumber="79">Taxon classification Animalia Lepidoptera Noctuidae</paragraph>
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</subSubSection>
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<subSubSection id="6A0C0FD714D96D2B150208D776B0537A" pageId="22" pageNumber="79" type="nomenclature">
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<paragraph id="0963A816C3A7A1A80AD322234DF6951D" pageId="22" pageNumber="79">
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Genus
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<taxonomicName id="13BCA2546E5D71BAB5FABD8B9C3F73F8" LSID="http://zoobank.org/E3418CA4-70AB-4F26-8EE6-ED770970B171" genus="Paraseptis" lsidName="Paraseptis" pageId="22" pageNumber="79" rank="genus">Paraseptis</taxonomicName>
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<taxonomicNameLabel id="A58877E149524352F5D6769B0DAB19BC" pageId="22" pageNumber="79">gen. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="4E055AF3F49CF2A37B515A16331933F0" pageId="22" pageNumber="79" type="type species">
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<paragraph id="897042003C63794A67795ADD93D02C77" pageId="22" pageNumber="79">Type species.</paragraph>
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<paragraph id="ADAC0FDC85D93069DBD1453E92F2C87D" pageId="22" pageNumber="79">
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<taxonomicName id="D74362C379BBDA8F86ED22D3198A07F8" class="Insecta" family="Noctuidae" genus="Hadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadena adnixa" order="Lepidoptera" pageId="22" pageNumber="79" phylum="Arthropoda" rank="species" species="adnixa">Hadena adnixa</taxonomicName>
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Grote.
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</paragraph>
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</subSubSection>
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<subSubSection id="9428C0E20FF6F900B738C0BE886AE729" pageId="22" pageNumber="79" type="etymology">
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<paragraph id="FF8BE2799FD7BD7DF3609EF9C72AA178" pageId="22" pageNumber="79">Etymology.</paragraph>
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<paragraph id="AC8E631DD458285F94E47C60F5491288" pageId="22" pageNumber="79">
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The name
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<taxonomicName id="7C281094039E6BBD2D03500B29439BB4" genus="Paraseptis" lsidName="Paraseptis" pageId="22" pageNumber="79" rank="genus">Paraseptis</taxonomicName>
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is derived from para meaning next to and septis by analogy to
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<taxonomicName id="7C929559C284E76861D342C920ACCAA0" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="22" pageNumber="79" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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. The name is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="0BE202F875DF71025EAAE82F60B6E905" lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79" type="diagnosis">
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<paragraph id="E3043D9B2DB03E2D80E6BA1A455DB92B" pageId="22" pageNumber="79">Diagnosis.</paragraph>
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<paragraph id="63B4C152A2FD538435BF6F8122B30025" lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79">
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<taxonomicName id="3B26890DC72B9082DEF266AEAC14C2B5" genus="Paraseptis" lsidName="Paraseptis" pageId="22" pageNumber="79" rank="genus">Paraseptis</taxonomicName>
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is a monotypic genus whose sole member occurs near the Pacific Coast of North America. It is mottled brown with typical noctuid wing markings, including a basal dash and dark wedges near the outer margin, and a pale off-white to ochre postreniform patch. The outer edge of the hindwing is concave focally between M1 and M3 as in
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<taxonomicName id="AAE3074F5961E0BB203414065255C02C" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="22" pageNumber="79" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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and
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<taxonomicName id="85DBB2AC6CBFC203C4C26ED803D0320E" genus="Viridiseptis" lsidName="Viridiseptis" pageId="22" pageNumber="79" rank="genus">Viridiseptis</taxonomicName>
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. The male genitalia resemble those of
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<taxonomicName id="001B2F3782C1ECC47D57FEA28E9F707E" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="22" pageNumber="79" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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but differ as follows: the valve has a much larger sacculus that extends above
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<pageBreakToken id="03E05EA63E58F621305B9AC1CEFB17FD" pageId="23" pageNumber="80" start="start">the</pageBreakToken>
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costal margin, a twisted upright ampulla, a rod-like digitus arising near the base of the cucullus from a longitudinal bar near the costa; a long curved aedeagus with distal spine patches of small spines; and a coiled vesica with basal and medial cornuti, but no long apical cornutus (occasional specimens with a minute apical cornutus). The female genitalia are also similar to those of
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<taxonomicName id="0C2E912D9A22C5E663014A33EF428A57" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="23" pageNumber="80" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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but differ in having more narrow papillae anales and a strongly sclerotized posterior ductus bursae. CO1 DNA barcodes of
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<taxonomicName id="78D0E56C6508B44DDF550A47EE403F47" genus="Paraseptis" lsidName="Paraseptis" pageId="23" pageNumber="80" rank="genus">Paraseptis</taxonomicName>
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are not similar to those of
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<taxonomicName id="3641BC0198EC58F8AB57AC0EB3B96808" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="23" pageNumber="80" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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and cluster variably with other genera in the
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<taxonomicName id="60B882DF07D28D9DD7B016D74AC4F9CE" lsidName="" pageId="23" pageNumber="80" rank="tribe" tribe="Xylenini">Xylenini</taxonomicName>
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when representatives from a large number of species are included in the sample set.
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</paragraph>
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</subSubSection>
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<subSubSection id="0191AED03DF2102EF2425CA7DFF2C151" lastPageId="24" lastPageNumber="81" pageId="23" pageNumber="80" type="description">
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<paragraph id="218D3CB877B5D6052EC786A031E0FA54" pageId="23" pageNumber="80">Description.</paragraph>
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<paragraph id="8C8E02D249A1A5D622B6D2D2DC712B04" lastPageId="24" lastPageNumber="81" pageId="23" pageNumber="80">
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Adults: Head: Eye rounded, normal sized. Antenna filiform in both sexes. Labial palpus unmodified with longer second segment and short third segment. Frons slightly convex, smooth. Thorax: Paired moderate-sized dorsal mesothoracic and smaller metathoracic tufts. Legs without tibial spines; tarsal segments with three rows of short spine-like setae. Abdomen: Male with coremata at base of abdomen, complete with lever, pocket, and
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<normalizedToken id="12662EA18F7D5192A64830BC750FE4F2" originalValue="Stobbe’s">Stobbe's</normalizedToken>
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gland; proximal segments with weak dorsal tufts. Forewing: Venation as typical for subfamily, approximately 0.6
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<normalizedToken id="34BC5A684ACE65715A7AEF441185A99A" originalValue="×">x</normalizedToken>
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as wide as long, with brownish, black, and off-white scales, appearing mottled brown with black typical noctuid markings including a basal dash. Hindwing: Venation trifine as typical for subfamily, M2 weak but usually visible, clustered close to M3 and CuA1; outer margin contour concave between veins M1 and M3. Male genitalia (Fig. 74): Tegumen narrow near base of uncus; penicillus large, quadrate. Uncus smoothly downcurved, narrow, tapering smoothly from base to acute tip. Juxta rectangular, circa 2/3
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<normalizedToken id="4AABEF4449472D4ADFFFEC53111D32F0" originalValue="×">x</normalizedToken>
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as wide as long. Valve weakly S-shaped, tapered from base to mid-portion then even in width to base of cucullus; sacculus strong, moderately sclerotized, 0.4
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<normalizedToken id="62F5EF6CAFF1A91C900D3E12B3FB98C9" originalValue="×">x</normalizedToken>
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valve length and 2
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<normalizedToken id="380B5242597D36224B7AC02894966ABE" originalValue="×">x</normalizedToken>
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valve width at base of ampulla, extending above costa; cucullus weakly constricted at base and expanded to 1.5
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<normalizedToken id="4205164CED75D5889575FDF9593EC9CE" originalValue="–1.7×">-1.7x</normalizedToken>
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valve width, slightly rounded with blunt apex, corona of circa 30 claw-like setae; clasper on mesial third, ampulla oriented perpendicular to valve and extending above costa, rod-like with slightly twist to mediolaterally flattened tip; digitus at distal end of an evenly-thick sclerotized ridge located slightly below costa from clasper to digitus origin near cucullus base, rod-like with blunt tip, oriented 45° to valve, ending near ventral cucullus. Aedeagus narrower and more robustly sclerotized than in
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<taxonomicName id="F63C8E3EB4C3D515ABFADF87231CD2F1" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="23" pageNumber="80" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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, 7
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<normalizedToken id="774E03025D5BCC35FF6973431617698B" originalValue="×">x</normalizedToken>
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as long as wide, distal half bent ventrad approximately 60°, small patches of small spines near dorsal and ventral apex; vesica slightly wider than aedeagus, coiled 360° to right and ventrad to end posterior, ventrad, and left of aedeagus tip, with subbasal patch of short cornuti on right, single spike-like diverticulum perpendicular to axis on anterior distal third, minute spike-like cornutus directed basad at apex in a few specimens, and very small dome-like mesial diverticulum. Female genitalia (Fig. 90): Papillae anales weakly sclerotized, asymmetrically cone-shaped with point near dorsum, circa 1.7
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<normalizedToken id="D104B76C2CBD08A99B1A6003648B85EB" originalValue="×">x</normalizedToken>
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as long as wide, covered posteriorly and apically with short thin setae that are slightly shorter near tip, lacking hair-like basal setae; apophyses moderately long, posterior apophysis 1.7
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<normalizedToken id="E80EE29A2943FBEE67EFACB775291A9A" originalValue="×">x</normalizedToken>
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anterior apophysis; ostium bursae membranous except for thin weak band in ventral wall; ductus bursae tubular, 1
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<normalizedToken id="3FE2C2E877A7D2A0F0799BF1A0FFE8F4" originalValue="×">x</normalizedToken>
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corpus bursae length, proximal 2/3 sclerotized with longitudinal ridge in dorsum,
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<pageBreakToken id="BA0EF9EF81C5B8D120C78B7FF7E75D7E" pageId="24" pageNumber="81" start="start">distal</pageBreakToken>
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1/3 membranous; corpus bursae ovoid, 0.75
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<normalizedToken id="140449231AAB3B038D26463D3F661981" originalValue="×">x</normalizedToken>
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as wide as long, with four long signa evenly spaced on anterior, posterior, and lateral sides; appendix bursae arising from right paramedial ventral posterior corpus bursae, moderately sclerotized, rugose, 0.75
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<normalizedToken id="BD347C6F58F92535012D1D58C53F8610" originalValue="×">x</normalizedToken>
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corpus bursae length, conical with 45-60° rightward bend to end ventral to, or slightly to right and ventral to, distal ductus bursae, with ductus seminalis near apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="AED6D50E8013B7FFEAF95CBD0D3E69DE" pageId="24" pageNumber="81" type="discussion">
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<paragraph id="3143F13ED44B2143EE7130963928FB96" pageId="24" pageNumber="81">Discussion.</paragraph>
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<paragraph id="9F58C863AF4F63B089CCEF214D8646AB" pageId="24" pageNumber="81">
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The structural differences of
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<taxonomicName id="C36D66EA28151F76623E13A6FFDA66BE" genus="Paraseptis" lsidName="Paraseptis" pageId="24" pageNumber="81" rank="genus">Paraseptis</taxonomicName>
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and
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<taxonomicName id="933174D7097E1EEF3C8ADF6FA2B68F1B" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="24" pageNumber="81" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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are surprising given the nearly identical habitus of
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<taxonomicName id="4317F6EB1CE5142B84CB00AF80D95F31" genus="Paraseptis" lsidName="Paraseptis adnixa" pageId="24" pageNumber="81" rank="species" species="adnixa">Paraseptis adnixa</taxonomicName>
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and
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<taxonomicName id="C4AFE0A75BBF9BC86E66C18F01925080" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis binotata" order="Lepidoptera" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="binotata">Aseptis binotata</taxonomicName>
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, which are often mixed in collections.
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</paragraph>
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<paragraph id="7BB6AAFF9EC4A5B1375BDCB9802BAC89" pageId="24" pageNumber="81">
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Several similar features of
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<taxonomicName id="771FCCEAA9CBC8898B876757EAA64DBA" genus="Paraseptis" lsidName="Paraseptis" pageId="24" pageNumber="81" rank="genus">Paraseptis</taxonomicName>
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and
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<taxonomicName id="51D40ACED77CF66D20A2DF4C4B78B516" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis" order="Lepidoptera" pageId="24" pageNumber="81" phylum="Arthropoda" rank="genus">Aseptis</taxonomicName>
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, including the hindwing shape and superficial resemblance, suggest that these genera are related closely. The hindwing shape is rare in other genera in the
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<taxonomicName id="AD386DF1E07B2CA3DBCD65F6B7DE7440" lsidName="" pageId="24" pageNumber="81" rank="tribe" tribe="Xylenini">Xylenini</taxonomicName>
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. It is a prominent feature of the Eurasian monotypic genus
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<taxonomicName id="A298FD0F11CECB12C7F753567355923B" class="Insecta" family="Noctuidae" genus="Atypha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atypha" order="Lepidoptera" pageId="24" pageNumber="81" phylum="Arthropoda" rank="genus">Atypha</taxonomicName>
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<normalizedToken id="F0FBBC1BF9CA88539EF845B61426F168" originalValue="Hübner">Huebner</normalizedToken>
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. The male genitalia of
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<taxonomicName id="D17003A6107AC68A9BD6B990EE1D87B5" class="Insecta" family="Noctuidae" genus="Atypha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atypha pulmonaris" order="Lepidoptera" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="pulmonaris">Atypha pulmonaris</taxonomicName>
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(Esper), illustrated by
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<bibRefCitation id="119BC3F50CBCDC2D6E93C254D6152D2E" author="Fibiger, M" journalOrPublisher="Soro, Denmark" pageId="31" pageNumber="88" title="Noctuidae Europaeae. Volume 9. Amphipyrinae-Xyleninae. Entomological Press." year="2007">Fibiger and Hacker (2007)</bibRefCitation>
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, are similar to those of
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<taxonomicName id="1904C22BB26F0B452CAFA86BC6678EB1" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis fumosa" order="Lepidoptera" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="fumosa">Aseptis fumosa</taxonomicName>
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and
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<taxonomicName id="4D55228483573EF9544ECEEEF588DD81" class="Insecta" family="Noctuidae" genus="Aseptis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aseptis characta" order="Lepidoptera" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="characta">Aseptis characta</taxonomicName>
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in having a simple valve, horizontal ampulla, and no digitus, suggesting that these three genera might share a common ancestor.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |