treatments-xml/data/04/19/7D/04197D19FFCDFF24FF59FB2E62F9F2EA.xml

<document id="3002204A8D1FFF77481723A16ED0E5CC" ID-CLB-Dataset="25162" ID-DOI="10.11646/zootaxa.4688.3.5" ID-GBIF-Dataset="62d23c6d-47a2-45b2-b066-f60dac27e9e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3516813" ID-ZooBank="DDA1FDD7-895E-4667-ACA7-4003B45DFF5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571832775124" checkinUser="plazi" docAuthor="Jewell, Tony R." docDate="2019" docId="04197D19FFCDFF24FF59FB2E62F9F2EA" docLanguage="en" docName="zootaxa.4688.3.5.pdf" docOrigin="Zootaxa 4688 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oligosoma pachysomaticum" docType="treatment" docVersion="8" lastPageNumber="395" masterDocId="F8200561FFCEFF22FFCEFF886142F345" masterDocTitle="New Zealand forest-dwelling skinks of the Oligosoma oliveri (McCann) species-complex (Reptilia: Scincidae): reinstatement of O. pachysomaticum (Robb) and an assessment of historical distribution ranges" masterLastPageNumber="398" masterPageNumber="382" pageNumber="392" updateTime="1698760724892" updateUser="ExternalLinkService">
<mods:mods id="837A0A502F513053420C379EFD272BD2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FF7A6986FCEB0A41B7023FE91415C7AD">
<mods:title id="625511E233788233F4F98FDBEC2E4ABA">New Zealand forest-dwelling skinks of the Oligosoma oliveri (McCann) species-complex (Reptilia: Scincidae): reinstatement of O. pachysomaticum (Robb) and an assessment of historical distribution ranges</mods:title>
</mods:titleInfo>
<mods:name id="DD43BCDCD408C86E13C6E8FB7B6620CC" type="personal">
<mods:role id="CBD08255A8F0C35A6F9805EB5F013177">
<mods:roleTerm id="5377D5D7365EFA2D610AC39AA8DFA592">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="67803B2F808895DC168BFB023B623B97">Jewell, Tony R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="568275CA7D4CC8E56FFC71DBBE78A382">text</mods:typeOfResource>
<mods:relatedItem id="98194E481A72A0EC22D7E4F23F8EF145" type="host">
<mods:titleInfo id="0AB32A38FCFFFF383A48131F3D885309">
<mods:title id="328A99F3813637793BFCDC0538EEDEC7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5422B0FA0E3F9E7ED798FA0FDFF278EF">
<mods:date id="75EF7562839724B20C1F9B6295048C28">2019</mods:date>
<mods:detail id="6DECF32461AEF2921BAC21A65E0C886C" type="pubDate">
<mods:number id="21558857526CDF6A7B519CF14189E5FA">2019-10-23</mods:number>
</mods:detail>
<mods:detail id="05C2CFBAE178850F2A052A30B632B6E6" type="volume">
<mods:number id="C456270F876FA5677EE239C710C0A847">4688</mods:number>
</mods:detail>
<mods:detail id="98CA2F20A67E90EE8C2AE75ECB18B24A" type="issue">
<mods:number id="A77FCB7EB9FB3F90A9CDF80E16F89C9E">3</mods:number>
</mods:detail>
<mods:extent id="0ED91850C5F1D7849773C8526DC9EB11" unit="page">
<mods:start id="E99984C006C56FFFEB326F7FB64FBC47">382</mods:start>
<mods:end id="D0FBC0D7FC5FF7EA2F7BA2B02858A2E6">398</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AEB875DC2F824AF827ED6D4E4D96879E">journal article</mods:classification>
<mods:identifier id="D851D08E7B39A9C23212A9B95E4C30EE" type="CLB-Dataset">25162</mods:identifier>
<mods:identifier id="7E26BCBE3788FBC59C3640ACBF7C238A" type="DOI">10.11646/zootaxa.4688.3.5</mods:identifier>
<mods:identifier id="724A52245DC97C6A19D59A6797BC2CBA" type="GBIF-Dataset">62d23c6d-47a2-45b2-b066-f60dac27e9e1</mods:identifier>
<mods:identifier id="2C1F5A82A25CB675326058BF46DF0E24" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B720095F8A9CFB86A44B658B0C1C70D4" type="Zenodo-Dep">3516813</mods:identifier>
<mods:identifier id="EC63F5F8B938D77F3BA39D254D597B74" type="ZooBank">DDA1FDD7-895E-4667-ACA7-4003B45DFF5D</mods:identifier>
</mods:mods>
<treatment id="04197D19FFCDFF24FF59FB2E62F9F2EA" ID-GBIF-Taxon="159812130" LSID="urn:lsid:plazi:treatment:04197D19FFCDFF24FF59FB2E62F9F2EA" httpUri="http://treatment.plazi.org/id/04197D19FFCDFF24FF59FB2E62F9F2EA" lastPageId="6" lastPageNumber="395" pageId="3" pageNumber="392">
<subSubSection id="C4AA9F84FFCDFF21FF59FB2E63D7F785" box="[151,661,1190,1217]" pageId="3" pageNumber="392" type="nomenclature">
<paragraph id="8C0FCC0FFFCDFF21FF59FB2E63D7F785" blockId="3.[151,661,1190,1252]" box="[151,661,1190,1217]" pageId="3" pageNumber="392">
<heading id="D7477B63FFCDFF21FF59FB2E63D7F785" bold="true" box="[151,661,1190,1217]" fontSize="11" level="1" pageId="3" pageNumber="392" reason="1">
<taxonomicName id="4BB0B78CFFCDFF21FF59FB2E63D7F785" authority="(Robb, 1975)" baseAuthorityName="Robb" baseAuthorityYear="1975" box="[151,661,1190,1217]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCDFF21FF59FB2E63D7F785" bold="true" box="[151,661,1190,1217]" pageId="3" pageNumber="392">
<emphasis id="BEC4101DFFCDFF21FF59FB2E60AFF784" bold="true" box="[151,493,1190,1217]" italics="true" pageId="3" pageNumber="392">Oligosoma pachysomaticum</emphasis>
(
<bibRefCitation id="E821B1FEFFCDFF21FE32FB2E63CFF785" author="Robb, J." box="[508,653,1190,1216]" pageId="3" pageNumber="392" pagination="477 - 484" refId="ref6122" refString="Robb, J. (1975) Two new skinks of the genus Leiolopisma from New Zealand. Proceedings of the Section of Sciences, Koninklijke Nederlandse Akademie van Wetenschappen, Series C, 78, 477 - 484." type="journal article" year="1975">Robb, 1975</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCDFF21FF59FB426047F662" pageId="3" pageNumber="392" type="description">
<paragraph id="8C0FCC0FFFCDFF21FF59FB4261B7F7A1" blockId="3.[151,661,1190,1252]" box="[151,245,1226,1252]" pageId="3" pageNumber="392">
<figureCitation id="148BD08AFFCDFF21FF59FB4261B7F7A1" box="[151,245,1226,1252]" captionStart="FIGURE 3" captionStartId="5.[151,250,1140,1165]" captionTargetBox="[151,1434,174,1121]" captionTargetId="figure@5.[151,1434,174,1121]" captionTargetPageId="5" captionText="FIGURE 3. Two examples of Oligosoma pachysomaticum (Robb) from Green Island in the Mercury Island group, east of Coromandel Peninsula (photos: Tony Whitaker)." figureDoi="http://doi.org/10.5281/zenodo.3516821" httpUri="https://zenodo.org/record/3516821/files/figure.png" pageId="3" pageNumber="392">Figure 3</figureCitation>
</paragraph>
<paragraph id="8C0FCC0FFFCDFF21FF59FA876047F662" blockId="3.[151,802,1295,1503]" box="[151,261,1295,1319]" pageId="3" pageNumber="392">Synonomy</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCDFF21FF59FAA56366F621" pageId="3" pageNumber="392" type="reference_group">
<paragraph id="8C0FCC0FFFCDFF21FF59FAA56300F603" blockId="3.[151,802,1295,1503]" box="[151,578,1325,1350]" pageId="3" pageNumber="392">
<taxonomicName id="4BB0B78CFFCDFF21FF59FAA56300F603" authority="Robb, 1975" authorityName="Robb" authorityYear="1975" box="[151,578,1325,1350]" class="Reptilia" family="Scincidae" genus="Leiolopisma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCDFF21FF59FAA56087F600" box="[151,453,1325,1349]" italics="true" pageId="3" pageNumber="392">Leiolopisma pachysomaticum</emphasis>
<bibRefCitation id="E821B1FEFFCDFF21FE05FAA56300F603" author="Robb, J." box="[459,578,1325,1350]" pageId="3" pageNumber="392" pagination="477 - 484" refId="ref6122" refString="Robb, J. (1975) Two new skinks of the genus Leiolopisma from New Zealand. Proceedings of the Section of Sciences, Koninklijke Nederlandse Akademie van Wetenschappen, Series C, 78, 477 - 484." type="journal article" year="1975">Robb, 1975</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8C0FCC0FFFCDFF21FF59FAC46366F621" blockId="3.[151,802,1295,1503]" box="[151,548,1355,1380]" pageId="3" pageNumber="392">
<taxonomicName id="4BB0B78CFFCDFF21FF59FAC4600AF621" authorityName="Hardy" authorityYear="1977" box="[151,328,1356,1380]" class="Squamata" family="Scincidae" genus="Cyclodina" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCDFF21FF59FAC4600AF621" box="[151,328,1356,1380]" italics="true" pageId="3" pageNumber="392">Cyclodina oliveri</emphasis>
</taxonomicName>
(in part) 
<bibRefCitation id="E821B1FEFFCDFF21FE69FAC36366F621" author="Hardy, G. S." box="[423,548,1355,1380]" pageId="3" pageNumber="392" pagination="221 - 325" refId="ref5756" refString="Hardy, G. S. (1977) The New Zealand Scincidae (Reptilia: Lacertilia); a taxonomic and Zoogeographic study. New Zealand Journal of Zoology, 4, 221 - 325. https: // doi. org / 10.1080 / 03014223.1977.9517956" type="journal article" year="1977">Hardy, 1977</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCDFF21FF59FAE36337F69A" pageId="3" pageNumber="392" type="description">
<paragraph id="8C0FCC0FFFCDFF21FF59FAE36260F6C6" blockId="3.[151,802,1295,1503]" box="[151,802,1386,1411]" pageId="3" pageNumber="392">
<taxonomicName id="4BB0B78CFFCDFF21FF59FAE360F2F6C6" authorityName="Wells &amp; Wellington" authorityYear="1985" box="[151,432,1387,1411]" class="Squamata" family="Scincidae" genus="Cyclodina" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCDFF21FF59FAE360F2F6C6" box="[151,432,1387,1411]" italics="true" pageId="3" pageNumber="392">Cyclodina pachysomaticum</emphasis>
</taxonomicName>
(in part?) 
<bibRefCitation id="E821B1FEFFCDFF21FDD4FAE26260F6C6" author="Wells, R. W. &amp; Wellington, C. R." box="[538,802,1386,1411]" pageId="3" pageNumber="392" pagination="62 - 64" refId="ref6449" refString="Wells, R. W. &amp; Wellington, C. R. (1985) A synopsis of the Amphibia and Reptilia of New Zealand. Australian Journal of Herpetology, Supplementary Series 1, 62 - 64. Available from: http: // www. iucn-tftsg. org / wp-content / uploads / file / Articles / Wells _ and _ Wellington _ 1985. pdf (Accessed 22 Oct. 2019)" type="book chapter" year="1985">
Wells &amp; 
<collectingRegion id="4E7402EDFFCDFF21FDBDFAE263A4F6C6" box="[627,742,1386,1411]" country="New Zealand" name="Wellington" pageId="3" pageNumber="392">Wellington</collectingRegion>
, 1985
</bibRefCitation>
</paragraph>
<paragraph id="8C0FCC0FFFCDFF21FF59FA01638CF6E4" blockId="3.[151,802,1295,1503]" box="[151,718,1417,1442]" pageId="3" pageNumber="392">
<taxonomicName id="4BB0B78CFFCDFF21FF59FA01600AF6E4" authorityName="Hardy" authorityYear="1977" box="[151,328,1417,1441]" class="Squamata" family="Scincidae" genus="Cyclodina" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCDFF21FF59FA01600AF6E4" box="[151,328,1417,1441]" italics="true" pageId="3" pageNumber="392">Cyclodina oliveri</emphasis>
</taxonomicName>
(in part) Patterson &amp; Daugherty, 1997
</paragraph>
<paragraph id="8C0FCC0FFFCDFF21FF59FA20633BF685" blockId="3.[151,802,1295,1503]" box="[151,633,1447,1472]" pageId="3" pageNumber="392">
<taxonomicName id="4BB0B78CFFCDFF21FF59FA20600AF685" authorityName="Hardy" authorityYear="1977" box="[151,328,1448,1472]" class="Squamata" family="Scincidae" genus="Cyclodina" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCDFF21FF59FA20600AF685" box="[151,328,1448,1472]" italics="true" pageId="3" pageNumber="392">Cyclodina oliveri</emphasis>
</taxonomicName>
(in part) 
<bibRefCitation id="E821B1FEFFCDFF21FE69FA2F633BF685" author="Chapple, D. G. &amp; Patterson, G. B. &amp; Gleeson, D. M. &amp; Daugherty, C. H. &amp; Ritchie, P. A." box="[423,633,1447,1472]" pageId="3" pageNumber="392" pagination="129 - 146" refId="ref5441" refString="Chapple, D. G., Patterson, G. B., Gleeson, D. M., Daugherty, C. H. &amp; Ritchie, P. A. (2008 b) Taxonomic revision of the marbled skink (Cyclodina oliveri, Reptilia: Scincidae) species complex, with a description of a new species. New Zealand Journal of Zoology, 35, 129 - 146. https: // doi. org / 10.1080 / 03014220809510110" type="journal article" year="2008">
Chapple 
<emphasis id="BEC4101DFFCDFF21FDCFFA216377F685" box="[513,565,1448,1472]" italics="true" pageId="3" pageNumber="392">et al.</emphasis>
2008b
</bibRefCitation>
</paragraph>
<paragraph id="8C0FCC0FFFCDFF21FF59FA4F6337F69A" blockId="3.[151,802,1295,1503]" box="[151,629,1478,1503]" pageId="3" pageNumber="392">
<taxonomicName id="4BB0B78CFFCDFF21FF59FA4F600DF69A" box="[151,335,1479,1503]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCDFF21FF59FA4F600DF69A" box="[151,335,1479,1503]" italics="true" pageId="3" pageNumber="392">Oligosoma oliveri</emphasis>
</taxonomicName>
(in part) 
<bibRefCitation id="E821B1FEFFCDFF21FE60FA4E6337F69A" author="Chapple, D. G. &amp; Ritchie, P. A. &amp; Daugherty, C. H." box="[430,629,1478,1503]" pageId="3" pageNumber="392" pagination="470 - 487" refId="ref5526" refString="Chapple, D. G., Ritchie, P. A. &amp; Daugherty, C. H. (2009) Origin, diversification and systematics of the New Zealand skink fauna (Reptilia: Scincidae). Molecular Phylogenetics and Evolution, 52, 470 - 487. https: // doi. org / 10.1016 / j. ympev. 2009.03.021" type="journal article" year="2009">
Chapple 
<emphasis id="BEC4101DFFCDFF21FDC7FA4F637EF69A" box="[521,572,1479,1503]" italics="true" pageId="3" pageNumber="392">et al.</emphasis>
2009
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCDFF21FF59F98560CCF50E" pageId="3" pageNumber="392" type="materials_examined">
<paragraph id="8C0FCC0FFFCDFF21FF59F98560CCF50E" blockId="3.[151,1436,1548,2044]" pageId="3" pageNumber="392">
<materialsCitation id="3CD8C652FFCDFF21FF59F98560CCF50E" ID-GBIF-Occurrence="2429072699" collectingDate="1972-11" collectionCode="AM" collectorName="A. H. Whitaker." location="Ruamahua-nui Island" municipality="Aldermans" pageId="3" pageNumber="392" specimenCode="H538" specimenCount="1" typeStatus="holotype">
<emphasis id="BEC4101DFFCDFF21FF59F985604FF562" bold="true" box="[151,269,1549,1575]" pageId="3" pageNumber="392">
<typeStatus id="530B72ADFFCDFF21FF59F985604AF562" box="[151,264,1549,1575]" pageId="3" pageNumber="392" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="EAA154CAFFCDFF21FED9F9856006F562" box="[279,324,1549,1575]" country="Australia" name="Australian Museum" pageId="3" pageNumber="392" type="Museum">AM</collectionCode>
<specimenCode id="DC166474FFCDFF21FE80F98460CFF562" box="[334,397,1548,1575]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="3" pageNumber="392" type="Museum">H538</specimenCode>
(author’s number R67), adult female, 
<location id="896F9AD4FFCDFF21FC82F9856500F562" LSID="urn:lsid:plazi:treatment:04197D19FFCDFF24FF59FB2E62F9F2EA:896F9AD4FFCDFF21FC82F9856500F562" box="[844,1090,1549,1575]" municipality="Aldermans" name="Ruamahua-nui Island" pageId="3" pageNumber="392">Ruamahua-nui Island</location>
, 
<collectingMunicipality id="6C6B5675FFCDFF21FB81F9856588F562" box="[1103,1226,1549,1575]" pageId="3" pageNumber="392">Aldermans</collectingMunicipality>
group, 
<date id="F80EEACFFFCDFF21FAE8F9856190F50E" pageId="3" pageNumber="392" value="1972-11">
<collectingDate id="E84A1327FFCDFF21FAE8F9856190F50E" pageId="3" pageNumber="392" value="1972-11">November 1972</collectingDate>
</date>
, 
<collectorName id="2145A9D9FFCDFF21FF12F9B960CCF50E" box="[220,398,1585,1612]" pageId="3" pageNumber="392">A. H. Whitaker.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCDFF21FF09F9DC65ECF4B9" pageId="3" pageNumber="392" type="diagnosis">
<paragraph id="8C0FCC0FFFCDFF21FF09F9DC65ECF4B9" blockId="3.[151,1436,1548,2044]" pageId="3" pageNumber="392">
<emphasis id="BEC4101DFFCDFF21FF09F9DC6000F52A" bold="true" box="[199,322,1620,1647]" pageId="3" pageNumber="392">Diagnosis.</emphasis>
An 
<taxonomicName id="4BB0B78CFFCDFF21FEBEF9DD60A9F52A" authorityName="Girard" authorityYear="1857" box="[368,491,1621,1647]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="genus">
<emphasis id="BEC4101DFFCDFF21FEBEF9DD60A9F52A" box="[368,491,1621,1647]" italics="true" pageId="3" pageNumber="392">Oligosoma</emphasis>
</taxonomicName>
with a squarish mid-body cross-section; opaque, divided palpebral disc; a prominent tear-drop mark beneath each eye; iris light brown to orange; mid-body scale rows 32–36; ventral scale rows 72–88; eye diameter&gt; distance from eye to mouth; adult SVL ≤ 
<quantity id="4B4861EAFFCDFF21FCC1F9156223F5FD" box="[783,865,1693,1720]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="3" pageNumber="392" unit="mm" value="84.0">84 mm</quantity>
; dorsal markings dominated by fine black flecking; belly cream and marked with numerous individual fine dark flecks. Distinguished from similar congeners (which share both the squarish body cross-section and the tear-drop subocular marking) as follows: 
<taxonomicName id="4BB0B78CFFCDFF21FB52F96D6594F5BA" baseAuthorityName="Robb" baseAuthorityYear="1970" box="[1180,1238,1765,1791]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="alani">
<emphasis id="BEC4101DFFCDFF21FB52F96D6594F5BA" box="[1180,1238,1765,1791]" italics="true" pageId="3" pageNumber="392">alani</emphasis>
</taxonomicName>
(sympatric) has a black iris, prominent dorsal blotches and is much larger at up to 
<quantity id="4B4861EAFFCDFF21FCABF8816280F461" box="[869,962,1801,1828]" metricMagnitude="-1" metricUnit="m" metricValue="1.42" pageId="3" pageNumber="392" unit="mm" value="142.0">142 mm</quantity>
SVL; 
<emphasis id="BEC4101DFFCDFF21FBC2F88265CCF466" box="[1036,1166,1802,1827]" italics="true" pageId="3" pageNumber="392">macgregori</emphasis>
has longitudinal streaks on the dorsum and a uniform belly; 
<emphasis id="BEC4101DFFCDFF21FDE6F8A663C5F402" box="[552,647,1838,1863]" italics="true" pageId="3" pageNumber="392">ornatum</emphasis>
(potentially sympatric) is smaller (SVL usually &lt;
<quantity id="4B4861EAFFCDFF21FB74F8A5644EF40D" box="[1210,1292,1837,1864]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="392" unit="mm" value="80.0">80 mm</quantity>
), usually has a yellow belly and usually has a tapering pale dorso-lateral stripe; 
<taxonomicName id="4BB0B78CFFCDFF21FC43F8D96294F42E" box="[909,982,1873,1899]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCDFF21FC43F8D96294F42E" box="[909,982,1873,1899]" italics="true" pageId="3" pageNumber="392">oliveri</emphasis>
</taxonomicName>
has 36–44 mid-body scale rows, 88–97 ventral scale rows, dorsal and ventral patterns dominated by pale blotches, and a smaller eye (diameter &lt;distance from eye to mouth); 
<taxonomicName id="4BB0B78CFFCDFF21FE49F812609DF4F6" box="[391,479,1946,1971]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="roimata">
<emphasis id="BEC4101DFFCDFF21FE49F812609DF4F6" box="[391,479,1946,1971]" italics="true" pageId="3" pageNumber="392">roimata</emphasis>
</taxonomicName>
is smaller at up to 
<quantity id="4B4861EAFFCDFF21FD71F8106252F4F1" box="[703,784,1944,1972]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="392" unit="mm" value="65.0">65 mm</quantity>
SVL and has only 1 primary temporal scale (versus 
<quantity id="4B4861EAFFCDFF21FAA1F81164DEF4F6" box="[1391,1436,1945,1971]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="392" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4BB0B78CFFCDFF21FF59F8356011F492" authorityName="Wells &amp; Wellington" authorityYear="1985" box="[151,339,1981,2007]" class="Squamata" family="Scincidae" genus="Cyclodina" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCDFF21FF59F8356011F492" box="[151,339,1981,2007]" italics="true" pageId="3" pageNumber="392">pachysomaticum</emphasis>
</taxonomicName>
); 
<taxonomicName id="4BB0B78CFFCDFF21FEA3F83660F4F492" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[365,438,1982,2007]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCDFF21FEA3F83660F4F492" box="[365,438,1982,2007]" italics="true" pageId="3" pageNumber="392">townsi</emphasis>
</taxonomicName>
usually has only 1 primary temporal scale and has few ventral markings; and 
<taxonomicName id="4BB0B78CFFCDFF21FAFCF83564DEF492" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[1330,1436,1981,2007]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCDFF21FAFCF83564DEF492" box="[1330,1436,1981,2007]" italics="true" pageId="3" pageNumber="392">whitakeri</emphasis>
</taxonomicName>
(sympatric) is larger at up to 
<quantity id="4B4861EAFFCDFF21FE12F869637BF4B9" box="[476,569,2017,2044]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="3" pageNumber="392" unit="mm" value="101.0">101 mm</quantity>
SVL, has a dark-brown iris and a yellow-orange venter.
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCAFF26FF09FF1161ABF3BF" pageId="4" pageNumber="393" type="etymology">
<paragraph id="8C0FCC0FFFCAFF26FF09FF1161ABF3BF" blockId="4.[151,1437,152,647]" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FF09FF11600AF3F6" bold="true" box="[199,328,153,179]" pageId="4" pageNumber="393">Etymology</emphasis>
. From the Greek 
<emphasis id="BEC4101DFFCAFF26FDC3FF106319F3F7" box="[525,603,152,178]" italics="true" pageId="4" pageNumber="393">pachys</emphasis>
(thick) and 
<emphasis id="BEC4101DFFCAFF26FD2FFF11625EF3F7" box="[737,796,153,178]" italics="true" pageId="4" pageNumber="393">soma</emphasis>
(body). In the absence of an established vernacular name for 
<taxonomicName id="4BB0B78CFFCAFF26FF71FF3460BFF393" box="[191,509,188,214]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FF71FF3460BFF393" box="[191,509,188,214]" italics="true" pageId="4" pageNumber="393">Oligosoma pachysomaticum</emphasis>
</taxonomicName>
I suggest ‘Coromandel skink’ to reflect the general location of the surviving populations.
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCAFF26FF09FE8C653BF222" pageId="4" pageNumber="393" type="distribution">
<paragraph id="8C0FCC0FFFCAFF26FF09FE8C653BF222" blockId="4.[151,1437,152,647]" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FF09FE8C6023F25A" bold="true" box="[199,353,260,287]" pageId="4" pageNumber="393">Distribution.</emphasis>
Known from a total of eight islands in the Mercury Group (Middle, Green and Korapuki Is.), Alderman Group (Half, Hongiora, Ruamahuanui and Ruamahuati Is.) and the Ohinau Group (Old Man Rock), all off the east coast of the Coromandel Peninsula (
<figureCitation id="148BD08AFFCAFF26FD61FEC46252F222" box="[687,784,332,359]" captionStart="FIGURE 1" captionStartId="1.[151,250,1525,1550]" captionTargetBox="[151,1436,301,1506]" captionTargetId="figure@1.[151,1436,301,1506]" captionTargetPageId="1" captionText="FIGURE 1. Map of upper North Island of New Zealand showing distribution records for extant populations of the four members of the Oligosoma oliveri species-complex. Bold outline = maximum expanse of land during the Otira Glaciation 18,000–20,000 years ago; thin outline = present extent of land; dotted line = former path of Waikato River; blue shading = suggested former distributional extent of O. townsi in this region; pink shading = suggested former distributional extent of O. pachysomaticum in this region." figureDoi="http://doi.org/10.5281/zenodo.3516817" httpUri="https://zenodo.org/record/3516817/files/figure.png" pageId="4" pageNumber="393">Figure 1</figureCitation>
; 
<tableCitation id="C132F9B4FFCAFF26FCD3FEC46233F222" box="[797,881,332,359]" captionStart="TABLE 1" captionStartId="4.[151,239,694,718]" captionTargetPageId="4" captionText="TABLE 1. Extant populations of the Oligosoma oliveri species-complex. Data from Towns (1999), Chapple et al. (2008b) and van Winkle et al. (2018)." httpUri="http://table.plazi.org/id/D8CF9C87FFCAFF26FF59FD3E608DF1B7" pageId="4" pageNumber="393" tableUuid="D8CF9C87FFCAFF26FF59FD3E608DF1B7">Table 1</tableCitation>
; 
<bibRefCitation id="E821B1FEFFCAFF26FCB1FEC4652FF222" author="Chapple, D. G. &amp; Patterson, G. B. &amp; Gleeson, D. M. &amp; Daugherty, C. H. &amp; Ritchie, P. A." box="[895,1133,332,359]" pageId="4" pageNumber="393" pagination="129 - 146" refId="ref5441" refString="Chapple, D. G., Patterson, G. B., Gleeson, D. M., Daugherty, C. H. &amp; Ritchie, P. A. (2008 b) Taxonomic revision of the marbled skink (Cyclodina oliveri, Reptilia: Scincidae) species complex, with a description of a new species. New Zealand Journal of Zoology, 35, 129 - 146. https: // doi. org / 10.1080 / 03014220809510110" type="journal article" year="2008">
Chapple 
<emphasis id="BEC4101DFFCAFF26FC2DFEC56559F223" box="[995,1051,332,358]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
2008b
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFCAFF24FF09FEF9636CF392" lastPageId="6" lastPageNumber="395" pageId="4" pageNumber="393" type="biology_ecology">
<paragraph id="8C0FCC0FFFCAFF26FF09FEF96442F1C2" blockId="4.[151,1437,152,647]" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FF09FEF9606EF2CE" bold="true" box="[199,300,369,395]" pageId="4" pageNumber="393">Ecology.</emphasis>
The following information is based on a study of the species on Green Island in the Mercury group by 
<bibRefCitation id="E821B1FEFFCAFF26FF74FE1C6028F2EA" author="Southey, I. C." box="[186,362,404,431]" pageId="4" pageNumber="393" refId="ref6314" refString="Southey, I. C. (1985) The ecology of three rare skinks on Middle Island, Mercury Island. Unpublished Thesis, University of Auckland, Auckland, 99 pp." type="book" year="1985">Southey (1985)</bibRefCitation>
. Lives primarily among deep leaf litter on the floor of coastal forest and scrub, being most abundant among regenerating scrub where the leaf litter is deepest. Nocturnal with peak activity in the first few hours of darkness, and only rarely sunbasks or forages by day. Co-exists in this habitat with two much larger skinks, 
<taxonomicName id="4BB0B78CFFCAFF26FA9DFE546056F15F" baseAuthorityName="Robb" baseAuthorityYear="1970" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="alani">
<emphasis id="BEC4101DFFCAFF26FA9DFE546056F15F" italics="true" pageId="4" pageNumber="393">Oligosoma alani</emphasis>
</taxonomicName>
(≤ 
<quantity id="4B4861EAFFCAFF26FEF2FD8960D8F15E" box="[316,410,513,539]" metricMagnitude="-1" metricUnit="m" metricValue="1.42" pageId="4" pageNumber="393" unit="mm" value="142.0">142 mm</quantity>
SVL) and 
<taxonomicName id="4BB0B78CFFCAFF26FDD7FD8963E4F15F" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[537,678,512,539]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCAFF26FDD7FD8963E4F15F" box="[537,678,512,539]" italics="true" pageId="4" pageNumber="393">O. whitakeri</emphasis>
</taxonomicName>
(≤ 
<quantity id="4B4861EAFFCAFF26FD00FD89626EF15E" box="[718,812,513,539]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="4" pageNumber="393" unit="mm" value="101.0">101 mm</quantity>
SVL) and is significantly less abundant than these, i.e. 
<taxonomicName id="4BB0B78CFFCAFF26FF59FDAD6034F17B" box="[151,374,548,575]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FF59FDAD6034F17B" box="[151,374,548,575]" italics="true" pageId="4" pageNumber="393">O. pachysomaticum</emphasis>
</taxonomicName>
number 46–91/ha, 
<taxonomicName id="4BB0B78CFFCAFF26FD9CFDAD63EDF17B" baseAuthorityName="Robb" baseAuthorityYear="1970" box="[594,687,548,575]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="alani">
<emphasis id="BEC4101DFFCAFF26FD9CFDAD63EDF17B" box="[594,687,548,575]" italics="true" pageId="4" pageNumber="393">O. alani</emphasis>
</taxonomicName>
277–779/ha and 
<taxonomicName id="4BB0B78CFFCAFF26FCBDFDAD6542F17B" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[883,1024,548,575]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCAFF26FCBDFDAD6542F17B" box="[883,1024,548,575]" italics="true" pageId="4" pageNumber="393">O. whitakeri</emphasis>
</taxonomicName>
161–903/ha. Also co-exists with the smaller forest/coastal scrub species 
<taxonomicName id="4BB0B78CFFCAFF26FDEAFDC163BDF126" authority="(Girard)" authorityName="Girard" baseAuthorityName="Girard" baseAuthorityYear="1858" box="[548,767,584,611]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="aeneum">
<emphasis id="BEC4101DFFCAFF26FDEAFDC163DFF127" box="[548,669,585,611]" italics="true" pageId="4" pageNumber="393">O. aeneum</emphasis>
(Girard)
</taxonomicName>
(
<quantity id="4B4861EAFFCAFF26FCC2FDC06222F126" box="[780,864,584,611]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="4" pageNumber="393" unit="mm" value="76.0">76 mm</quantity>
). Between 2–4 young are produced in April. A wide variety of invertebrates are preyed on as well as some fruit. Predators include centipedes and tuatara.
</paragraph>
<caption id="D8CF9C87FFCAFF26FF59FD3E608DF1B7" ID-Table-UUID="D8CF9C87FFCAFF26FF59FD3E608DF1B7" httpUri="http://table.plazi.org/id/D8CF9C87FFCAFF26FF59FD3E608DF1B7" pageId="4" pageNumber="393" targetBox="[166,1344,765,2000]" targetIsTable="true" targetPageId="4">
<paragraph id="8C0FCC0FFFCAFF26FF59FD3E608DF1B7" blockId="4.[151,1436,694,754]" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FF59FD3E604BF18B" bold="true" box="[151,265,694,718]" pageId="4" pageNumber="393">TABLE 1.</emphasis>
Extant populations of the 
<taxonomicName id="4BB0B78CFFCAFF26FDD3FD3E639FF18B" box="[541,733,694,718]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDD3FD3E639FF18B" box="[541,733,694,718]" italics="true" pageId="4" pageNumber="393">Oligosoma oliveri</emphasis>
</taxonomicName>
species-complex. Data from 
<bibRefCitation id="E821B1FEFFCAFF26FBDFFD3F65E4F18B" author="Towns, D. R." box="[1041,1190,694,718]" pageId="4" pageNumber="393" refId="ref6348" refString="Towns, D. R. (1999) Cyclodina spp. skink recovery plan, 1999 - 2004. New Zealand Department of Conservation, Wellington, 69 pp. Available from: https: // www. doc. govt. nz / Documents / conservation / native-animals / reptiles-and-frogs / Cyclodinaskinkrecoveryplan. pdf (Accessed 22 Oct. 2019)" type="book" year="1999">Towns (1999)</bibRefCitation>
, 
<bibRefCitation id="E821B1FEFFCAFF26FB7EFD3E64DEF18B" author="Chapple, D. G. &amp; Patterson, G. B. &amp; Gleeson, D. M. &amp; Daugherty, C. H. &amp; Ritchie, P. A." box="[1200,1436,694,718]" pageId="4" pageNumber="393" pagination="129 - 146" refId="ref5441" refString="Chapple, D. G., Patterson, G. B., Gleeson, D. M., Daugherty, C. H. &amp; Ritchie, P. A. (2008 b) Taxonomic revision of the marbled skink (Cyclodina oliveri, Reptilia: Scincidae) species complex, with a description of a new species. New Zealand Journal of Zoology, 35, 129 - 146. https: // doi. org / 10.1080 / 03014220809510110" type="journal article" year="2008">
Chapple 
<emphasis id="BEC4101DFFCAFF26FAC0FD3F6401F18B" box="[1294,1347,694,718]" italics="true" pageId="4" pageNumber="393">et al.</emphasis>
(2008b)
</bibRefCitation>
and 
<bibRefCitation id="E821B1FEFFCAFF26FF0AFD53608EF1B7" author="van Winkle, D. &amp; Baling, M. &amp; Hitchmough, R." box="[196,460,730,754]" pageId="4" pageNumber="393" refId="ref6411" refString="van Winkle, D., Baling, M. &amp; Hitchmough, R. (2018) Reptiles and amphibians of New Zealand, a field guide. Auckland University Press, Auckland, 366 pp." type="book" year="2018">
van Winkle 
<emphasis id="BEC4101DFFCAFF26FE8BFD536036F1B7" box="[325,372,730,754]" italics="true" pageId="4" pageNumber="393">et al</emphasis>
. (2018)
</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8C0FCC0FFFCAFF26FF68FD7662ABF495" pageId="4" pageNumber="393">
<table id="FEB03EAFFFCA00DDFF68FD756402F495" box="[166,1344,765,2000]" gridcols="4" gridrows="33" pageId="4" pageNumber="393">
<tr id="3280CE4DFFCA00DDFF68FD756402F053" box="[166,1344,765,790]" gridrow="0" pageId="4" pageNumber="393">
<th id="7151A731FFCA00DDFF68FD756006F053" box="[166,324,765,790]" gridcol="0" gridrow="0" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FF68FD766072F053" bold="true" box="[166,304,766,790]" pageId="4" pageNumber="393">Island group</emphasis>
</th>
<th id="7151A731FFCA00DDFE4AFD756364F053" box="[388,550,765,790]" gridcol="1" gridrow="0" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FE4AFD766353F053" bold="true" box="[388,529,765,790]" pageId="4" pageNumber="393">Island or site</emphasis>
</th>
<th id="7151A731FFCA00DDFDA0FD75625AF053" box="[622,792,765,790]" gridcol="2" gridrow="0" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FDA0FD7563FFF053" bold="true" box="[622,701,765,790]" pageId="4" pageNumber="393">Species</emphasis>
</th>
<th id="7151A731FFCA00DDFCAAFD756402F053" box="[868,1344,765,790]" gridcol="3" gridrow="0" pageId="4" pageNumber="393">
<emphasis id="BEC4101DFFCAFF26FCAAFD7662E2F053" bold="true" box="[868,928,766,790]" pageId="4" pageNumber="393">Notes</emphasis>
</th>
</tr>
<tr id="3280CE4DFFCA00DDFF68FCAC6402F078" box="[166,1344,804,829]" gridrow="1" pageId="4" pageNumber="393" rowspan-3="1">
<th id="7151A731FFCA00DDFF68FCAC6006F078" box="[166,324,804,829]" gridcol="0" gridrow="1" pageId="4" pageNumber="393">Alderman</th>
<td id="7151A731FFCA00DDFE4AFCAC6364F078" box="[388,550,804,829]" gridcol="1" gridrow="1" pageId="4" pageNumber="393">Half I.</td>
<td id="7151A731FFCA00DDFDA0FCAC625AF078" box="[622,792,804,829]" gridcol="2" gridrow="1" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FCAC625AF079" box="[622,792,804,828]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FCAC625AF079" box="[622,792,804,828]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FCC16402F027" box="[166,1344,841,866]" gridrow="2" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AFCC16364F027" box="[388,550,841,866]" gridcol="1" gridrow="2" pageId="4" pageNumber="393">Hongiora I.</td>
<td id="7151A731FFCA00DDFDA0FCC1625AF027" box="[622,792,841,866]" gridcol="2" gridrow="2" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FCC2625AF027" box="[622,792,842,866]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FCC2625AF027" box="[622,792,842,866]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FCE76402F0CD" box="[166,1344,879,904]" gridrow="3" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AFCE76364F0CD" box="[388,550,879,904]" gridcol="1" gridrow="3" pageId="4" pageNumber="393">Ruamahuanui I.</td>
<td id="7151A731FFCA00DDFDA0FCE7625AF0CD" box="[622,792,879,904]" gridcol="2" gridrow="3" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FCF8625AF0CD" box="[622,792,880,904]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FCF8625AF0CD" box="[622,792,880,904]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFCE76402F0CD" box="[868,1344,879,904]" gridcol="3" gridrow="3" pageId="4" pageNumber="393">Type locality</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FC1D6402F0EB" box="[166,1344,917,942]" gridrow="4" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AFC1D6364F0EB" box="[388,550,917,942]" gridcol="1" gridrow="4" pageId="4" pageNumber="393">Ruamahuaiti I.</td>
<td id="7151A731FFCA00DDFDA0FC1D625AF0EB" box="[622,792,917,942]" gridcol="2" gridrow="4" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FC1E625AF0EB" box="[622,792,918,942]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FC1E625AF0EB" box="[622,792,918,942]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FC336402F091" box="[166,1344,955,980]" gridrow="5" pageId="4" pageNumber="393" rowspan-3="1">
<th id="7151A731FFCA00DDFF68FC336006F091" box="[166,324,955,980]" gridcol="0" gridrow="5" pageId="4" pageNumber="393">Barrier</th>
<td id="7151A731FFCA00DDFE4AFC336364F091" box="[388,550,955,980]" gridcol="1" gridrow="5" pageId="4" pageNumber="393">Great Barrier I.</td>
<td id="7151A731FFCA00DDFDA0FC33625AF091" box="[622,792,955,980]" gridcol="2" gridrow="5" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FC3463F2F096" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,956,979]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FC3463F2F096" box="[622,688,956,979]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FC696402F0BF" box="[166,1344,993,1018]" gridrow="6" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AFC696364F0BF" box="[388,550,993,1018]" gridcol="1" gridrow="6" pageId="4" pageNumber="393">Little Barrier I.</td>
<td id="7151A731FFCA00DDFDA0FC69625AF0BF" box="[622,792,993,1018]" gridcol="2" gridrow="6" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FC6A63F2F0BC" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,994,1017]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FC6A63F2F0BC" box="[622,688,994,1017]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FB8E6402F75A" box="[166,1344,1030,1055]" gridrow="7" pageId="4" pageNumber="393">
<th id="7151A731FFCA00DDFF68FB8E6006F75A" box="[166,324,1030,1055]" gridcol="0" gridrow="7" pageId="4" pageNumber="393">Hen &amp; Chicken</th>
<td id="7151A731FFCA00DDFE4AFB8E6364F75A" box="[388,550,1030,1055]" gridcol="1" gridrow="7" pageId="4" pageNumber="393">Coppermine I.</td>
<td id="7151A731FFCA00DDFDA0FB8E625AF75A" box="[622,792,1030,1055]" gridcol="2" gridrow="7" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FB8063F2F75A" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1032,1055]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FB8063F2F75A" box="[622,688,1032,1055]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFB8E6402F75A" box="[868,1344,1030,1055]" gridcol="3" gridrow="7" pageId="4" pageNumber="393">ex. Middle Stack, 2002</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FBA46402F700" box="[166,1344,1068,1093]" gridrow="8" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AFBA46364F700" box="[388,550,1068,1093]" gridcol="1" gridrow="8" pageId="4" pageNumber="393">Lady Alice</td>
<td id="7151A731FFCA00DDFDA0FBA4625AF700" box="[622,792,1068,1093]" gridcol="2" gridrow="8" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FBA663F2F700" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1070,1093]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FBA663F2F700" box="[622,688,1070,1093]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFBA46402F700" box="[868,1344,1068,1093]" gridcol="3" gridrow="8" pageId="4" pageNumber="393">ex Muriwhenua I., 1997 &amp; 2005</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FBDA6402F72E" box="[166,1344,1106,1131]" gridrow="9" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AFBDA6364F72E" box="[388,550,1106,1131]" gridcol="1" gridrow="9" pageId="4" pageNumber="393">Muriwhenua I.</td>
<td id="7151A731FFCA00DDFDA0FBDA625AF72E" box="[622,792,1106,1131]" gridcol="2" gridrow="9" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FBDB63F2F72F" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1107,1130]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FBDB63F2F72F" box="[622,688,1107,1130]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FBF06402F7D4" box="[166,1344,1144,1169]" gridrow="10" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AFBF06364F7D4" box="[388,550,1144,1169]" gridcol="1" gridrow="10" pageId="4" pageNumber="393">Middle Stack I.</td>
<td id="7151A731FFCA00DDFDA0FBF0625AF7D4" box="[622,792,1144,1169]" gridcol="2" gridrow="10" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FBF163F2F7D5" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1145,1168]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FBF163F2F7D5" box="[622,688,1145,1168]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FB166402F7F2" box="[166,1344,1182,1207]" gridrow="11" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AFB166364F7F2" box="[388,550,1182,1207]" gridcol="1" gridrow="11" pageId="4" pageNumber="393">Pupuha I.</td>
<td id="7151A731FFCA00DDFDA0FB16625AF7F2" box="[622,792,1182,1207]" gridcol="2" gridrow="11" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FB1763F2F7F3" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1183,1206]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FB1763F2F7F3" box="[622,688,1183,1206]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFB166402F7F2" box="[868,1344,1182,1207]" gridcol="3" gridrow="11" pageId="4" pageNumber="393">Type locality</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FB4B6402F799" box="[166,1344,1219,1244]" gridrow="12" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AFB4B6364F799" box="[388,550,1219,1244]" gridcol="1" gridrow="12" pageId="4" pageNumber="393">Whareware I.</td>
<td id="7151A731FFCA00DDFDA0FB4B625AF799" box="[622,792,1219,1244]" gridcol="2" gridrow="12" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FB4D63F2F799" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1221,1244]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FB4D63F2F799" box="[622,688,1221,1244]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FB616402F647" box="[166,1344,1257,1282]" gridrow="13" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AFB616364F647" box="[388,550,1257,1282]" gridcol="1" gridrow="13" pageId="4" pageNumber="393">Whatpuki I.</td>
<td id="7151A731FFCA00DDFDA0FB61625AF647" box="[622,792,1257,1282]" gridcol="2" gridrow="13" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FB6363F2F647" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1259,1282]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0FB6363F2F647" box="[622,688,1259,1282]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFB616402F647" box="[868,1344,1257,1282]" gridcol="3" gridrow="13" pageId="4" pageNumber="393">ex. Middle Stack, 2000</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FA876402F66D" box="[166,1344,1295,1320]" gridrow="14" pageId="4" pageNumber="393">
<th id="7151A731FFCA00DDFF68FA876006F66D" box="[166,324,1295,1320]" gridcol="0" gridrow="14" pageId="4" pageNumber="393">North I.</th>
<td id="7151A731FFCA00DDFE4AFA876364F66D" box="[388,550,1295,1320]" gridcol="1" gridrow="14" pageId="4" pageNumber="393">Pukarua Bay</td>
<td id="7151A731FFCA00DDFDA0FA87625AF66D" box="[622,792,1295,1320]" gridcol="2" gridrow="14" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FA98638CF66D" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[622,718,1296,1320]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCAFF26FDA0FA98638CF66D" box="[622,718,1296,1320]" italics="true" pageId="4" pageNumber="393">whitakeri</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFA876402F66D" box="[868,1344,1295,1320]" gridcol="3" gridrow="14" pageId="4" pageNumber="393">small captive colony (probably extinct in wild)</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FABD6402F60B" box="[166,1344,1333,1358]" gridrow="15" pageId="4" pageNumber="393" rowspan-3="1">
<th id="7151A731FFCA00DDFF68FABD6006F60B" box="[166,324,1333,1358]" gridcol="0" gridrow="15" pageId="4" pageNumber="393">Mercury</th>
<td id="7151A731FFCA00DDFE4AFABD6364F60B" box="[388,550,1333,1358]" gridcol="1" gridrow="15" pageId="4" pageNumber="393">Middle I.</td>
<td id="7151A731FFCA00DDFDA0FABD625AF60B" box="[622,792,1333,1358]" gridcol="2" gridrow="15" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FABD625AF608" box="[622,792,1333,1357]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FABD625AF608" box="[622,792,1333,1357]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FAD36402F631" box="[166,1344,1371,1396]" gridrow="16" pageId="4" pageNumber="393" rowspan-0="1" rowspan-1="1">
<td id="7151A731FFCA00DDFDA0FAD3625AF631" box="[622,792,1371,1396]" gridcol="2" gridrow="16" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FAD3638CF631" box="[622,718,1371,1396]" pageId="4" pageNumber="393" rank="species" species="whitakeri">whitakeri</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFAD36402F631" box="[868,1344,1371,1396]" gridcol="3" gridrow="16" pageId="4" pageNumber="393">Type locality</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FA096402F6DC" box="[166,1344,1409,1433]" gridrow="17" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AFA096364F6DC" box="[388,550,1409,1433]" gridcol="1" gridrow="17" pageId="4" pageNumber="393">Green I.</td>
<td id="7151A731FFCA00DDFDA0FA09625AF6DC" box="[622,792,1409,1433]" gridcol="2" gridrow="17" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FA09625AF6DC" box="[622,792,1409,1433]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FA09625AF6DC" box="[622,792,1409,1433]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FA2E6402F6FA" box="[166,1344,1446,1471]" gridrow="18" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AFA2E6364F6FA" box="[388,550,1446,1471]" gridcol="1" gridrow="18" pageId="4" pageNumber="393">Korapuki I.</td>
<td id="7151A731FFCA00DDFDA0FA2E625AF6FA" box="[622,792,1446,1471]" gridcol="2" gridrow="18" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FA2F625AF6FA" box="[622,792,1447,1471]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FA2F625AF6FA" box="[622,792,1447,1471]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFA2E6402F6FA" box="[868,1344,1446,1471]" gridcol="3" gridrow="18" pageId="4" pageNumber="393">ex Green I., 1988</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FA446402F6A0" box="[166,1344,1484,1509]" gridrow="19" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AFA446364F6A0" box="[388,550,1484,1509]" gridcol="1" gridrow="19" pageId="4" pageNumber="393">Red Mercury I.</td>
<td id="7151A731FFCA00DDFDA0FA44625AF6A0" box="[622,792,1484,1509]" gridcol="2" gridrow="19" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FA45625AF6A0" box="[622,792,1485,1509]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0FA45625AF6A0" box="[622,792,1485,1509]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFA446402F6A0" box="[868,1344,1484,1509]" gridcol="3" gridrow="19" pageId="4" pageNumber="393">survival doubtful</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68FA7A6402F54E" box="[166,1344,1522,1547]" gridrow="20" pageId="4" pageNumber="393" rowspan-0="1" rowspan-1="1">
<td id="7151A731FFCA00DDFDA0FA7A625AF54E" box="[622,792,1522,1547]" gridcol="2" gridrow="20" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0FA7A638CF54F" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[622,718,1522,1546]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCAFF26FDA0FA7A638CF54F" box="[622,718,1522,1546]" italics="true" pageId="4" pageNumber="393">whitakeri</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAFA7A6402F54E" box="[868,1344,1522,1547]" gridcol="3" gridrow="20" pageId="4" pageNumber="393">ex. Middle Island</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F9906402F574" box="[166,1344,1560,1585]" gridrow="21" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AF9906364F574" box="[388,550,1560,1585]" gridcol="1" gridrow="21" pageId="4" pageNumber="393">Stanely I.</td>
<td id="7151A731FFCA00DDFDA0F990625AF574" box="[622,792,1560,1585]" gridcol="2" gridrow="21" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F990638CF575" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[622,718,1560,1584]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCAFF26FDA0F990638CF575" box="[622,718,1560,1584]" italics="true" pageId="4" pageNumber="393">whitakeri</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAF9906402F574" box="[868,1344,1560,1585]" gridcol="3" gridrow="21" pageId="4" pageNumber="393">ex. Middle Island</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F9B56402F513" box="[166,1344,1597,1622]" gridrow="22" pageId="4" pageNumber="393" rowspan-3="1">
<th id="7151A731FFCA00DDFF68F9B56006F513" box="[166,324,1597,1622]" gridcol="0" gridrow="22" pageId="4" pageNumber="393">nr Coromandel</th>
<td id="7151A731FFCA00DDFE4AF9B56364F513" box="[388,550,1597,1622]" gridcol="1" gridrow="22" pageId="4" pageNumber="393">Castle I.</td>
<td id="7151A731FFCA00DDFDA0F9B5625AF513" box="[622,792,1597,1622]" gridcol="2" gridrow="22" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F9B6638CF513" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[622,718,1598,1622]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCAFF26FDA0F9B6638CF513" box="[622,718,1598,1622]" italics="true" pageId="4" pageNumber="393">whitakeri</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F9EB6402F539" box="[166,1344,1635,1660]" gridrow="23" pageId="4" pageNumber="393" rowspan-3="1">
<th id="7151A731FFCA00DDFF68F9EB6006F539" box="[166,324,1635,1660]" gridcol="0" gridrow="23" pageId="4" pageNumber="393">Mokohinau</th>
<td id="7151A731FFCA00DDFE4AF9EB6364F539" box="[388,550,1635,1660]" gridcol="1" gridrow="23" pageId="4" pageNumber="393">Tatapihi I.</td>
<td id="7151A731FFCA00DDFDA0F9EB625AF539" box="[622,792,1635,1660]" gridcol="2" gridrow="23" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F9ED63F2F539" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1637,1660]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0F9ED63F2F539" box="[622,688,1637,1660]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F9016402F5E7" box="[166,1344,1673,1698]" gridrow="24" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AF9016364F5E7" box="[388,550,1673,1698]" gridcol="1" gridrow="24" pageId="4" pageNumber="393">Stack ‘H’</td>
<td id="7151A731FFCA00DDFDA0F901625AF5E7" box="[622,792,1673,1698]" gridcol="2" gridrow="24" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F90263F2F5E4" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[622,688,1674,1697]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFCAFF26FDA0F90263F2F5E4" box="[622,688,1674,1697]" italics="true" pageId="4" pageNumber="393">townsi</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F9276402F58D" box="[166,1344,1711,1736]" gridrow="25" pageId="4" pageNumber="393" rowspan-3="1">
<th id="7151A731FFCA00DDFF68F9276006F58D" box="[166,324,1711,1736]" gridcol="0" gridrow="25" pageId="4" pageNumber="393">Ohinau</th>
<td id="7151A731FFCA00DDFE4AF9276364F58D" box="[388,550,1711,1736]" gridcol="1" gridrow="25" pageId="4" pageNumber="393">Old Man Rk</td>
<td id="7151A731FFCA00DDFDA0F927625AF58D" box="[622,792,1711,1736]" gridcol="2" gridrow="25" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F927625AF582" box="[622,792,1711,1735]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCAFF26FDA0F927625AF582" box="[622,792,1711,1735]" italics="true" pageId="4" pageNumber="393">pachysomaticum</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F95D6402F5AB" box="[166,1344,1749,1774]" gridrow="26" pageId="4" pageNumber="393" rowspan-3="1">
<th id="7151A731FFCA00DDFF68F95D6006F5AB" box="[166,324,1749,1774]" gridcol="0" gridrow="26" pageId="4" pageNumber="393">Poor Knights</th>
<td id="7151A731FFCA00DDFE4AF95D6364F5AB" box="[388,550,1749,1774]" gridcol="1" gridrow="26" pageId="4" pageNumber="393">Aorangi I.</td>
<td id="7151A731FFCA00DDFDA0F95D625AF5AB" box="[622,792,1749,1774]" gridcol="2" gridrow="26" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F95D63F2F5A8" box="[622,688,1749,1773]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDA0F95D63F2F5A8" box="[622,688,1749,1773]" italics="true" pageId="4" pageNumber="393">oliveri</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F9726402F456" box="[166,1344,1786,1811]" gridrow="27" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AF9726364F456" box="[388,550,1786,1811]" gridcol="1" gridrow="27" pageId="4" pageNumber="393">Archway I.</td>
<td id="7151A731FFCA00DDFDA0F972625AF456" box="[622,792,1786,1811]" gridcol="2" gridrow="27" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F97363F2F456" box="[622,688,1787,1811]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDA0F97363F2F456" box="[622,688,1787,1811]" italics="true" pageId="4" pageNumber="393">oliveri</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F8A86402F47C" box="[166,1344,1824,1849]" gridrow="28" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AF8A86364F47C" box="[388,550,1824,1849]" gridcol="1" gridrow="28" pageId="4" pageNumber="393">Motu Kapiti I.</td>
<td id="7151A731FFCA00DDFDA0F8A8625AF47C" box="[622,792,1824,1849]" gridcol="2" gridrow="28" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F8A963F2F47C" box="[622,688,1825,1849]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDA0F8A963F2F47C" box="[622,688,1825,1849]" italics="true" pageId="4" pageNumber="393">oliveri</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F8CE6402F41A" box="[166,1344,1862,1887]" gridrow="29" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AF8CE6364F41A" box="[388,550,1862,1887]" gridcol="1" gridrow="29" pageId="4" pageNumber="393">“Stack A”</td>
<td id="7151A731FFCA00DDFDA0F8CE625AF41A" box="[622,792,1862,1887]" gridcol="2" gridrow="29" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F8CF63F2F41A" box="[622,688,1863,1887]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDA0F8CF63F2F41A" box="[622,688,1863,1887]" italics="true" pageId="4" pageNumber="393">oliveri</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F8E46402F4C0" box="[166,1344,1900,1925]" gridrow="30" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AF8E46364F4C0" box="[388,550,1900,1925]" gridcol="1" gridrow="30" pageId="4" pageNumber="393">“Stack B”</td>
<td id="7151A731FFCA00DDFDA0F8E4625AF4C0" box="[622,792,1900,1925]" gridcol="2" gridrow="30" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F8E463F2F4C1" box="[622,688,1900,1924]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDA0F8E463F2F4C1" box="[622,688,1900,1924]" italics="true" pageId="4" pageNumber="393">oliveri</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F8196402F4EF" box="[166,1344,1937,1962]" gridrow="31" pageId="4" pageNumber="393" rowspan-0="1" rowspan-3="1">
<td id="7151A731FFCA00DDFE4AF8196364F4EF" box="[388,550,1937,1962]" gridcol="1" gridrow="31" pageId="4" pageNumber="393">“Stack C”</td>
<td id="7151A731FFCA00DDFDA0F819625AF4EF" box="[622,792,1937,1962]" gridcol="2" gridrow="31" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F81A63F2F4EF" box="[622,688,1938,1962]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDA0F81A63F2F4EF" box="[622,688,1938,1962]" italics="true" pageId="4" pageNumber="393">oliveri</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3280CE4DFFCA00DDFF68F83F6402F495" box="[166,1344,1975,2000]" gridrow="32" pageId="4" pageNumber="393" rowspan-0="1">
<td id="7151A731FFCA00DDFE4AF83F6364F495" box="[388,550,1975,2000]" gridcol="1" gridrow="32" pageId="4" pageNumber="393">Tawhiti Rahi I.</td>
<td id="7151A731FFCA00DDFDA0F83F625AF495" box="[622,792,1975,2000]" gridcol="2" gridrow="32" pageId="4" pageNumber="393">
<taxonomicName id="4BB0B78CFFCAFF26FDA0F83063F2F495" box="[622,688,1976,2000]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCAFF26FDA0F83063F2F495" box="[622,688,1976,2000]" italics="true" pageId="4" pageNumber="393">oliveri</emphasis>
</taxonomicName>
</td>
<td id="7151A731FFCA00DDFCAAF83F6402F495" box="[868,1344,1975,2000]" gridcol="3" gridrow="32" pageId="4" pageNumber="393">Type locality</td>
</tr>
</table>
</paragraph>
<caption id="D8CF9C87FFCBFF27FF59FBFC63C6F7F4" ID-DOI="http://doi.org/10.5281/zenodo.3516821" ID-Zenodo-Dep="3516821" httpUri="https://zenodo.org/record/3516821/files/figure.png" pageId="5" pageNumber="394" startId="5.[151,250,1140,1165]" targetBox="[151,1434,174,1121]" targetPageId="5">
<paragraph id="8C0FCC0FFFCBFF27FF59FBFC63C6F7F4" blockId="5.[151,1436,1140,1201]" pageId="5" pageNumber="394">
<emphasis id="BEC4101DFFCBFF27FF59FBFC6054F7C8" bold="true" box="[151,278,1140,1165]" pageId="5" pageNumber="394">FIGURE 3.</emphasis>
Two examples of 
<taxonomicName id="4BB0B78CFFCBFF27FE15FBFC620DF7C8" authority="(Robb)" authorityName="Robb" baseAuthorityName="Robb" box="[475,847,1140,1165]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FE15FBFC63BCF7C9" box="[475,766,1140,1164]" italics="true" pageId="5" pageNumber="394">Oligosoma pachysomaticum</emphasis>
(Robb)
</taxonomicName>
from Green Island in the Mercury Island group, east of Coromandel Peninsula (photos: Tony Whitaker).
</paragraph>
</caption>
<paragraph id="8C0FCC0FFFCBFF27FF09FB566512F694" blockId="5.[151,1437,1246,2029]" pageId="5" pageNumber="394">
<emphasis id="BEC4101DFFCBFF27FF09FB566031F7BC" bold="true" box="[199,371,1246,1273]" pageId="5" pageNumber="394">Biogeography.</emphasis>
The islands on which 
<taxonomicName id="4BB0B78CFFCBFF27FDBCFB5762F2F7BC" box="[626,944,1247,1273]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FDBCFB5762F2F7BC" box="[626,944,1247,1273]" italics="true" pageId="5" pageNumber="394">Oligosoma pachysomaticum</emphasis>
</taxonomicName>
occurs were connected to the 
<collectingCountry id="F4A78C9FFFCBFF27FAC8FB5764DEF7BC" box="[1286,1436,1247,1273]" name="New Zealand" pageId="5" pageNumber="394">New Zealand</collectingCountry>
mainland during the Otira glacial maximum 18–20,000 years B.P., when sea-levels fell to 
<quantity id="4B4861EAFFCBFF27FB4CFA8B658AF658" box="[1154,1224,1283,1309]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="394" unit="m" value="110.0">110 m</quantity>
below present levels (Haywood 1991; 
<bibRefCitation id="E821B1FEFFCBFF27FE4EFAAF6306F604" author="Kirkpatrick, R." box="[384,580,1319,1345]" pageId="5" pageNumber="394" refId="ref6070" refString="Kirkpatrick, R. (1999) Contempory atlas of New Zealand-the shapes of our nation. David Bateman, Auckland, 150 pp." type="book" year="1999">Kirkpatrick 1999</bibRefCitation>
). As such, 
<taxonomicName id="4BB0B78CFFCBFF27FD72FAAF62D8F604" box="[700,922,1319,1345]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FD72FAAF62D8F604" box="[700,922,1319,1345]" italics="true" pageId="5" pageNumber="394">O. pachysomaticum</emphasis>
</taxonomicName>
will likely have been present on the mainland until displaced by the introduction of predatory mammals by people, a process which began between 700–800 years ago (
<bibRefCitation id="E821B1FEFFCBFF27FF03FAE76090F6CC" author="Wilmshurst, J. M. &amp; Atholl, J. &amp; Anderson, A. J. &amp; Thomas, F. G. &amp; Higham, T. F. G. &amp; Worthy, T. H." box="[205,466,1391,1417]" pageId="5" pageNumber="394" pagination="7676 - 7680" refId="ref6525" refString="Wilmshurst, J. M., Atholl, J., Anderson, A. J., Thomas, F. G., Higham, T. F. G. &amp; Worthy, T. H. (2008) Dating the late prehistoric dispersal of Polynesians to New Zealand using the commensal Pacific rat. PNAS, 105 (22), 7676 - 7680. https: // doi. org / 10.1073 / pnas. 0801507105" type="journal article" year="2008">
Wilmshurst 
<emphasis id="BEC4101DFFCBFF27FE99FAF860CDF6CC" box="[343,399,1391,1417]" italics="true" pageId="5" pageNumber="394">et al.</emphasis>
2008
</bibRefCitation>
). Subfossil remains attributable to the 
<taxonomicName id="4BB0B78CFFCBFF27FCB5FAE762A7F6CC" box="[891,997,1391,1417]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCBFF27FCB5FAE762A7F6CC" box="[891,997,1391,1417]" italics="true" pageId="5" pageNumber="394">O. oliveri</emphasis>
</taxonomicName>
species-complex (but not 
<taxonomicName id="4BB0B78CFFCBFF27FAC4FAE764D7F6CC" baseAuthorityName="Hardy" baseAuthorityYear="1977" box="[1290,1429,1391,1417]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="whitakeri">
<emphasis id="BEC4101DFFCBFF27FAC4FAE764D7F6CC" box="[1290,1429,1391,1417]" italics="true" pageId="5" pageNumber="394">O. whitakeri</emphasis>
</taxonomicName>
, which is distinguishable) have been found on the mainland in the nearby 
<collectingRegion id="4E7402EDFFCBFF27FC0BFA1B651DF6E8" box="[965,1119,1427,1453]" country="New Zealand" name="Bay of Plenty" pageId="5" pageNumber="394">Bay of Plenty</collectingRegion>
area (
<bibRefCitation id="E821B1FEFFCBFF27FB6FFA1B64D2F6EB" author="Chapple, D. G. &amp; Patterson, G. B. &amp; Gleeson, D. M. &amp; Daugherty, C. H. &amp; Ritchie, P. A." box="[1185,1424,1427,1454]" pageId="5" pageNumber="394" pagination="129 - 146" refId="ref5441" refString="Chapple, D. G., Patterson, G. B., Gleeson, D. M., Daugherty, C. H. &amp; Ritchie, P. A. (2008 b) Taxonomic revision of the marbled skink (Cyclodina oliveri, Reptilia: Scincidae) species complex, with a description of a new species. New Zealand Journal of Zoology, 35, 129 - 146. https: // doi. org / 10.1080 / 03014220809510110" type="journal article" year="2008">
Chapple 
<emphasis id="BEC4101DFFCBFF27FAC9FA1C647AF6E8" box="[1287,1336,1427,1453]" italics="true" pageId="5" pageNumber="394">et al</emphasis>
. 2008b
</bibRefCitation>
); based on proximity to extant populations these were most likely 
<taxonomicName id="4BB0B78CFFCBFF27FCA4FA3F650BF694" box="[874,1097,1463,1489]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FCA4FA3F650BF694" box="[874,1097,1463,1489]" italics="true" pageId="5" pageNumber="394">O. pachysomaticum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8C0FCC0FFFCBFF27FF09FA5364C0F450" blockId="5.[151,1437,1246,2029]" pageId="5" pageNumber="394">
<emphasis id="BEC4101DFFCBFF27FF09FA53600BF6B0" bold="true" box="[199,329,1499,1525]" pageId="5" pageNumber="394">Phylogeny.</emphasis>
The two sampled island groups of 
<taxonomicName id="4BB0B78CFFCBFF27FD2EFA536562F6B0" box="[736,1056,1499,1525]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FD2EFA536562F6B0" box="[736,1056,1499,1525]" italics="true" pageId="5" pageNumber="394">Oligosoma pachysomaticum</emphasis>
</taxonomicName>
(i.e. Alderman and Mercury) exhibit a similar level of divergence among mitochondrial DNA sequences as they each do to 
<emphasis id="BEC4101DFFCBFF27FB57FA776449F55C" box="[1177,1291,1535,1561]" italics="true" pageId="5" pageNumber="394">
<taxonomicName id="4BB0B78CFFCBFF27FB57FA776445F55C" box="[1177,1287,1535,1561]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="oliveri">O. oliveri</taxonomicName>
,
</emphasis>
which could indicate a split between 1.07–1.43 million years ago (
<bibRefCitation id="E821B1FEFFCBFF27FD2AF9AB628FF578" author="Chapple, D. G. &amp; Daugherty, C. H. &amp; Ritchie, P. A." box="[740,973,1571,1597]" pageId="5" pageNumber="394" pagination="129 - 146" refId="ref5369" refString="Chapple, D. G., Daugherty, C. H. &amp; Ritchie, P. A. (2008 a) Comparative phylogeography reveals pre-decline population structure of New Zealand Cyclodina (Reptilia: Scincidae) species, Biological Journal of the Linnean Society, 95 (2), 129 - 146 https: // doi. org / 10.1111 / j. 1095 - 8312.2008.01062. x" type="journal article" year="2008">
Chapple 
<emphasis id="BEC4101DFFCBFF27FC87F9AC623BF578" box="[841,889,1571,1597]" italics="true" pageId="5" pageNumber="394">et al</emphasis>
. 2008a
</bibRefCitation>
, b). This is much earlier than the actual (&lt;20,000 year) split between their islands and as such could point to cryptic speciation within 
<taxonomicName id="4BB0B78CFFCBFF27FB42F9CF642BF524" box="[1164,1385,1607,1633]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FB42F9CF642BF524" box="[1164,1385,1607,1633]" italics="true" pageId="5" pageNumber="394">O. pachysomaticum</emphasis>
</taxonomicName>
as it is here defined. However, there is yet no corroboration for such a timeframe and if anything, the biogeographic and morphological evidence must cast doubt on this interpretation. Alternative explanations to be considered including a mitochondrial split much older than the actual split of the skink populations, an accelerated rate of genetic drift in one or other population, and genetic introgression from an unknown and probably now extinct lineage. For this reason the age of the divergence between the Alderman and Mercury populations is best regarded as unresolved.
</paragraph>
<paragraph id="8C0FCC0FFFCBFF24FF09F896636CF392" blockId="5.[151,1437,1246,2029]" lastBlockId="6.[151,1437,153,431]" lastPageId="6" lastPageNumber="395" pageId="5" pageNumber="394">
<emphasis id="BEC4101DFFCBFF27FF09F8966014F47C" bold="true" box="[199,342,1822,1849]" pageId="5" pageNumber="394">Recognition</emphasis>
. 
<taxonomicName id="4BB0B78CFFCBFF27FEAAF89763E0F47C" box="[356,674,1823,1849]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FEAAF89763E0F47C" box="[356,674,1823,1849]" italics="true" pageId="5" pageNumber="394">Oligosoma pachysomaticum</emphasis>
</taxonomicName>
is here reinstated as a species separate from 
<taxonomicName id="4BB0B78CFFCBFF27FB58F8976443F47C" box="[1174,1281,1823,1849]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCBFF27FB58F8976443F47C" box="[1174,1281,1823,1849]" italics="true" pageId="5" pageNumber="394">O. oliveri</emphasis>
</taxonomicName>
because multiple lines of evidence suggest a geographical isolation that has precluded interaction for at least 1–2 million years (see ‘Biogeography’ above and ‘Phylogeny’ for 
<taxonomicName id="4BB0B78CFFCBFF27FD0AF8EF6270F4C4" box="[708,818,1895,1921]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFCBFF27FD0AF8EF6270F4C4" box="[708,818,1895,1921]" italics="true" pageId="5" pageNumber="394">O. oliveri</emphasis>
</taxonomicName>
), in combination with a distinct niche (as defined by their respective sympatric competitors, see ‘Ecology’ for each species, above), and also morphological differentiation (
<tableCitation id="C132F9B4FFCBFF27FF1DF8276068F48C" box="[211,298,1967,1993]" captionStart="TABLE 2" captionStartId="6.[151,239,478,502]" captionTargetPageId="6" captionText="TABLE 2. Phenotypic and ecological contrasts between Oligosoma oliveri (McCann) and O. pachysomaticum (Robb)." httpUri="http://table.plazi.org/id/D8CF9C87FFC8FF24FF59FE5664CFF2B2" pageId="5" pageNumber="394" tableUuid="D8CF9C87FFC8FF24FF59FE5664CFF2B2">Table 2</tableCitation>
) which demonstrates that during this isolation the two lineages have been evolving, to a significant degree, along separate evolutionary trajectories. 
<taxonomicName id="4BB0B78CFFCBFF27FD7AF85B62D3F4A8" box="[692,913,2003,2029]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFCBFF27FD7AF85B62D3F4A8" box="[692,913,2003,2029]" italics="true" pageId="5" pageNumber="394">O. pachysomaticum</emphasis>
</taxonomicName>
meets a number of the species definitions (e.g. ‘Simpsons’, ‘niche’, ‘phylogenetic III’, ‘phenetic’, ‘genotypic’) within the encompassing ‘general-lineage’ speciesconcept of 
<bibRefCitation id="E821B1FEFFC8FF24FEDAFF356098F392" author="de Queiroz, K." box="[276,474,189,215]" pageId="6" pageNumber="395" pagination="57 - 75" refId="ref5649" refString="de Queiroz, K. (1998) The general lineage concept of species, species criteria, and the process of speciation: a conceptual unification and terminological recommendations. In: Howard, D. J. &amp; Berlocher, S. H. (Eds.), Endless forms: species and speciation. Oxford University Press, New York, pp. 57 - 75." type="book chapter" year="1998">de Queiroz (1998</bibRefCitation>
, 
<bibRefCitation id="E821B1FEFFC8FF24FE28FF356361F392" author="de Queiroz, K." box="[486,547,189,215]" pageId="6" pageNumber="395" pagination="879 - 886" refId="ref5720" refString="de Queiroz, K. (2007) Species Concepts and Species Delimitation. Systematic Biology, 56 (6), 879 - 886. https: // doi. org / 10.1080 / 10635150701701083" type="journal article" year="2007">2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4AA9F84FFC8FF24FF09FF6962F9F2EA" pageId="6" pageNumber="395" type="discussion">
<paragraph id="8C0FCC0FFFC8FF24FF09FF6962F9F2EA" blockId="6.[151,1437,153,431]" pageId="6" pageNumber="395">
<emphasis id="BEC4101DFFC8FF24FF09FF696079F3BE" bold="true" box="[199,315,225,251]" pageId="6" pageNumber="395">Remarks.</emphasis>
<bibRefCitation id="E821B1FEFFC8FF24FE8FFF6960A8F3BE" author="Hardy, G. S." box="[321,490,225,251]" pageId="6" pageNumber="395" pagination="221 - 325" refId="ref5756" refString="Hardy, G. S. (1977) The New Zealand Scincidae (Reptilia: Lacertilia); a taxonomic and Zoogeographic study. New Zealand Journal of Zoology, 4, 221 - 325. https: // doi. org / 10.1080 / 03014223.1977.9517956" type="journal article" year="1977">Hardy’s (1977)</bibRefCitation>
decision to place 
<taxonomicName id="4BB0B78CFFC8FF24FD7FFF6962ACF3BE" box="[689,1006,225,251]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="395" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFC8FF24FD7FFF6962ACF3BE" box="[689,1006,225,251]" italics="true" pageId="6" pageNumber="395">Oligosoma pachysomaticum</emphasis>
</taxonomicName>
in the synonymy of 
<taxonomicName id="4BB0B78CFFC8FF24FB1EFF696478F3BE" box="[1232,1338,225,251]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="395" phylum="Chordata" rank="species" species="oliveri">
<emphasis id="BEC4101DFFC8FF24FB1EFF696478F3BE" box="[1232,1338,225,251]" italics="true" pageId="6" pageNumber="395">O. oliveri</emphasis>
</taxonomicName>
was justified by the discovery of populations that appeared to be intermediate in phylogeny, morphology and biogeography. However, the decision by 
<bibRefCitation id="E821B1FEFFC8FF24FE78FEA163ECF201" author="Chapple, D. G. &amp; Patterson, G. B. &amp; Gleeson, D. M. &amp; Daugherty, C. H. &amp; Ritchie, P. A." box="[438,686,297,324]" pageId="6" pageNumber="395" pagination="129 - 146" refId="ref5441" refString="Chapple, D. G., Patterson, G. B., Gleeson, D. M., Daugherty, C. H. &amp; Ritchie, P. A. (2008 b) Taxonomic revision of the marbled skink (Cyclodina oliveri, Reptilia: Scincidae) species complex, with a description of a new species. New Zealand Journal of Zoology, 35, 129 - 146. https: // doi. org / 10.1080 / 03014220809510110" type="journal article" year="2008">
Chapple 
<emphasis id="BEC4101DFFC8FF24FDD7FEA26312F206" box="[537,592,297,323]" italics="true" pageId="6" pageNumber="395">et al.</emphasis>
(2008b)
</bibRefCitation>
to retain them as a singular entity, after removing all of the intermediate populations again as 
<taxonomicName id="4BB0B78CFFC8FF24FE0EFEC56369F222" baseAuthorityName="Chapple, Patterson, Gleeson, Daugherty &amp; Ritchie" baseAuthorityYear="2008" box="[448,555,333,359]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="395" phylum="Chordata" rank="species" species="townsi">
<emphasis id="BEC4101DFFC8FF24FE0EFEC56369F222" box="[448,555,333,359]" italics="true" pageId="6" pageNumber="395">O. townsi</emphasis>
</taxonomicName>
, was less convincing because the phenotypic and biogeographic evidence now showed a substantial differentiation between the two. 
<taxonomicName id="4BB0B78CFFC8FF24FD32FEF9629EF2CE" box="[764,988,369,395]" class="Reptilia" family="Scincidae" genus="Oligosoma" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="395" phylum="Chordata" rank="species" species="pachysomaticum">
<emphasis id="BEC4101DFFC8FF24FD32FEF9629EF2CE" box="[764,988,369,395]" italics="true" pageId="6" pageNumber="395">O. pachysomaticum</emphasis>
</taxonomicName>
is here returned to its original concept, i.e. that of 
<bibRefCitation id="E821B1FEFFC8FF24FEDEFE1D60E2F2EA" author="Robb, J." box="[272,416,404,431]" pageId="6" pageNumber="395" pagination="477 - 484" refId="ref6122" refString="Robb, J. (1975) Two new skinks of the genus Leiolopisma from New Zealand. Proceedings of the Section of Sciences, Koninklijke Nederlandse Akademie van Wetenschappen, Series C, 78, 477 - 484." type="journal article" year="1975">Robb (1975)</bibRefCitation>
, who provides a full morphological description.
</paragraph>
</subSubSection>
</treatment>
</document>