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<document id="06C08E54D9557C7C22EFC3D1805D323B" ID-CLB-Dataset="274285" ID-DOI="10.5852/ejt.2023.893.2285" ID-GBIF-Dataset="e441bc9d-bb44-4ee6-b29f-a6724b96592b" ID-ISSN="2118-9773" ID-Zenodo-Dep="8385630" ID-ZooBank="D9DA4879-BC1D-4947-96E8-7D574DE334BA" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1695891225180" checkinUser="plazi" docAuthor="Leng, Zhaoqi, Tran, Anh Duc &amp; Ye, Zhen" docDate="2023" docId="045E878DFF9D29793A3DF9E673680B87" docLanguage="en" docName="EJT.2023.893.1-73.pdf" docOrigin="European Journal of Taxonomy 893 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2285/9863" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhyacobates zetteli Tran &amp; Nguyen 2016" docType="treatment" docVersion="5" lastPageNumber="67" masterDocId="F867FFF5FFDC293B381FFFB270530F1A" masterDocTitle="Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="66" updateTime="1699690649759" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="25864CFA0E4BCA8F13CFCB7D807E4183">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:namePart id="718D127203E098BADA83BF36F5B734C0">Leng, Zhaoqi</mods:namePart>
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<mods:affiliation id="57E4FD44E9E5BC101D70BD21684B0C4D">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
<mods:nameIdentifier id="935DF58077AD6FC7B1614D406236525F" type="email">zhaoqileng0712@gmail.com</mods:nameIdentifier>
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<mods:namePart id="61923C993CF70F8A10D3880DE4984A4F">Tran, Anh Duc</mods:namePart>
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<mods:affiliation id="8DF22F021A74A253D7EF30CA84665B8D">Faculty of Biology, VNU University of Science, Vietnam National University, Hanoi, Nguyen Trai, Thanh Xuan, Hanoi, Vietnam.</mods:affiliation>
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<mods:namePart id="C08416A4B4A3A11F2D784D1DFD3C0640">Ye, Zhen</mods:namePart>
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<mods:affiliation id="464D2192DB0920E07D6F9537F24A57DE">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:title id="C4A3BC91C9AFD9D22047C165AFD861A4">European Journal of Taxonomy</mods:title>
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<subSubSection id="C4ED6510FF9D297A3A3DF9E674410975" box="[546,1042,1620,1647]" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="8C48369BFF9D297A3A3DF9E674410975" blockId="65.[546,1042,1620,1683]" box="[546,1042,1620,1647]" pageId="65" pageNumber="66">
<heading id="D70081F7FF9D297A3A3DF9E674410975" box="[546,1042,1620,1647]" centered="true" fontSize="11" level="2" pageId="65" pageNumber="66" reason="2">
<taxonomicName id="4BF74D18FF9D297A3A3DF9E674410975" ID-CoL="4SNHX" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[546,1042,1620,1647]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="zetteli">
<emphasis id="BE83EA89FF9D297A3A3DF9E6735B0975" bold="true" box="[546,776,1620,1647]" italics="true" pageId="65" pageNumber="66">Rhyacobates zetteli</emphasis>
<bibRefCitation id="E8664B6AFF9D297A3B0FF9E774410975" author="Tran &amp; Nguyen" box="[784,1042,1621,1647]" firstAuthor="Tran" pageId="65" pageNumber="66" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C4ED6510FF9D297A3AA6F9CA73290988" box="[697,890,1656,1683]" pageId="65" pageNumber="66" type="description">
<paragraph id="8C48369BFF9D297A3AA6F9CA73290988" blockId="65.[546,1042,1620,1683]" box="[697,890,1656,1683]" pageId="65" pageNumber="66">
<figureCitation id="14CC2A1EFF9D297A3AA6F9CA734B0988" box="[697,792,1656,1682]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="65" pageNumber="66">Figs 3R</figureCitation>
,
<figureCitation id="14CC2A1EFF9D297A3B3BF9CA73190989" box="[804,842,1656,1683]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="65" pageNumber="66">5Q</figureCitation>
,
<figureCitation id="14CC2A1EFF9D297A3B49F9CA73290988" box="[854,890,1656,1682]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="65" pageNumber="66">7H</figureCitation>
</paragraph>
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<paragraph id="8C48369BFF9D297A38A2F90B74E309C9" blockId="65.[189,1200,1721,1748]" box="[189,1200,1721,1748]" pageId="65" pageNumber="66">
<treatmentCitationGroup id="ACE711B5FF9D297A38A2F90B74E309C9" box="[189,1200,1721,1748]" pageId="65" pageNumber="66">
<treatmentCitation id="0D56108AFF9D297A38A2F90B737109CE" ID-DOI="http://doi.org/10.5281/zenodo.4509706" ID-Zenodo-Dep="4509706" author="Tran A. D. &amp; Nguyen X. Q." box="[189,802,1721,1748]" httpUri="http://treatment.plazi.org/id/4F67DC7C68669F66FF00FF30182DFA36" page="508" pageId="65" pageNumber="66" year="2016">
<taxonomicName id="4BF74D18FF9D297A38A2F90B737109CE" ID-CoL="4SNHX" authority="Tran &amp; Nguyen, 2016: 505 - 508" authorityName="Tran &amp; Nguyen" authorityPageNumber="505 - 508" authorityYear="2016" box="[189,802,1721,1748]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="zetteli">
<emphasis id="BE83EA89FF9D297A38A2F90B71F309C9" box="[189,416,1721,1747]" italics="true" pageId="65" pageNumber="66">Rhyacobates zetteli</emphasis>
<bibRefCitation id="E8664B6AFF9D297A39B8F908737109CE" author="Tran &amp; Nguyen" box="[423,802,1721,1748]" firstAuthor="Tran" page="505 - 508" pageId="65" pageNumber="66" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016: 505–508</bibRefCitation>
</taxonomicName>
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, figs 11–23 (original description).
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C4ED6510FF9D297938A2F948726A0D00" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="8C48369BFF9D297A38A2F9487165080E" blockId="65.[189,310,1786,1812]" box="[189,310,1786,1812]" pageId="65" pageNumber="66">
<heading id="D70081F7FF9D297A38A2F9487165080E" bold="true" box="[189,310,1786,1812]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="BE83EA89FF9D297A38A2F9487165080E" bold="true" box="[189,310,1786,1812]" pageId="65" pageNumber="66">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFF9D297938A2F89B726A0D00" blockId="65.[187,1401,1833,2000]" lastBlockId="66.[189,1401,265,538]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
Body length of apterous females 9.80–10.60, of apterous males 7.30–7.70. Color (
<figureCitation id="14CC2A1EFF9D297A3C95F89B74BB0859" box="[1162,1256,1833,1859]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="65" pageNumber="66">Fig. 3R</figureCitation>
): dorsum of head mainly yellow with small black markings; pronotum mainly yellow with narrow black marking on anterior margin and sometimes laterally; mesonotum with broad median marking from anterior to posterior margin (median marking broader in females than in males); metanotum with laterally expanded yellow marking (more extensive in females). Female: abdomen elongate and straight; ventral length of pregenital abdomen about half of body length (
<figureCitation id="14CC2A1EFF9E29793B3BFEBB73D50E3E" box="[804,902,265,292]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="66" pageNumber="67">Fig. 5Q</figureCitation>
); connexivum of segments I–VI narrow; connexival corners of segment VI with small processes; abdominal segment VII tapering towards apex; mediotergite VII with small median process on posterior margin; sternum VII with bilobate posterior margin, connexival corners with small blunt processes which bend mesad. Male: middle trochanter with 6–7 spines; middle femur with spines but not in distinct row; length of middle tibia about equal to length of hind tibia; pygophore simple, with straight apical margin; proctiger with subtrapezoidal lobes laterally (
<figureCitation id="14CC2A1EFF9E2979396DFE6E71830EEC" box="[370,464,476,502]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="66" pageNumber="67">Fig. 7H</figureCitation>
); paramere relatively long and slender, curved at basal third, distal part tapering towards narrowly rounded apex.
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FF9E297938A2FDF273350C83" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="8C48369BFF9E297938A2FDF271F90D40" blockId="66.[189,426,576,602]" box="[189,426,576,602]" pageId="66" pageNumber="67">
<heading id="D70081F7FF9E297938A2FDF271F90D40" bold="true" box="[189,426,576,602]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="BE83EA89FF9E297938A2FDF271F90D40" bold="true" box="[189,426,576,602]" pageId="66" pageNumber="67">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFF9E297938F5FDDD71FD0D93" blockId="66.[189,516,623,685]" box="[234,430,623,649]" pageId="66" pageNumber="67">
<emphasis id="BE83EA89FF9E297938F5FDDD71FD0D93" bold="true" box="[234,430,623,649]" pageId="66" pageNumber="67">
<typeStatus id="534C8839FF9E297938F5FDDD71740D93" box="[234,295,623,649]" pageId="66" pageNumber="67">Type</typeStatus>
specimens
</emphasis>
</paragraph>
<paragraph id="8C48369BFF9E297938A2FD2172570DB6" blockId="66.[189,516,623,685]" box="[189,516,658,685]" pageId="66" pageNumber="67">
See
<bibRefCitation id="E8664B6AFF9E297938F1FD2172530DB6" author="Tran &amp; Nguyen" box="[238,512,658,685]" firstAuthor="Tran" pageId="66" pageNumber="67" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="8C48369BFF9E297938F5FD6671B70DF4" blockId="66.[189,1398,724,856]" box="[234,484,724,750]" pageId="66" pageNumber="67">
<emphasis id="BE83EA89FF9E297938F5FD6671B70DF4" bold="true" box="[234,484,724,750]" pageId="66" pageNumber="67">Non-type specimens</emphasis>
</paragraph>
<paragraph id="8C48369BFF9E297938A2FD4572B10C42" blockId="66.[189,1398,724,856]" pageId="66" pageNumber="67">
<materialsCitation id="3C9F3CC6FF9E297938A2FD4574FD0C2F" ID-GBIF-Occurrence="4429869337" collectingDate="2020-10-25" collectionCode="ZVNU" collectorName="Nui X &amp; A. D. Tran" country="Vietnam" latitude="22.348446" location="Vang" longLatPrecision="1" longitude="103.770004" municipality="Sa Pa" pageId="66" pageNumber="67" specimenCount="3" specimenCount-female="3" stateProvince="Lao Cai">
<collectingCountry id="F4E0760BFF9E297938A2FD4571170C0B" box="[189,324,759,785]" name="Vietnam" pageId="66" pageNumber="67">VIETNAM</collectingCountry>
•
<specimenCount id="9AF1FD12FF9E2979397CFD4571FB0C08" box="[355,424,759,786]" pageId="66" pageNumber="67" type="female">3 ♀♀</specimenCount>
(apterous);
<collectingRegion id="4E33F879FF9E29793A23FD4573410C0B" box="[572,786,759,785]" country="Vietnam" name="Lao Cai" pageId="66" pageNumber="67">Lào Cai Province</collectingRegion>
,
<collectingMunicipality id="6C2CACE1FF9E29793B3EFD45733A0C08" box="[801,873,759,786]" pageId="66" pageNumber="67">Sa Pa</collectingMunicipality>
,
<collectorName id="2102534DFF9E29793B67FD4573970C08" box="[888,964,759,786]" pageId="66" pageNumber="67">Núi X</collectorName>
ẻ area,
<location id="89286040FF9E29793C03FD45740F0C0B" LSID="urn:lsid:plazi:treatment:045E878DFF9D29793A3DF9E673680B87:89286040FF9E29793C03FD45740F0C0B" box="[1052,1116,759,785]" country="Vietnam" latitude="22.348446" longLatPrecision="1" longitude="103.770004" municipality="Sa Pa" name="Vang" pageId="66" pageNumber="67" stateProvince="Lao Cai">Vàng</location>
stream and its feeders;
<geoCoordinate id="E9C3505CFF9E297938A2FCAB713B0C2F" box="[189,360,793,821]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="66" pageNumber="67" precision="1" seconds="54.4" value="22.348446">22°20′54.4″ N</geoCoordinate>
,
<geoCoordinate id="E9C3505CFF9E2979396CFCAB72790C2F" box="[371,554,793,821]" degrees="103" direction="east" minutes="46" orientation="longitude" pageId="66" pageNumber="67" precision="1" seconds="12.0" value="103.770004">103°46′12.0″ E</geoCoordinate>
;
<date id="F849105BFF9E29793A2AFCA872830C2E" box="[565,720,794,821]" pageId="66" pageNumber="67" value="2020-10-25">
<collectingDate id="E80DE9B3FF9E29793A2AFCA872830C2E" box="[565,720,794,821]" pageId="66" pageNumber="67" value="2020-10-25">25 Oct. 2020</collectingDate>
</date>
;
<collectorName id="2102534DFF9E29793AC6FCA9731C0C2F" box="[729,847,795,821]" pageId="66" pageNumber="67">A.D. Tran</collectorName>
<emphasis id="BE83EA89FF9E29793B4AFCA973C30C2E" box="[853,912,794,820]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
leg.; TAD20-19;
<collectionCode id="EAE6AE5EFF9E29793C42FCA974FD0C2F" box="[1117,1198,795,821]" pageId="66" pageNumber="67">ZVNU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3C9F3CC6FF9E29793CDAFCA8728D0C42" ID-GBIF-Occurrence="4429869313" collectingDate="2020-10-25" collectionCode="NKUM" collectorName="Nui X &amp; A. D. Tran" country="Vietnam" latitude="22.348446" location="Vang" longLatPrecision="1" longitude="103.770004" municipality="Sa Pa" pageId="66" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai">
<specimenCount id="9AF1FD12FF9E29793CDAFCA874A20C2F" box="[1221,1265,794,821]" pageId="66" pageNumber="67" type="female">1 ♀</specimenCount>
(apterous); same collection data as for preceding;
<collectionCode id="EAE6AE5EFF9E29793A9FFC8C728D0C42" box="[640,734,830,856]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="66" pageNumber="67">NKUM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8C48369BFF9E297938A2FCCD73350C83" blockId="66.[189,870,895,922]" box="[189,870,895,922]" pageId="66" pageNumber="67">
GPS data of previous records: see
<bibRefCitation id="E8664B6AFF9E29793A4FFC3273300C83" author="Tran &amp; Nguyen" box="[592,867,895,922]" firstAuthor="Tran" pageId="66" pageNumber="67" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FF9E297938A2FC7272F80B13" box="[189,683,960,1034]" pageId="66" pageNumber="67" type="distribution">
<paragraph id="8C48369BFF9E297938A2FC7271050CC0" blockId="66.[189,342,960,986]" box="[189,342,960,986]" pageId="66" pageNumber="67">
<heading id="D70081F7FF9E297938A2FC7271050CC0" bold="true" box="[189,342,960,986]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="BE83EA89FF9E297938A2FC7271050CC0" bold="true" box="[189,342,960,986]" pageId="66" pageNumber="67">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFF9E297938A2FC5D72F80B13" blockId="66.[189,683,1007,1034]" box="[189,683,1007,1034]" pageId="66" pageNumber="67">
<collectingCountry id="F4E0760BFF9E297938A2FC5D71760B13" box="[189,293,1007,1033]" name="Vietnam" pageId="66" pageNumber="67">Vietnam</collectingCountry>
:
<collectingRegion id="4E33F879FF9E2979392FFC5D71DD0B13" box="[304,398,1007,1033]" country="Vietnam" name="Lao Cai" pageId="66" pageNumber="67">Lào Cai</collectingRegion>
(
<bibRefCitation id="E8664B6AFF9E29793982FC5D72CC0B13" author="Tran &amp; Nguyen" box="[413,671,1007,1034]" firstAuthor="Tran" pageId="66" pageNumber="67" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FF9E297938A2FB8273680B87" pageId="66" pageNumber="67" type="discussion">
<paragraph id="8C48369BFF9E297938A2FB8271F90B51" blockId="66.[189,426,1072,1099]" box="[189,426,1072,1099]" pageId="66" pageNumber="67">
<heading id="D70081F7FF9E297938A2FB8271F90B51" bold="true" box="[189,426,1072,1099]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="BE83EA89FF9E297938A2FB8271F90B51" bold="true" box="[189,426,1072,1099]" pageId="66" pageNumber="67">Comparative notes</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFF9E297938A2FBED73680B87" blockId="66.[189,1398,1119,1181]" pageId="66" pageNumber="67">
<taxonomicName id="4BF74D18FF9E297938A2FBED71F10B63" ID-CoL="4SNHX" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[189,418,1119,1145]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="zetteli">
<emphasis id="BE83EA89FF9E297938A2FBED71F10B63" box="[189,418,1119,1145]" italics="true" pageId="66" pageNumber="67">Rhyacobates zetteli</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4BF74D18FF9E29793A97FBD273430B63" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[648,784,1120,1145]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="BE83EA89FF9E29793A97FBD273430B63" box="[648,784,1120,1145]" italics="true" pageId="66" pageNumber="67">R. angustus</emphasis>
</taxonomicName>
, as discussed by
<treatmentCitation id="0D56108AFF9E29793BFCFBED757D0B60" ID-DOI="http://doi.org/10.5281/zenodo.4509706" ID-Zenodo-Dep="4509706" author="Tran A. D. &amp; Nguyen X. Q." box="[995,1326,1119,1146]" httpUri="http://treatment.plazi.org/id/4F67DC7C68669F66FF00FF30182DFA36" page="503" pageId="66" pageNumber="67" year="2016">
<bibRefCitation id="E8664B6AFF9E29793BFCFBED757D0B60" author="Tran &amp; Nguyen" box="[995,1326,1119,1146]" firstAuthor="Tran" page="503" pageId="66" pageNumber="67" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen (2016: 503</bibRefCitation>
</treatmentCitation>
, 507– 508). Also see comparative notes under
<taxonomicName id="4BF74D18FF9E29793A8CFB31728A0B86" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[659,729,1154,1180]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bui" status="sp. nov.">
<emphasis id="BE83EA89FF9E29793A8CFB31728A0B86" box="[659,729,1154,1180]" italics="true" pageId="66" pageNumber="67">R. bui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FF9E29793AFFFB3173680B87" box="[736,827,1155,1181]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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