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<document id="D2339D4408B8D611C5CB2E235415D103" ID-CLB-Dataset="26107" ID-DOI="10.11646/zootaxa.4651.1.8" ID-GBIF-Dataset="0e1cc894-7613-41e5-bc48-b5224cce0670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3359237" ID-ZooBank="EB7EE06F-0CA3-4AA5-A152-31645179F419" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564781000653" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2019" docId="087A87B4FFD6FFFEBD88FD213C49FD25" docLanguage="en" docName="zootaxa.4651.1.8.pdf" docOrigin="Zootaxa 4651 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Salmoneus serratidigitus" docType="treatment" docVersion="14" lastPageNumber="134" masterDocId="F443FFCCFFD1FFF7BD1FFFDC3E32FFA2" masterDocTitle="The eastern Pacific species of Salmoneus Holthuis, 1955, with description of a remarkable new species from Las Perlas Archipelago, Panama (Malacostraca: Decapoda: Alpheidae)" masterLastPageNumber="140" masterPageNumber="125" pageNumber="132" updateTime="1698748658439" updateUser="ExternalLinkService">
<mods:mods id="EFF6E115A6C3E7590482B057C75A5C9F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1273E8A92580CF501327FF2EA41095EE">The eastern Pacific species of Salmoneus Holthuis, 1955, with description of a remarkable new species from Las Perlas Archipelago, Panama (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<mods:namePart id="323AF5AABEDB98F75AD63A96A1AE9264">Anker, Arthur</mods:namePart>
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<mods:typeOfResource id="08AEACD11341DED09ED4C6969A6B00FA">text</mods:typeOfResource>
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<mods:title id="7555412234DFCE376A10D11ADECF2191">Zootaxa</mods:title>
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<mods:part id="F207019006017DB97B24560D5E937CDA">
<mods:date id="6885B2FADB2E8C6C13E221930FA17720">2019</mods:date>
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<mods:number id="FA7870598C86C3FFD398138B2694F20E">2019-08-02</mods:number>
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<mods:detail id="A46C2CF7B42775DC69D8421E9FEC02C3" type="volume">
<mods:number id="6C0A70F2E92A813E3C0619A7FED941E7">4651</mods:number>
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<mods:classification id="A88E996F87231930E566625AB78602A8">journal article</mods:classification>
<mods:identifier id="3C730A59BE9F5A216098516FD63C2687" type="CLB-Dataset">26107</mods:identifier>
<mods:identifier id="58455D4E04742563DA9425845D17D6DA" type="DOI">10.11646/zootaxa.4651.1.8</mods:identifier>
<mods:identifier id="AB19101A4C50923720941DA1248DC879" type="GBIF-Dataset">0e1cc894-7613-41e5-bc48-b5224cce0670</mods:identifier>
<mods:identifier id="8B68DD9ACDD2C028FC602A10C90ECB07" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D3E55BE25B26D613B2CA0C83AFB23C2A" type="Zenodo-Dep">3359237</mods:identifier>
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<treatment id="087A87B4FFD6FFFEBD88FD213C49FD25" ID-GBIF-Taxon="158956248" LSID="urn:lsid:plazi:treatment:087A87B4FFD6FFFEBD88FD213C49FD25" httpUri="http://treatment.plazi.org/id/087A87B4FFD6FFFEBD88FD213C49FD25" lastPageId="9" lastPageNumber="134" pageId="7" pageNumber="132">
<subSubSection id="C8C96529FFD6FFF0BD88FD213CA7FCB5" box="[151,661,764,792]" pageId="7" pageNumber="132" type="nomenclature">
<paragraph id="806C36A2FFD6FFF0BD88FD213CA7FCB5" blockId="7.[151,661,764,827]" box="[151,661,764,792]" pageId="7" pageNumber="132">
<heading id="DB2481CEFFD6FFF0BD88FD213CA7FCB5" bold="true" box="[151,661,764,792]" fontSize="11" level="1" pageId="7" pageNumber="132" reason="1">
<taxonomicName id="47D34D21FFD6FFF0BD88FD213CA7FCB5" ID-CoL="85Y2S" authority="(Coutiere, 1897)" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[151,661,764,792]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD6FFF0BD88FD213CA7FCB5" bold="true" box="[151,661,764,792]" pageId="7" pageNumber="132">
<emphasis id="B2A7EAB0FFD6FFF0BD88FD213FF6FCBA" bold="true" box="[151,452,765,792]" italics="true" pageId="7" pageNumber="132">Salmoneus serratidigitus</emphasis>
(
<bibRefCitation id="E4424B53FFD6FFF0BCCCFD203CBEFCB5" author="Coutiere, H." box="[467,652,764,791]" pageId="7" pageNumber="132" pagination="380 - 386" refId="ref8928" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8C96529FFD6FFF0BD88FCFD3AD6FB97" pageId="7" pageNumber="132" type="description">
<paragraph id="806C36A2FFD6FFF0BD88FCFD3EEBFC99" blockId="7.[151,661,764,827]" box="[151,217,800,827]" pageId="7" pageNumber="132">
<figureCitation id="18E82A27FFD6FFF0BD88FCFD3EEBFC99" box="[151,217,800,827]" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1743]" captionTargetBox="[151,1436,704,1698]" captionTargetId="figure@8.[151,1436,704,1698]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus serratidigitus (Coutière, 1897): ovigerous specimen (cl 6.1 mm) from Las Perlas, Panama, MZUSP 33810 [a]; non-ovigerous specimen (cl 4.9 mm, parasitised by a pair of bopyrid isopods) from Las Perlas, Panama, MNHN- IU-2014-20291 [b, c]; a—living individual, chelipeds folded under the body, lateral view; b—same, chelipeds folded under the body, dorsal view; c—same, major cheliped extended, lateral view." httpUri="https://zenodo.org/record/3359247/files/figure.png" pageId="7" pageNumber="132">Fig. 5</figureCitation>
</paragraph>
<paragraph id="806C36A2FFD6FFF0BD88FCB83AD6FB97" blockId="7.[151,1437,868,1077]" pageId="7" pageNumber="132">
(Selected synonymy)
<taxonomicName id="47D34D21FFD6FFF0BD88FC593C48FC3E" authority="Coutiere 1897: 382" authorityName="Coutiere" authorityPageNumber="382" authorityYear="1897" box="[151,634,899,924]" class="Malacostraca" family="Alpheidae" genus="Jousseaumea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD6FFF0BD88FC593F9CFC3E" box="[151,430,900,924]" italics="true" pageId="7" pageNumber="132">Jousseaumea serratidigitus</emphasis>
<bibRefCitation id="E4424B53FFD6FFF0BCACFC5F3C48FC3E" author="Coutiere, H." box="[435,634,899,924]" pageId="7" pageNumber="132" pagination="380 - 386" refId="ref8928" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière 1897: 382</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="E4424B53FFD6FFF0BF9BFC5F3D78FC3E" author="Coutiere, H." box="[644,842,899,924]" pageId="7" pageNumber="132" pagination="1 - 559" refId="ref8963" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière 1899: 463</bibRefCitation>
, 544 (records from the Gulf of
<collectingRegion id="4217F840FFD6FFF0B998FC5F3AC1FC3E" box="[1159,1267,899,924]" country="United States of America" name="California" pageId="7" pageNumber="132">California</collectingRegion>
).
<taxonomicName id="47D34D21FFD6FFF0BD88FC7E3C89FC19" authority="- Banner &amp; Banner 1981: 58" authorityName="- Banner &amp; Banner" authorityPageNumber="58" authorityYear="1981" box="[151,699,930,955]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD6FFF0BD88FC7E3FA1FC18" box="[151,403,930,954]" italics="true" pageId="7" pageNumber="132">Salmoneus serratidigitus</emphasis>
—
<bibRefCitation id="E4424B53FFD6FFF0BCB2FC7F3C89FC19" author="Banner, D. M. &amp; Banner, A. H." box="[429,699,930,955]" pageId="7" pageNumber="132" pagination="1 - 99" refId="ref8791" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 58</bibRefCitation>
</taxonomicName>
(part.), figs. 7a–g, 8;
<bibRefCitation id="E4424B53FFD6FFF0BE8CFC7E3A60FC19" author="Wicksten, M. K." box="[915,1106,930,955]" pageId="7" pageNumber="132" pagination="1 - 59" refId="ref9935" refString="Wicksten, M. K. (1983) A monograph on the shallow water caridean shrimps of the Gulf of California, Mexico. Alan Hancock Monographs in Marine Biology, 13, 1 - 59." type="journal article" year="1983">Wicksten 1983: 40</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0B944FC7E3BA5FC19" author="Wicksten, M. K. &amp; Hendrickx, M. E." box="[1115,1431,930,955]" pageId="7" pageNumber="132" pagination="1 - 11" refId="ref10089" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (1992) Checklist of penaeoid and caridean shrimps (Decapoda: Penaeoidea, Caridea) from the eastern tropical Pacific. Proceedings of San Diego Society of Natural History, 9, 1 - 11." type="journal article" year="1992">Wicksten &amp; Hendrickx 1992: 6</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0BDD8FC1D3FA4FC78" author="Wicksten, M. K." box="[199,406,961,986]" pageId="7" pageNumber="132" pagination="147 - 156" refId="ref10020" refString="Wicksten, M. K. (1993) Caridean and stenopodid shrimp of the Galapagos Islands. In: James, M. J. (Ed.), Galapagos Marine Invertebrates. Chapter 6. Plenum, New York, pp. 147 - 156. https: // doi. org / 10.1007 / 978 - 1 - 4899 - 0646 - 5 _ 7" type="book chapter" year="1993">Wicksten 1993: 151</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0BCBEFC1E3C7EFC7B" author="Bruce, A. J." box="[417,588,961,985]" pageId="7" pageNumber="132" pagination="329 - 343" refId="ref8831" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (Ed.), The marine fauna and flora of Hong Kong and southern China III. Proceedings of the Fourth International Marine Biological Workshop. The Marine Fauna and Flora of Hong Kong and southern China, Hong Kong University Press, Hong Kong, pp. 329 - 343." type="book chapter" year="1992">Bruce 1992: 340</bibRefCitation>
, fig. 7; Hernández Aguilera &amp; Martínez Guzmán 1992: 4;
<bibRefCitation id="E4424B53FFD6FFF0B9BEFC1D3BAEFC78" author="Hickman, C. P. &amp; Zimmerman, T. L." box="[1185,1436,961,986]" pageId="7" pageNumber="132" refId="ref9392" refString="Hickman, C. P. &amp; Zimmerman, T. L. (2000) A field guide to crustaceans of Galapagos. Galapagos marine life series. Sugar Spring Press, Lexington, 156 pp." type="book" year="2000">Hickman &amp; Zimmerman</bibRefCitation>
2000: 43;
<bibRefCitation id="E4424B53FFD6FFF0BC30FC033C4CFC5A" author="Wicksten, M. K. &amp; Hendrickx, M. E." box="[303,638,991,1016]" pageId="7" pageNumber="132" pagination="49 - 76" refId="ref10139" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (2003) An updated checklist of benthic marine and brackish water shrimps (Decapoda: Penaeoidea, Stenopodidea, Caridea) from the eastern tropical Pacific. In: Hendrickx, M. E. (Ed.), Contributions to the study of East Pacific crustaceans / Contribuciones al estudio de los crustaceos del Pacifico Este. Instituto de Ciencias del Mar y Limnologia, UNAM, Mexico City, pp. 49 - 76." type="book chapter" year="2003">Wicksten &amp; Hendrickx 2003: 66</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0BF96FC033A69FC5A" author="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R." box="[649,1115,991,1016]" pageId="7" pageNumber="132" pagination="221 - 239" refId="ref9512" refString="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R. (2007) Crustaceos (Crustacea: Sessilia, Stomatopoda, Isopoda, Amphipoda, Decapoda) de Bahia Malaga, Valle del Cauca (Pacifico colombiano). Biota Colombiana, 8, 221 - 239." type="journal article" year="2007">Lazarus-Agudelo &amp; Cantera-Kintz 2007: 228</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0B979FC033B5CFC5A" author="Duris, Z. &amp; Horka, I." box="[1126,1390,991,1016]" pageId="7" pageNumber="132" pagination="773 - 793" refId="ref9127" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016: 785</bibRefCitation>
, fig. 13. Not
<emphasis id="B2A7EAB0FFD6FFF0BDDBFBC13FE5FB97" box="[196,471,1053,1077]" italics="true" pageId="7" pageNumber="132">
<taxonomicName id="47D34D21FFD6FFF0BDDBFBC13F00FB97" authorityName="Holthuis" authorityYear="1955" box="[196,306,1053,1077]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="genus">Salmoneus</taxonomicName>
serratidigitus—
</emphasis>
<bibRefCitation id="E4424B53FFD6FFF0BCC8FBC03C9CFB97" author="Hendrickx, M. E." box="[471,686,1052,1077]" pageId="7" pageNumber="132" pagination="395 - 397" refId="ref9299" refString="Hendrickx, M. E. (2001) New distribution, size and habitats records of decapod crustaceans from the eastern tropical Pacific. Revista de Biologia Tropical, 49, 395 - 397." type="journal article" year="2001">Hendrickx 2001: 395</bibRefCitation>
(=
<taxonomicName id="47D34D21FFD6FFF0BFCEFBC13A54FB97" authority="Ramos-Tafur, 2018" authorityName="Ramos-Tafur" authorityYear="2018" box="[721,1126,1052,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="luiszapatai">
<emphasis id="B2A7EAB0FFD6FFF0BFCEFBC13DAAFB97" box="[721,920,1053,1077]" italics="true" pageId="7" pageNumber="132">Alpheus luiszapatai</emphasis>
<bibRefCitation id="E4424B53FFD6FFF0BE80FBC03A54FB97" author="Ramos-Tafur, G. E." box="[927,1126,1052,1077]" pageId="7" pageNumber="132" pagination="540 - 556" refId="ref9661" refString="Ramos-Tafur, G. E. (2018) Alpheus luiszapatai, a new species of rare and colorful deep water alpheid shrimp (Crustacea: Decapoda: Alpheidae) from Arusi, Choco Department, Pacific Coast of Colombia. Zootaxa, 4403 (3), 540 - 556. https: // doi. org / 10.11646 / zootaxa. 4403.3.7" type="journal article" year="2018">Ramos-Tafur, 2018</bibRefCitation>
</taxonomicName>
; see below).
</paragraph>
</subSubSection>
<subSubSection id="C8C96529FFD6FFFEBD88FBBF3AA5FEE1" lastPageId="9" lastPageNumber="134" pageId="7" pageNumber="132" type="materials_examined">
<paragraph id="806C36A2FFD6FFF0BD88FBBF3B6EF938" blockId="7.[151,1437,1123,2014]" pageId="7" pageNumber="132">
<emphasis id="B2A7EAB0FFD6FFF0BD88FBBF3F48FBDC" bold="true" box="[151,378,1123,1150]" pageId="7" pageNumber="132">Material examined</emphasis>
. 1 non-ovigerous specimen (cl
<quantity id="472B9B47FFD6FFF0BFC2FBB83D04FBDC" box="[733,822,1124,1150]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="132" unit="mm" value="4.9">4.9 mm</quantity>
, parasitised by a pair of bopyrid isopods), MNHN-IU- 2014-20291,
<collectingCountry id="F8C47632FFD6FFF0BC2FFB543FBEFB00" box="[304,396,1160,1186]" name="Panama" pageId="7" pageNumber="132">Panama</collectingCountry>
, Las Perlas Archipelago, small rocky islet off
<collectingCountry id="F8C47632FFD6FFF0BEB8FB5B3DFDFB03" box="[935,975,1159,1185]" name="Iceland" pageId="7" pageNumber="132">Isla</collectingCountry>
Contadora, under rocks at extreme low tide, leg. A. Anker
<emphasis id="B2A7EAB0FFD6FFF0BC75FB703F93FB64" box="[362,417,1196,1222]" italics="true" pageId="7" pageNumber="132">et al.</emphasis>
,
<date id="F46D1062FFD6FFF0BCB0FB773C1EFB67" box="[431,556,1195,1221]" pageId="7" pageNumber="132" value="2006-03-31">31.03.2006</date>
[fcn 06-348]; 1 ovigerous specimen (cl
<quantity id="472B9B47FFD6FFF0BEF4FB773A77FB64" box="[1003,1093,1195,1222]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="7" pageNumber="132" unit="mm" value="6.1">6.1 mm</quantity>
),
<collectionCode id="E6C2AE67FFD6FFF0B948FB703A81FB64" box="[1111,1203,1196,1222]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="132">MZUSP</collectionCode>
33810,
<collectingCountry id="F8C47632FFD6FFF0B811FB703B58FB64" box="[1294,1386,1196,1222]" name="Panama" pageId="7" pageNumber="132">Panama</collectingCountry>
, Las Perlas Archipelago, small rocky islet off north-eastern coast of
<collectingCountry id="F8C47632FFD6FFF0BE4CFB133D49FB4B" box="[851,891,1231,1257]" name="Iceland" pageId="7" pageNumber="132">Isla</collectingCountry>
Saboga, rock-sand bottom, depth at low tide:
<quantity id="472B9B47FFD6FFF0B866FB133E9DFAAC" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="132" unit="m" value="0.5">0.5 m</quantity>
, strong current passage, under large rock on coarse sand, leg. A. Anker,
<date id="F46D1062FFD6FFF0BEFAFB2F3A55FAAC" box="[997,1127,1267,1294]" pageId="7" pageNumber="132" value="2015-04-21">21.04.2015</date>
; 1 non-ovigerous specimen (cl
<quantity id="472B9B47FFD6FFF0BDA2FACB3F2AFA90" box="[189,280,1303,1330]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="7" pageNumber="132" unit="mm" value="6.8">6.8 mm</quantity>
),
<collectionCode id="E6C2AE67FFD6FFF0BC33FAC43FBCFA90" box="[300,398,1304,1330]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="7" pageNumber="132" type="Museum">FLMNH</collectionCode>
<collectionCode id="E6C2AE67FFD6FFF0BC89FAC43F88FA90" box="[406,442,1304,1330]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="7" pageNumber="132" type="Museum">UF</collectionCode>
51902,
<collectingCountry id="F8C47632FFD6FFF0BF08FAC43C41FA90" box="[535,627,1304,1330]" name="Panama" pageId="7" pageNumber="132">Panama</collectingCountry>
, Las Perlas Archipelago,
<collectingCountry id="F8C47632FFD6FFF0BE8BFACB3D8EFA93" box="[916,956,1303,1329]" name="Iceland" pageId="7" pageNumber="132">Isla</collectingCountry>
Saboga, small rocky-sandy island in front of Saboga village,
<geoCoordinate id="E5E75065FFD6FFF0BC75FAE03FC9FAF4" box="[362,507,1340,1366]" degrees="8" direction="north" minutes="37" orientation="latitude" pageId="7" pageNumber="132" precision="1" seconds="42.4" value="8.628445">8°37’42.4”N</geoCoordinate>
<geoCoordinate id="E5E75065FFD6FFF0BF1DFAE03C94FAF4" box="[514,678,1340,1366]" degrees="79" direction="west" minutes="03" orientation="longitude" pageId="7" pageNumber="132" precision="1" seconds="24.4" value="-79.05678">79°03’24.4”W</geoCoordinate>
, intertidal, leg. P.P.G. Pachelle &amp; M. Leray,
<date id="F46D1062FFD6FFF0B987FAE03B24FAF4" box="[1176,1302,1340,1366]" pageId="7" pageNumber="132" value="2019-03-23">23.03.2019</date>
[fcn PP 19- 077]; 1 ovigerous specimen (cl
<quantity id="472B9B47FFD6FFF0BCEAFA833C7DFAD8" box="[501,591,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="132" unit="mm" value="5.6">5.6 mm</quantity>
),
<collectionCode id="E6C2AE67FFD6FFF0BF7EFABC3CF1FAD8" box="[609,707,1376,1402]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="7" pageNumber="132" type="Museum">FLMNH</collectionCode>
<collectionCode id="E6C2AE67FFD6FFF0BFD6FABC3CDFFAD8" box="[713,749,1376,1402]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="7" pageNumber="132" type="Museum">UF</collectionCode>
51903, same collection data as for previous specimen [fcn PP 19-078]; 1 ovigerous specimen (cl
<quantity id="472B9B47FFD6FFF0BF07FA5F3C42FA3C" box="[536,624,1411,1438]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="132" unit="mm" value="7.6">7.6 mm</quantity>
),
<collectionCode id="E6C2AE67FFD6FFF0BF9DFA583CD6FA3C" box="[642,740,1412,1438]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="7" pageNumber="132" type="Museum">FLMNH</collectionCode>
<collectionCode id="E6C2AE67FFD6FFF0BFF5FA583D3CFA3C" box="[746,782,1412,1438]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="7" pageNumber="132" type="Museum">UF</collectionCode>
51904, Las Perlas Archipelago,
<collectingCountry id="F8C47632FFD6FFF0B96CFA5F3AA9FA3F" box="[1139,1179,1411,1437]" name="Iceland" pageId="7" pageNumber="132">Isla</collectingCountry>
Bartolomé, rocky reef,
<geoCoordinate id="E5E75065FFD6FFF0BD88FA7B3F1AFA60" box="[151,296,1447,1474]" degrees="8" direction="north" minutes="38" orientation="latitude" pageId="7" pageNumber="132" precision="1" seconds="51.8" value="8.647722">8°38’51.8”N</geoCoordinate>
<geoCoordinate id="E5E75065FFD6FFF0BC31FA743FE1FA60" box="[302,467,1448,1474]" degrees="79" direction="west" minutes="02" orientation="longitude" pageId="7" pageNumber="132" precision="1" seconds="17.3" value="-79.03814">79°02’17.3”W</geoCoordinate>
, intertidal, leg. P.P.G. Pachelle &amp; M. Leray,
<date id="F46D1062FFD6FFF0BEA0FA743A0FFA60" box="[959,1085,1448,1474]" pageId="7" pageNumber="132" value="2019-03-24">24.03.2019</date>
[fcn PP 19-080]; 1 non-ovigerous specimen (cl
<quantity id="472B9B47FFD6FFF0BC46FA103F80FA44" box="[345,434,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="132" unit="mm" value="3.4">3.4 mm</quantity>
),
<collectionCode id="E6C2AE67FFD6FFF0BCDAFA103C15FA44" box="[453,551,1484,1510]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="7" pageNumber="132" type="Museum">FLMNH</collectionCode>
<collectionCode id="E6C2AE67FFD6FFF0BF32FA103C63FA44" box="[557,593,1484,1510]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="7" pageNumber="132" type="Museum">UF</collectionCode>
51905,
<collectingCountry id="F8C47632FFD6FFF0BFB4FA103D35FA44" box="[683,775,1484,1510]" name="Panama" pageId="7" pageNumber="132">Panama</collectingCountry>
, Coiba Archipelago, Coiba, small rocky island off Bahía de Santa Cruz,
<geoCoordinate id="E5E75065FFD6FFF0BC3FFA333F82F9A8" box="[288,432,1519,1546]" degrees="7" direction="north" minutes="37" orientation="latitude" pageId="7" pageNumber="132" precision="1" seconds="53.4" value="7.6315002">7°37’53.4”N</geoCoordinate>
<geoCoordinate id="E5E75065FFD6FFF0BCA8FA333C6EF9AB" box="[439,604,1519,1545]" degrees="81" direction="west" minutes="46" orientation="longitude" pageId="7" pageNumber="132" precision="1" seconds="44.9" value="-81.779144">81°46’44.9”W</geoCoordinate>
, exposed intertidal, in rock crevices, leg. P.P.G. Pachelle
<emphasis id="B2A7EAB0FFD6FFF0B9C6FA2C3B23F9A8" box="[1241,1297,1520,1546]" italics="true" pageId="7" pageNumber="132">et al.</emphasis>
,
<date id="F46D1062FFD6FFF0B801FA2C3BAEF9A8" box="[1310,1436,1520,1546]" pageId="7" pageNumber="132" value="2019-02-20">20.02.2019</date>
[fcn PAN-156]; 1 ovigerous specimen (cl
<quantity id="472B9B47FFD6FFF0BF76F9CF3CF0F98C" box="[617,706,1555,1582]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="132" unit="mm" value="5.7">5.7 mm</quantity>
),
<collectionCode id="E6C2AE67FFD6FFF0BFCAF9C83D0DF98C" box="[725,831,1556,1582]" pageId="7" pageNumber="132">OUMNH</collectionCode>
.
<collectionCode id="E6C2AE67FFD6FFF0BE5DF9C83D5EF98C" box="[834,876,1556,1582]" country="Austria" name="Zoological Collection, University of Vienna" pageId="7" pageNumber="132" type="University or college">ZC</collectionCode>
. 2015-08-033,
<collectingCountry id="F8C47632FFD6FFF0B90AF9C83A43F98C" box="[1045,1137,1556,1582]" name="Panama" pageId="7" pageNumber="132">Panama</collectingCountry>
, Río Mar, in rock crevices, low tide, leg. M. Torchin,
<date id="F46D1062FFD6FFF0BCA1F9E43C09F9F0" box="[446,571,1592,1618]" pageId="7" pageNumber="132" value="2007-02-20">20.02.2007</date>
[fcn 07-083]; 1 ovigerous specimen (cl 5.0 mm),
<collectionCode id="E6C2AE67FFD6FFF0B970F9E43AF9F9F0" box="[1135,1227,1592,1618]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="132">MZUSP</collectionCode>
34072,
<collectingCountry id="F8C47632FFD6FFF0B838F9EB3BAAF9F3" box="[1319,1432,1591,1617]" name="Colombia" pageId="7" pageNumber="132">Colombia</collectingCountry>
, Bahía Málaga, Los Negritos,
<geoCoordinate id="E5E75065FFD6FFF0BCFEF9873C4DF9D4" box="[481,639,1627,1654]" degrees="03" direction="north" minutes="53" orientation="latitude" pageId="7" pageNumber="132" precision="1" seconds="47.6" value="3.8965554">03°53’47.6”N</geoCoordinate>
<geoCoordinate id="E5E75065FFD6FFF0BF99F9803D18F9D4" box="[646,810,1628,1654]" degrees="77" direction="west" minutes="24" orientation="longitude" pageId="7" pageNumber="132" precision="1" seconds="09.3" value="-77.40259">77°24’09.3”W</geoCoordinate>
, small rocky island with large boulders, some cemented with sponges, under large rocks on coarse sand, depth:
<quantity id="472B9B47FFD6FFF0BE1EF9A33D6DF938" box="[769,863,1663,1690]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="7" pageNumber="132" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">0–0.5 m</quantity>
, leg. A. Anker,
<date id="F46D1062FFD6FFF0B912F9A33AB1F93B" box="[1037,1155,1663,1689]" pageId="7" pageNumber="132" value="2009-04-27">27.iv.2009</date>
[fcn COL 00-244].
</paragraph>
<paragraph id="806C36A2FFD6FFF0BDD8F97F3C2BF940" blockId="7.[151,1437,1123,2014]" pageId="7" pageNumber="132">
<emphasis id="B2A7EAB0FFD6FFF0BDD8F97F3F63F91C" bold="true" box="[199,337,1699,1726]" pageId="7" pageNumber="132">Description</emphasis>
. See
<bibRefCitation id="E4424B53FFD6FFF0BC8BF97F3C70F91C" author="Coutiere, H." box="[404,578,1699,1726]" pageId="7" pageNumber="132" pagination="380 - 386" refId="ref8928" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière (1897</bibRefCitation>
,
<bibRefCitation id="E4424B53FFD6FFF0BF50F9783CBAF91C" author="Coutiere, H." box="[591,648,1700,1726]" pageId="7" pageNumber="132" pagination="1 - 559" refId="ref8963" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">1899</bibRefCitation>
) and
<bibRefCitation id="E4424B53FFD6FFF0BFD3F9783DDCF91C" author="Banner, D. M. &amp; Banner, A. H." box="[716,1006,1700,1726]" pageId="7" pageNumber="132" pagination="1 - 99" refId="ref8791" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner (1981)</bibRefCitation>
; for additional illustrations see
<bibRefCitation id="E4424B53FFD6FFF0B846F9783ED0F940" author="Bruce, A. J." pageId="7" pageNumber="132" pagination="329 - 343" refId="ref8831" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (Ed.), The marine fauna and flora of Hong Kong and southern China III. Proceedings of the Fourth International Marine Biological Workshop. The Marine Fauna and Flora of Hong Kong and southern China, Hong Kong University Press, Hong Kong, pp. 329 - 343." type="book chapter" year="1992">Bruce (1992)</bibRefCitation>
and
<bibRefCitation id="E4424B53FFD6FFF0BC07F91B3C27F940" author="Duris, Z. &amp; Horka, I." box="[280,533,1735,1762]" pageId="7" pageNumber="132" pagination="773 - 793" refId="ref9127" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="806C36A2FFD6FFF0BDD8F9373E87F888" blockId="7.[151,1437,1123,2014]" pageId="7" pageNumber="132">
<emphasis id="B2A7EAB0FFD6FFF0BDD8F9373F6AF8A4" bold="true" box="[199,344,1771,1798]" pageId="7" pageNumber="132">Colouration</emphasis>
. Uniformly bright yellow-orange (
<bibRefCitation id="E4424B53FFD6FFF0BFF7F9373DBDF8A4" author="Coutiere, H." box="[744,911,1771,1798]" pageId="7" pageNumber="132" pagination="1 - 559" refId="ref8963" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière 1899</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0BE81F9373ACCF8A4" author="Hickman, C. P. &amp; Zimmerman, T. L." box="[926,1278,1771,1798]" pageId="7" pageNumber="132" refId="ref9392" refString="Hickman, C. P. &amp; Zimmerman, T. L. (2000) A field guide to crustaceans of Galapagos. Galapagos marine life series. Sugar Spring Press, Lexington, 156 pp." type="book" year="2000">Hickman &amp; Zimmerman 2000</bibRefCitation>
; see also
<figureCitation id="18E82A27FFD6FFF0B86FF9373E95F888" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1743]" captionTargetBox="[151,1436,704,1698]" captionTargetId="figure@8.[151,1436,704,1698]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus serratidigitus (Coutière, 1897): ovigerous specimen (cl 6.1 mm) from Las Perlas, Panama, MZUSP 33810 [a]; non-ovigerous specimen (cl 4.9 mm, parasitised by a pair of bopyrid isopods) from Las Perlas, Panama, MNHN- IU-2014-20291 [b, c]; a—living individual, chelipeds folded under the body, lateral view; b—same, chelipeds folded under the body, dorsal view; c—same, major cheliped extended, lateral view." httpUri="https://zenodo.org/record/3359247/files/figure.png" pageId="7" pageNumber="132">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="806C36A2FFD6FFF0BDD8F8E83FE1F8EF" blockId="7.[151,1437,1123,2014]" box="[199,467,1843,1870]" pageId="7" pageNumber="132">
<materialsCitation id="30BB3CFFFFD6FFF0BDD8F8E83FFDF8EF" ID-GBIF-Occurrence="2302332017" box="[199,463,1843,1870]" country="Djibouti" location="Djibouti" pageId="7" pageNumber="132" specimenCount="1" typeStatus="holotype">
<emphasis id="B2A7EAB0FFD6FFF0BDD8F8E83F52F8EC" bold="true" box="[199,352,1843,1870]" pageId="7" pageNumber="132">
<typeStatus id="5F688800FFD6FFF0BDD8F8E83F30F8EC" box="[199,258,1844,1870]" pageId="7" pageNumber="132">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F8C47632FFD6FFF0BC71F8EF3FFDF8EF" box="[366,463,1843,1869]" name="Djibouti" pageId="7" pageNumber="132">Djibouti</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="806C36A2FFD6FFF0BDD8F88B3CF3F87C" blockId="7.[151,1437,1123,2014]" pageId="7" pageNumber="132">
<emphasis id="B2A7EAB0FFD6FFF0BDD8F88B3F6BF8D0" bold="true" box="[199,345,1879,1906]" pageId="7" pageNumber="132">Distribution</emphasis>
. Indo-Pacific, from the Red Sea and Gulf of Aden to
<collectingCountry id="F8C47632FFD6FFF0BEB8F88B3A02F8D3" box="[935,1072,1879,1905]" name="Madagascar" pageId="7" pageNumber="132">Madagascar</collectingCountry>
,
<collectingCountry id="F8C47632FFD6FFF0B924F8843A4CF8D0" box="[1083,1150,1880,1906]" name="Japan" pageId="7" pageNumber="132">Japan</collectingCountry>
,
<collectingCountry id="F8C47632FFD6FFF0B999F88B3AC2F8D3" box="[1158,1264,1879,1905]" name="Australia" pageId="7" pageNumber="132">Australia</collectingCountry>
,
<collectingCountry id="F8C47632FFD6FFF0B9E5F88B3B60F8D3" box="[1274,1362,1879,1905]" name="Mexico" pageId="7" pageNumber="132">Mexico</collectingCountry>
,
<collectingCountry id="F8C47632FFD6FFF0B843F8843E8EF834" name="Panama" pageId="7" pageNumber="132">Panama</collectingCountry>
,
<collectingCountry id="F8C47632FFD6FFF0BDD8F8A73F07F837" box="[199,309,1915,1941]" name="Colombia" pageId="7" pageNumber="132">Colombia</collectingCountry>
and Galapagos (
<bibRefCitation id="E4424B53FFD6FFF0BCF0F8A03CC4F834" author="Banner, D. M. &amp; Banner, A. H." box="[495,758,1916,1942]" pageId="7" pageNumber="132" pagination="1 - 99" refId="ref8791" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0BE1DF8A03DB4F834" author="Bruce, A. J." box="[770,902,1916,1942]" pageId="7" pageNumber="132" pagination="329 - 343" refId="ref8831" refString="Bruce, A. J. (1992) Additions to the marine caridean fauna of Hong Kong, with a description of a new species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (Ed.), The marine fauna and flora of Hong Kong and southern China III. Proceedings of the Fourth International Marine Biological Workshop. The Marine Fauna and Flora of Hong Kong and southern China, Hong Kong University Press, Hong Kong, pp. 329 - 343." type="book chapter" year="1992">Bruce 1992</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD6FFF0BE8EF8A73A49F837" author="Duris, Z. &amp; Horka, I." box="[913,1147,1915,1942]" pageId="7" pageNumber="132" pagination="773 - 793" refId="ref9127" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
; present study). The exact range of
<taxonomicName id="47D34D21FFD6FFF0BDE4F87C3F9DF818" authorityName="- Banner &amp; Banner" authorityYear="1981" box="[251,431,1952,1978]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD6FFF0BDE4F87C3F9DF818" box="[251,431,1952,1978]" italics="true" pageId="7" pageNumber="132">S. serratidigitus</emphasis>
</taxonomicName>
needs further studies as several records of the species were apparently based on misidentifications (
<bibRefCitation id="E4424B53FFD6FFF0BC09F81F3C30F87F" author="Duris, Z. &amp; Horka, I." box="[278,514,1987,2014]" pageId="7" pageNumber="132" pagination="773 - 793" refId="ref9127" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
; see also below).
</paragraph>
<paragraph id="806C36A2FFD9FFFFBDD8FF453D39FF75" blockId="8.[151,1437,152,683]" pageId="8" pageNumber="133">
<emphasis id="B2A7EAB0FFD9FFFFBDD8FF453F14FF11" bold="true" box="[199,294,153,179]" pageId="8" pageNumber="133">Ecology</emphasis>
. Intertidal and shallow subtidal (
<quantity id="472B9B47FFD9FFFFBFBFFF453CDFFF11" box="[672,749,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="8" pageNumber="133" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
); typically under rocks and coral rubble, often in rubble and rock crevices, coralline algae etc. (A. Anker, pers. obs.).
</paragraph>
<paragraph id="806C36A2FFD9FFFFBDD8FF3D3F83FD08" blockId="8.[151,1437,152,683]" pageId="8" pageNumber="133">
<emphasis id="B2A7EAB0FFD9FFFFBDD8FF3D3F06FF59" bold="true" box="[199,308,225,251]" pageId="8" pageNumber="133">Remarks</emphasis>
.
<taxonomicName id="47D34D21FFD9FFFFBC5BFF3C3C6FFF58" authorityName="- Banner &amp; Banner" authorityYear="1981" box="[324,605,224,250]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD9FFFFBC5BFF3C3C6FFF58" box="[324,605,224,250]" italics="true" pageId="8" pageNumber="133">Salmoneus serratidigitus</emphasis>
</taxonomicName>
is currently seen as a widespread Indo-Pacific species, ranging from the Red Sea to the tropical eastern Pacific, where it ranges from
<collectingCountry id="F8C47632FFD9FFFFBE2BFED83DB8FEBC" box="[820,906,260,286]" name="Mexico" pageId="8" pageNumber="133">Mexico</collectingCountry>
(Cabo San Lucas in
<collectingRegion id="4217F840FFD9FFFFB970FED83B7BFEBD" box="[1135,1353,260,287]" country="Mexico" name="Baja California Sur" pageId="8" pageNumber="133">Baja California Sur</collectingRegion>
, Teacapan in
<collectingRegion id="4217F840FFD9FFFFBDFCFEF43F08FEE0" box="[227,314,296,322]" country="Mexico" name="Nayarit" pageId="8" pageNumber="133">Nayarit</collectingRegion>
,
<collectingCountry id="F8C47632FFD9FFFFBC5AFEF43F5FFEE0" box="[325,365,296,322]" name="Iceland" pageId="8" pageNumber="133">Isla</collectingCountry>
Clarion) to
<collectingCountry id="F8C47632FFD9FFFFBCE9FEF43C56FEE0" box="[502,612,296,322]" name="Colombia" pageId="8" pageNumber="133">Colombia</collectingCountry>
and the Galapagos Archipelago (
<bibRefCitation id="E4424B53FFD9FFFFBEC7FEF53AD0FEE1" author="Banner, D. M. &amp; Banner, A. H." box="[984,1250,297,323]" pageId="8" pageNumber="133" pagination="1 - 99" refId="ref8791" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD9FFFFB9F2FEF43BA5FEE1" author="Wicksten, M. K." box="[1261,1431,296,323]" pageId="8" pageNumber="133" pagination="1 - 59" refId="ref9935" refString="Wicksten, M. K. (1983) A monograph on the shallow water caridean shrimps of the Gulf of California, Mexico. Alan Hancock Monographs in Marine Biology, 13, 1 - 59." type="journal article" year="1983">Wicksten 1983</bibRefCitation>
; Hernández &amp; Martínez 1992;
<bibRefCitation id="E4424B53FFD9FFFFBCEDFE903D72FEC5" author="Wicksten, M. K. &amp; Hendrickx, M. E." box="[498,832,332,359]" pageId="8" pageNumber="133" pagination="49 - 76" refId="ref10139" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (2003) An updated checklist of benthic marine and brackish water shrimps (Decapoda: Penaeoidea, Stenopodidea, Caridea) from the eastern tropical Pacific. In: Hendrickx, M. E. (Ed.), Contributions to the study of East Pacific crustaceans / Contribuciones al estudio de los crustaceos del Pacifico Este. Instituto de Ciencias del Mar y Limnologia, UNAM, Mexico City, pp. 49 - 76." type="book chapter" year="2003">Wicksten &amp; Hendrickx 2003</bibRefCitation>
;
<bibRefCitation id="E4424B53FFD9FFFFBE50FE903B12FEC5" author="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R." box="[847,1312,332,359]" pageId="8" pageNumber="133" pagination="221 - 239" refId="ref9512" refString="Lazarus-Agudelo, J. F. &amp; Cantera-Kintz, J. R. (2007) Crustaceos (Crustacea: Sessilia, Stomatopoda, Isopoda, Amphipoda, Decapoda) de Bahia Malaga, Valle del Cauca (Pacifico colombiano). Biota Colombiana, 8, 221 - 239." type="journal article" year="2007">Lazarus-Agudelo &amp; Cantera-Kintz 2007</bibRefCitation>
). The only previous published record of
<taxonomicName id="47D34D21FFD9FFFFBCFCFEAD3CA5FE28" authorityName="- Banner &amp; Banner" authorityYear="1981" box="[483,663,368,395]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD9FFFFBCFCFEAD3CA5FE28" box="[483,663,368,395]" italics="true" pageId="8" pageNumber="133">S. serratidigitus</emphasis>
</taxonomicName>
from
<collectingCountry id="F8C47632FFD9FFFFBFC2FEAC3D79FE28" box="[733,843,368,394]" name="Colombia" pageId="8" pageNumber="133">Colombia</collectingCountry>
was a listing of the species in a table of crustaceans of Bahía Málaga, based on material listed in unpublished dissertations or PhD theses (Lazarus-Agudelo &amp; Cantera- Kintz 2007 and references therein). Thus, the present specimen from Bahía Málaga confirms the occurrence of this species in
<collectingCountry id="F8C47632FFD9FFFFBC0FFE003FB0FE54" box="[272,386,476,502]" name="Colombia" pageId="8" pageNumber="133">Colombia</collectingCountry>
. In addition, the material from Las Perlas, Coiba and Río Mar represents the first record of
<taxonomicName id="47D34D21FFD9FFFFBD88FDDD3F79FDB8" authorityName="- Banner &amp; Banner" authorityYear="1981" box="[151,331,512,539]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD9FFFFBD88FDDD3F79FDB8" box="[151,331,512,539]" italics="true" pageId="8" pageNumber="133">S. serratidigitus</emphasis>
</taxonomicName>
from the Pacific coast of
<collectingCountry id="F8C47632FFD9FFFFBF6AFDDD3CE3FDB9" box="[629,721,513,539]" name="Panama" pageId="8" pageNumber="133">Panama</collectingCountry>
.
<bibRefCitation id="E4424B53FFD9FFFFBFC2FDDC3A75FDB9" author="Hickman, C. P. &amp; Zimmerman, T. L." box="[733,1095,512,539]" pageId="8" pageNumber="133" refId="ref9392" refString="Hickman, C. P. &amp; Zimmerman, T. L. (2000) A field guide to crustaceans of Galapagos. Galapagos marine life series. Sugar Spring Press, Lexington, 156 pp." type="book" year="2000">Hickman &amp; Zimmerman (2000)</bibRefCitation>
published an excellent colour photograph of
<taxonomicName id="47D34D21FFD9FFFFBC22FDF93FC3FD9C" authorityName="- Banner &amp; Banner" authorityYear="1981" box="[317,497,548,575]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD9FFFFBC22FDF93FC3FD9C" box="[317,497,548,575]" italics="true" pageId="8" pageNumber="133">S. serratidigitus</emphasis>
</taxonomicName>
from Galapagos, which corresponds well with the description of the colouration in
<bibRefCitation id="E4424B53FFD9FFFFBD88FD943F78FDC1" author="Coutiere, H." box="[151,330,584,611]" pageId="8" pageNumber="133" pagination="1 - 559" refId="ref8963" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière (1899)</bibRefCitation>
, as well as with the colour of the Panamanian specimens (
<figureCitation id="18E82A27FFD9FFFFBED5FD943A3FFDC1" box="[970,1037,584,611]" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1743]" captionTargetBox="[151,1436,704,1698]" captionTargetId="figure@8.[151,1436,704,1698]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus serratidigitus (Coutière, 1897): ovigerous specimen (cl 6.1 mm) from Las Perlas, Panama, MZUSP 33810 [a]; non-ovigerous specimen (cl 4.9 mm, parasitised by a pair of bopyrid isopods) from Las Perlas, Panama, MNHN- IU-2014-20291 [b, c]; a—living individual, chelipeds folded under the body, lateral view; b—same, chelipeds folded under the body, dorsal view; c—same, major cheliped extended, lateral view." httpUri="https://zenodo.org/record/3359247/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
). Despite the overall morphological homogeneity of
<taxonomicName id="47D34D21FFD9FFFFBC50FDB13C30FD24" authorityName="- Banner &amp; Banner" authorityYear="1981" box="[335,514,620,647]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD9FFFFBC50FDB13C30FD24" box="[335,514,620,647]" italics="true" pageId="8" pageNumber="133">S. serratidigitus</emphasis>
</taxonomicName>
, a phylogeographic analysis of this taxon across its wide Indo-Pacific distribution range is highly desirable.
</paragraph>
<caption id="D4AC662AFFD9FFFFBD88F96A3D0DF899" httpUri="https://zenodo.org/record/3359247/files/figure.png" pageId="8" pageNumber="133" startId="8.[151,250,1718,1743]" targetBox="[151,1436,704,1698]" targetPageId="8">
<paragraph id="806C36A2FFD9FFFFBD88F96A3D0DF899" blockId="8.[151,1437,1718,1851]" pageId="8" pageNumber="133">
<emphasis id="B2A7EAB0FFD9FFFFBD88F96A3F3DF96C" bold="true" box="[151,271,1718,1743]" pageId="8" pageNumber="133">FIGURE 5</emphasis>
.
<taxonomicName id="47D34D21FFD9FFFFBC01F96A3CE1F96C" authority="(Coutiere, 1897)" authorityName="Coutiere" authorityYear="1897" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[286,723,1718,1743]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD9FFFFBC01F96A3C2FF96C" box="[286,541,1718,1742]" italics="true" pageId="8" pageNumber="133">Salmoneus serratidigitus</emphasis>
(
<bibRefCitation id="E4424B53FFD9FFFFBF33F96A3CF8F96C" author="Coutiere, H." box="[556,714,1718,1743]" pageId="8" pageNumber="133" pagination="380 - 386" refId="ref8928" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
)
</taxonomicName>
: ovigerous specimen (cl 6.1 mm) from Las Perlas, Panama, MZUSP 33810 [a]; non-ovigerous specimen (cl 4.9 mm, parasitised by a pair of bopyrid isopods) from Las Perlas, Panama, MNHN- IU-2014-20291 [b, c]; a—living individual, chelipeds folded under the body, lateral view; b—same, chelipeds folded under the body, dorsal view; c—same, major cheliped extended, lateral view.
</paragraph>
</caption>
<paragraph id="806C36A2FFD9FFFEBDD8F8513AA5FEE1" blockId="8.[151,1437,1932,2031]" lastBlockId="9.[151,1437,153,647]" lastPageId="9" lastPageNumber="134" pageId="8" pageNumber="133">
Noteworthy, two ovigerous specimens from Las Perlas (
<collectionCode id="E6C2AE67FFD9FFFFBE26F8513DAFF805" box="[825,925,1933,1959]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="133" type="Museum">FLMNH</collectionCode>
<collectionCode id="E6C2AE67FFD9FFFFBEBCF8513DF5F805" box="[931,967,1933,1959]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="133" type="Museum">UF</collectionCode>
51903 and 51904) seem to be much more setose than the others. In these individuals, the carapace and some selected areas of the pleon are covered with numerous, very conspicuous, erect setae, approaching the condition observed in the western-central Atlantic
<taxonomicName id="47D34D21FFD9FFFEB82FF8093F80FF11" authority="Manning &amp; Chace, 1990" authorityName="Manning &amp; Chace" authorityYear="1990" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" lastPageId="9" lastPageNumber="134" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B2A7EAB0FFD9FFFFB82FF8093BAEF84C" box="[1328,1436,2005,2031]" italics="true" pageId="8" pageNumber="133">S. setosus</emphasis>
<bibRefCitation id="E4424B53FFD8FFFEBD88FF453F80FF11" author="Manning, R. B. &amp; Chace, F. A. Jr." box="[151,434,153,179]" pageId="9" pageNumber="134" pagination="1 - 91" refId="ref9563" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, 1 - 91. https: // doi. org / 10.5479 / si. 00810282.503" type="journal article" year="1990">Manning &amp; Chace, 1990</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E4424B53FFD8FFFEBCDCFF453CEFFF11" author="Manning, R. B. &amp; Chace, F. A. Jr." box="[451,733,153,179]" pageId="9" pageNumber="134" pagination="1 - 91" refId="ref9563" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, 1 - 91. https: // doi. org / 10.5479 / si. 00810282.503" type="journal article" year="1990">Manning &amp; Chace 1990</bibRefCitation>
). Since the hirsuteness of the carapace appears to be variable in
<taxonomicName id="47D34D21FFD8FFFEBDABFF613F5AFF75" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[180,360,189,215]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD8FFFEBDABFF613F5AFF75" box="[180,360,189,215]" italics="true" pageId="9" pageNumber="134">S. serratidigitus</emphasis>
</taxonomicName>
, it may also be found variable in the possibly related
<taxonomicName id="47D34D21FFD8FFFEBEDBFF613A03FF75" authorityName="Manning &amp; Chace" authorityYear="1990" box="[964,1073,189,215]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B2A7EAB0FFD8FFFEBEDBFF613A03FF75" box="[964,1073,189,215]" italics="true" pageId="9" pageNumber="134">S. setosus</emphasis>
</taxonomicName>
. This in turn would result in the questioning of the validity of
<taxonomicName id="47D34D21FFD8FFFEBCC0FF3D3CFBFF59" authority="Anker, 2007" authorityName="Anker" authorityYear="2007" box="[479,713,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="rocas">
<emphasis id="B2A7EAB0FFD8FFFEBCC0FF3D3C05FF59" box="[479,567,225,251]" italics="true" pageId="9" pageNumber="134">S. rocas</emphasis>
<bibRefCitation id="E4424B53FFD8FFFEBF23FF3D3CFBFF59" author="Anker, A." box="[572,713,225,251]" pageId="9" pageNumber="134" pagination="21 - 39" refId="ref8513" refString="Anker, A. (2007) New species and records of alpheid shrimps, genera Salmoneus Holthuis and Parabetaeus Coutiere, from the tropical western Atlantic (Decapoda, Caridea). Zootaxa, 1653 (1), 21 - 39. https: // doi. org / 10.11646 / zootaxa. 1653.1.2" type="journal article" year="2007">Anker, 2007</bibRefCitation>
</taxonomicName>
, presently only known from the
<typeStatus id="5F688800FFD8FFFEB930FF3D3AA3FF59" box="[1071,1169,225,251]" pageId="9" pageNumber="134" type="holotype">holotype</typeStatus>
collected at the isolated Atol das Rocas, off northeastern
<collectingCountry id="F8C47632FFD8FFFEBF1DFED93C74FEBD" box="[514,582,261,287]" name="Brazil" pageId="9" pageNumber="134">Brazil</collectingCountry>
(
<bibRefCitation id="E4424B53FFD8FFFEBF4BFED93CECFEBD" author="Anker, A." box="[596,734,261,287]" pageId="9" pageNumber="134" pagination="21 - 39" refId="ref8513" refString="Anker, A. (2007) New species and records of alpheid shrimps, genera Salmoneus Holthuis and Parabetaeus Coutiere, from the tropical western Atlantic (Decapoda, Caridea). Zootaxa, 1653 (1), 21 - 39. https: // doi. org / 10.11646 / zootaxa. 1653.1.2" type="journal article" year="2007">Anker 2007</bibRefCitation>
), since the two species are separated mainly by the presence (in
<taxonomicName id="47D34D21FFD8FFFEBD88FEF53F36FEE1" authorityName="Manning &amp; Chace" authorityYear="1990" box="[151,260,297,323]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B2A7EAB0FFD8FFFEBD88FEF53F36FEE1" box="[151,260,297,323]" italics="true" pageId="9" pageNumber="134">S. setosus</emphasis>
</taxonomicName>
) and absence (in
<taxonomicName id="47D34D21FFD8FFFEBCD5FEF53C16FEE1" authorityName="Anker" authorityYear="2007" box="[458,548,297,323]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="rocas">
S.
<emphasis id="B2A7EAB0FFD8FFFEBCF8FEF63C16FEE1" box="[487,548,298,323]" italics="true" pageId="9" pageNumber="134">rocas</emphasis>
</taxonomicName>
) of the thickened erect setae on the carapace and pleon.
</paragraph>
</subSubSection>
<subSubSection id="C8C96529FFD8FFFEBDD8FE913C49FD25" pageId="9" pageNumber="134" type="reference_group">
<paragraph id="806C36A2FFD8FFFEBDD8FE913C49FD25" blockId="9.[151,1437,153,647]" pageId="9" pageNumber="134">
<taxonomicName id="47D34D21FFD8FFFEBDD8FE913FEDFEC5" ID-CoL="85Y2S" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[199,479,333,359]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD8FFFEBDD8FE913FEDFEC5" box="[199,479,333,359]" italics="true" pageId="9" pageNumber="134">Salmoneus serratidigitus</emphasis>
</taxonomicName>
is a small (cl &lt;
<quantity id="472B9B47FFD8FFFEBF8AFE913CEAFEC5" box="[661,728,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="134" unit="mm" value="7.0">7 mm</quantity>
) intertidal and shallow subtidal alpheid shrimp, often dwelling under rocks and coral rubble, or in various crevices (
<bibRefCitation id="E4424B53FFD8FFFEBFFFFEAD3DB6FE29" author="Coutiere, H." box="[736,900,369,395]" pageId="9" pageNumber="134" pagination="1 - 559" refId="ref8963" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière 1899</bibRefCitation>
; present study). This is why
<bibRefCitation id="E4424B53FFD8FFFEB9DDFEAD3BAEFE29" author="Hendrickx, M. E." box="[1218,1436,369,395]" pageId="9" pageNumber="134" pagination="395 - 397" refId="ref9299" refString="Hendrickx, M. E. (2001) New distribution, size and habitats records of decapod crustaceans from the eastern tropical Pacific. Revista de Biologia Tropical, 49, 395 - 397." type="journal article" year="2001">Hendrickx’s (2001)</bibRefCitation>
record of a single, unusually large specimen (cl
<quantity id="472B9B47FFD8FFFEBFDCFE493D27FE0D" box="[707,789,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="134" unit="mm" value="17.0">17 mm</quantity>
) of
<taxonomicName id="47D34D21FFD8FFFEBE58FE493DCFFE0D" ID-CoL="85Y2S" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[839,1021,405,431]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B2A7EAB0FFD8FFFEBE58FE493DCFFE0D" box="[839,1021,405,431]" italics="true" pageId="9" pageNumber="134">S. serratidigitus</emphasis>
</taxonomicName>
from a “very low dissolved oxygen concentration (0.44 ml/l)” habitat and depth of
<quantity id="472B9B47FFD8FFFEBFA0FE653D14FE71" box="[703,806,441,467]" metricMagnitude="1" metricUnit="m" metricValue="6.15" metricValueMax="6.2" metricValueMin="6.1" pageId="9" pageNumber="134" unit="m" value="61.5" valueMax="62.0" valueMin="61.0">61–62 m</quantity>
off Teacapan,
<collectingRegion id="4217F840FFD8FFFEBECBFE653A19FE71" box="[980,1067,441,467]" country="Mexico" name="Sinaloa" pageId="9" pageNumber="134">Sinaloa</collectingRegion>
,
<collectingCountry id="F8C47632FFD8FFFEB925FE653AA0FE71" box="[1082,1170,441,467]" name="Mexico" pageId="9" pageNumber="134">Mexico</collectingCountry>
, appeared to be highly suspicious. Therefore, this specimen, deposited in the collection of the Instituto de Ciencias del Mar y Limnologia, Unidad Académica Mazatlán (formerly Estación Mazatlán / UNAM), was re-examined and photographed by Dr. Michel E. Hendrickx upon the author’s request. The photographs sent by Dr. Hendrickx clearly show a specimen of the genus
<taxonomicName id="47D34D21FFD8FFFEBC15FD953C2FFDC1" ID-CoL="84LB5" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[266,541,585,611]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B2A7EAB0FFD8FFFEBC15FD953F56FDC1" box="[266,356,585,611]" italics="true" pageId="9" pageNumber="134">Alpheus</emphasis>
Fabricius, 1798
</taxonomicName>
, with high probability the recently described
<taxonomicName id="47D34D21FFD8FFFEB904FD953BAAFDC1" ID-CoL="85LKY" authority="Ramos-Tafur, 2018" authorityName="Ramos-Tafur" authorityYear="2018" box="[1051,1432,585,611]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="luiszapatai">
<emphasis id="B2A7EAB0FFD8FFFEB904FD953A87FDC1" box="[1051,1205,585,611]" italics="true" pageId="9" pageNumber="134">A. luiszapatai</emphasis>
<bibRefCitation id="E4424B53FFD8FFFEB9A2FD953BAAFDC1" author="Ramos-Tafur, G. E." box="[1213,1432,585,611]" pageId="9" pageNumber="134" pagination="540 - 556" refId="ref9661" refString="Ramos-Tafur, G. E. (2018) Alpheus luiszapatai, a new species of rare and colorful deep water alpheid shrimp (Crustacea: Decapoda: Alpheidae) from Arusi, Choco Department, Pacific Coast of Colombia. Zootaxa, 4403 (3), 540 - 556. https: // doi. org / 10.11646 / zootaxa. 4403.3.7" type="journal article" year="2018">Ramos-Tafur, 2018</bibRefCitation>
</taxonomicName>
, previously unknown from Mexican waters.
</paragraph>
</subSubSection>
</treatment>
</document>
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