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<document id="729F20E83780E6931E514BC96A6133E5" ID-CLB-Dataset="26107" ID-DOI="10.11646/zootaxa.4651.1.8" ID-GBIF-Dataset="0e1cc894-7613-41e5-bc48-b5224cce0670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3359237" ID-ZooBank="EB7EE06F-0CA3-4AA5-A152-31645179F419" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564781000653" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2019" docId="087A87B4FFD8FFFBBD88F90E3C13FEE1" docLanguage="en" docName="zootaxa.4651.1.8.pdf" docOrigin="Zootaxa 4651 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Salmoneus excavatus Anker 2011" docType="treatment" docVersion="13" lastPageNumber="137" masterDocId="F443FFCCFFD1FFF7BD1FFFDC3E32FFA2" masterDocTitle="The eastern Pacific species of Salmoneus Holthuis, 1955, with description of a remarkable new species from Las Perlas Archipelago, Panama (Malacostraca: Decapoda: Alpheidae)" masterLastPageNumber="140" masterPageNumber="125" pageNumber="134" updateTime="1698748658439" updateUser="ExternalLinkService">
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<mods:title id="6BAB73C72FC9D530EDCDF2DF3B8464E8">The eastern Pacific species of Salmoneus Holthuis, 1955, with description of a remarkable new species from Las Perlas Archipelago, Panama (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<treatment id="087A87B4FFD8FFFBBD88F90E3C13FEE1" ID-DOI="http://doi.org/10.5281/zenodo.3511377" ID-GBIF-Taxon="158956247" ID-Zenodo-Dep="3511377" LSID="urn:lsid:plazi:treatment:087A87B4FFD8FFFBBD88F90E3C13FEE1" httpUri="http://treatment.plazi.org/id/087A87B4FFD8FFFBBD88F90E3C13FEE1" lastPageId="12" lastPageNumber="137" pageId="9" pageNumber="134">
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<taxonomicName id="47D34D21FFD8FFFEBD88F90E3C04F94E" ID-CoL="85Y27" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[151,566,1746,1773]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B2A7EAB0FFD8FFFEBD88F90E3C04F94E" bold="true" box="[151,566,1746,1773]" pageId="9" pageNumber="134">
<emphasis id="B2A7EAB0FFD8FFFEBD88F90E3FA9F94F" bold="true" box="[151,411,1746,1773]" italics="true" pageId="9" pageNumber="134">Salmoneus excavatus</emphasis>
<bibRefCitation id="E4424B53FFD8FFFEBCBEF90E3C04F94E" author="Anker, A." box="[417,566,1746,1772]" pageId="9" pageNumber="134" pagination="39 - 50" refId="ref8626" refString="Anker, A. (2011) Two new species of Salmoneus Holthuis, 1955 with a deep dorsal depression on the carapace (Crustacea, Decapoda, Alpheidae). Zootaxa, 3041 (1), 39 - 50. https: // doi. org / 10.11646 / zootaxa. 3041.1.2" type="journal article" year="2011">Anker, 2011</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8C96529FFD8FFFEBD88F92A3EEBF8B3" box="[151,217,1782,1809]" pageId="9" pageNumber="134" type="description">
<paragraph id="806C36A2FFD8FFFEBD88F92A3EEBF8B3" blockId="9.[151,566,1746,1809]" box="[151,217,1782,1809]" pageId="9" pageNumber="134">
<figureCitation id="18E82A27FFD8FFFEBD88F92A3EEBF8B3" box="[151,217,1782,1809]" captionStart="FIGURE 7" captionStartId="11.[151,250,1924,1949]" captionTargetBox="[151,1432,166,1904]" captionTargetId="figure@11.[151,1432,166,1904]" captionTargetPageId="11" captionText="FIGURE 7. Salmoneus excavatus Anker, 2011: non-ovigerous specimen (cl 5.0 mm) from Las Perlas, Panama, MZUSP 33663 [a, b]; ovigerous specimen (cl 4.9 mm), from the same locality, OUMNH.ZC. 2015-08-014 [c]; a—living individual, dorsal view; b—same, lateral view; c—living individual, lateral view." httpUri="https://zenodo.org/record/3359251/files/figure.png" pageId="9" pageNumber="134">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8C96529FFD8FFFEBD88F8E93C4DF8EF" box="[151,639,1845,1870]" pageId="9" pageNumber="134" type="reference_group">
<paragraph id="806C36A2FFD8FFFEBD88F8E93C4DF8EF" blockId="9.[151,639,1845,1870]" box="[151,639,1845,1870]" pageId="9" pageNumber="134">
<treatmentCitationGroup id="A0C3118CFFD8FFFEBD88F8E93C4DF8EF" box="[151,639,1845,1870]" pageId="9" pageNumber="134">
<taxonomicName id="47D34D21FFD8FFFEBD88F8E93C24F8EF" ID-CoL="85Y27" authority="Anker 2011: 40" authorityName="Anker" authorityPageNumber="40" authorityYear="2011" box="[151,534,1845,1870]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B2A7EAB0FFD8FFFEBD88F8E93F5DF8EF" box="[151,367,1845,1869]" italics="true" pageId="9" pageNumber="134">Salmoneus excavatus</emphasis>
<treatmentCitation id="017210B3FFD8FFFEBC6AF8E93C24F8EF" ID-DOI="http://doi.org/10.5281/zenodo.3506273" ID-Zenodo-Dep="3506273" author="Anker, A." box="[373,534,1845,1870]" httpUri="http://treatment.plazi.org/id/AF0887E75E0EFFE6FF1DD04AFD614D41" page="40" pageId="9" pageNumber="134" year="2011">
<bibRefCitation id="E4424B53FFD8FFFEBC6AF8E93C24F8EF" author="Anker, A." box="[373,534,1845,1870]" pageId="9" pageNumber="134" pagination="39 - 50" refId="ref8626" refString="Anker, A. (2011) Two new species of Salmoneus Holthuis, 1955 with a deep dorsal depression on the carapace (Crustacea, Decapoda, Alpheidae). Zootaxa, 3041 (1), 39 - 50. https: // doi. org / 10.11646 / zootaxa. 3041.1.2" type="journal article" year="2011">Anker 2011: 40</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, figs. 13.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8C96529FFD8FFFBBD88F8A93C13FEE1" lastPageId="12" lastPageNumber="137" pageId="9" pageNumber="134" type="materials_examined">
<paragraph id="806C36A2FFD8FFFEBD88F8A93F3BF85E" blockId="9.[151,1436,1909,2044]" pageId="9" pageNumber="134">
<emphasis id="B2A7EAB0FFD8FFFEBD88F8A93FB6F832" bold="true" box="[151,388,1909,1936]" pageId="9" pageNumber="134">Material examined.</emphasis>
1 non-ovigerous specimen (cl 5.0 mm),
<collectionCode id="E6C2AE67FFD8FFFEBE7CF8AA3D8DF832" box="[867,959,1910,1936]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="134">MZUSP</collectionCode>
33663,
<collectingCountry id="F8C47632FFD8FFFEB93EF8AB3A4FF833" box="[1057,1149,1911,1937]" name="Panama" pageId="9" pageNumber="134">Panama</collectingCountry>
, Las Perlas Archipelago,
<collectingCountry id="F8C47632FFD8FFFEBD88F8463E8DF816" box="[151,191,1946,1972]" name="Iceland" pageId="9" pageNumber="134">Isla</collectingCountry>
Casayeta, shallow muddy bay fringed by mangroves, under muddy intertidal rocks at low tide, leg. A. Anker,
<date id="F46D1062FFD8FFFEBD88F8613F2BF87A" box="[151,281,1981,2008]" pageId="9" pageNumber="134" value="2015-04-20">20.04.2015</date>
; 1 ovigerous specimen (cl
<quantity id="472B9B47FFD8FFFEBF53F8623C9AF87B" box="[588,680,1982,2009]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="134" unit="mm" value="4.9">4.9 mm</quantity>
),
<collectionCode id="E6C2AE67FFD8FFFEBFA2F8623D15F87A" box="[701,807,1982,2008]" pageId="9" pageNumber="134">OUMNH</collectionCode>
.
<collectionCode id="E6C2AE67FFD8FFFEBE35F8623D66F87A" box="[810,852,1982,2008]" country="Austria" name="Zoological Collection, University of Vienna" pageId="9" pageNumber="134" type="University or college">ZC</collectionCode>
. 2015-08-014, same collection data as for previous specimen.
</paragraph>
<caption id="D4AC662AFFDBFFFDBD88F9FE3F8BF921" httpUri="https://zenodo.org/record/3359249/files/figure.png" pageId="10" pageNumber="135" startId="10.[151,250,1570,1595]" targetBox="[151,1436,181,1551]" targetPageId="10">
<paragraph id="806C36A2FFDBFFFDBD88F9FE3F8BF921" blockId="10.[151,1436,1570,1667]" pageId="10" pageNumber="135">
<emphasis id="B2A7EAB0FFDBFFFDBD88F9FE3F3EF999" bold="true" box="[151,268,1570,1595]" pageId="10" pageNumber="135">FIGURE 6</emphasis>
.
<taxonomicName id="47D34D21FFDBFFFDBC07F9FF3CC1F999" authority="Anker &amp; Lazarus, 2015" authorityName="Anker &amp; Lazarus" authorityYear="2015" box="[280,755,1570,1595]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="135" phylum="Arthropoda" rank="species" species="malagensis">
<emphasis id="B2A7EAB0FFDBFFFDBC07F9FF3FCFF999" box="[280,509,1571,1595]" italics="true" pageId="10" pageNumber="135">Salmoneus malagensis</emphasis>
<bibRefCitation id="E4424B53FFDBFFFDBF1DF9FE3CC1F999" author="Anker, A. &amp; Lazarus, J. F." box="[514,755,1570,1595]" pageId="10" pageNumber="135" pagination="520 - 534" refId="ref8680" refString="Anker, A. &amp; Lazarus, J. F. (2015) On two new species of the shrimp genus Salmoneus Holthuis, 1955 (Decapoda, Caridea, Alpheidae) from the tropical eastern Pacific. Zootaxa, 3957 (5), 520 - 534. https: // doi. org / 10.11646 / zootaxa. 3957.5.2" type="journal article" year="2015">Anker &amp; Lazarus, 2015</bibRefCitation>
</taxonomicName>
: non-ovigerous specimen (cl 3.4 mm, missing major cheliped) from Puerto Cañas, Azuero Peninsula, Panama, FLMNH UF 51907; a—living individual (covered with some mud particles), dorsal view; b—same, lateral view.
</paragraph>
</caption>
<paragraph id="806C36A2FFDBFFFDBDD8F9083C19F94D" blockId="10.[151,1437,1748,2028]" box="[199,555,1748,1775]" pageId="10" pageNumber="135">
<emphasis id="B2A7EAB0FFDBFFFDBDD8F9083F6BF94D" bold="true" box="[199,345,1748,1775]" pageId="10" pageNumber="135">Description.</emphasis>
See
<bibRefCitation id="E4424B53FFDBFFFDBC91F9093C15F94D" author="Anker, A." box="[398,551,1749,1775]" pageId="10" pageNumber="135" pagination="39 - 50" refId="ref8626" refString="Anker, A. (2011) Two new species of Salmoneus Holthuis, 1955 with a deep dorsal depression on the carapace (Crustacea, Decapoda, Alpheidae). Zootaxa, 3041 (1), 39 - 50. https: // doi. org / 10.11646 / zootaxa. 3041.1.2" type="journal article" year="2011">Anker (2011)</bibRefCitation>
.
</paragraph>
<paragraph id="806C36A2FFDBFFFDBDD8F9243C05F89A" blockId="10.[151,1437,1748,2028]" pageId="10" pageNumber="135">
<emphasis id="B2A7EAB0FFDBFFFDBDD8F9243F6AF8B1" bold="true" box="[199,344,1784,1811]" pageId="10" pageNumber="135">Colouration</emphasis>
. Semitransparent creamy whitish, sometimes with slight yellowish tinge; gonads or fresh eggs yellow (
<bibRefCitation id="E4424B53FFDBFFFDBDEDF8C13F49F895" author="Anker, A." box="[242,379,1821,1847]" pageId="10" pageNumber="135" pagination="39 - 50" refId="ref8626" refString="Anker, A. (2011) Two new species of Salmoneus Holthuis, 1955 with a deep dorsal depression on the carapace (Crustacea, Decapoda, Alpheidae). Zootaxa, 3041 (1), 39 - 50. https: // doi. org / 10.11646 / zootaxa. 3041.1.2" type="journal article" year="2011">Anker 2011</bibRefCitation>
; see also
<figureCitation id="18E82A27FFDBFFFDBCFAF8C13C1BF89A" box="[485,553,1821,1848]" captionStart="FIGURE 7" captionStartId="11.[151,250,1924,1949]" captionTargetBox="[151,1432,166,1904]" captionTargetId="figure@11.[151,1432,166,1904]" captionTargetPageId="11" captionText="FIGURE 7. Salmoneus excavatus Anker, 2011: non-ovigerous specimen (cl 5.0 mm) from Las Perlas, Panama, MZUSP 33663 [a, b]; ovigerous specimen (cl 4.9 mm), from the same locality, OUMNH.ZC. 2015-08-014 [c]; a—living individual, dorsal view; b—same, lateral view; c—living individual, lateral view." httpUri="https://zenodo.org/record/3359251/files/figure.png" pageId="10" pageNumber="135">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="806C36A2FFDBFFFDBDD8F89E3A97F8FE" blockId="10.[151,1437,1748,2028]" box="[199,1189,1856,1884]" pageId="10" pageNumber="135">
<materialsCitation id="30BB3CFFFFDBFFFDBDD8F89E3A93F8FE" ID-GBIF-Occurrence="2302332022" box="[199,1185,1856,1884]" country="Panama" location="Southwest" municipality="Pacific" pageId="10" pageNumber="135" specimenCount="1" stateProvince="Panama" typeStatus="holotype">
<emphasis id="B2A7EAB0FFDBFFFDBDD8F89E3F52F8F9" bold="true" box="[199,352,1856,1884]" pageId="10" pageNumber="135">
<typeStatus id="5F688800FFDBFFFDBDD8F89E3F30F8FE" box="[199,258,1858,1884]" pageId="10" pageNumber="135">Type</typeStatus>
locality
</emphasis>
.
<location id="850C6079FFDBFFFDBC71F89D3FD1F8F9" LSID="urn:lsid:plazi:treatment:087A87B4FFD8FFFBBD88F90E3C13FEE1:850C6079FFDBFFFDBC71F89D3FD1F8F9" box="[366,483,1857,1883]" country="Panama" municipality="Pacific" name="Southwest" pageId="10" pageNumber="135" stateProvince="Panama">Southwest</location>
of
<collectingRegion id="4217F840FFDBFFFDBF17F89D3CE8F8F9" box="[520,730,1857,1883]" country="Panama" name="Panama" pageId="10" pageNumber="135">Ciudad de Panamá</collectingRegion>
(
<collectingRegion id="4217F840FFDBFFFDBFF7F89D3D4CF8FE" box="[744,894,1857,1884]" country="Panama" name="Panama" pageId="10" pageNumber="135">Panama City</collectingRegion>
),
<collectingMunicipality id="6008ACD8FFDBFFFDBE8EF89D3DEDF8F9" box="[913,991,1857,1883]" pageId="10" pageNumber="135">Pacific</collectingMunicipality>
coast of
<collectingCountry id="F8C47632FFDBFFFDB95AF89E3A93F8FE" box="[1093,1185,1858,1884]" name="Panama" pageId="10" pageNumber="135">Panama</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="806C36A2FFDBFFFDBDD8F8B83ED1F806" blockId="10.[151,1437,1748,2028]" pageId="10" pageNumber="135">
<emphasis id="B2A7EAB0FFDBFFFDBDD8F8B83F6BF8DD" bold="true" box="[199,345,1892,1919]" pageId="10" pageNumber="135">Distribution</emphasis>
. Eastern Pacific:
<collectingCountry id="F8C47632FFDBFFFDBF33F8B93CA8F8DD" box="[556,666,1893,1919]" name="Colombia" pageId="10" pageNumber="135">Colombia</collectingCountry>
(Bahía Málaga) and
<collectingCountry id="F8C47632FFDBFFFDBE86F8BA3DC0F822" box="[921,1010,1894,1920]" name="Panama" pageId="10" pageNumber="135">Panama</collectingCountry>
(Chumical, Veracruz, Las Perlas Islands).
</paragraph>
<paragraph id="806C36A2FFDBFFFDBDD8F8723FE2F849" blockId="10.[151,1437,1748,2028]" pageId="10" pageNumber="135">
<emphasis id="B2A7EAB0FFDBFFFDBDD8F8723F14F86A" bold="true" box="[199,294,1966,1992]" pageId="10" pageNumber="135">Ecology</emphasis>
. Intertidal and very shallow subtidal; rocky and mixed rocky-sandy shores, sometimes close to mangroves; under muddy rocks.
</paragraph>
<caption id="D4AC662AFFDAFFFCBD88F8583D25F847" httpUri="https://zenodo.org/record/3359251/files/figure.png" pageId="11" pageNumber="136" startId="11.[151,250,1924,1949]" targetBox="[151,1432,166,1904]" targetPageId="11">
<paragraph id="806C36A2FFDAFFFCBD88F8583D25F847" blockId="11.[151,1436,1924,2021]" pageId="11" pageNumber="136">
<emphasis id="B2A7EAB0FFDAFFFCBD88F8583F3FF83F" bold="true" box="[151,269,1924,1949]" pageId="11" pageNumber="136">FIGURE 7</emphasis>
.
<taxonomicName id="47D34D21FFDAFFFCBC06F8593C44F83F" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[281,630,1924,1949]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="136" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B2A7EAB0FFDAFFFCBC06F8593FB5F83F" box="[281,391,1925,1949]" italics="true" pageId="11" pageNumber="136">Salmoneus</emphasis>
<emphasis id="B2A7EAB0FFDAFFFCBC92F85A3FC3F83F" box="[397,497,1926,1949]" italics="true" pageId="11" pageNumber="136">excavatus</emphasis>
<bibRefCitation id="E4424B53FFDAFFFCBCE9F8583C44F83F" author="Anker, A." box="[502,630,1924,1949]" pageId="11" pageNumber="136" pagination="39 - 50" refId="ref8626" refString="Anker, A. (2011) Two new species of Salmoneus Holthuis, 1955 with a deep dorsal depression on the carapace (Crustacea, Decapoda, Alpheidae). Zootaxa, 3041 (1), 39 - 50. https: // doi. org / 10.11646 / zootaxa. 3041.1.2" type="journal article" year="2011">Anker, 2011</bibRefCitation>
</taxonomicName>
: non-ovigerous specimen (cl 5.0 mm) from Las Perlas, Panama, MZUSP 33663 [a, b]; ovigerous specimen (cl 4.9 mm), from the same locality, OUMNH.ZC. 2015-08-014 [c]; a—living individual, dorsal view; b—same, lateral view; c—living individual, lateral view.
</paragraph>
</caption>
<paragraph id="806C36A2FFDDFFFBBDD8FF453C13FEE1" blockId="12.[151,1437,152,323]" pageId="12" pageNumber="137">
<emphasis id="B2A7EAB0FFDDFFFBBDD8FF453F06FF11" bold="true" box="[199,308,153,179]" pageId="12" pageNumber="137">Remarks</emphasis>
. Among the eastern Pacific
<taxonomicName id="47D34D21FFDDFFFBBF78FF443CEFFF10" box="[615,733,152,178]" class="Malacostraca" family="Alpheidae" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="137" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
,
<taxonomicName id="47D34D21FFDDFFFBBFF8FF443DE6FF10" authorityName="Anker" authorityYear="2011" box="[743,980,152,178]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="137" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B2A7EAB0FFDDFFFBBFF8FF443DE6FF10" box="[743,980,152,178]" italics="true" pageId="12" pageNumber="137">Salmoneus excavatus</emphasis>
</taxonomicName>
is the only species with a dorsally deeply indented carapace (
<bibRefCitation id="E4424B53FFDDFFFBBC6CFF603FCCFF75" author="Anker, A." box="[371,510,188,215]" pageId="12" pageNumber="137" pagination="39 - 50" refId="ref8626" refString="Anker, A. (2011) Two new species of Salmoneus Holthuis, 1955 with a deep dorsal depression on the carapace (Crustacea, Decapoda, Alpheidae). Zootaxa, 3041 (1), 39 - 50. https: // doi. org / 10.11646 / zootaxa. 3041.1.2" type="journal article" year="2011">Anker 2011</bibRefCitation>
; dorsal depression visible as a thin longitudinal line in
<figureCitation id="18E82A27FFDDFFFBB965FF603AFFFF75" box="[1146,1229,188,215]" captionStart="FIGURE 7" captionStartId="11.[151,250,1924,1949]" captionTargetBox="[151,1432,166,1904]" captionTargetId="figure@11.[151,1432,166,1904]" captionTargetPageId="11" captionText="FIGURE 7. Salmoneus excavatus Anker, 2011: non-ovigerous specimen (cl 5.0 mm) from Las Perlas, Panama, MZUSP 33663 [a, b]; ovigerous specimen (cl 4.9 mm), from the same locality, OUMNH.ZC. 2015-08-014 [c]; a—living individual, dorsal view; b—same, lateral view; c—living individual, lateral view." httpUri="https://zenodo.org/record/3359251/files/figure.png" pageId="12" pageNumber="137">Fig. 7c</figureCitation>
). The species was described based on only four specimens, three from the Pacific coast of
<collectingCountry id="F8C47632FFDDFFFBBEFEFF3D3A08FF59" box="[993,1082,225,251]" name="Panama" pageId="12" pageNumber="137">Panama</collectingCountry>
(Chumical and Veracruz areas west of
<collectingRegion id="4217F840FFDDFFFBBDF0FED93FB3FEBD" box="[239,385,260,287]" country="Panama" name="Panama" pageId="12" pageNumber="137">Panama City</collectingRegion>
) and one from
<collectingCountry id="F8C47632FFDDFFFBBF34FED83CABFEBC" box="[555,665,260,286]" name="Colombia" pageId="12" pageNumber="137">Colombia</collectingCountry>
(Bahía Málaga). It is now recorded for the first time from Las Perlas Archipelago in the Bay of
<collectingCountry id="F8C47632FFDDFFFBBCDEFEF53C2FFEE1" box="[449,541,297,323]" name="Panama" pageId="12" pageNumber="137">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>