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<document id="BC6DF2E4745919521220CE6838D7BA52" ID-CLB-Dataset="274285" ID-DOI="10.5852/ejt.2023.893.2285" ID-GBIF-Dataset="e441bc9d-bb44-4ee6-b29f-a6724b96592b" ID-ISSN="2118-9773" ID-Zenodo-Dep="8385630" ID-ZooBank="D9DA4879-BC1D-4947-96E8-7D574DE334BA" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1695891225180" checkinUser="plazi" docAuthor="Leng, Zhaoqi, Tran, Anh Duc &amp; Ye, Zhen" docDate="2023" docId="045E878DFFE729053A01F8B6756D0A41" docLanguage="en" docName="EJT.2023.893.1-73.pdf" docOrigin="European Journal of Taxonomy 893 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2285/9863" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhyacobates svenhedini" docType="treatment" docVersion="9" lastPageNumber="63" masterDocId="F867FFF5FFDC293B381FFFB270530F1A" masterDocTitle="Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="60" updateTime="1699690649759" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="687EE40269744E4B76D25B19EFBCA065">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:namePart id="EEAA7C01794B52FEBE045D363AA2A9AC">Leng, Zhaoqi</mods:namePart>
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<heading id="D70081F7FFE729003A01F8B674460804" box="[542,1045,1796,1823]" centered="true" fontSize="11" level="2" pageId="59" pageNumber="60" reason="2">
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<emphasis id="BE83EA89FFE729003A01F8B673130805" bold="true" box="[542,832,1796,1823]" italics="true" pageId="59" pageNumber="60">Rhyacobates svenhedini</emphasis>
(
<bibRefCitation id="E8664B6AFFE729003B4FF8B6745E0804" author="Lundblad" box="[848,1037,1796,1822]" firstAuthor="Lundblad" pageId="59" pageNumber="60" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934</bibRefCitation>
)
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<subSubSection id="C4ED6510FFE729003A52F89573B50858" box="[589,998,1831,1858]" pageId="59" pageNumber="60" type="description">
<paragraph id="8C48369BFFE729003A52F89573B50858" blockId="59.[542,1045,1796,1858]" box="[589,998,1831,1858]" pageId="59" pageNumber="60">
<figureCitation id="14CC2A1EFFE729003A52F89572F90858" box="[589,682,1831,1858]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[336,1266,297,1835]" captionTargetId="figure-14@4.[255,1333,265,1851]" captionTargetPageId="4" captionText="Fig. 2. Habitus of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise. A. R. abdominalisAndersen &amp; Chen, 1995.B. R. chinensis Hungerford &amp; Matsuda, 1959.C. R. lundbladi (Hungerford, 1957). D. R. scorpio Andersen &amp; Chen, 1995. E. R. svenhedini (Lundblad, 1934). F. R. takahashii Esaki, 1923, dealated macropterous female. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385634" httpUri="https://zenodo.org/record/8385634/files/figure.png" pageId="59" pageNumber="60">Figs 2E</figureCitation>
, 
<figureCitation id="14CC2A1EFFE729003AA9F89A72870858" box="[694,724,1832,1858]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="59" pageNumber="60">3P</figureCitation>
, 
<figureCitation id="14CC2A1EFFE729003AFEF89A73630858" box="[737,816,1832,1858]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. gongvo Tran &amp; Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen &amp; Chen, 1995. K. R. recurvus Andersen &amp; Chen, 1995. L. R. scorpio Andersen &amp; Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="59" pageNumber="60">4M–N</figureCitation>
, 
<figureCitation id="14CC2A1EFFE729003B23F89573310858" box="[828,866,1831,1858]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="59" pageNumber="60">5O</figureCitation>
, 
<figureCitation id="14CC2A1EFFE729003B71F89A73DE0858" box="[878,909,1832,1858]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="59" pageNumber="60">7F</figureCitation>
, 
<figureCitation id="14CC2A1EFFE729003B85F89A73EE0858" box="[922,957,1832,1858]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen &amp; Chen, 1995. J. R. recurvus Andersen &amp; Chen, 1995. K. R. scorpio Andersen &amp; Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="59" pageNumber="60">8L</figureCitation>
, 
<figureCitation id="14CC2A1EFFE729003BD6F89A73B50858" box="[969,998,1832,1858]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="59" pageNumber="60">34</figureCitation>
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<treatmentCitation id="0D56108AFFE7290038A2F8DB72B7089E" author="Lundblad O. M." box="[189,740,1897,1924]" page="25" pageId="59" pageNumber="60" year="1934">
<taxonomicName id="4BF74D18FFE7290038A2F8DB72B7089E" ID-CoL="6GTHT" authority="Lundblad, 1934: 23 - 25" authorityName="Lundblad" authorityPageNumber="23 - 25" authorityYear="1934" box="[189,740,1897,1924]" class="Insecta" family="Gerridae" genus="Esakobates" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFE7290038A2F8DB71940899" box="[189,455,1897,1923]" italics="true" pageId="59" pageNumber="60">Esakobates svenhedini</emphasis>
<bibRefCitation id="E8664B6AFFE7290039D0F8DB72B7089E" author="Lundblad" box="[463,740,1897,1924]" firstAuthor="Lundblad" page="23 - 25" pageId="59" pageNumber="60" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934: 23–25</bibRefCitation>
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, pl. 2 fig. 10 (original description).
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<emphasis id="BE83EA89FFE7290038A2F818718D08DE" box="[189,478,1962,1988]" italics="true" pageId="59" pageNumber="60">Rhyacobates svenhedini</emphasis>
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– 
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<bibRefCitation id="E8664B6AFFE729003A14F818738908DE" author="Hungerford &amp; Matsuda" box="[523,986,1962,1989]" firstAuthor="Hungerford" page="69 - 72" pageId="59" pageNumber="60" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda 1959: 69–72</bibRefCitation>
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. — 
<treatmentCitation id="0D56108AFFE729003C07F818751408DE" author="Matsuda R." box="[1048,1351,1962,1988]" page="273" pageId="59" pageNumber="60" year="1960">
<bibRefCitation id="E8664B6AFFE729003C07F818751408DE" author="Matsuda" box="[1048,1351,1962,1988]" firstAuthor="Matsuda" page="273 - 276" pageId="59" pageNumber="60" pagination="25 - 632" refId="ref33267" refString="Matsuda R. 1960. Morphology, evolution and a classification of the Gerridae (Hemiptera-Heteroptera). The University of Kansas Science Bulletin 41: 25 - 632. https: // doi. org / 10.5962 / bhl. part. 15602" type="journal article" year="1960">Matsuda 1960: 273–276</bibRefCitation>
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. — 
<treatmentCitation id="0D56108AFFE7290038F5F87F726308FD" author="Andersen N. M. &amp; Chen P. P." box="[234,560,1997,2024]" page="61" pageId="59" pageNumber="60" year="1995">
<bibRefCitation id="E8664B6AFFE7290038F5F87F726308FD" author="Andersen &amp; Chen" box="[234,560,1997,2024]" firstAuthor="Andersen" page="61" pageId="59" pageNumber="60" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen 1995: 61</bibRefCitation>
</treatmentCitation>
, figs 24–25 (with remarks).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFE0290738A2FEBA75130DAB" pageId="60" pageNumber="61" type="diagnosis">
<paragraph id="8C48369BFFE0290738A2FEBA71650E38" blockId="60.[189,310,264,290]" box="[189,310,264,290]" pageId="60" pageNumber="61">
<heading id="D70081F7FFE0290738A2FEBA71650E38" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="BE83EA89FFE0290738A2FEBA71650E38" bold="true" box="[189,310,264,290]" pageId="60" pageNumber="61">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFE0290738A2FE8575130DAB" blockId="60.[187,1400,311,690]" pageId="60" pageNumber="61">
Body length of apterous females 10.30–11.59, of apterous males 7.21–7.90. Female: pronotum mainly black with a median brownish-yellow spot, mesonotum with a median brownish-yellow stripe, metanotum medially with a subtriangular yellow marking and with a sharp apex angle (
<figureCitation id="14CC2A1EFFE029073CFBFECF75160E82" box="[1252,1349,381,408]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[336,1266,297,1835]" captionTargetId="figure-14@4.[255,1333,265,1851]" captionTargetPageId="4" captionText="Fig. 2. Habitus of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise. A. R. abdominalisAndersen &amp; Chen, 1995.B. R. chinensis Hungerford &amp; Matsuda, 1959.C. R. lundbladi (Hungerford, 1957). D. R. scorpio Andersen &amp; Chen, 1995. E. R. svenhedini (Lundblad, 1934). F. R. takahashii Esaki, 1923, dealated macropterous female. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385634" httpUri="https://zenodo.org/record/8385634/files/figure.png" pageId="60" pageNumber="61">Figs 2E</figureCitation>
, 
<figureCitation id="14CC2A1EFFE029073D4BFECC75210E82" box="[1364,1394,382,408]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="60" pageNumber="61">3P</figureCitation>
, 
<figureCitation id="14CC2A1EFFE0290738A2FE1370A70EA1" box="[189,244,417,443]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">34A</figureCitation>
); posterior margin of abdominal mediotergite VII with a short median process; posterior margin of abdominal segment VII with four distinct processes dorsally, including two lobe-like processes on each connexival corner (
<figureCitation id="14CC2A1EFFE0290739C3FE5572390D1B" box="[476,618,487,513]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34D–F</figureCitation>
): one long, posteriorly directed outer process and a smaller, shorter, pointed inner process (in some individuals, the inner process is very small or indistinct); posterior margin of sternum VII with a small angular median process (
<figureCitation id="14CC2A1EFFE029073B3AFD9F73DC0D52" box="[805,911,557,584]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34E</figureCitation>
). Male: middle trochanter without spines; middle femur with scattered small spines, not arranged in distinct row (
<figureCitation id="14CC2A1EFFE029073C1FFDE374370D71" box="[1024,1124,593,619]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34J</figureCitation>
); length of middle tibia ca 1.3 times length of hind tibia; proctiger relatively large, with angular lobes laterally (
<figureCitation id="14CC2A1EFFE029073CD5FDC675760D94" box="[1226,1317,628,654]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="60" pageNumber="61">Figs 7F</figureCitation>
, 
<figureCitation id="14CC2A1EFFE029073D2DFDC6753A0D94" box="[1330,1385,628,654]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">34K</figureCitation>
); paramere strongly curved at basal third, distal part slender, with hook-shaped apex (
<figureCitation id="14CC2A1EFFE029073C8AFD2574A00DAB" box="[1173,1267,663,689]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen &amp; Chen, 1995. J. R. recurvus Andersen &amp; Chen, 1995. K. R. scorpio Andersen &amp; Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="60" pageNumber="61">Figs 8L</figureCitation>
, 
<figureCitation id="14CC2A1EFFE029073CE0FD2575600DAB" box="[1279,1331,663,689]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">34L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFE0290738A2FD6A73DB0CB4" pageId="60" pageNumber="61" type="materials_examined">
<paragraph id="8C48369BFFE0290738A2FD6A71F90DE8" blockId="60.[189,426,728,754]" box="[189,426,728,754]" pageId="60" pageNumber="61">
<heading id="D70081F7FFE0290738A2FD6A71F90DE8" bold="true" box="[189,426,728,754]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="BE83EA89FFE0290738A2FD6A71F90DE8" bold="true" box="[189,426,728,754]" pageId="60" pageNumber="61">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFE0290738F5FCB571B70C3B" blockId="60.[189,1398,775,942]" box="[234,484,775,801]" pageId="60" pageNumber="61">
<emphasis id="BE83EA89FFE0290738F5FCB571B70C3B" bold="true" box="[234,484,775,801]" pageId="60" pageNumber="61">Non-type specimens</emphasis>
</paragraph>
<paragraph id="8C48369BFFE0290738A2FC9873DB0CB4" blockId="60.[189,1398,775,942]" pageId="60" pageNumber="61">
<materialsCitation id="3C9F3CC6FFE0290738A2FC98754D0C72" ID-GBIF-Occurrence="4429869378" collectingDate="2016-07-16" collectionCode="NKUM" collectorName="Chen-guang Zheng" country="China" county="Qing-chuan County" elevation="1077" latitude="32.52364" location="Guang-yuan City" longLatPrecision="1" longitude="104.82325" pageId="60" pageNumber="61" specimenCount="52" specimenCount-female="52" stateProvince="Sichuan">
<collectingCountry id="F4E0760BFFE0290738A2FC98714B0C5E" box="[189,280,810,836]" name="China" pageId="60" pageNumber="61">CHINA</collectingCountry>
– 
<emphasis id="BE83EA89FFE029073932FC9872560C5E" bold="true" box="[301,517,810,836]" pageId="60" pageNumber="61">
<collectingRegion id="4E33F879FFE029073932FC9872560C5E" box="[301,517,810,836]" country="China" name="Sichuan" pageId="60" pageNumber="61">Sichuan Province</collectingRegion>
</emphasis>
• 31 ♁♁, 
<specimenCount id="9AF1FD12FFE029073A6AFC9872900C5F" box="[629,707,810,837]" pageId="60" pageNumber="61" type="female">52 ♀♀</specimenCount>
(apterous); 
<location id="89286040FFE029073B54FC9874470C5E" LSID="urn:lsid:plazi:treatment:045E878DFFE729053A01F8B6756D0A41:89286040FFE029073B54FC9874470C5E" box="[843,1044,810,836]" country="China" county="Qing-chuan County" latitude="32.52364" longLatPrecision="1" longitude="104.82325" name="Guang-yuan City" pageId="60" pageNumber="61" stateProvince="Sichuan">Guang-yuan City</location>
, 
<collectingCounty id="65294E17FFE029073C02FC9875570C5E" box="[1053,1284,810,836]" pageId="60" pageNumber="61">Qing-chuan County</collectingCounty>
, Tang-jia-he; 
<geoCoordinate id="E9C3505CFFE0290738F8FCFE71C00C72" box="[231,403,844,872]" degrees="32" direction="north" minutes="31" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="25.1" value="32.52364">32°31′25.1″ N</geoCoordinate>
, 
<geoCoordinate id="E9C3505CFFE029073981FCFE72070C72" box="[414,596,844,872]" degrees="104" direction="east" minutes="49" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="23.7" value="104.82325">104°49′23.7″ E</geoCoordinate>
; 
<elevation id="07DAD1A8FFE029073A7FFCFC72AB0C7D" box="[608,760,845,872]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="60" pageNumber="61" unit="m" value="1077.0">
<quantity id="4B0F9B7EFFE029073A7FFCFC72EA0C72" box="[608,697,846,872]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="60" pageNumber="61" unit="m" value="1077.0">1077 m</quantity>
a.s.l.
</elevation>
; 
<date id="F849105BFFE029073B1BFCFF73C40C7D" box="[772,919,845,871]" pageId="60" pageNumber="61" value="2016-07-16">
<collectingDate id="E80DE9B3FFE029073B1BFCFF73C40C7D" box="[772,919,845,871]" pageId="60" pageNumber="61" value="2016-07-16">16 Jul. 2016</collectingDate>
</date>
; 
<collectorName id="2102534DFFE029073BBDFCFF74D00C7D" box="[930,1155,845,871]" pageId="60" pageNumber="61">Chen-guang Zheng</collectorName>
leg.; 
<collectionCode id="EAE6AE5EFFE029073CDBFCFC754D0C72" box="[1220,1310,846,872]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="60" pageNumber="61">NKUM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3C9F3CC6FFE029073D2AFCFC73D60CB4" ID-GBIF-Occurrence="4429869384" collectingDate="2016-07-16" collectionCode="NKUM" collectorName="Chen-guang Zheng" country="China" county="Qing-chuan County" elevation="1077" latitude="32.52364" location="Guang-yuan City" longLatPrecision="1" longitude="104.82325" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Sichuan">
2 ♁♁ (macropterous); 
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, 
<collectingCounty id="65294E17FFE029073A7DFCC373030C91" box="[610,848,881,907]" pageId="60" pageNumber="61">Qing-chuan County</collectingCounty>
, Tang-jia-he; 
<geoCoordinate id="E9C3505CFFE029073BE7FCDD74F40C91" box="[1016,1191,879,907]" degrees="32" direction="north" minutes="31" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="25.1" value="32.52364">32°31′25.1″ N</geoCoordinate>
, 
<geoCoordinate id="E9C3505CFFE029073CA9FCDD75220C91" box="[1206,1393,879,907]" degrees="104" direction="east" minutes="49" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="23.7" value="104.82325">104°49′23.7″ E</geoCoordinate>
; 
<elevation id="07DAD1A8FFE0290738A2FC2671050CB4" box="[189,342,916,942]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="60" pageNumber="61" unit="m" value="1077.0">
<quantity id="4B0F9B7EFFE0290738A2FC2671450CB4" box="[189,278,916,942]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="60" pageNumber="61" unit="m" value="1077.0">1077 m</quantity>
a.s.l.
</elevation>
; 
<date id="F849105BFFE02907397CFC2671A40CB4" box="[355,503,916,942]" pageId="60" pageNumber="61" value="2016-07-16">
<collectingDate id="E80DE9B3FFE02907397CFC2671A40CB4" box="[355,503,916,942]" pageId="60" pageNumber="61" value="2016-07-16">16 Jul. 2016</collectingDate>
</date>
; 
<collectorName id="2102534DFFE029073A1CFC2672B60CB4" box="[515,741,916,942]" pageId="60" pageNumber="61">Chen-guang Zheng</collectorName>
leg.; 
<collectionCode id="EAE6AE5EFFE029073B38FC2673D60CB4" box="[807,901,916,942]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="60" pageNumber="61">NKUM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFE0290538A2FC6771AA0B60" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61" type="description">
<paragraph id="8C48369BFFE0290738A2FC6771AE0CF5" blockId="60.[189,509,981,1007]" box="[189,509,981,1007]" pageId="60" pageNumber="61">
<heading id="D70081F7FFE0290738A2FC6771AE0CF5" bold="true" box="[189,509,981,1007]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="BE83EA89FFE0290738A2FC6771AE0CF5" bold="true" box="[189,509,981,1007]" pageId="60" pageNumber="61">Supplemental description</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFE0290738F5FBB171EB0B07" blockId="60.[186,1400,1027,1266]" box="[234,440,1027,1053]" pageId="60" pageNumber="61">
<emphasis id="BE83EA89FFE0290738F5FBB171EB0B07" bold="true" box="[234,440,1027,1053]" pageId="60" pageNumber="61">Apterous female</emphasis>
</paragraph>
<paragraph id="8C48369BFFE0290738A2FB9A75150BEB" blockId="60.[186,1400,1027,1266]" pageId="60" pageNumber="61">
<smallCapsWord id="8AAEA047FFE0290738A2FB9A71380B5A" baselines="1083,1083" box="[189,363,1064,1090]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="60" pageNumber="61">MEASUREMENTS</smallCapsWord>
. Body length 10.30–11.59, width 2.70–3.22, head width 1.56, interocular width 0.76, eye length (dorsal view) 0.61; relative lengths of antennal segments I–IV: 5.02: 1.22:1.34:1.23; pronotum: length 0.91,width 1.94;mesonotum: length2.59, width 3.16; metanotum:length 0.82, width 2.68;abdomen length (ventral view) 6.80; abdominal sternum VII: length 2.03, width 1.41; abdominal mediotergite I: length 0.54, width 1.36; relative lengths of leg segments (femur:tibia: tarsal segment I:tarsal segment II): fore leg: 5.19: 4.63:2.48:1.04, middle leg: 13.91:7.10: 3.32: 0.51, hind leg: 13.61:5.56:0.15:0.24.
</paragraph>
<paragraph id="8C48369BFFE0290738A2FAAB754E0AA5" blockId="60.[186,1399,1304,1472]" pageId="60" pageNumber="61">
<smallCapsWord id="8AAEA047FFE0290738A2FAAB711B0A2B" baselines="1324,1324" box="[189,328,1305,1331]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="60" pageNumber="61">COLORATION</smallCapsWord>
. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum mainly black with a median brownish-yellow stripe, metanotum medially with a subtriangular yellow marking and with a sharp apex angle (
<figureCitation id="14CC2A1EFFE029073C13FAED74340A63" box="[1036,1127,1375,1401]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[336,1266,297,1835]" captionTargetId="figure-14@4.[255,1333,265,1851]" captionTargetPageId="4" captionText="Fig. 2. Habitus of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise. A. R. abdominalisAndersen &amp; Chen, 1995.B. R. chinensis Hungerford &amp; Matsuda, 1959.C. R. lundbladi (Hungerford, 1957). D. R. scorpio Andersen &amp; Chen, 1995. E. R. svenhedini (Lundblad, 1934). F. R. takahashii Esaki, 1923, dealated macropterous female. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385634" httpUri="https://zenodo.org/record/8385634/files/figure.png" pageId="60" pageNumber="61">Figs 2E</figureCitation>
, 
<figureCitation id="14CC2A1EFFE029073C6FFAED74DD0A63" box="[1136,1166,1375,1401]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="60" pageNumber="61">3P</figureCitation>
, 
<figureCitation id="14CC2A1EFFE029073C88FAED749C0A63" box="[1175,1231,1375,1401]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">34A</figureCitation>
). Connexivum dorsally brownish-yellow. Abdominal tergum I with a median yellow marking. Mesosternum chiefly blackish with a median subtriangular yellowish spot (
<figureCitation id="14CC2A1EFFE029073B2FFA1773CF0AA5" box="[816,924,1445,1471]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34B</figureCitation>
). Abdominal venter light yellow.
</paragraph>
<paragraph id="8C48369BFFE0290738A2FA5574AF09ED" blockId="60.[189,1399,1511,1784]" pageId="60" pageNumber="61">
<smallCapsWord id="8AAEA047FFE0290738A2FA55717F0AE5" baselines="1530,1530" box="[189,300,1511,1537]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="60" pageNumber="61">ABDOMEN</smallCapsWord>
. Abdomen relatively long, caudal part moderately curved dorsad (
<figureCitation id="14CC2A1EFFE029073C7AFA557485091B" box="[1125,1238,1511,1537]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34C</figureCitation>
). Connexival margin of abdominal segments I–II erect, of III–VII reflexed (
<figureCitation id="14CC2A1EFFE029073BAAF9B87475093E" box="[949,1062,1546,1572]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34A</figureCitation>
), meeting each other above terminal mediotergites (
<figureCitation id="14CC2A1EFFE0290739C1F99F722B095D" box="[478,632,1581,1607]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34A, D</figureCitation>
). Posterior margin of abdominal mediotergite VII with a short median process. Abdominal segment VII elongate, about as long as two preceding abdominal segments together (
<figureCitation id="14CC2A1EFFE02907392DF9C171CA0994" box="[306,409,1651,1678]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="60" pageNumber="61">Figs 5O</figureCitation>
, 
<figureCitation id="14CC2A1EFFE0290739B5F9C6718C0994" box="[426,479,1652,1678]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">34C</figureCitation>
). Posterior margin of abdominal segment VII with four distinct processes, including two lobe-like processes on each connexival corner (
<figureCitation id="14CC2A1EFFE029073BDFF925740609AB" box="[960,1109,1687,1713]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34D–F</figureCitation>
): one long, posteriorly-directed outer process and a smaller pointed inner process (in some individuals, the inner process very small or indistinct), posterior margin of sternum VII with a small median process (
<figureCitation id="14CC2A1EFFE029073C9AF96F74A309ED" box="[1157,1264,1757,1783]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="60" pageNumber="61">Fig. 34E</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFE0290738F5F8AC71F20823" blockId="60.[186,1400,1822,1990]" box="[234,417,1822,1849]" pageId="60" pageNumber="61">
<emphasis id="BE83EA89FFE0290738F5F8AC71F20823" bold="true" box="[234,417,1822,1849]" pageId="60" pageNumber="61">Apterous male</emphasis>
</paragraph>
<paragraph id="8C48369BFFE0290538A2F8F075640E5D" blockId="60.[186,1400,1822,1990]" lastBlockId="62.[187,1398,265,327]" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61">
<smallCapsWord id="8AAEA047FFE0290738A2F8F071380840" baselines="1877,1877" box="[189,363,1858,1884]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="60" pageNumber="61">MEASUREMENTS</smallCapsWord>
. Body length 7.21–7.90, width 1.70–2.22, head width 1.36, interocular width 0.72, eye length (dorsal view) 0.51; relative lengths of antennal segments I–IV: 3.42: 1.08:1.23:0.93; pronotum: length 0.69, width 1.45;mesonotum: length2.22, width 2.22; metanotum:length 0.66, width 2.09;abdomen length (ventral view) 3.53; abdominal sternum VII: length 0.54, width 1.04; abdominal mediotergite I: length 0.27, width 0.98; relative lengths of leg segments (femur:tibia: tarsal segment I:tarsal segment II): fore leg: 4.44: 3.84:1.29:0.78, middle leg: 12.90:6.85: 2.83: 0.51, hind leg: 13.3: 4.23: 0.21: 0.15.
</paragraph>
<caption id="D8886613FFE1290638A2F94E72C808F0" ID-DOI="http://doi.org/10.5281/zenodo.8385708" ID-Zenodo-Dep="8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="61" pageNumber="62" startId="61.[189,232,1788,1814]" targetBox="[312,1309,275,1739]" targetPageId="61">
<paragraph id="8C48369BFFE1290638A2F94E72C808F0" blockId="61.[187,1399,1788,2026]" pageId="61" pageNumber="62">
<emphasis id="BE83EA89FFE1290638A2F94E714A080C" bold="true" box="[189,281,1788,1814]" pageId="61" pageNumber="62">Fig. 34.</emphasis>
Morphological features of 
<taxonomicName id="4BF74D18FFE129063A4DF94E746D080C" authority="(Lundblad, 1934)" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[594,1086,1788,1814]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFE129063A4DF94E7338080C" box="[594,875,1788,1814]" italics="true" pageId="61" pageNumber="62">Rhyacobates svenhedini</emphasis>
(
<bibRefCitation id="E8664B6AFFE129063B64F94E7465080C" author="Lundblad" box="[891,1078,1788,1814]" firstAuthor="Lundblad" pageId="61" pageNumber="62" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934</bibRefCitation>
)
</taxonomicName>
. 
<emphasis id="BE83EA89FFE129063C53F94E7432080C" bold="true" box="[1100,1121,1788,1814]" pageId="61" pageNumber="62">A</emphasis>
. Body of female, dorsal view. 
<emphasis id="BE83EA89FFE12906391CF89271440820" bold="true" box="[259,279,1824,1850]" pageId="61" pageNumber="62">B</emphasis>
. Body of female, excluding head and prothorax, ventral view. 
<emphasis id="BE83EA89FFE129063BE3F8AD74420823" bold="true" box="[1020,1041,1823,1849]" pageId="61" pageNumber="62">C</emphasis>
. Body of female, lateral view. 
<emphasis id="BE83EA89FFE1290638A1F8F170800847" bold="true" box="[190,211,1859,1885]" pageId="61" pageNumber="62">D</emphasis>
. Abdominal end of female, dorsal view. 
<emphasis id="BE83EA89FFE129063AB4F8F172EC0847" bold="true" box="[683,703,1859,1885]" pageId="61" pageNumber="62">E</emphasis>
. Abdominal end of female, ventral view. 
<emphasis id="BE83EA89FFE129063CBDF8F174E70847" bold="true" box="[1186,1204,1859,1885]" pageId="61" pageNumber="62">F</emphasis>
. Abdominal end of female, lateral view. 
<emphasis id="BE83EA89FFE1290639D2F8D47259089A" bold="true" box="[461,522,1894,1920]" pageId="61" pageNumber="62">G–H</emphasis>
. Body of male, apterous form (G) and macropterous form (H), dorsal view. 
<emphasis id="BE83EA89FFE1290638A3F83B709408B9" bold="true" box="[188,199,1929,1955]" pageId="61" pageNumber="62">I</emphasis>
. Body of male, excluding head and prothorax, ventral view. 
<emphasis id="BE83EA89FFE129063B89F83B73F708B9" bold="true" box="[918,932,1929,1955]" pageId="61" pageNumber="62">J</emphasis>
. Basal part of right middle leg of male, ventral view. 
<emphasis id="BE83EA89FFE12906397EF81E712508DC" bold="true" box="[353,374,1964,1990]" pageId="61" pageNumber="62">K</emphasis>
. Proctiger of male, dorsal view. 
<emphasis id="BE83EA89FFE129063B1EF81E734008DC" bold="true" box="[769,787,1964,1990]" pageId="61" pageNumber="62">L</emphasis>
. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8C48369BFFE2290538A6FEDC727F0D0E" blockId="62.[185,1399,366,533]" pageId="62" pageNumber="63">
C OLORATION. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum mainly black with a median brownish-yellow stripe. Metanotum chiefly blackish, without yellow markings (
<figureCitation id="14CC2A1EFFE229053AAFFE0673470ED4" box="[688,788,436,462]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. gongvo Tran &amp; Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen &amp; Chen, 1995. K. R. recurvus Andersen &amp; Chen, 1995. L. R. scorpio Andersen &amp; Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="62" pageNumber="63">Figs 4M</figureCitation>
, 
<figureCitation id="14CC2A1EFFE229053B01FE0673060ED4" box="[798,853,436,462]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="62" pageNumber="63">34G</figureCitation>
). Connexivum dorsally blackish. Mesosternum chiefly blackish with a median subtriangular yellowish spot, narrower than that of female (
<figureCitation id="14CC2A1EFFE229053D1AFE65753A0EEB" box="[1285,1385,471,497]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="62" pageNumber="63">Fig. 34I</figureCitation>
). Abdominal venter light yellow.
</paragraph>
<paragraph id="8C48369BFFE2290538A2FD8E71390D63" blockId="62.[187,1398,572,634]" pageId="62" pageNumber="63">
<smallCapsWord id="8AAEA047FFE2290538A2FD8E70B80D4E" baselines="591,591" box="[189,235,572,598]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="62" pageNumber="63">LEG</smallCapsWord>
. Middle trochanter without spines; middle femur with scattered small spines, not arranged in distinct row (
<figureCitation id="14CC2A1EFFE2290538E6FDED710E0D63" box="[249,349,607,633]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="62" pageNumber="63">Fig. 34J</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFE2290538A2FD1373350C1B" blockId="62.[189,1399,672,769]" pageId="62" pageNumber="63">
<smallCapsWord id="8AAEA047FFE2290538A2FD1371620DA3" baselines="692,692" box="[189,305,673,699]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="62" pageNumber="63">GENITALIA</smallCapsWord>
. Abdominal segment VIII ventro-laterally impressed. Pygophore large, ovate. Proctiger relatively large, with angular lobes laterally (
<figureCitation id="14CC2A1EFFE229053AC3FD76736A0DC4" box="[732,825,708,734]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="62" pageNumber="63">Figs 7F</figureCitation>
, 
<figureCitation id="14CC2A1EFFE229053B57FD76732D0DC4" box="[840,894,708,734]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="62" pageNumber="63">34K</figureCitation>
). Paramere strongly curved at basal third, distal part slender, with hook-shaped apex (
<figureCitation id="14CC2A1EFFE229053AA4FD55734A0C1B" box="[699,793,743,769]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen &amp; Chen, 1995. J. R. recurvus Andersen &amp; Chen, 1995. K. R. scorpio Andersen &amp; Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="62" pageNumber="63">Figs 8L</figureCitation>
, 
<figureCitation id="14CC2A1EFFE229053B3AFD55730A0C1B" box="[805,857,743,769]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="62" pageNumber="63">34L</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFE2290538F5FC9A718D0C58" blockId="62.[188,1398,808,940]" box="[234,478,808,834]" pageId="62" pageNumber="63">
<emphasis id="BE83EA89FFE2290538F5FC9A718D0C58" bold="true" box="[234,478,808,834]" pageId="62" pageNumber="63">Macropterous male</emphasis>
</paragraph>
<paragraph id="8C48369BFFE2290538A2FCF974150CB6" blockId="62.[188,1398,808,940]" pageId="62" pageNumber="63">
Similar to apterous female in general structure and coloration with following exceptions: thorax with a pronotal lobe, anterior part with a subrhombic yellow marking, posterior part elongate, covering most of mesonotum; posterior margin broadly rounded and brownish (
<figureCitation id="14CC2A1EFFE229053B8AFC2073A50CB6" box="[917,1014,914,940]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. gongvo Tran &amp; Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen &amp; Chen, 1995. K. R. recurvus Andersen &amp; Chen, 1995. L. R. scorpio Andersen &amp; Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="62" pageNumber="63">Figs 4N</figureCitation>
, 
<figureCitation id="14CC2A1EFFE229053C1CFC20746A0CB6" box="[1027,1081,914,940]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. G–H. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="62" pageNumber="63">34H</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFE2290538A2FC6671AA0B60" blockId="62.[185,1399,979,1147]" pageId="62" pageNumber="63">
<smallCapsWord id="8AAEA047FFE2290538A2FC6671380CF6" baselines="999,999" box="[189,363,980,1006]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="62" pageNumber="63">MEASUREMENTS</smallCapsWord>
. Body length 7.35, width 2.11, head width 1.30, interocular width 0.62, eye length (dorsal view) 0.51;relative lengths of antennal segments I–IV: 3.88:1.05: 1.18: 0.99; pronotum: length2.71,width 2.14; mesonotum width 2.11; metanotum width 1.80; relative lengths of leg segments (femur:tibia: tarsal segment I: tarsal segment II): fore leg: 3.96: 3.04:1.23:0.61, middle leg: 11.36:6.31: 3.05: 0.75, hind leg: 11.67: 4.99: 0.25: 0.21.
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFE2290538A2FB1373500BF1" box="[189,771,1185,1259]" pageId="62" pageNumber="63" type="distribution">
<paragraph id="8C48369BFFE2290538A2FB1371050BA1" blockId="62.[189,342,1185,1211]" box="[189,342,1185,1211]" pageId="62" pageNumber="63">
<heading id="D70081F7FFE2290538A2FB1371050BA1" bold="true" box="[189,342,1185,1211]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="BE83EA89FFE2290538A2FB1371050BA1" bold="true" box="[189,342,1185,1211]" pageId="62" pageNumber="63">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFE2290538A2FB6273500BF1" blockId="62.[189,771,1232,1259]" box="[189,771,1232,1259]" pageId="62" pageNumber="63">
<collectingCountry id="F4E0760BFFE2290538A2FB62715B0BF0" box="[189,264,1232,1258]" name="China" pageId="62" pageNumber="63">China</collectingCountry>
: 
<collectingRegion id="4E33F879FFE22905390CFB6271320BF0" box="[275,353,1232,1258]" country="China" name="Fujian" pageId="62" pageNumber="63">Fujian</collectingRegion>
, 
<collectingRegion id="4E33F879FFE229053973FB6271980BF0" box="[364,459,1232,1258]" country="China" name="Sichuan" pageId="62" pageNumber="63">Sichuan</collectingRegion>
(
<bibRefCitation id="E8664B6AFFE2290539C5FB6272A50BF1" author="Andersen &amp; Chen" box="[474,758,1232,1259]" firstAuthor="Andersen" pageId="62" pageNumber="63" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFE2290538A2FAA3756D0A41" pageId="62" pageNumber="63" type="discussion">
<paragraph id="8C48369BFFE2290538A2FAA371F90A36" blockId="62.[189,426,1297,1324]" box="[189,426,1297,1324]" pageId="62" pageNumber="63">
<heading id="D70081F7FFE2290538A2FAA371F90A36" bold="true" box="[189,426,1297,1324]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="BE83EA89FFE2290538A2FAA371F90A36" bold="true" box="[189,426,1297,1324]" pageId="62" pageNumber="63">Comparative notes</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFE2290538A2FAF2756D0A41" blockId="62.[189,1342,1344,1371]" box="[189,1342,1344,1371]" pageId="62" pageNumber="63">
<taxonomicName id="4BF74D18FFE2290538A2FAF271840A40" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[189,471,1344,1370]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFE2290538A2FAF271840A40" box="[189,471,1344,1370]" italics="true" pageId="62" pageNumber="63">Rhyacobates svenhedini</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4BF74D18FFE229053AAEFAF373110A40" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[689,834,1344,1370]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFE229053AAEFAF373110A40" box="[689,834,1344,1370]" italics="true" pageId="62" pageNumber="63">R. lundbladi</emphasis>
</taxonomicName>
; see comparative notes under 
<taxonomicName id="4BF74D18FFE229053CB9FAF375640A40" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[1190,1335,1344,1370]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFE229053CB9FAF375640A40" box="[1190,1335,1344,1370]" italics="true" pageId="62" pageNumber="63">R. lundbladi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>