1242 lines
194 KiB
XML
1242 lines
194 KiB
XML
<document id="33E0F130128F2F6D3094F1105C0E1A75" ID-CLB-Dataset="83504" ID-DOI="10.1080/00222933.2022.2071179" ID-GBIF-Dataset="f559a5e8-864a-459b-8fa2-83bfd99862f3" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758474" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1656311937750" checkinUser="plazi" docAuthor="Humara-Gil, Karla J., Granja-Fernández, Rebeca, Bautista-Guerrero, Eric & Rodríguez-Troncoso, Alma P." docDate="2022" docId="03DF230B0A0A947EFE016824FD7EB54D" docLanguage="en" docName="JNATHIST.56.5-8.365-384.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 56 (5 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2022.2071179" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Ophioderma occultum Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso 2022, sp. nov." docType="treatment" docVersion="10" lastPageNumber="380" masterDocId="FFE65B730A0E9471FFCA6A44FFD8B442" masterDocTitle="Overlooked for over a century: Ophioderma occultum sp. nov. (Echinodermata), a new species of brittle star from the Eastern Pacific" masterLastPageNumber="384" masterPageNumber="365" pageNumber="369" updateTime="1702002689895" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6F7EF33CE6B34364BA5C77AD2768DF2F">Overlooked for over a century: Ophioderma occultum sp. nov. (Echinodermata), a new species of brittle star from the Eastern Pacific</mods:title>
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<mods:namePart id="6BE137DC2F2721895E6BE45454D70BD5">Humara-Gil, Karla J.</mods:namePart>
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<mods:namePart id="D7EE9284E1CEFA1B7D8A92063BB61B35">Bautista-Guerrero, Eric</mods:namePart>
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<mods:title id="2AF6CC0719C093CDF647CAB8FC47E6E7">Journal of Natural History</mods:title>
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<mods:date id="9EE7CEE907EE1BFD216B9878BE4AB75C">2022</mods:date>
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<treatment id="03DF230B0A0A947EFE016824FD7EB54D" ID-DOI="http://doi.org/10.5281/zenodo.6772313" ID-GBIF-Taxon="196365692" ID-Zenodo-Dep="6772313" LSID="urn:lsid:plazi:treatment:03DF230B0A0A947EFE016824FD7EB54D" httpUri="http://treatment.plazi.org/id/03DF230B0A0A947EFE016824FD7EB54D" lastPageId="15" lastPageNumber="380" pageId="4" pageNumber="369">
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<paragraph id="8BC9921D0A0A9475FE016824FC81B639" blockId="4.[459,857,608,670]" box="[459,857,608,635]" pageId="4" pageNumber="369">
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<heading id="D08125710A0A9475FE016824FC81B639" bold="true" box="[459,857,608,635]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="369" reason="3">
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<emphasis id="B9024E0F0A0A9475FE016824FC81B639" bold="true" box="[459,857,608,635]" pageId="4" pageNumber="369">
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<taxonomicName id="4C76E99E0A0A9475FE016824FD31B639" ID-CoL="9C7NQ" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[459,745,608,635]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="4" pageNumber="369" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
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<emphasis id="B9024E0F0A0A9475FE016824FD31B639" bold="true" box="[459,745,608,635]" italics="true" pageId="4" pageNumber="369">Ophioderma occultum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A231F3740A0A9475FD3B6824FC81B639" box="[753,857,608,635]" pageId="4" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C36CC1960A0A9475FE3168C7FCF0B6DC" box="[507,808,643,670]" pageId="4" pageNumber="369" type="description">
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<paragraph id="8BC9921D0A0A9475FE3168C7FCF0B6DC" blockId="4.[459,857,608,670]" box="[507,808,643,670]" pageId="4" pageNumber="369">
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<heading id="D08125710A0A9475FE3168C7FCF0B6DC" box="[507,808,643,670]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="369" reason="2">
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<emphasis id="B9024E0F0A0A9475FE3168C7FCF0B6DC" bold="true" box="[507,808,643,670]" pageId="4" pageNumber="369">
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(
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<figureCitation id="134D8E980A0A9475FDCF68C7FD75B6DC" box="[517,685,643,670]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="5.[160,225,1629,1652]" captionStartId-1="6.[160,225,1629,1652]" captionStartId-2="7.[160,226,1629,1652]" captionTargetBox-0="[164,1152,143,1596]" captionTargetBox-1="[164,1152,143,1596]" captionTargetBox-2="[164,1152,143,1596]" captionTargetId-0="figure-3@5.[163,1153,142,1597]" captionTargetId-1="figure-3@6.[163,1153,142,1597]" captionTargetId-2="figure-3@7.[163,1153,142,1597]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." captionText-1="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." captionText-2="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi-0="http://doi.org/10.5281/zenodo.6758478" figureDoi-1="http://doi.org/10.5281/zenodo.6758480" figureDoi-2="http://doi.org/10.5281/zenodo.6758482" httpUri-0="https://zenodo.org/record/6758478/files/figure.png" httpUri-1="https://zenodo.org/record/6758480/files/figure.png" httpUri-2="https://zenodo.org/record/6758482/files/figure.png" pageId="4" pageNumber="369">Figures 1 – 3</figureCitation>
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||
;
|
||
<tableCitation id="C6F4A7A60A0A9475FD7768C7FCC6B6DC" box="[701,798,643,670]" captionStart="Table 1" captionStartId="8.[126,182,439,462]" captionTargetPageId="8" captionText="Table 1. Main characters that distinguish Eastern Pacific Ophioderma species." httpUri="http://table.plazi.org/id/DF09C2950A069479FFB46BF3FCA2B58C" pageId="4" pageNumber="369" tableUuid="DF09C2950A069479FFB46BF3FCA2B58C">Table 1</tableCitation>
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||
)
|
||
</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36CC1960A0A9475FF6A689FFD8EB73D" pageId="4" pageNumber="369" type="reference_group">
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<paragraph id="8BC9921D0A0A9475FF6A689FFD66B6B6" blockId="4.[160,1155,731,895]" box="[160,702,731,756]" pageId="4" pageNumber="369">
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<treatmentCitationGroup id="AB66B5330A0A9475FF6A689FFD66B6B6" box="[160,702,731,756]" pageId="4" pageNumber="369">
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<taxonomicName id="4C76E99E0A0A9475FF6A689FFE5AB6B6" baseAuthorityName="Lütken" baseAuthorityYear="1859" box="[160,386,731,756]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="369" phylum="Echinodermata" rank="species" species="panamensis">
|
||
<emphasis id="B9024E0F0A0A9475FF6A689FFE5AB6B6" box="[160,386,731,756]" italics="true" pageId="4" pageNumber="369">Ophiura panamensis</emphasis>
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</taxonomicName>
|
||
. –
|
||
<treatmentCitation id="0AD7B40C0A0A9475FE6E689FFD86B6B6" author="Ives JE" box="[420,606,731,756]" page="76" pageId="4" pageNumber="369" year="1889">
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<bibRefCitation id="EFE7EFEC0A0A9475FE6E689FFD86B6B6" author="Ives JE" box="[420,606,731,756]" pageId="4" pageNumber="369" pagination="76 - 77" refId="ref9999" refString="Ives JE. 1889. Variation in Ophiura panamensis and Ophiura teres. Proc Acad Nat Sci Phila. 41: 76 - 77." type="journal article" year="1889">Ives 1889: 76–77</bibRefCitation>
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</treatmentCitation>
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(partim)
|
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC9921D0A0A9475FF6A68BAFD4AB71E" blockId="4.[160,1155,731,895]" pageId="4" pageNumber="369">
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<taxonomicName id="4C76E99E0A0A9475FF6A68BAFE69B755" ID-CoL="7W5PK" authorityName="Lutken" authorityYear="1859" box="[160,433,766,791]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="4" pageNumber="369" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A0A9475FF6A68BAFE69B755" box="[160,433,766,791]" italics="true" pageId="4" pageNumber="369">Ophioderma panamense</emphasis>
|
||
</taxonomicName>
|
||
. –
|
||
<treatmentCitation id="0AD7B40C0A0A9475FE1368BAFD2EB755" author="Ziesenhenne FC" box="[473,758,766,791]" page="194" pageId="4" pageNumber="369" year="1955">
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<bibRefCitation id="EFE7EFEC0A0A9475FE1368BAFD63B755" author="Ziesenhenne FC" box="[473,699,766,791]" pageId="4" pageNumber="369" pagination="185 - 201" refId="ref11224" refString="Ziesenhenne FC. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation fo Scientific Research, editor. Essays in the Natural Sciences in honor of Captain Allan Hancock. Los Angeles: University of Southern California Press; p. 185 - 201. doi: 10.5962 / bhl. title. 6088." type="book chapter" year="1955">Ziesenhenne (1955)</bibRefCitation>
|
||
: 194
|
||
</treatmentCitation>
|
||
, 197 (partim). –
|
||
<bibRefCitation id="EFE7EFEC0A0A9475FC7368BAFB5BB755" author="Brusca RC & Smith CR" box="[953,1155,766,791]" pageId="4" pageNumber="369" pagination="304 - 343" refId="ref9370" refString="Brusca RC, Smith CR. 1973. Echinodermata (Starfish, etc.). In: Brusca RC, editor. A handbook to the common intertidal invertebrates of the Gulf of California. Tucson: University of Arizona Press; p. 304 - 343." type="book chapter" year="1973">Brusca and Smith</bibRefCitation>
|
||
(1973): 318, 320, figs 12.15–12.16 (partim). –
|
||
<bibRefCitation id="EFE7EFEC0A0A9475FD786965FCA9B778" author="Brusca RC" box="[690,881,801,826]" pageId="4" pageNumber="369" pagination="398 - 422" refId="ref9419" refString="Brusca RC. 1980. 26. Echinodermata. In: Brusca RC, editor. Common intertidal invertebrates of the Gulf of California. Tucson: The University of Arizona Press; p. 398 - 422." type="book chapter" year="1980">Brusca 1980: 407</bibRefCitation>
|
||
, fig. 26.15a–b (partim). –
|
||
<bibRefCitation id="EFE7EFEC0A0A9475FF0A6907FE25B71E" author="Kerstitch A & Bertsch H" box="[192,509,835,860]" pageId="4" pageNumber="369" refId="ref10024" refString="Kerstitch A, Bertsch H. 2007. Sea of Cortez. Marine invertebrates. A guide for the Pacific Coast, Mexico to Peru. 2 nd ed. California: Sea Challengers; p. 124." type="journal volume" year="2007">Kerstitch and Bertsch (2007)</bibRefCitation>
|
||
: 113 (partim)
|
||
</paragraph>
|
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<paragraph id="8BC9921D0A0A9475FF6A6922FD8EB73D" blockId="4.[160,1155,731,895]" box="[160,598,870,895]" pageId="4" pageNumber="369">
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<taxonomicName id="4C76E99E0A0A9475FF6A6922FEEBB73D" authorityName="Lyman" authorityYear="1860" box="[160,307,870,895]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="369" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A0A9475FF6A6922FEEBB73D" box="[160,307,870,895]" italics="true" pageId="4" pageNumber="369">Ophiura teres</emphasis>
|
||
</taxonomicName>
|
||
. –
|
||
<bibRefCitation id="EFE7EFEC0A0A9475FE9F6922FE17B73D" author="Ives JE" box="[341,463,870,895]" pageId="4" pageNumber="369" pagination="76 - 77" refId="ref9999" refString="Ives JE. 1889. Variation in Ophiura panamensis and Ophiura teres. Proc Acad Nat Sci Phila. 41: 76 - 77." type="journal article" year="1889">Ives (1889)</bibRefCitation>
|
||
: 77 (partim)
|
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</paragraph>
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</subSubSection>
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<subSubSection id="C36CC1960A0A9475FF6A69EEFD27B126" pageId="4" pageNumber="369" type="materials_examined">
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<paragraph id="8BC9921D0A0A9475FF6A69EEFEA7B786" blockId="4.[160,1157,938,1311]" box="[160,383,938,964]" pageId="4" pageNumber="369">
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<emphasis id="B9024E0F0A0A9475FF6A69EEFEA7B786" bold="true" box="[160,383,938,964]" italics="true" pageId="4" pageNumber="369">Material examined</emphasis>
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</paragraph>
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<paragraph id="8BC9921D0A0A9475FF6A6989FE4CB15D" blockId="4.[160,1157,938,1311]" pageId="4" pageNumber="369">
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<emphasis id="B9024E0F0A0A9475FF6A6989FE94B7A5" bold="true" box="[160,332,973,999]" italics="true" pageId="4" pageNumber="369">Type material.</emphasis>
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<materialsCitation id="3B1E98400A0A9475FE96698AFCBFB030" ID-GBIF-Occurrence="3819603308" accessionNumber="OK076949, OK078012" collectingDate="2019-11-07" collectionCode="COI" collectorName="K. J. Humara-Gil & R. Granja-Fernandez & B. Maya-Alvarado & A. Hernandez-Morales & M. A. Medina-Lopez" country="Mexico" elevation="2" latitude="24.285" location="Punta Pichilingue" longLatPrecision="74" longitude="-110.334" municipality="Gulf of California" pageId="4" pageNumber="369" specimenCode="ICML-EMU 12867" specimenCount="1" stateProvince="Baja California Sur" typeStatus="holotype">
|
||
<typeStatus id="54CD2CBF0A0A9475FE96698AFE1DB7A5" box="[348,453,974,999]" pageId="4" pageNumber="369" type="holotype">Holotype</typeStatus>
|
||
: ICML-UNAM 18429;
|
||
<collectingCountry id="F361D28D0A0A9475FD7F698AFCD0B7A5" box="[693,776,974,999]" name="Mexico" pageId="4" pageNumber="369">Mexico</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BAD08670A0A9475FCD9698AFC0FB7A5" box="[787,983,974,999]" pageId="4" pageNumber="369">Gulf of California</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49B25CFF0A0A9475FC28698AFF10B048" country="Mexico" name="Baja California Sur" pageId="4" pageNumber="369">Baja California Sur</collectingRegion>
|
||
,
|
||
<location id="8EA9C4C60A0A9475FF2969B5FE65B048" LSID="urn:lsid:plazi:treatment:03DF230B0A0A947EFE016824FD7EB54D:8EA9C4C60A0A9475FF2969B5FE65B048" box="[227,445,1009,1034]" country="Mexico" latitude="24.285" longLatPrecision="74" longitude="-110.334" municipality="Gulf of California" name="Punta Pichilingue" pageId="4" pageNumber="369" stateProvince="Baja California Sur">Punta Pichilingue</location>
|
||
(
|
||
<geoCoordinate id="EE42F4DA0A0A9475FE1369B5FD9DB048" box="[473,581,1009,1034]" degrees="24.285" direction="north" orientation="latitude" pageId="4" pageNumber="369" precision="55" value="24.285">24.285°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE42F4DA0A0A9475FD9569B5FD06B048" box="[607,734,1009,1034]" degrees="110.334" direction="west" orientation="longitude" pageId="4" pageNumber="369" precision="55" value="-110.334">110.334°W</geoCoordinate>
|
||
);
|
||
<quantity id="4C8E3FF80A0A9475FD3469B5FCA6B048" box="[766,894,1009,1034]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="4" pageNumber="369" unit="m" value="1.5" valueMax="2.5" valueMin="0.5">
|
||
<elevation id="005B752E0A0A9475FD3469B5FCA6B048" box="[766,894,1009,1034]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="4" pageNumber="369" unit="m" value="1.5" valueMax="2.5" valueMin="0.5">0.5–2.5 m</elevation>
|
||
</quantity>
|
||
; sand under rocks;
|
||
<date id="FFC8B4DD0A0A9475FF6A6E57FE5BB06E" box="[160,387,1043,1068]" pageId="4" pageNumber="369" value="2019-11-07">
|
||
<collectingDate id="EF8C4D350A0A9475FF6A6E57FE5BB06E" box="[160,387,1043,1068]" pageId="4" pageNumber="369" value="2019-11-07">7 November 2019</collectingDate>
|
||
</date>
|
||
; coll.
|
||
<collectorName id="2683F7CB0A0A9475FE1C6E57FD48B06E" box="[470,656,1043,1068]" pageId="4" pageNumber="369">K.J. Humara-Gil</collectorName>
|
||
,
|
||
<collectorName id="2683F7CB0A0A9475FD6E6E57FC41B06E" box="[676,921,1043,1068]" pageId="4" pageNumber="369">R. Granja-Fernández</collectorName>
|
||
,
|
||
<collectorName id="2683F7CB0A0A9475FC666E57FB58B06E" box="[940,1152,1043,1068]" pageId="4" pageNumber="369">B. Maya-Alvarado</collectorName>
|
||
,
|
||
<collectorName id="2683F7CB0A0A9475FF6A6E72FE78B00D" box="[160,416,1078,1103]" pageId="4" pageNumber="369">A. Hernández-Morales</collectorName>
|
||
,
|
||
<collectorName id="2683F7CB0A0A9475FE656E72FD56B00D" box="[431,654,1078,1103]" pageId="4" pageNumber="369">M.A. Medina-López</collectorName>
|
||
; preserved in 96% ethanol;
|
||
<collectionCode id="ED670AD80A0A9475FC146E72FBDEB00D" box="[990,1030,1078,1103]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="4" pageNumber="369" type="Herbarium">COI</collectionCode>
|
||
sequence:
|
||
<accessionNumber id="94250FFE0A0A9475FF6A6E1DFECEB030" box="[160,278,1113,1138]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OK076949" pageId="4" pageNumber="369">OK076949</accessionNumber>
|
||
(GenBank); 16S sequence:
|
||
<accessionNumber id="94250FFE0A0A9475FDAC6E1DFD04B030" box="[614,732,1113,1138]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OK078012" pageId="4" pageNumber="369">OK078012</accessionNumber>
|
||
(GenBank).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1E98400A0A9475FCBE6E1DFC9CB0F5" ID-GBIF-Occurrence="3819603305" accessionNumber="OK076949, OK078012" collectingDate="2019-11-07" collectionCode="COI" country="Mexico" elevation="2" latitude="24.285" location="Punta Pichilingue" longLatPrecision="74" longitude="-110.334" municipality="Gulf of California" pageId="4" pageNumber="369" specimenCode="ICML-EMU 12867" specimenCount="5" stateProvince="Baja California Sur" typeStatus="paratype">
|
||
<typeStatus id="54CD2CBF0A0A9475FCBE6E1DFC3FB030" box="[884,999,1113,1138]" pageId="4" pageNumber="369" type="paratype">Paratypes</typeStatus>
|
||
: ICML-UNAM 18430,
|
||
<specimenCount id="9D7059940A0A9475FF3A6E3FFE98B0D6" box="[240,320,1147,1172]" pageId="4" pageNumber="369" type="generic">5 spec.</specimenCount>
|
||
; same data as
|
||
<typeStatus id="54CD2CBF0A0A9475FE146E3FFD9DB0D6" box="[478,581,1147,1172]" pageId="4" pageNumber="369" type="holotype">holotype</typeStatus>
|
||
; preserved in 96% ethanol (1 used for DNA analysis, labelled as ‘OO4’;
|
||
<collectionCode id="ED670AD80A0A9475FE4C6EDAFE77B0F5" box="[390,431,1182,1207]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="4" pageNumber="369" type="Herbarium">COI</collectionCode>
|
||
sequence:
|
||
<accessionNumber id="94250FFE0A0A9475FD886EDAFD6FB0F5" box="[578,695,1182,1207]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OK076948" pageId="4" pageNumber="369">OK076948</accessionNumber>
|
||
[GenBank];
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1E98400A0A9475FC986EDAFDB8B098" ID-GBIF-Occurrence="3819603302" accessionNumber="OK076949, OK078012" collectingDate="2019-11-07" collectionCode="R, COI" country="Mexico" elevation="2" latitude="24.285" location="Punta Pichilingue" longLatPrecision="74" longitude="-110.334" municipality="Gulf of California" pageId="4" pageNumber="369" specimenCode="ICML-EMU 12867" specimenCount="3" stateProvince="Baja California Sur" typeStatus="holotype">
|
||
16S sequence:
|
||
<accessionNumber id="94250FFE0A0A9475FBC46EDAFB5CB0F5" box="[1038,1156,1182,1207]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OK078011" pageId="4" pageNumber="369">OK078011</accessionNumber>
|
||
[GenBank]). ICML-UNAM 18431,
|
||
<specimenCount id="9D7059940A0A9475FDC16E85FD85B098" box="[523,605,1217,1242]" pageId="4" pageNumber="369" type="generic">3 spec.</specimenCount>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1E98400A0A9475FDAD6E85FF07B0BE" ID-GBIF-Occurrence="3819603301" accessionNumber="OK076949, OK078012" collectingDate="2019-11-07" collectionCode="R, COI" country="Mexico" elevation="2" latitude="24.285" location="Punta Pichilingue" longLatPrecision="74" longitude="-110.334" municipality="Gulf of California" pageId="4" pageNumber="369" specimenCode="ICML-EMU 12867" specimenCount="5" stateProvince="Baja California Sur" typeStatus="holotype">
|
||
same data as
|
||
<typeStatus id="54CD2CBF0A0A9475FCC96E85FCB2B098" box="[771,874,1217,1242]" pageId="4" pageNumber="369" type="holotype">holotype</typeStatus>
|
||
; dry.
|
||
<specimenCode id="DBD03A660A0A9475FC6D6E85FBB2B098" box="[935,1130,1217,1242]" collectionCode="ICML-EMU" pageId="4" pageNumber="369">ICML-EMU 12867</specimenCode>
|
||
,
|
||
<specimenCount id="9D7059940A0A9475FBBF6E85FF04B0BE" pageId="4" pageNumber="369" type="generic">5 spec.</specimenCount>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1E98400A0A9475FF2C6EA7FE48B15D" ID-GBIF-Occurrence="3819603309" accessionNumber="OK076949, OK078012" collectingDate="2019-11-07" collectionCode="R, COI" country="Mexico" elevation="2" latitude="24.285" location="Punta Pichilingue" longLatPrecision="74" longitude="-110.334" municipality="Gulf of California" pageId="4" pageNumber="369" specimenCode="ICML-EMU 12868" specimenCount="3" stateProvince="Baja California Sur" typeStatus="holotype">
|
||
same data as
|
||
<typeStatus id="54CD2CBF0A0A9475FE4B6EA7FE30B0BE" box="[385,488,1251,1276]" pageId="4" pageNumber="369" type="holotype">holotype</typeStatus>
|
||
; preserved in 96% ethanol.
|
||
<specimenCode id="DBD03A660A0A9475FCEA6EA7FC3BB0BE" box="[800,995,1251,1276]" collectionCode="ICML-EMU" pageId="4" pageNumber="369">ICML-EMU 12868</specimenCode>
|
||
,
|
||
<specimenCount id="9D7059940A0A9475FC276EA7FBE7B0BE" box="[1005,1087,1251,1276]" pageId="4" pageNumber="369" type="generic">3 spec.</specimenCount>
|
||
; same data as
|
||
<typeStatus id="54CD2CBF0A0A9475FF326F42FE87B15D" box="[248,351,1286,1311]" pageId="4" pageNumber="369" type="holotype">holotype</typeStatus>
|
||
; dry
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A0A9475FF6A6F0EFD27B126" blockId="4.[160,767,1354,1380]" box="[160,767,1354,1380]" pageId="4" pageNumber="369">
|
||
<emphasis id="B9024E0F0A0A9475FF6A6F0EFE56B126" bold="true" box="[160,398,1354,1380]" italics="true" pageId="4" pageNumber="369">Additional material.</emphasis>
|
||
See the Supplemental material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36CC1960A0A9475FF6A6FD4FECBB215" pageId="4" pageNumber="369" type="diagnosis">
|
||
<paragraph id="8BC9921D0A0A9475FF6A6FD4FECCB1E8" blockId="4.[160,1155,1424,1623]" box="[160,276,1424,1450]" pageId="4" pageNumber="369">
|
||
<emphasis id="B9024E0F0A0A9475FF6A6FD4FECCB1E8" bold="true" box="[160,276,1424,1450]" italics="true" pageId="4" pageNumber="369">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A0A9475FF6A6FF7FECBB215" blockId="4.[160,1155,1424,1623]" pageId="4" pageNumber="369">Radial shields and adoral shields covered with granules. DAPs divided into two (typically) or more irregular pieces for about 2/3 of the arm. Colour pattern dark grey to brown (fixed specimens), or dark olive green to brown (in vivo specimens); arms with subtle lighter transverse bands, and parallel longitudinal white bands over some median and distal arm segments.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF09C2950A0B9474FF6A6C19FC06B2AB" ID-DOI="http://doi.org/10.5281/zenodo.6758478" ID-Zenodo-Dep="6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="5" pageNumber="370" startId="5.[160,225,1629,1652]" targetBox="[164,1152,143,1596]" targetPageId="5">
|
||
<paragraph id="8BC9921D0A0B9474FF6A6C19FC06B2AB" blockId="5.[160,1157,1629,1769]" pageId="5" pageNumber="370">
|
||
<emphasis id="B9024E0F0A0B9474FF6A6C19FF21B236" bold="true" box="[160,249,1629,1652]" pageId="5" pageNumber="370">Figure 1.</emphasis>
|
||
<taxonomicName id="4C76E99E0A0B9474FF356C19FE0BB236" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[255,467,1629,1652]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="5" pageNumber="370" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A0B9474FF356C19FE0BB236" box="[255,467,1629,1652]" italics="true" pageId="5" pageNumber="370">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A0B9474FE126C19FDFFB236" box="[472,551,1629,1652]" pageId="5" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF09C2950A089477FF6A6C19FDDEB345" ID-DOI="http://doi.org/10.5281/zenodo.6758480" ID-Zenodo-Dep="6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="6" pageNumber="371" startId="6.[160,225,1629,1652]" targetBox="[164,1152,143,1596]" targetPageId="6">
|
||
<paragraph id="8BC9921D0A089477FF6A6C19FDDEB345" blockId="6.[160,1157,1629,1799]" pageId="6" pageNumber="371">
|
||
<emphasis id="B9024E0F0A089477FF6A6C19FF22B236" bold="true" box="[160,250,1629,1652]" pageId="6" pageNumber="371">Figure 2.</emphasis>
|
||
<taxonomicName id="4C76E99E0A089477FECA6C19FE0EB236" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[256,470,1629,1652]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="6" pageNumber="371" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A089477FECA6C19FE0EB236" box="[256,470,1629,1652]" italics="true" pageId="6" pageNumber="371">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A089477FE166C19FDF3B236" box="[476,555,1629,1652]" pageId="6" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF09C2950A099476FF6A6C19FE33B366" ID-DOI="http://doi.org/10.5281/zenodo.6758482" ID-Zenodo-Dep="6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="7" pageNumber="372" startId="7.[160,226,1629,1652]" targetBox="[164,1152,143,1596]" targetPageId="7">
|
||
<paragraph id="8BC9921D0A099476FF6A6C19FE33B366" blockId="7.[159,1156,1629,1828]" pageId="7" pageNumber="372">
|
||
<emphasis id="B9024E0F0A099476FF6A6C19FF27B236" bold="true" box="[160,255,1629,1652]" pageId="7" pageNumber="372">Figure 3.</emphasis>
|
||
Morphological variations recorded in
|
||
<taxonomicName id="4C76E99E0A099476FDB76C19FC97B236" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[637,847,1629,1652]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="7" pageNumber="372" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A099476FDB76C19FC97B236" box="[637,847,1629,1652]" italics="true" pageId="7" pageNumber="372">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A099476FC926C19FC7FB236" box="[856,935,1629,1652]" pageId="7" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(a) Disc with partially uncovered radial shields (arrowheads;CE-UAM 390,DD = 15.4 mm); (b) white-spotted oral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679, DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF09C2950A069479FFB46BF3FCA2B58C" ID-Table-UUID="DF09C2950A069479FFB46BF3FCA2B58C" box="[126,890,439,462]" httpUri="http://table.plazi.org/id/DF09C2950A069479FFB46BF3FCA2B58C" pageId="8" pageNumber="373" startId="8.[126,182,439,462]" targetBox="[136,1680,475,1448]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="8BC9921D0A069479FFB46BF3FCA2B58C" blockId="8.[126,890,439,462]" box="[126,890,439,462]" pageId="8" pageNumber="373">
|
||
<emphasis id="B9024E0F0A069479FFB46BF3FF17B58C" bold="true" box="[126,207,439,462]" pageId="8" pageNumber="373">Table 1.</emphasis>
|
||
Main characters that distinguish Eastern Pacific
|
||
<taxonomicName id="4C76E99E0A069479FD656BF3FCA2B58C" box="[687,890,439,462]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="undetermined">
|
||
<emphasis id="B9024E0F0A069479FD656BF3FCFEB58C" box="[687,806,439,462]" italics="true" pageId="8" pageNumber="373">Ophioderma</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9921D0A069479FE026B9FFA32B1EA" pageId="8" pageNumber="373">
|
||
<table id="F97660BD0A066B8EFF426B9FF948B1EA" box="[136,1680,475,1448]" gridcols="6" gridrows="40" pageId="8" pageNumber="373">
|
||
<tr id="3546905F0A066B8EFF426B9FF948B5AF" box="[136,1680,475,493]" gridrow="0" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<th id="7697F9230A066B8EFEA66B9FFD67B5AF" box="[364,703,475,493]" gridcol="1" gridrow="0" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FE026B9FFDFAB5AF" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[456,546,475,493]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A069479FE026B9FFDFAB5AF" box="[456,546,475,493]" italics="true" pageId="8" pageNumber="373">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A069479FDE26B9FFDBFB5AF" box="[552,615,475,493]" pageId="8" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="7697F9230A066B8EFD076B9FFC4AB5AF" box="[717,914,475,493]" gridcol="2" gridrow="0" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FCCC6B9FFC81B5AF" box="[774,857,475,493]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="hendleri">
|
||
<emphasis id="B9024E0F0A069479FCCC6B9FFC81B5AF" box="[774,857,475,493]" italics="true" pageId="8" pageNumber="373">O. hendleri</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7697F9230A066B8EFC556B9FFB69B5AF" box="[927,1201,475,493]" gridcol="3" gridrow="0" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FC3B6B9FFB86B5AF" authorityName="Lutken" authorityYear="1859" box="[1009,1118,475,493]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A069479FC3B6B9FFB86B5AF" box="[1009,1118,475,493]" italics="true" pageId="8" pageNumber="373">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7697F9230A066B8EFB776B9FFA48B5AF" box="[1213,1424,475,493]" gridcol="4" gridrow="0" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FB2F6B9FFAB2B5AF" authorityName="H. L. Clark" authorityYear="1917" box="[1253,1386,475,493]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="pentacanthum">
|
||
<emphasis id="B9024E0F0A069479FB2F6B9FFAB2B5AF" box="[1253,1386,475,493]" italics="true" pageId="8" pageNumber="373">O. pentacanthum</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="7697F9230A066B8EFA556B9FF948B5AF" box="[1439,1680,475,493]" gridcol="5" gridrow="0" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FA276B9FF989B5AF" box="[1517,1617,475,493]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="peruanum">
|
||
<emphasis id="B9024E0F0A069479FA276B9FF989B5AF" box="[1517,1617,475,493]" italics="true" pageId="8" pageNumber="373">O. peruanum</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426BB8F948B64C" box="[136,1680,508,526]" gridrow="1" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<td id="7697F9230A066B8EFEA66BB8FD67B64C" box="[364,703,508,526]" gridcol="1" gridrow="1" pageId="8" pageNumber="373">Figures 1 – 3</td>
|
||
<td id="7697F9230A066B8EFD076BB8FC4AB64C" box="[717,914,508,526]" gridcol="2" gridrow="1" pageId="8" pageNumber="373">Figure 4(a–b)</td>
|
||
<td id="7697F9230A066B8EFC556BB8FB69B64C" box="[927,1201,508,526]" gridcol="3" gridrow="1" pageId="8" pageNumber="373">Figure 4(c)</td>
|
||
<td id="7697F9230A066B8EFB776BB8FA48B64C" box="[1213,1424,508,526]" gridcol="4" gridrow="1" pageId="8" pageNumber="373">Figure 4(d–e)</td>
|
||
<td id="7697F9230A066B8EFA556BB8F948B64C" box="[1439,1680,508,526]" gridcol="5" gridrow="1" pageId="8" pageNumber="373">Figure 4(f)</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF42685FF948B66F" box="[136,1680,539,557]" gridrow="2" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF42685FFE89B66F" box="[136,337,539,557]" gridcol="0" gridrow="2" pageId="8" pageNumber="373">Disc granules</th>
|
||
<td id="7697F9230A066B8EFEA6685FFD67B66F" box="[364,703,539,557]" gridcol="1" gridrow="2" pageId="8" pageNumber="373">Rounded</td>
|
||
<td id="7697F9230A066B8EFD07685FFC4AB66F" box="[717,914,539,557]" gridcol="2" gridrow="2" pageId="8" pageNumber="373">Rounded</td>
|
||
<td id="7697F9230A066B8EFC55685FFB69B66F" box="[927,1201,539,557]" gridcol="3" gridrow="2" pageId="8" pageNumber="373">Rounded</td>
|
||
<td id="7697F9230A066B8EFB77685FFA48B66F" box="[1213,1424,539,557]" gridcol="4" gridrow="2" pageId="8" pageNumber="373">Rounded</td>
|
||
<td id="7697F9230A066B8EFA55685FF948B66F" box="[1439,1680,539,557]" gridcol="5" gridrow="2" pageId="8" pageNumber="373">Rounded</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426876F948B606" box="[136,1680,562,580]" gridrow="3" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426876FE89B606" box="[136,337,562,580]" gridcol="0" gridrow="3" pageId="8" pageNumber="373">Radial shields</th>
|
||
<td id="7697F9230A066B8EFEA66876FD67B606" box="[364,703,562,580]" gridcol="1" gridrow="3" pageId="8" pageNumber="373">Usually covered</td>
|
||
<td id="7697F9230A066B8EFD076876FC4AB606" box="[717,914,562,580]" gridcol="2" gridrow="3" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFC556876FB69B606" box="[927,1201,562,580]" gridcol="3" gridrow="3" pageId="8" pageNumber="373">Naked</td>
|
||
<td id="7697F9230A066B8EFB776876FA48B606" box="[1213,1424,562,580]" gridcol="4" gridrow="3" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFA556876F948B606" box="[1439,1680,562,580]" gridcol="5" gridrow="3" pageId="8" pageNumber="373">Covered/naked</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF42680DF948B619" box="[136,1680,585,603]" gridrow="4" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF42680DFE89B619" box="[136,337,585,603]" gridcol="0" gridrow="4" pageId="8" pageNumber="373">Oral shields</th>
|
||
<td id="7697F9230A066B8EFEA6680DFD67B619" box="[364,703,585,603]" gridcol="1" gridrow="4" pageId="8" pageNumber="373">Wider than long, subtriangular</td>
|
||
<td id="7697F9230A066B8EFD07680DFC4AB619" box="[717,914,585,603]" gridcol="2" gridrow="4" pageId="8" pageNumber="373">Wider than long,</td>
|
||
<td id="7697F9230A066B8EFC55680DFB69B619" box="[927,1201,585,603]" gridcol="3" gridrow="4" pageId="8" pageNumber="373">Wider than long, subtriangular</td>
|
||
<td id="7697F9230A066B8EFB77680DFA48B619" box="[1213,1424,585,603]" gridcol="4" gridrow="4" pageId="8" pageNumber="373">Wider than long,</td>
|
||
<td id="7697F9230A066B8EFA55680DF948B619" box="[1439,1680,585,603]" gridcol="5" gridrow="4" pageId="8" pageNumber="373">Wider than long, subtriangular</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426825F948B631" box="[136,1680,609,627]" gridrow="5" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFD076825FC4AB631" box="[717,914,609,627]" gridcol="2" gridrow="5" pageId="8" pageNumber="373">subtriangular</td>
|
||
<td id="7697F9230A066B8EFB776825FA48B631" box="[1213,1424,609,627]" gridcol="4" gridrow="5" pageId="8" pageNumber="373">subtriangular</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF42683CF948B6C8" box="[136,1680,632,650]" gridrow="6" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF42683CFE89B6C8" box="[136,337,632,650]" gridcol="0" gridrow="6" pageId="8" pageNumber="373">Adoral shields</th>
|
||
<td id="7697F9230A066B8EFEA6683CFD67B6C8" box="[364,703,632,650]" gridcol="1" gridrow="6" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFD07683CFC4AB6C8" box="[717,914,632,650]" gridcol="2" gridrow="6" pageId="8" pageNumber="373">Naked</td>
|
||
<td id="7697F9230A066B8EFC55683CFB69B6C8" box="[927,1201,632,650]" gridcol="3" gridrow="6" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFB77683CFA48B6C8" box="[1213,1424,632,650]" gridcol="4" gridrow="6" pageId="8" pageNumber="373">Naked</td>
|
||
<td id="7697F9230A066B8EFA55683CF948B6C8" box="[1439,1680,632,650]" gridcol="5" gridrow="6" pageId="8" pageNumber="373">Covered</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4268CBF948B6E3" box="[136,1680,655,673]" gridrow="7" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF4268CBFE89B6E3" box="[136,337,655,673]" gridcol="0" gridrow="7" pageId="8" pageNumber="373">Distal genital slit</th>
|
||
<td id="7697F9230A066B8EFEA668CBFD67B6E3" box="[364,703,655,673]" gridcol="1" gridrow="7" pageId="8" pageNumber="373">Scales and granules</td>
|
||
<td id="7697F9230A066B8EFD0768CBFC4AB6E3" box="[717,914,655,673]" gridcol="2" gridrow="7" pageId="8" pageNumber="373">Scales and granules</td>
|
||
<td id="7697F9230A066B8EFC5568CBFB69B6E3" box="[927,1201,655,673]" gridcol="3" gridrow="7" pageId="8" pageNumber="373">Scales and granules</td>
|
||
<td id="7697F9230A066B8EFB7768CBFA48B6E3" box="[1213,1424,655,673]" gridcol="4" gridrow="7" pageId="8" pageNumber="373">Scales and granules</td>
|
||
<td id="7697F9230A066B8EFA5568CBF948B6E3" box="[1439,1680,655,673]" gridcol="5" gridrow="7" pageId="8" pageNumber="373">Scales and granules</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4268E3F948B6FB" box="[136,1680,679,697]" gridrow="8" pageId="8" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="7697F9230A066B8EFF4268E3FE89B6FB" box="[136,337,679,697]" gridcol="0" gridrow="8" pageId="8" pageNumber="373">ornamentation</th>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4268FAF948B692" box="[136,1680,702,720]" gridrow="9" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF4268FAFE89B692" box="[136,337,702,720]" gridcol="0" gridrow="9" pageId="8" pageNumber="373">Dorsal arm plates</th>
|
||
<td id="7697F9230A066B8EFEA668FAFD67B692" box="[364,703,702,720]" gridcol="1" gridrow="9" pageId="8" pageNumber="373">Divided (typically 2 pieces, up to 5)</td>
|
||
<td id="7697F9230A066B8EFD0768FAFC4AB692" box="[717,914,702,720]" gridcol="2" gridrow="9" pageId="8" pageNumber="373">Entire</td>
|
||
<td id="7697F9230A066B8EFC5568FAFB69B692" box="[927,1201,702,720]" gridcol="3" gridrow="9" pageId="8" pageNumber="373">Mostly entire (can present some</td>
|
||
<td id="7697F9230A066B8EFB7768FAFA48B692" box="[1213,1424,702,720]" gridcol="4" gridrow="9" pageId="8" pageNumber="373">Entire</td>
|
||
<td id="7697F9230A066B8EFA5568FAF948B692" box="[1439,1680,702,720]" gridcol="5" gridrow="9" pageId="8" pageNumber="373">Divided (up to 8 pieces)</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426892F948B6AA" box="[136,1680,726,744]" gridrow="10" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFC556892FB69B6AA" box="[927,1201,726,744]" gridcol="3" gridrow="10" pageId="8" pageNumber="373">DAPs divided into 2 pieces)</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4268A9F948B6BD" box="[136,1680,749,767]" gridrow="11" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF4268A9FE89B6BD" box="[136,337,749,767]" gridcol="0" gridrow="11" pageId="8" pageNumber="373">Maximum number of arm</th>
|
||
<td id="7697F9230A066B8EFEA668A9FD67B6BD" box="[364,703,749,767]" gridcol="1" gridrow="11" pageId="8" pageNumber="373">11</td>
|
||
<td id="7697F9230A066B8EFD0768A9FC4AB6BD" box="[717,914,749,767]" gridcol="2" gridrow="11" pageId="8" pageNumber="373">11</td>
|
||
<td id="7697F9230A066B8EFC5568A9FB69B6BD" box="[927,1201,749,767]" gridcol="3" gridrow="11" pageId="8" pageNumber="373">12</td>
|
||
<td id="7697F9230A066B8EFB7768A9FA48B6BD" box="[1213,1424,749,767]" gridcol="4" gridrow="11" pageId="8" pageNumber="373">6</td>
|
||
<td id="7697F9230A066B8EFA5568A9F948B6BD" box="[1439,1680,749,767]" gridcol="5" gridrow="11" pageId="8" pageNumber="373">11</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426941F948B755" box="[136,1680,773,791]" gridrow="12" pageId="8" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="7697F9230A066B8EFF426941FE89B755" box="[136,337,773,791]" gridcol="0" gridrow="12" pageId="8" pageNumber="373">spines</th>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426958F948B76C" box="[136,1680,796,814]" gridrow="13" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426958FE89B76C" box="[136,337,796,814]" gridcol="0" gridrow="13" pageId="8" pageNumber="373">Dorsal colour pattern</th>
|
||
<td id="7697F9230A066B8EFEA66958FD67B76C" box="[364,703,796,814]" gridcol="1" gridrow="13" pageId="8" pageNumber="373">In vivo: disc dark olive green to brown.</td>
|
||
<td id="7697F9230A066B8EFD076958FC4AB76C" box="[717,914,796,814]" gridcol="2" gridrow="13" pageId="8" pageNumber="373">In vivo: disc dark brown,</td>
|
||
<td id="7697F9230A066B8EFC556958FB69B76C" box="[927,1201,796,814]" gridcol="3" gridrow="13" pageId="8" pageNumber="373">In vivo: disc green, brown, or grey,</td>
|
||
<td id="7697F9230A066B8EFB776958FA48B76C" box="[1213,1424,796,814]" gridcol="4" gridrow="13" pageId="8" pageNumber="373">In vivo: disc orange with</td>
|
||
<td id="7697F9230A066B8EFA556958F948B76C" box="[1439,1680,796,814]" gridcol="5" gridrow="13" pageId="8" pageNumber="373">In vivo: disc and arms brown,</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426970F948B704" box="[136,1680,820,838]" gridrow="14" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<td id="7697F9230A066B8EFEA66970FD67B704" box="[364,703,820,838]" gridcol="1" gridrow="14" pageId="8" pageNumber="373">Arms with lighter transverse bands, and</td>
|
||
<td id="7697F9230A066B8EFD076970FC4AB704" box="[717,914,820,838]" gridcol="2" gridrow="14" pageId="8" pageNumber="373">mottled with white.</td>
|
||
<td id="7697F9230A066B8EFC556970FB69B704" box="[927,1201,820,838]" gridcol="3" gridrow="14" pageId="8" pageNumber="373">mottled with yellow, orange,</td>
|
||
<td id="7697F9230A066B8EFB776970FA48B704" box="[1213,1424,820,838]" gridcol="4" gridrow="14" pageId="8" pageNumber="373">dark brown blotches.</td>
|
||
<td id="7697F9230A066B8EFA556970F948B704" box="[1439,1680,820,838]" gridcol="5" gridrow="14" pageId="8" pageNumber="373">speckled with beige</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426908F948B71C" box="[136,1680,844,862]" gridrow="15" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<td id="7697F9230A066B8EFEA66908FD67B71C" box="[364,703,844,862]" gridcol="1" gridrow="15" pageId="8" pageNumber="373">parallel white bands in the median and</td>
|
||
<td id="7697F9230A066B8EFD076908FC4AB71C" box="[717,914,844,862]" gridcol="2" gridrow="15" pageId="8" pageNumber="373">Arms with transverse</td>
|
||
<td id="7697F9230A066B8EFC556908FB69B71C" box="[927,1201,844,862]" gridcol="3" gridrow="15" pageId="8" pageNumber="373">red, pink or white. Arms with</td>
|
||
<td id="7697F9230A066B8EFB776908FA48B71C" box="[1213,1424,844,862]" gridcol="4" gridrow="15" pageId="8" pageNumber="373">Arms with transverse</td>
|
||
<td id="7697F9230A066B8EFA556908F948B71C" box="[1439,1680,844,862]" gridcol="5" gridrow="15" pageId="8" pageNumber="373">Preserved: as in vivo.</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426921F948B735" box="[136,1680,869,887]" gridrow="16" pageId="8" pageNumber="373" rowspan-0="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFEA66921FD67B735" box="[364,703,869,887]" gridcol="1" gridrow="16" pageId="8" pageNumber="373">distal arm sections</td>
|
||
<td id="7697F9230A066B8EFD076921FC4AB735" box="[717,914,869,887]" gridcol="2" gridrow="16" pageId="8" pageNumber="373">white bands</td>
|
||
<td id="7697F9230A066B8EFC556921FB69B735" box="[927,1201,869,887]" gridcol="3" gridrow="16" pageId="8" pageNumber="373">transverse bands</td>
|
||
<td id="7697F9230A066B8EFB776921FA48B735" box="[1213,1424,869,887]" gridcol="4" gridrow="16" pageId="8" pageNumber="373">brown bands</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426938F948B7CC" box="[136,1680,892,910]" gridrow="17" pageId="8" pageNumber="373" rowspan-0="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFEA66938FD67B7CC" box="[364,703,892,910]" gridcol="1" gridrow="17" pageId="8" pageNumber="373">Preserved: disc dark grey to brown. Arms as</td>
|
||
<td id="7697F9230A066B8EFD076938FC4AB7CC" box="[717,914,892,910]" gridcol="2" gridrow="17" pageId="8" pageNumber="373">Preserved: as in vivo.</td>
|
||
<td id="7697F9230A066B8EFC556938FB69B7CC" box="[927,1201,892,910]" gridcol="3" gridrow="17" pageId="8" pageNumber="373">Preserved: disc with variable</td>
|
||
<td id="7697F9230A066B8EFB776938FA48B7CC" box="[1213,1424,892,910]" gridcol="4" gridrow="17" pageId="8" pageNumber="373">Preserved: disc light</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4269D0F948B7E4" box="[136,1680,916,934]" gridrow="18" pageId="8" pageNumber="373" rowspan-0="1" rowspan-2="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFEA669D0FD67B7E4" box="[364,703,916,934]" gridcol="1" gridrow="18" pageId="8" pageNumber="373">in vivo, but with subtle lighter transverse</td>
|
||
<td id="7697F9230A066B8EFC5569D0FB69B7E4" box="[927,1201,916,934]" gridcol="3" gridrow="18" pageId="8" pageNumber="373">shades of brown. Arms as</td>
|
||
<td id="7697F9230A066B8EFB7769D0FA48B7E4" box="[1213,1424,916,934]" gridcol="4" gridrow="18" pageId="8" pageNumber="373">brown, with the centre</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4269E8F948B7FC" box="[136,1680,940,958]" gridrow="19" pageId="8" pageNumber="373" rowspan-0="1" rowspan-2="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFEA669E8FD67B7FC" box="[364,703,940,958]" gridcol="1" gridrow="19" pageId="8" pageNumber="373">bands.</td>
|
||
<td id="7697F9230A066B8EFC5569E8FB69B7FC" box="[927,1201,940,958]" gridcol="3" gridrow="19" pageId="8" pageNumber="373">in vivo.</td>
|
||
<td id="7697F9230A066B8EFB7769E8FA48B7FC" box="[1213,1424,940,958]" gridcol="4" gridrow="19" pageId="8" pageNumber="373">mottled with dark</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426981F948B795" box="[136,1680,965,983]" gridrow="20" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFB776981FA48B795" box="[1213,1424,965,983]" gridcol="4" gridrow="20" pageId="8" pageNumber="373">brown. Arms as in vivo.</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4269A1F948B7B5" box="[136,1680,997,1015]" gridrow="21" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<td id="7697F9230A066B8EFEA669A1FD67B7B5" box="[364,703,997,1015]" gridcol="1" gridrow="21" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FE1769A1FD97B7B5" authorityName="Caso" authorityYear="1986" box="[477,591,997,1015]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="sodipallaresi">
|
||
<emphasis id="B9024E0F0A069479FE1769A1FD97B7B5" box="[477,591,997,1015]" italics="true" pageId="8" pageNumber="373">O. sodipallaresi</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="7697F9230A066B8EFD0769A1FC4AB7B5" box="[717,914,997,1015]" gridcol="2" gridrow="21" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FCD969A1FC94B7B5" authorityName="Lyman" authorityYear="1860" box="[787,844,997,1015]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A069479FCD969A1FC94B7B5" box="[787,844,997,1015]" italics="true" pageId="8" pageNumber="373">O. teres</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="7697F9230A066B8EFC5569A1FB69B7B5" box="[927,1201,997,1015]" gridcol="3" gridrow="21" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FC2069A1FBBDB7B5" box="[1002,1125,997,1015]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="subSpecies" species="teres" subSpecies="unicolor">
|
||
<emphasis id="B9024E0F0A069479FC2069A1FBBDB7B5" box="[1002,1125,997,1015]" italics="true" pageId="8" pageNumber="373">O. teres unicolor</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="7697F9230A066B8EFB7769A1FA48B7B5" box="[1213,1424,997,1015]" gridcol="4" gridrow="21" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FB3669A1FA8AB7B5" authorityName="Hendler" authorityYear="1996" box="[1276,1362,997,1015]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="vansyoci">
|
||
<emphasis id="B9024E0F0A069479FB3669A1FA8AB7B5" box="[1276,1362,997,1015]" italics="true" pageId="8" pageNumber="373">O. vansyoci</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="7697F9230A066B8EFA5569A1F948B7B5" box="[1439,1680,997,1015]" gridcol="5" gridrow="21" pageId="8" pageNumber="373">
|
||
<taxonomicName id="4C76E99E0A069479FA2369A1F98DB7B5" authorityName="Lutken" authorityYear="1856" box="[1513,1621,997,1015]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="8" pageNumber="373" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B9024E0F0A069479FA2369A1F98DB7B5" box="[1513,1621,997,1015]" italics="true" pageId="8" pageNumber="373">O. variegatum</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF4269B8F948B04C" box="[136,1680,1020,1038]" gridrow="22" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<td id="7697F9230A066B8EFEA669B8FD67B04C" box="[364,703,1020,1038]" gridcol="1" gridrow="22" pageId="8" pageNumber="373">Figure 4(g–i)</td>
|
||
<td id="7697F9230A066B8EFD0769B8FC4AB04C" box="[717,914,1020,1038]" gridcol="2" gridrow="22" pageId="8" pageNumber="373">Figure 4(j)</td>
|
||
<td id="7697F9230A066B8EFC5569B8FB69B04C" box="[927,1201,1020,1038]" gridcol="3" gridrow="22" pageId="8" pageNumber="373">Figure 4(k)</td>
|
||
<td id="7697F9230A066B8EFB7769B8FA48B04C" box="[1213,1424,1020,1038]" gridcol="4" gridrow="22" pageId="8" pageNumber="373">Figure 4(l–m)</td>
|
||
<td id="7697F9230A066B8EFA5569B8F948B04C" box="[1439,1680,1020,1038]" gridcol="5" gridrow="22" pageId="8" pageNumber="373">Figure 4(n–o)</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426E58F948B06C" box="[136,1680,1052,1070]" gridrow="23" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426E58FE89B06C" box="[136,337,1052,1070]" gridcol="0" gridrow="23" pageId="8" pageNumber="373">Disc granules</th>
|
||
<td id="7697F9230A066B8EFEA66E58FD67B06C" box="[364,703,1052,1070]" gridcol="1" gridrow="23" pageId="8" pageNumber="373">Rounded</td>
|
||
<td id="7697F9230A066B8EFD076E58FC4AB06C" box="[717,914,1052,1070]" gridcol="2" gridrow="23" pageId="8" pageNumber="373">Rounded</td>
|
||
<td id="7697F9230A066B8EFC556E58FB69B06C" box="[927,1201,1052,1070]" gridcol="3" gridrow="23" pageId="8" pageNumber="373">Rounded</td>
|
||
<td id="7697F9230A066B8EFB776E58FA48B06C" box="[1213,1424,1052,1070]" gridcol="4" gridrow="23" pageId="8" pageNumber="373">Flattened</td>
|
||
<td id="7697F9230A066B8EFA556E58F948B06C" box="[1439,1680,1052,1070]" gridcol="5" gridrow="23" pageId="8" pageNumber="373">Rounded</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426E77F948B007" box="[136,1680,1075,1093]" gridrow="24" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426E77FE89B007" box="[136,337,1075,1093]" gridcol="0" gridrow="24" pageId="8" pageNumber="373">Radial shields</th>
|
||
<td id="7697F9230A066B8EFEA66E77FD67B007" box="[364,703,1075,1093]" gridcol="1" gridrow="24" pageId="8" pageNumber="373">Covered/naked</td>
|
||
<td id="7697F9230A066B8EFD076E77FC4AB007" box="[717,914,1075,1093]" gridcol="2" gridrow="24" pageId="8" pageNumber="373">Covered/naked</td>
|
||
<td id="7697F9230A066B8EFC556E77FB69B007" box="[927,1201,1075,1093]" gridcol="3" gridrow="24" pageId="8" pageNumber="373">Naked</td>
|
||
<td id="7697F9230A066B8EFB776E77FA48B007" box="[1213,1424,1075,1093]" gridcol="4" gridrow="24" pageId="8" pageNumber="373">Naked</td>
|
||
<td id="7697F9230A066B8EFA556E77F948B007" box="[1439,1680,1075,1093]" gridcol="5" gridrow="24" pageId="8" pageNumber="373">Covered</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426E0FF948B01F" box="[136,1680,1099,1117]" gridrow="25" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426E0FFE89B01F" box="[136,337,1099,1117]" gridcol="0" gridrow="25" pageId="8" pageNumber="373">Oral shields</th>
|
||
<td id="7697F9230A066B8EFEA66E0FFD67B01F" box="[364,703,1099,1117]" gridcol="1" gridrow="25" pageId="8" pageNumber="373">Wider than long, rounded</td>
|
||
<td id="7697F9230A066B8EFD076E0FFC4AB01F" box="[717,914,1099,1117]" gridcol="2" gridrow="25" pageId="8" pageNumber="373">Wider than long,</td>
|
||
<td id="7697F9230A066B8EFC556E0FFB69B01F" box="[927,1201,1099,1117]" gridcol="3" gridrow="25" pageId="8" pageNumber="373">Wider than long, subtriangular</td>
|
||
<td id="7697F9230A066B8EFB776E0FFA48B01F" box="[1213,1424,1099,1117]" gridcol="4" gridrow="25" pageId="8" pageNumber="373">Wider than long,</td>
|
||
<td id="7697F9230A066B8EFA556E0FF948B01F" box="[1439,1680,1099,1117]" gridcol="5" gridrow="25" pageId="8" pageNumber="373">Longer than wide, oval</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426E26F948B036" box="[136,1680,1122,1140]" gridrow="26" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
|
||
<td id="7697F9230A066B8EFD076E26FC4AB036" box="[717,914,1122,1140]" gridcol="2" gridrow="26" pageId="8" pageNumber="373">subtriangular</td>
|
||
<td id="7697F9230A066B8EFB776E26FA48B036" box="[1213,1424,1122,1140]" gridcol="4" gridrow="26" pageId="8" pageNumber="373">subtriangular</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426E3EF948B0CE" box="[136,1680,1146,1164]" gridrow="27" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426E3EFE89B0CE" box="[136,337,1146,1164]" gridcol="0" gridrow="27" pageId="8" pageNumber="373">Adoral shields</th>
|
||
<td id="7697F9230A066B8EFEA66E3EFD67B0CE" box="[364,703,1146,1164]" gridcol="1" gridrow="27" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFD076E3EFC4AB0CE" box="[717,914,1146,1164]" gridcol="2" gridrow="27" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFC556E3EFB69B0CE" box="[927,1201,1146,1164]" gridcol="3" gridrow="27" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFB776E3EFA48B0CE" box="[1213,1424,1146,1164]" gridcol="4" gridrow="27" pageId="8" pageNumber="373">Covered</td>
|
||
<td id="7697F9230A066B8EFA556E3EF948B0CE" box="[1439,1680,1146,1164]" gridcol="5" gridrow="27" pageId="8" pageNumber="373">Naked</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426ED5F948B0E1" box="[136,1680,1169,1187]" gridrow="28" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426ED5FE89B0E1" box="[136,337,1169,1187]" gridcol="0" gridrow="28" pageId="8" pageNumber="373">Distal genital slit</th>
|
||
<td id="7697F9230A066B8EFEA66ED5FD67B0E1" box="[364,703,1169,1187]" gridcol="1" gridrow="28" pageId="8" pageNumber="373">Granules</td>
|
||
<td id="7697F9230A066B8EFD076ED5FC4AB0E1" box="[717,914,1169,1187]" gridcol="2" gridrow="28" pageId="8" pageNumber="373">Scales and granules</td>
|
||
<td id="7697F9230A066B8EFC556ED5FB69B0E1" box="[927,1201,1169,1187]" gridcol="3" gridrow="28" pageId="8" pageNumber="373">Scales and granules</td>
|
||
<td id="7697F9230A066B8EFB776ED5FA48B0E1" box="[1213,1424,1169,1187]" gridcol="4" gridrow="28" pageId="8" pageNumber="373">Granules</td>
|
||
<td id="7697F9230A066B8EFA556ED5F948B0E1" box="[1439,1680,1169,1187]" gridcol="5" gridrow="28" pageId="8" pageNumber="373">Granules</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426EEDF948B0F9" box="[136,1680,1193,1211]" gridrow="29" pageId="8" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="7697F9230A066B8EFF426EEDFE89B0F9" box="[136,337,1193,1211]" gridcol="0" gridrow="29" pageId="8" pageNumber="373">ornamentation</th>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426E84F948B090" box="[136,1680,1216,1234]" gridrow="30" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426E84FE89B090" box="[136,337,1216,1234]" gridcol="0" gridrow="30" pageId="8" pageNumber="373">Dorsal arm plates</th>
|
||
<td id="7697F9230A066B8EFEA66E84FD67B090" box="[364,703,1216,1234]" gridcol="1" gridrow="30" pageId="8" pageNumber="373">Divided (up to 3 pieces)</td>
|
||
<td id="7697F9230A066B8EFD076E84FC4AB090" box="[717,914,1216,1234]" gridcol="2" gridrow="30" pageId="8" pageNumber="373">Divided (up to 5 pieces)</td>
|
||
<td id="7697F9230A066B8EFC556E84FB69B090" box="[927,1201,1216,1234]" gridcol="3" gridrow="30" pageId="8" pageNumber="373">Divided (up to 6 pieces)</td>
|
||
<td id="7697F9230A066B8EFB776E84FA48B090" box="[1213,1424,1216,1234]" gridcol="4" gridrow="30" pageId="8" pageNumber="373">Divided (up to 8 pieces)</td>
|
||
<td id="7697F9230A066B8EFA556E84F948B090" box="[1439,1680,1216,1234]" gridcol="5" gridrow="30" pageId="8" pageNumber="373">Entire</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426E93F948B0AB" box="[136,1680,1239,1257]" gridrow="31" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426E93FE89B0AB" box="[136,337,1239,1257]" gridcol="0" gridrow="31" pageId="8" pageNumber="373">Maximum number of arm</th>
|
||
<td id="7697F9230A066B8EFEA66E93FD67B0AB" box="[364,703,1239,1257]" gridcol="1" gridrow="31" pageId="8" pageNumber="373">10</td>
|
||
<td id="7697F9230A066B8EFD076E93FC4AB0AB" box="[717,914,1239,1257]" gridcol="2" gridrow="31" pageId="8" pageNumber="373">11</td>
|
||
<td id="7697F9230A066B8EFC556E93FB69B0AB" box="[927,1201,1239,1257]" gridcol="3" gridrow="31" pageId="8" pageNumber="373">13</td>
|
||
<td id="7697F9230A066B8EFB776E93FA48B0AB" box="[1213,1424,1239,1257]" gridcol="4" gridrow="31" pageId="8" pageNumber="373">9</td>
|
||
<td id="7697F9230A066B8EFA556E93F948B0AB" box="[1439,1680,1239,1257]" gridcol="5" gridrow="31" pageId="8" pageNumber="373">9</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426EABF948B143" box="[136,1680,1263,1281]" gridrow="32" pageId="8" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="7697F9230A066B8EFF426EABFE89B143" box="[136,337,1263,1281]" gridcol="0" gridrow="32" pageId="8" pageNumber="373">spines</th>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426F42F948B15A" box="[136,1680,1286,1304]" gridrow="33" pageId="8" pageNumber="373">
|
||
<th id="7697F9230A066B8EFF426F42FE89B15A" box="[136,337,1286,1304]" gridcol="0" gridrow="33" pageId="8" pageNumber="373">Dorsal colour pattern</th>
|
||
<td id="7697F9230A066B8EFEA66F42FD67B15A" box="[364,703,1286,1304]" gridcol="1" gridrow="33" pageId="8" pageNumber="373">In vivo: not documented.</td>
|
||
<td id="7697F9230A066B8EFD076F42FC4AB15A" box="[717,914,1286,1304]" gridcol="2" gridrow="33" pageId="8" pageNumber="373">In vivo:not documented.</td>
|
||
<td id="7697F9230A066B8EFC556F42FB69B15A" box="[927,1201,1286,1304]" gridcol="3" gridrow="33" pageId="8" pageNumber="373">In vivo:</td>
|
||
<td id="7697F9230A066B8EFB776F42FA48B15A" box="[1213,1424,1286,1304]" gridcol="4" gridrow="33" pageId="8" pageNumber="373">In vivo: disc and arms dark</td>
|
||
<td id="7697F9230A066B8EFA556F42F948B15A" box="[1439,1680,1286,1304]" gridcol="5" gridrow="33" pageId="8" pageNumber="373">In vivo: disc olive green with</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426F5AF948B172" box="[136,1680,1310,1328]" gridrow="34" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<td id="7697F9230A066B8EFEA66F5AFD67B172" box="[364,703,1310,1328]" gridcol="1" gridrow="34" pageId="8" pageNumber="373">Preserved: disc and arms brown, speckled</td>
|
||
<td id="7697F9230A066B8EFD076F5AFC4AB172" box="[717,914,1310,1328]" gridcol="2" gridrow="34" pageId="8" pageNumber="373">Preserved: disc and arms</td>
|
||
<td id="7697F9230A066B8EFC556F5AFB69B172" box="[927,1201,1310,1328]" gridcol="3" gridrow="34" pageId="8" pageNumber="373">not documented.</td>
|
||
<td id="7697F9230A066B8EFB776F5AFA48B172" box="[1213,1424,1310,1328]" gridcol="4" gridrow="34" pageId="8" pageNumber="373">brown.</td>
|
||
<td id="7697F9230A066B8EFA556F5AF948B172" box="[1439,1680,1310,1328]" gridcol="5" gridrow="34" pageId="8" pageNumber="373">dark pink or red. Arms with</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426F72F948B10A" box="[136,1680,1334,1352]" gridrow="35" pageId="8" pageNumber="373" rowspan-0="1">
|
||
<td id="7697F9230A066B8EFEA66F72FD67B10A" box="[364,703,1334,1352]" gridcol="1" gridrow="35" pageId="8" pageNumber="373">with beige.</td>
|
||
<td id="7697F9230A066B8EFD076F72FC4AB10A" box="[717,914,1334,1352]" gridcol="2" gridrow="35" pageId="8" pageNumber="373">brown, speckled with</td>
|
||
<td id="7697F9230A066B8EFC556F72FB69B10A" box="[927,1201,1334,1352]" gridcol="3" gridrow="35" pageId="8" pageNumber="373">Preserved: disc and arms dark</td>
|
||
<td id="7697F9230A066B8EFB776F72FA48B10A" box="[1213,1424,1334,1352]" gridcol="4" gridrow="35" pageId="8" pageNumber="373">Preserved: as in vivo.</td>
|
||
<td id="7697F9230A066B8EFA556F72F948B10A" box="[1439,1680,1334,1352]" gridcol="5" gridrow="35" pageId="8" pageNumber="373">transverse olive green and</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426F0AF948B122" box="[136,1680,1358,1376]" gridrow="36" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-4="1">
|
||
<td id="7697F9230A066B8EFD076F0AFC4AB122" box="[717,914,1358,1376]" gridcol="2" gridrow="36" pageId="8" pageNumber="373">beige.</td>
|
||
<td id="7697F9230A066B8EFC556F0AFB69B122" box="[927,1201,1358,1376]" gridcol="3" gridrow="36" pageId="8" pageNumber="373">brown.</td>
|
||
<td id="7697F9230A066B8EFA556F0AF948B122" box="[1439,1680,1358,1376]" gridcol="5" gridrow="36" pageId="8" pageNumber="373">dark pink/red bands</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426F22F948B13A" box="[136,1680,1382,1400]" gridrow="37" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<td id="7697F9230A066B8EFA556F22F948B13A" box="[1439,1680,1382,1400]" gridcol="5" gridrow="37" pageId="8" pageNumber="373">Preserved: light brown. Arms</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426F3AF948B1D2" box="[136,1680,1406,1424]" gridrow="38" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<td id="7697F9230A066B8EFA556F3AF948B1D2" box="[1439,1680,1406,1424]" gridcol="5" gridrow="38" pageId="8" pageNumber="373">with transverse dark brown</td>
|
||
</tr>
|
||
<tr id="3546905F0A066B8EFF426FD2F948B1EA" box="[136,1680,1430,1448]" gridrow="39" pageId="8" pageNumber="373" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<td id="7697F9230A066B8EFA556FD2F948B1EA" box="[1439,1680,1430,1448]" gridcol="5" gridrow="39" pageId="8" pageNumber="373">bands.</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C36CC1960A079478FF6A6ACAFEFEB4EA" box="[160,294,142,168]" pageId="9" pageNumber="374" type="description">
|
||
<paragraph id="8BC9921D0A079478FF6A6ACAFEFEB4EA" blockId="9.[160,1156,142,1659]" box="[160,294,142,168]" pageId="9" pageNumber="374">
|
||
<emphasis id="B9024E0F0A079478FF6A6ACAFEFEB4EA" bold="true" box="[160,294,142,168]" italics="true" pageId="9" pageNumber="374">Description</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36CC1960A07947AFF6A6AF5FF33B755" lastPageId="11" lastPageNumber="376" pageId="9" pageNumber="374" type="materials_examined">
|
||
<paragraph id="8BC9921D0A079478FF6A6AF5FEEEB5A2" blockId="9.[160,1156,142,1659]" pageId="9" pageNumber="374">
|
||
<emphasis id="B9024E0F0A079478FF6A6AF5FECAB489" bold="true" box="[160,274,177,203]" italics="true" pageId="9" pageNumber="374">
|
||
<typeStatus id="54CD2CBF0A079478FF6A6AF5FED5B489" box="[160,269,177,203]" pageId="9" pageNumber="374" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
DD =
|
||
<quantity id="4C8E3FF80A079478FEA26AF6FE17B489" box="[360,463,178,203]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="9" pageNumber="374" unit="mm" value="15.9">15.9 mm</quantity>
|
||
, AL =
|
||
<quantity id="4C8E3FF80A079478FDD16AF6FD5AB489" box="[539,642,178,203]" metricMagnitude="-2" metricUnit="m" metricValue="4.93" pageId="9" pageNumber="374" unit="mm" value="49.3">49.3 mm</quantity>
|
||
. Disc nearly rounded, covered with rounded granules slightly separated from each other; size of the granules increasing from the disc centre towards the periphery. Dorsal disc granule density 52 per mm2. Radial shields covered with granules (
|
||
<figureCitation id="134D8E980A079478FE666B5EFDDDB571" box="[428,517,282,307]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(a)). Interradii covered with granules similar to those on the dorsal side. Four genital slits per interradius. Proximal genital slits oval, in contact with the distal section of the oral shield and the 1st LAP; reaching up to the 2nd VAP. Distal genital slits oval, twice longer than proximal genital slits, placed between 5th and 7th arm segments; surrounded by granules and, only next to the arm, imbricated scales (
|
||
<figureCitation id="134D8E980A079478FF626B83FEDDB5A2" box="[168,261,455,480]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(b)).
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A079478FF0A6BAEFBB0B779" blockId="9.[160,1156,142,1659]" pageId="9" pageNumber="374">
|
||
Oral shields broader than long, subtriangular; proximal edges convex forming a rounded apex; rounded lateral edges; straight distal edge. Madreporite with oval, central depression. Adoral shields covered with scattered granules, larger than those on the interradii. Jaws with 7–8 oral papillae: LyOs two times longer than broad, angled upwards; AdShSp the largest, subquadrangular; 2°AdShSp similar to AdShSp, smaller; LOPas 3–4 (rarely two), conical, pointed and elongated, slightly separated; IPa similar in shape to LOPas; TPa 1–2 at the apex of jaw, robust and pointed. Teeth, five: vT robust, subtriangular, larger than TPa; median teeth quadrangular; the dorsalmost the smallest, triangular and pointed. One OPRSp at each side of jaw, large, prominent. Oral plates covered with elongated, separated granules, larger than those on the adoral shields (
|
||
<figureCitation id="134D8E980A079478FC0C6966FBC5B779" box="[966,1053,802,827]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(b–c)).
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A079478FF0A6900FCC4B1E9" blockId="9.[160,1156,142,1659]" pageId="9" pageNumber="374">
|
||
Five arms flattened, tapering distally; one cut for molecular analysis, one complete probably regenerated from the midsection, one regenerating close to tip, two broken near tip. Dorsal arm base covered with granules and scales located laterally over the first three DAPs (
|
||
<figureCitation id="134D8E980A079478FEE669E8FE5DB787" box="[300,389,940,965]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(d)). DAPs wider than long, divided typically into two and up to five (only one DAP in all five arms) irregular pieces in approximately 2/3 of the arm; the number of pieces decreases distalwards (
|
||
<figureCitation id="134D8E980A079478FD4A69B6FD03B049" box="[640,731,1010,1035]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(d–f)). Distalmost DAPs entire, small, trapezoidal (
|
||
<figureCitation id="134D8E980A079478FEE16E50FE5AB06F" box="[299,386,1044,1069]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(f)). First VAP small, 1.5 as wide as long, with rounded edges (
|
||
<figureCitation id="134D8E980A079478FBE36E50FB5BB06F" box="[1065,1155,1044,1069]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(b)). Subsequent VAPs quadrangular, as long as wide; proximal edge straight, distal edge convex (
|
||
<figureCitation id="134D8E980A079478FF346E1EFE8DB031" box="[254,341,1114,1139]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(g–h)). Distalmost VAPs pentagonal, longer than wide (
|
||
<figureCitation id="134D8E980A079478FC776E1EFBCFB031" box="[957,1047,1114,1139]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(i)). A pair of pores are visible between the two proximalmost VAPs in all five arms (
|
||
<figureCitation id="134D8E980A079478FC086E38FBC1B0D7" box="[962,1049,1148,1173]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(b)). LAPs conspicuous, wider than long. LAP with up to eight arm spines, rarely nine. Arm spine sequence of the longest arm (right side): first 10 segments, 2, 2, 3, 3, 4, 4, 5, 6, 8, 8; 11th– 20th, eight arm spines; 21st–30th, 7–8; 31st–40th, 6–8; 41st–50th, 6–8; 51st–60th, 5–6; 61st– 70th, 3–5; 71st–75th, two. Arm spines conical with blunt tips, length 2/3 of LAP. Dorsalmost arm spine the shortest; ventralmost the longest and more robust, in contact with abradial tentacle scale of the following segment (
|
||
<figureCitation id="134D8E980A079478FD426F08FD3CB127" box="[648,740,1356,1381]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(d–j)). Two tentacle scales; adradial tentacle scale oval, elongated, 2/3 the length of VAP; abradial tentacle scale shorter, 2/3 the length of adradial one, subtriangular (
|
||
<figureCitation id="134D8E980A079478FDB26FD6FD08B1E9" box="[632,720,1426,1451]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(g–h)).
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A07947BFF0A6FF0FD7DB4AF" blockId="9.[160,1156,142,1659]" lastBlockId="10.[160,1155,142,237]" lastPageId="10" lastPageNumber="375" pageId="9" pageNumber="374">
|
||
General colouration dark grey (specimen in ethanol;
|
||
<figureCitation id="134D8E980A079478FCC76FF0FCBCB18F" box="[781,868,1460,1485]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(k)). Dorsal side: disc dark grey, with scattered white granules (
|
||
<figureCitation id="134D8E980A079478FDF36F93FD57B1B2" box="[569,655,1495,1520]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(a)). Arms grey with subtle lighter transverse bands; 1–3 adjacent DAPs and LAPs with a pair of parallel white blotches every 3–15 segments in the median and distal arm sections, macroscopically perceived as parallel longitudinal white bands (
|
||
<figureCitation id="134D8E980A079478FE0F6C7BFDC3B21A" box="[453,539,1599,1624]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(e, j–k)). Ventral side: interradii dark grey with scattered white granules (
|
||
<figureCitation id="134D8E980A079478FE9E6C26FE74B239" box="[340,428,1634,1659]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(b)). Jaws and arms light grey, oral papillae beige (
|
||
<figureCitation id="134D8E980A079478FC296C26FBE1B239" box="[995,1081,1634,1659]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="9" pageNumber="374">Figure 1</figureCitation>
|
||
(b–c)). Some VAPs with a white spot located randomly (
|
||
<figureCitation id="134D8E980A04947BFD136ACAFCECB4E5" box="[729,820,142,167]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="10" pageNumber="375">Figure 1</figureCitation>
|
||
(b, h)). LAPs grey, some with a dorsal white blotch. Dorsalmost arm spines grey, becoming clearer to the ventralmost side, with a white spot on the tip (
|
||
<figureCitation id="134D8E980A04947BFDEF6A90FDA6B4AF" box="[549,638,212,237]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="10" pageNumber="375">Figure 1</figureCitation>
|
||
(j)).
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A04947BFF6A6B5CFDE8B570" blockId="10.[160,1156,280,1727]" box="[160,560,280,306]" pageId="10" pageNumber="375">
|
||
<emphasis id="B9024E0F0A04947BFF6A6B5CFDE8B570" bold="true" box="[160,560,280,306]" italics="true" pageId="10" pageNumber="375">
|
||
<typeStatus id="54CD2CBF0A04947BFF6A6B5CFED2B570" box="[160,266,280,306]" pageId="10" pageNumber="375" type="paratype">Paratype</typeStatus>
|
||
and non-type variations
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A04947BFF6A6B78FC5CB7E0" blockId="10.[160,1156,280,1727]" pageId="10" pageNumber="375">
|
||
Specimens in the
|
||
<typeStatus id="54CD2CBF0A04947BFEAE6B78FE4EB517" box="[356,406,316,341]" pageId="10" pageNumber="375">type</typeStatus>
|
||
series varied from
|
||
<quantity id="4C8E3FF80A04947BFDAF6B78FCD0B517" box="[613,776,316,341]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" metricValueMax="2.11" metricValueMin="0.37" pageId="10" pageNumber="375" unit="mm" value="12.4" valueMax="21.1" valueMin="3.7">3.7 to 21.1 mm</quantity>
|
||
in disc diameter size. The smallest specimen (DD =
|
||
<quantity id="4C8E3FF80A04947BFEAD6B1AFE19B535" box="[359,449,350,375]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="10" pageNumber="375" unit="mm" value="3.7">3.7 mm</quantity>
|
||
,
|
||
<figureCitation id="134D8E980A04947BFE056B1AFDF7B535" box="[463,559,350,375]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
) presented a disc granule density of 329 per mm2 (
|
||
<figureCitation id="134D8E980A04947BFF626BC5FEDDB5D8" box="[168,261,385,410]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(a)). Its jaws were covered with granules, including oral shields (
|
||
<figureCitation id="134D8E980A04947BFC2B6BC5FBE7B5D8" box="[993,1087,385,410]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(b–c)). Only one oral shield, presumably the madreporite, was partially uncovered; it presented a dark grey colour and a minute depression on the centre (
|
||
<figureCitation id="134D8E980A04947BFCFD6B82FC4CB59D" box="[823,916,454,479]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(b)). This is consistent with what
|
||
<bibRefCitation id="EFE7EFEC0A04947BFEDD6BADFE2DB640" author="Ziesenhenne FC" box="[279,501,489,514]" pageId="10" pageNumber="375" pagination="185 - 201" refId="ref11224" refString="Ziesenhenne FC. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation fo Scientific Research, editor. Essays in the Natural Sciences in honor of Captain Allan Hancock. Los Angeles: University of Southern California Press; p. 185 - 201. doi: 10.5962 / bhl. title. 6088." type="book chapter" year="1955">Ziesenhenne (1955)</bibRefCitation>
|
||
observed in
|
||
<taxonomicName id="4C76E99E0A04947BFD4C6BADFCFAB640" authorityName="Lutken" authorityYear="1859" box="[646,802,489,514]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="10" pageNumber="375" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A04947BFD4C6BADFCFAB640" box="[646,802,489,514]" italics="true" pageId="10" pageNumber="375">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
(DD =
|
||
<quantity id="4C8E3FF80A04947BFCB86BADFC6DB640" box="[882,949,489,514]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="375" unit="mm" value="4.0">4 mm</quantity>
|
||
), in which the first oral shield to be uncovered was the madreporite. Distal genital slits only presented granules (
|
||
<figureCitation id="134D8E980A04947BFEC7686AFEB1B605" box="[269,361,558,583]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(b)), in comparison with those of larger specimens which also presented scales (
|
||
<figureCitation id="134D8E980A04947BFF396815FE95B628" box="[243,333,593,618]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="10" pageNumber="375">Figure 1</figureCitation>
|
||
(b)). Similarly, the specimen presented only granules on the dorsal arm base, extending over two arm segments (
|
||
<figureCitation id="134D8E980A04947BFD5A6830FD2AB6CF" box="[656,754,628,653]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(d)). All the DAPs were entire and surrounded by granules (
|
||
<figureCitation id="134D8E980A04947BFE7468D2FDC3B6ED" box="[446,539,662,687]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(d–e)); distal ones were completely covered with granules (
|
||
<figureCitation id="134D8E980A04947BFF1668FDFEE1B690" box="[220,313,697,722]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(f)). Some proximal VAPs were partially covered with granules (
|
||
<figureCitation id="134D8E980A04947BFC3668FDFB81B690" box="[1020,1113,697,722]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(b)), the rest were naked (
|
||
<figureCitation id="134D8E980A04947BFE476898FE33B6B7" box="[397,491,732,757]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(g–i)). All the LAPs were partially covered with granules and presented up to five arm spines (
|
||
<figureCitation id="134D8E980A04947BFDFB68BAFD4CB755" box="[561,660,766,791]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(j)). The presence of granules on the disc, including jaws, and arms (DAPs, VAPs, LAPs) is common among
|
||
<taxonomicName id="4C76E99E0A04947BFC446965FBCEB778" authorityName="Muller and Troschel" authorityYear="1840" box="[910,1046,801,826]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="10" pageNumber="375" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9024E0F0A04947BFC446965FBCEB778" box="[910,1046,801,826]" italics="true" pageId="10" pageNumber="375">Ophioderma</emphasis>
|
||
</taxonomicName>
|
||
juveniles (e.g.
|
||
<taxonomicName id="4C76E99E0A04947BFF106900FD46B71F" authority="Lutken, 1856" authorityName="Lutken" authorityYear="1856" box="[218,670,836,861]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="10" pageNumber="375" phylum="Echinodermata" rank="species" species="brevicaudum">
|
||
<emphasis id="B9024E0F0A04947BFF106900FE24B71F" box="[218,508,836,861]" italics="true" pageId="10" pageNumber="375">Ophioderma brevicaudum</emphasis>
|
||
Lütken, 1856
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation id="EFE7EFEC0A04947BFD666900FCFEB71F" author="Clark AH" box="[684,806,836,861]" pageId="10" pageNumber="375" pagination="441 - 456" refId="ref9560" refString="Clark AH. 1939. Echinoderms of the Smithsonian-Hartford expedition, 1937, with other West Indian records. Proc US Natl Mus. 86 (3056): 441 - 456. doi: 10.5479 / si. 00963801.86 - 3056.441." type="journal article" year="1939">Clark 1939</bibRefCitation>
|
||
;
|
||
<taxonomicName id="4C76E99E0A04947BFCFD6900FEC5B73D" authority="Granja-Fernandez et al. 2020" authorityName="Granja-Fernandez" authorityYear="2020" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="10" pageNumber="375" phylum="Echinodermata" rank="species" species="hendleri">
|
||
<emphasis id="B9024E0F0A04947BFCFD6900FC69B71F" box="[823,945,836,861]" italics="true" pageId="10" pageNumber="375">O. hendleri</emphasis>
|
||
<bibRefCitation id="EFE7EFEC0A04947BFC776900FEC5B73D" author="Granja-Fernandez R & Pineda-Enriquez T & Solis-Marin FA & Laguarda-Figueras A" pageId="10" pageNumber="375" pagination="11 - 41" refId="ref9695" refString="Granja-Fernandez R, Pineda-Enriquez T, Solis-Marin FA, Laguarda-Figueras A. 2020. Ophioderma hendleri sp. nov. (Echinodermata: Ophiuroidea: Ophiodermatidae) and its congeners from the Eastern Pacific. Eur J Taxon. 729: 11 - 41. doi: 10.5852 / ejt. 2020.729.1187" type="journal article" year="2020">
|
||
Granja-Fernández
|
||
<emphasis id="B9024E0F0A04947BFF6A6922FF0CB73D" box="[160,212,870,895]" italics="true" pageId="10" pageNumber="375">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C76E99E0A04947BFEE46922FD69B73D" authority="Ziesenhenne 1955" authorityName="Ziesenhenne" authorityYear="1955" box="[302,689,870,895]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="10" pageNumber="375" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A04947BFEE46922FE09B73D" box="[302,465,870,895]" italics="true" pageId="10" pageNumber="375">O. panamense</emphasis>
|
||
<bibRefCitation id="EFE7EFEC0A04947BFE166922FD69B73D" author="Ziesenhenne FC" box="[476,689,870,895]" pageId="10" pageNumber="375" pagination="185 - 201" refId="ref11224" refString="Ziesenhenne FC. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation fo Scientific Research, editor. Essays in the Natural Sciences in honor of Captain Allan Hancock. Los Angeles: University of Southern California Press; p. 185 - 201. doi: 10.5962 / bhl. title. 6088." type="book chapter" year="1955">Ziesenhenne 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C76E99E0A04947BFD086922FB59B73D" authority="Lutken, 1856" authorityName="Lutken" authorityYear="1856" box="[706,1153,870,895]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="10" pageNumber="375" phylum="Echinodermata" rank="species" species="rubicundum">
|
||
<emphasis id="B9024E0F0A04947BFD086922FC04B73D" box="[706,988,870,895]" italics="true" pageId="10" pageNumber="375">Ophioderma rubicundum</emphasis>
|
||
Lütken, 1856
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation id="EFE7EFEC0A04947BFF6A69CDFEEFB7E0" author="Hendler G" box="[160,311,905,930]" pageId="10" pageNumber="375" pagination="189 - 311" refId="ref9803" refString="Hendler G. 2018. Armed to the teeth: a new paradigm for the buccal skeleton of brittle stars (Echinodermata: Ophiuroidea). Contrib Sci. 526: 189 - 311. doi: 10.5962 / p. 324539." type="journal article" year="2018">Hendler 2018</bibRefCitation>
|
||
). One arm segment presented three tentacle scales.
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A04947BFF0A69E8FC89B250" blockId="10.[160,1156,280,1727]" pageId="10" pageNumber="375">
|
||
As individuals became larger (DD =
|
||
<quantity id="4C8E3FF80A04947BFD9C69E8FD75B787" box="[598,685,940,965]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="10" pageNumber="375" unit="mm" value="5.7">5.7 mm</quantity>
|
||
), some scales on the dorsal arm base and on the distal genital slits were visible; oral shields started to uncover but still presented some granules, and the maximum number of arm spines increased to six. A specimen of DD =
|
||
<quantity id="4C8E3FF80A04947BFF2F6E50FE91B06F" box="[229,329,1044,1069]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="10" pageNumber="375" unit="mm" value="11.2">11.2 mm</quantity>
|
||
showed oral shields totally naked, but less elongated than those of the
|
||
<typeStatus id="54CD2CBF0A04947BFF6A6E72FEDFB00D" box="[160,263,1078,1103]" pageId="10" pageNumber="375" type="holotype">holotype</typeStatus>
|
||
, and its DAPs were mostly entire as in smaller specimens. This specimen, as well as the
|
||
<typeStatus id="54CD2CBF0A04947BFF216E1DFE97B030" box="[235,335,1113,1138]" pageId="10" pageNumber="375" type="holotype">holotype</typeStatus>
|
||
(DD =
|
||
<quantity id="4C8E3FF80A04947BFE6B6E1DFDDEB030" box="[417,518,1113,1138]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="10" pageNumber="375" unit="mm" value="15.9">15.9 mm</quantity>
|
||
), presented 1–6 abnormal arm spines in some segments along the arms; these were twice as wide as typical and variable in shape (e.g. bifurcated, flattened), mainly present in the proximal and median section of the arm. A specimen of DD =
|
||
<quantity id="4C8E3FF80A04947BFF2C6E85FE92B098" box="[230,330,1217,1242]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="10" pageNumber="375" unit="mm" value="13.3">13.3 mm</quantity>
|
||
presented most DAPs divided into two pieces; some median and distal DAPs were entire. Specimens of similar sizes to the
|
||
<typeStatus id="54CD2CBF0A04947BFCD86EA0FCAFB0BF" box="[786,887,1252,1277]" pageId="10" pageNumber="375" type="holotype">holotype</typeStatus>
|
||
or larger (DD =
|
||
<quantity id="4C8E3FF80A04947BFB8D6EA0FED9B15D" metricMagnitude="-2" metricUnit="m" metricValue="1.81" metricValueMax="2.11" metricValueMin="1.51" pageId="10" pageNumber="375" unit="mm" value="18.1" valueMax="21.1" valueMin="15.1">15.1– 21.1 mm</quantity>
|
||
) presented a maximum of nine arm spines, and sporadically three tentacle scales in 1–3 arm segments (
|
||
<figureCitation id="134D8E980A04947BFE5E6F6DFE36B100" box="[404,494,1321,1346]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="10" pageNumber="375">Figure 3</figureCitation>
|
||
(e)). One specimen (DD =
|
||
<quantity id="4C8E3FF80A04947BFCC26F6DFCB1B100" box="[776,873,1321,1346]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="10" pageNumber="375" unit="mm" value="17.5">17.5 mm</quantity>
|
||
) presented some median DAPs entire, which were 2.5 times as wide as long. Regarding colouration, the extent of white granules on the dorsal disc varied in every specimen; they could be: (1) a few scattered on the disc as in the
|
||
<typeStatus id="54CD2CBF0A04947BFE336FD5FD85B1E8" box="[505,605,1425,1450]" pageId="10" pageNumber="375" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="134D8E980A04947BFDA66FD5FD1BB1E8" box="[620,707,1425,1450]" captionStart="Figure 1" captionStartId="5.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@5.[163,1153,142,1597]" captionTargetPageId="5" captionText="Figure 1. Ophioderma occultum sp. nov., holotype (ICML-UNAM 18429, DD = 15.9 mm). (a) Dorsal disc; (b) ventral disc; (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost);(g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2° AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758478" httpUri="https://zenodo.org/record/6758478/files/figure.png" pageId="10" pageNumber="375">Figure 1</figureCitation>
|
||
(a)), (2) arranged in larger groups at the margin and centre of the disc (
|
||
<figureCitation id="134D8E980A04947BFE386FF0FD95B18F" box="[498,589,1460,1485]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(a)), or (3) not present at all; the smallest specimens presented a greater extension of white granules on the dorsal disc (
|
||
<figureCitation id="134D8E980A04947BFC796F92FBCAB1AD" box="[947,1042,1494,1519]" captionStart="Figure 2" captionStartId="6.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@6.[163,1153,142,1597]" captionTargetPageId="6" captionText="Figure 2. Ophioderma occultum sp. nov., paratype (ICML-UNAM 18430, DD = 3.7 mm). (a) Dorsal disc; (b) ventral disc (insert: jaw with the partially uncovered oral shield); (c) close-up of a jaw; (d–f) dorsal arm (proximalmost, median, distalmost); (g–i) ventral arm (proximalmost, median, distalmost); (j) lateral arm plates and arm spines; (k) dorsal view. 2°AdShSp, secondary adoral shield spine; AdShSp, adoral shield spine; IPa, infradental papilla; LOPas, lateral oral papillae; LyOs, Lyman’s ossicle; TPa, tooth papilla; vT, ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.6758480" httpUri="https://zenodo.org/record/6758480/files/figure.png" pageId="10" pageNumber="375">Figure 2</figureCitation>
|
||
(a)). Some specimens presented white-spotted oral shields (
|
||
<figureCitation id="134D8E980A04947BFD036FBDFCFFB250" box="[713,807,1529,1554]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="10" pageNumber="375">Figure 3</figureCitation>
|
||
(b)).
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A04947AFF0A6C58FED7B517" blockId="10.[160,1156,280,1727]" lastBlockId="11.[160,1155,142,479]" lastPageId="11" lastPageNumber="376" pageId="10" pageNumber="375">
|
||
Additionally, some variations were noticed in non-type specimens (DD sizes up to
|
||
<quantity id="4C8E3FF80A04947BFF6A6C7AFEDEB215" box="[160,262,1598,1623]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="10" pageNumber="375" unit="mm" value="26.7">26.7 mm</quantity>
|
||
): (1)
|
||
<specimenCount id="9D7059940A04947BFEF76C7AFE0EB215" box="[317,470,1598,1623]" pageId="10" pageNumber="375" type="generic">16 specimens</specimenCount>
|
||
presented 1–3 partially naked radial shields (
|
||
<figureCitation id="134D8E980A04947BFC136C7AFBEEB215" box="[985,1078,1598,1623]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="10" pageNumber="375">Figure 3</figureCitation>
|
||
(a)); (2)
|
||
<specimenCount id="9D7059940A04947BFF6A6C25FEE7B238" box="[160,319,1633,1658]" pageId="10" pageNumber="375" type="generic">one specimen</specimenCount>
|
||
presented white oral shields with a grey madreporite (
|
||
<figureCitation id="134D8E980A04947BFC7F6C25FBCAB238" box="[949,1042,1633,1658]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="10" pageNumber="375">Figure 3</figureCitation>
|
||
(c)); (3)
|
||
<specimenCount id="9D7059940A04947BFBA26C25FECEB2DF" pageId="10" pageNumber="375" type="generic">13 specimens</specimenCount>
|
||
(DD =
|
||
<quantity id="4C8E3FF80A04947BFEA36CC0FDD0B2DF" box="[361,520,1668,1693]" metricMagnitude="-2" metricUnit="m" metricValue="2.315" metricValueMax="2.55" metricValueMin="2.08" pageId="10" pageNumber="375" unit="mm" value="23.15" valueMax="25.5" valueMin="20.8">20.8–25.5 mm</quantity>
|
||
) presented up to 11 arm spines; (4) one large specimen (DD =
|
||
<quantity id="4C8E3FF80A04947BFF226CE2FE93B2FD" box="[232,331,1702,1727]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="10" pageNumber="375" unit="mm" value="26.7">26.7 mm</quantity>
|
||
) presented few granules over the six proximalmost DAPs (
|
||
<figureCitation id="134D8E980A04947BFC136CE2FBEEB2FD" box="[985,1078,1702,1727]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="10" pageNumber="375">Figure 3</figureCitation>
|
||
(d)); (5)
|
||
<specimenCount id="9D7059940A05947AFF6A6ACAFE83B4E5" box="[160,347,142,167]" pageId="11" pageNumber="376" type="generic">eight specimens</specimenCount>
|
||
(DD =
|
||
<quantity id="4C8E3FF80A05947AFE7D6ACAFD81B4E5" box="[439,601,142,167]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" metricValueMax="1.85" metricValueMin="1.29" pageId="11" pageNumber="376" unit="mm" value="15.7" valueMax="18.5" valueMin="12.9">12.9–18.5 mm</quantity>
|
||
) presented some abnormal arm spines such as those observed in the
|
||
<typeStatus id="54CD2CBF0A05947AFE576AF5FDD9B488" box="[413,513,177,202]" pageId="11" pageNumber="376" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D7059940A05947AFDF16AF5FD09B488" box="[571,721,177,202]" pageId="11" pageNumber="376" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
(
|
||
<figureCitation id="134D8E980A05947AFD156AF5FCE5B488" box="[735,829,177,202]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="11" pageNumber="376">Figure 3</figureCitation>
|
||
(f)); (6) some fixed specimens presented a brown general colouration instead of dark grey (
|
||
<figureCitation id="134D8E980A05947AFCB16A90FC06B4AF" box="[891,990,212,237]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="11" pageNumber="376">Figure 3</figureCitation>
|
||
(g)), while live specimens showed a dark olive green to brown colour (
|
||
<figureCitation id="134D8E980A05947AFCC56AB2FCB5B54D" box="[783,877,246,271]" captionStart="Figure 3" captionStartId="7.[160,226,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@7.[163,1153,142,1597]" captionTargetPageId="7" captionText="Figure 3. Morphological variations recorded in Ophioderma occultum sp. nov. (a) Disc with partially uncoveredradial shields (arrowheads;CE-UAM 390,DD = 15.4mm);(b) white-spottedoral shields (paratype: ICML-EMU 12868, DD = 19.9 mm); (c) white oral shields (CE-UAM 214, DD = 7.3 mm); (d) granules on the proximalmost DAPs (ICML-UNAM 11593, DD = 26.7 mm); (e) arm segment with three tentacle scales (arrowhead; paratype: ICML-UNAM 18430, DD = 17.2 mm); (f) abnormal arm spines (arrowheads; CE-UAM 679,DD = 16.3 mm); (g) brown-coloured specimen (CE-UAM 673, DD = 23.5 mm); (h–i) in vivo colouration (Photographs: M.A. Medina-López)." figureDoi="http://doi.org/10.5281/zenodo.6758482" httpUri="https://zenodo.org/record/6758482/files/figure.png" pageId="11" pageNumber="376">Figure 3</figureCitation>
|
||
(h–i)); and (7) the parallel white blotches on the arms of some specimens were present in up to 18 contiguous DAPs and LAPs.
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A05947AFF0A6B1AFD81B59D" blockId="11.[160,1155,142,479]" pageId="11" pageNumber="376">
|
||
Of the variations observed in
|
||
<taxonomicName id="4C76E99E0A05947AFDC36B1AFD53B535" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[521,651,350,375]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A05947AFDC36B1AFD53B535" box="[521,651,350,375]" italics="true" pageId="11" pageNumber="376">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A05947AFD5B6B1AFD32B535" box="[657,746,350,375]" pageId="11" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, features such as partially uncovered radial shields, granules on DAPs of large specimens, abnormal arm spines, and arm segments with three tentacle scales are considered rare, as they were observed only in <7% of the total specimens examined.
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A05947AFF6A684FFE18B667" blockId="11.[160,1156,523,791]" box="[160,448,523,549]" pageId="11" pageNumber="376">
|
||
<emphasis id="B9024E0F0A05947AFF6A684FFE18B667" bold="true" box="[160,448,523,549]" italics="true" pageId="11" pageNumber="376">Habitat and distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A05947AFF6A686AFF33B755" blockId="11.[160,1156,523,791]" pageId="11" pageNumber="376">
|
||
<taxonomicName id="4C76E99E0A05947AFF6A686AFE40B605" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[160,408,558,583]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A05947AFF6A686AFE40B605" box="[160,408,558,583]" italics="true" pageId="11" pageNumber="376">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A05947AFE68686AFE25B605" box="[418,509,558,583]" pageId="11" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is distributed in shallow waters, from intertidal to
|
||
<quantity id="4C8E3FF80A05947AFB9E686AFB5BB605" box="[1108,1155,558,583]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="376" unit="m" value="6.0">6 m</quantity>
|
||
depth, and is typically found in rocky reefs and tide pools, on sediment under mediumsized rocks. The species has only been found in the
|
||
<location id="8EA9C4C60A05947AFD2F6830FC24B6CF" LSID="urn:lsid:plazi:treatment:03DF230B0A0A947EFE016824FD7EB54D:8EA9C4C60A05947AFD2F6830FC24B6CF" box="[741,1020,628,653]" name="northern Mexican Pacific" pageId="11" pageNumber="376" stateProvince="California">northern Mexican Pacific</location>
|
||
, particularly in the Gulf of
|
||
<collectingRegion id="49B25CFF0A05947AFE8A68D2FE72B6ED" box="[320,426,662,687]" country="United States of America" name="California" pageId="11" pageNumber="376">California</collectingRegion>
|
||
(in the States of
|
||
<collectingRegion id="49B25CFF0A05947AFDA568D2FC36B6ED" box="[623,1006,662,687]" isEnumeration="true" pageId="11" pageNumber="376">Baja California, Baja California Sur</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49B25CFF0A05947AFBE768D2FBA6B6ED" box="[1069,1150,662,687]" country="Mexico" name="Sonora" pageId="11" pageNumber="376">Sonora</collectingRegion>
|
||
) and islas Revillagigedo (isla Roca Partida).
|
||
<collectorName id="2683F7CB0A05947AFDB268FDFD4AB690" box="[632,658,697,722]" pageId="11" pageNumber="376">Its</collectorName>
|
||
northernmost record corresponds to
|
||
<collectorName id="2683F7CB0A05947AFBF168FDFB58B690" box="[1083,1152,697,722]" pageId="11" pageNumber="376">Polilla</collectorName>
|
||
,
|
||
<collectorName id="2683F7CB0A05947AFF6A6898FF05B6B7" box="[160,221,732,757]" pageId="11" pageNumber="376">Bahía</collectorName>
|
||
de los
|
||
<collectorName id="2683F7CB0A05947AFEF06898FE4CB6B7" box="[314,404,732,757]" pageId="11" pageNumber="376">Ángeles</collectorName>
|
||
(
|
||
<geoCoordinate id="EE42F4DA0A05947AFE6F6898FE05B6B7" box="[421,477,732,757]" degrees="29" direction="north" orientation="latitude" pageId="11" pageNumber="376" precision="55555" value="29.0">29°N</geoCoordinate>
|
||
) and the southernmost to
|
||
<location id="8EA9C4C60A05947AFCD46898FC6EB6B7" LSID="urn:lsid:plazi:treatment:03DF230B0A0A947EFE016824FD7EB54D:8EA9C4C60A05947AFCD46898FC6EB6B7" box="[798,950,732,757]" latitude="19.0" name="bahia Eclipse" pageId="11" pageNumber="376" stateProvince="California">bahía Eclipse</location>
|
||
, isla
|
||
<collectorName id="2683F7CB0A05947AFC3E6898FB5BB6B7" box="[1012,1155,732,757]" pageId="11" pageNumber="376">Roca Partida</collectorName>
|
||
(
|
||
<geoCoordinate id="EE42F4DA0A05947AFF6D68BAFF38B755" box="[167,224,766,791]" degrees="19" direction="north" orientation="latitude" pageId="11" pageNumber="376" precision="55555" value="19.0">19°N</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36CC1960A05947AFF6A6907FB58B7A5" pageId="11" pageNumber="376" type="etymology">
|
||
<paragraph id="8BC9921D0A05947AFF6A6907FEC5B71F" blockId="11.[160,1155,835,999]" box="[160,285,835,861]" pageId="11" pageNumber="376">
|
||
<emphasis id="B9024E0F0A05947AFF6A6907FEC5B71F" bold="true" box="[160,285,835,861]" italics="true" pageId="11" pageNumber="376">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A05947AFF6A6922FB58B7A5" blockId="11.[160,1155,835,999]" pageId="11" pageNumber="376">
|
||
The species name
|
||
<taxonomicName id="4C76E99E0A05947AFEA56922FE0CB73D" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[367,468,870,895]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="occultum">
|
||
<emphasis id="B9024E0F0A05947AFEA56922FE0CB73D" box="[367,468,870,895]" italics="true" pageId="11" pageNumber="376">occultum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B9024E0F0A05947AFE296922FDFEB73D" box="[483,550,870,895]" italics="true" pageId="11" pageNumber="376">-us, -a</emphasis>
|
||
) is a Latin singular neuter adjective in the nominative case, meaning ‘hidden’.
|
||
<taxonomicName id="4C76E99E0A05947AFE7869CDFD7FB7E0" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[434,679,905,930]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A05947AFE7869CDFD7FB7E0" box="[434,679,905,930]" italics="true" pageId="11" pageNumber="376">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A05947AFD6569CDFCDFB7E0" box="[687,775,905,930]" pageId="11" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been collected and recorded since the nineteenth century in the north of
|
||
<collectingCountry id="F361D28D0A05947AFD5169E8FD36B787" box="[667,750,940,965]" name="Mexico" pageId="11" pageNumber="376">Mexico</collectingCountry>
|
||
; however, it was mistakenly considered a colour phase of
|
||
<taxonomicName id="4C76E99E0A05947AFE6F698AFD9AB7A5" authorityName="Ziesenhenne" authorityYear="1955" box="[421,578,974,999]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A05947AFE6F698AFD9AB7A5" box="[421,578,974,999]" italics="true" pageId="11" pageNumber="376">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
. It remained hidden in plain sight, hence its name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36CC1960A05947EFF6A6E57FD7EB54D" lastPageId="15" lastPageNumber="380" pageId="11" pageNumber="376" type="discussion">
|
||
<paragraph id="8BC9921D0A05947AFF6A6E57FEDEB06F" blockId="11.[160,1156,1043,1519]" box="[160,262,1043,1069]" pageId="11" pageNumber="376">
|
||
<emphasis id="B9024E0F0A05947AFF6A6E57FEDEB06F" bold="true" box="[160,262,1043,1069]" italics="true" pageId="11" pageNumber="376">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A05947AFF6A6E72FB95B1E8" blockId="11.[160,1156,1043,1519]" pageId="11" pageNumber="376">
|
||
<taxonomicName id="4C76E99E0A05947AFF6A6E72FE40B00D" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[160,408,1078,1103]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A05947AFF6A6E72FE40B00D" box="[160,408,1078,1103]" italics="true" pageId="11" pageNumber="376">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A05947AFE686E72FE25B00D" box="[418,509,1078,1103]" pageId="11" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
|
||
most resembles
|
||
<taxonomicName id="4C76E99E0A05947AFD0F6E72FCBEB00D" authorityName="Ziesenhenne" authorityYear="1955" box="[709,870,1078,1103]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A05947AFD0F6E72FCBEB00D" box="[709,870,1078,1103]" italics="true" pageId="11" pageNumber="376">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
, with which it has been confused for over a century (
|
||
<bibRefCitation id="EFE7EFEC0A05947AFE156E1DFD92B030" author="Ives JE" box="[479,586,1113,1138]" pageId="11" pageNumber="376" pagination="76 - 77" refId="ref9999" refString="Ives JE. 1889. Variation in Ophiura panamensis and Ophiura teres. Proc Acad Nat Sci Phila. 41: 76 - 77." type="journal article" year="1889">Ives 1889</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE7EFEC0A05947AFD9F6E1DFCFBB030" author="Ziesenhenne FC" box="[597,803,1113,1138]" pageId="11" pageNumber="376" pagination="185 - 201" refId="ref11224" refString="Ziesenhenne FC. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation fo Scientific Research, editor. Essays in the Natural Sciences in honor of Captain Allan Hancock. Los Angeles: University of Southern California Press; p. 185 - 201. doi: 10.5962 / bhl. title. 6088." type="book chapter" year="1955">Ziesenhenne 1955</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE7EFEC0A05947AFCE56E1DFBF6B030" author="Brusca RC & Smith CR" box="[815,1070,1113,1138]" pageId="11" pageNumber="376" pagination="304 - 343" refId="ref9370" refString="Brusca RC, Smith CR. 1973. Echinodermata (Starfish, etc.). In: Brusca RC, editor. A handbook to the common intertidal invertebrates of the Gulf of California. Tucson: University of Arizona Press; p. 304 - 343." type="book chapter" year="1973">Brusca and Smith 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE7EFEC0A05947AFBF06E1DFF0FB0D7" author="Brusca RC" pageId="11" pageNumber="376" pagination="398 - 422" refId="ref9419" refString="Brusca RC. 1980. 26. Echinodermata. In: Brusca RC, editor. Common intertidal invertebrates of the Gulf of California. Tucson: The University of Arizona Press; p. 398 - 422." type="book chapter" year="1980">Brusca 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE7EFEC0A05947AFF2E6E38FDC8B0D7" author="Kerstitch A & Bertsch H" box="[228,528,1148,1173]" pageId="11" pageNumber="376" refId="ref10024" refString="Kerstitch A, Bertsch H. 2007. Sea of Cortez. Marine invertebrates. A guide for the Pacific Coast, Mexico to Peru. 2 nd ed. California: Sea Challengers; p. 124." type="journal volume" year="2007">Kerstitch and Bertsch 2007</bibRefCitation>
|
||
). Both species present covered adoral shields, scales on the base of the arms, scales and granules ornamenting the distal genital slit, and banded arms. However,
|
||
<taxonomicName id="4C76E99E0A05947AFE9D6E85FE2DB098" authorityName="Ziesenhenne" authorityYear="1955" box="[343,501,1217,1242]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A05947AFE9D6E85FE2DB098" box="[343,501,1217,1242]" italics="true" pageId="11" pageNumber="376">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
has naked radial shields, DAPs usually entire, can display different colours on the dorsal disc (green, brown, grey mottled with yellow, orange, or pink), and its bands on the arms are conspicuous (
|
||
<figureCitation id="134D8E980A05947AFD136F42FCEEB15D" box="[729,822,1286,1311]" captionStart="Figure 4" captionStartId="12.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@12.[163,1153,142,1597]" captionTargetPageId="12" captionText="Figure 4. Some morphological characters of Eastern Pacific Ophioderma that differ from those of Ophioderma occultum sp. nov. Ophioderma hendleri (holotype: ICML-UNAM 18315, DD = 14.8 mm): (a) close-up of a jaw; (b) dorsal arm. Ophioderma panamense (holotype: NHMD-107679, DD = 21.9 mm): (c) portion of dorsal disc and arm. Ophioderma pentacanthum (paratype: MCZ OPH-4519," figureDoi="http://doi.org/10.5281/zenodo.6758484" httpUri="https://zenodo.org/record/6758484/files/figure.png" pageId="11" pageNumber="376">Figure 4</figureCitation>
|
||
(c);
|
||
<tableCitation id="C6F4A7A60A05947AFC946F42FC68B15D" box="[862,944,1286,1311]" captionStart="Table 1" captionStartId="8.[126,182,439,462]" captionTargetPageId="8" captionText="Table 1. Main characters that distinguish Eastern Pacific Ophioderma species." httpUri="http://table.plazi.org/id/DF09C2950A069479FFB46BF3FCA2B58C" pageId="11" pageNumber="376" tableUuid="DF09C2950A069479FFB46BF3FCA2B58C">Table 1</tableCitation>
|
||
;
|
||
<bibRefCitation id="EFE7EFEC0A05947AFC776F42FECDB100" author="Granja-Fernandez R & Herrero-Perezrul MD & Lopez-Perez RA & Hernandez L & Rodriguez-Zaragoza FA & Jones RW & Pineda-Lopez R" pageId="11" pageNumber="376" pagination="101 - 145" refId="ref9645" refString="Granja-Fernandez R, Herrero-Perezrul MD, Lopez-Perez RA, Hernandez L, Rodriguez-Zaragoza FA, Jones RW, Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys. 406: 101 - 145. doi: 10.3897 / zookeys. 4066306" type="journal article" year="2014">Granja-Fernández et al. 2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE7EFEC0A05947AFEE86F6DFE81B100" author="Granja-Fernandez R & Pineda-Enriquez T & Solis-Marin FA & Laguarda-Figueras A" box="[290,345,1321,1346]" pageId="11" pageNumber="376" pagination="11 - 41" refId="ref9695" refString="Granja-Fernandez R, Pineda-Enriquez T, Solis-Marin FA, Laguarda-Figueras A. 2020. Ophioderma hendleri sp. nov. (Echinodermata: Ophiuroidea: Ophiodermatidae) and its congeners from the Eastern Pacific. Eur J Taxon. 729: 11 - 41. doi: 10.5852 / ejt. 2020.729.1187" type="journal article" year="2020">2020</bibRefCitation>
|
||
), while
|
||
<taxonomicName id="4C76E99E0A05947AFE7B6F6DFDEDB100" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[433,565,1321,1346]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A05947AFE7B6F6DFDEDB100" box="[433,565,1321,1346]" italics="true" pageId="11" pageNumber="376">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A05947AFDF66F6DFD4BB100" box="[572,659,1321,1346]" pageId="11" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
|
||
presents radial shields usually covered with granules, DAPs divided into two (typically) or more pieces, subtle lighter bands on the arms, and characteristic parallel white blotches on some DAPs and LAPs in the median and distal arm sections, giving the appearance of parallel longitudinal white bands.
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A05947AFF0A6FF0FC27B1AD" blockId="11.[160,1156,1043,1519]" pageId="11" pageNumber="376">
|
||
Some authors noticed the aforementioned characters but considered them intraspecific variations of
|
||
<taxonomicName id="4C76E99E0A05947AFEAF6F92FDDAB1AD" authorityName="Ziesenhenne" authorityYear="1955" box="[357,514,1494,1519]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A05947AFEAF6F92FDDAB1AD" box="[357,514,1494,1519]" italics="true" pageId="11" pageNumber="376">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFE7EFEC0A05947AFDC56F92FD59B1AD" author="Ives JE" box="[527,641,1494,1519]" pageId="11" pageNumber="376" pagination="76 - 77" refId="ref9999" refString="Ives JE. 1889. Variation in Ophiura panamensis and Ophiura teres. Proc Acad Nat Sci Phila. 41: 76 - 77." type="journal article" year="1889">Ives (1889</bibRefCitation>
|
||
, 76–77) mentioned the following:
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A05947AFF0A6C48FC35B2F7" blockId="11.[192,1124,1548,1717]" pageId="11" pageNumber="376">
|
||
“Mr. Lockington records a
|
||
<taxonomicName id="4C76E99E0A05947AFE196C48FD95B260" box="[467,589,1548,1570]" pageId="11" pageNumber="376" rank="variety" variety="with">variety with</taxonomicName>
|
||
a brown disk and olive green, arms without bands except at the tips” and “The darkest variety of
|
||
<taxonomicName id="4C76E99E0A05947AFD476C6DFD0DB27D" authorityName="Lyman" authorityYear="1860" box="[653,725,1577,1599]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A05947AFD476C6DFD0DB27D" box="[653,725,1577,1599]" italics="true" pageId="11" pageNumber="376">O. teres</emphasis>
|
||
</taxonomicName>
|
||
closely resembles the darkest variety of
|
||
<taxonomicName id="4C76E99E0A05947AFF0A6C03FE89B21F" baseAuthorityName="Lütken" baseAuthorityYear="1859" box="[192,337,1607,1629]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="11" pageNumber="376" phylum="Echinodermata" rank="species" species="panamensis">
|
||
<emphasis id="B9024E0F0A05947AFF0A6C03FE89B21F" box="[192,337,1607,1629]" italics="true" pageId="11" pageNumber="376">O. Panamensis</emphasis>
|
||
</taxonomicName>
|
||
[sic]. In both forms some of the upper arm-plates of the outer portion of the arm have white markings, which form segments of two broken parallel longitudinal lines, one on each side of the arm. Each segment occupies one or more plates and is separated from the next segment by an also variable number of plates without any such markings”.
|
||
</paragraph>
|
||
<caption id="DF09C2950A02947DFF6A6C19FB5DB28E" ID-DOI="http://doi.org/10.5281/zenodo.6758484" ID-Zenodo-Dep="6758484" httpUri="https://zenodo.org/record/6758484/files/figure.png" pageId="12" pageNumber="377" startId="12.[160,225,1629,1652]" targetBox="[164,1152,143,1596]" targetPageId="12">
|
||
<paragraph id="8BC9921D0A02947DFF6A6C19FB5DB28E" blockId="12.[160,1157,1629,1740]" pageId="12" pageNumber="377">
|
||
<emphasis id="B9024E0F0A02947DFF6A6C19FF25B236" bold="true" box="[160,253,1629,1652]" pageId="12" pageNumber="377">Figure 4.</emphasis>
|
||
Some morphological characters of Eastern Pacific
|
||
<taxonomicName id="4C76E99E0A02947DFCCD6C19FCA6B236" authorityName="Muller and Troschel" authorityYear="1840" box="[775,894,1629,1652]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="12" pageNumber="377" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9024E0F0A02947DFCCD6C19FCA6B236" box="[775,894,1629,1652]" italics="true" pageId="12" pageNumber="377">Ophioderma</emphasis>
|
||
</taxonomicName>
|
||
that differ from those of
|
||
<taxonomicName id="4C76E99E0A02947DFF6A6C3EFEADB2D3" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[160,373,1658,1681]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="12" pageNumber="377" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A02947DFF6A6C3EFEADB2D3" box="[160,373,1658,1681]" italics="true" pageId="12" pageNumber="377">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A02947DFEB06C3EFE1FB2D3" box="[378,455,1658,1681]" pageId="12" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<taxonomicName id="4C76E99E0A02947DFE016C3EFD4DB2D3" box="[459,661,1658,1681]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="12" pageNumber="377" phylum="Echinodermata" rank="species" species="hendleri">
|
||
<emphasis id="B9024E0F0A02947DFE016C3EFD4DB2D3" box="[459,661,1658,1681]" italics="true" pageId="12" pageNumber="377">Ophioderma hendleri</emphasis>
|
||
</taxonomicName>
|
||
(holotype: ICML-UNAM 18315, DD = 14.8 mm): (a) close-up of a jaw; (b) dorsal arm.
|
||
<taxonomicName id="4C76E99E0A02947DFE266CDCFD0FB2ED" authorityName="Ziesenhenne" authorityYear="1955" box="[492,727,1688,1711]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="12" pageNumber="377" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A02947DFE266CDCFD0FB2ED" box="[492,727,1688,1711]" italics="true" pageId="12" pageNumber="377">Ophioderma panamense</emphasis>
|
||
</taxonomicName>
|
||
(holotype: NHMD-107679, DD = 21.9 mm): (c) portion of dorsal disc and arm.
|
||
<taxonomicName id="4C76E99E0A02947DFDF66CF1FC8EB28E" authorityName="H. L. Clark" authorityYear="1917" box="[572,854,1717,1740]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="12" pageNumber="377" phylum="Echinodermata" rank="species" species="pentacanthum">
|
||
<emphasis id="B9024E0F0A02947DFDF66CF1FC8EB28E" box="[572,854,1717,1740]" italics="true" pageId="12" pageNumber="377">Ophioderma pentacanthum</emphasis>
|
||
</taxonomicName>
|
||
(paratype: MCZ OPH-4519,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9921D0A03947CFF6A6ACAFB5BB488" blockId="13.[160,1157,142,202]" pageId="13" pageNumber="378">
|
||
This coincides with the colour pattern observed in
|
||
<taxonomicName id="4C76E99E0A03947CFCD26ACAFC7CB4E5" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[792,932,142,167]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFCD26ACAFC7CB4E5" box="[792,932,142,167]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFC7B6ACAFBC8B4E5" box="[945,1040,142,167]" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Likewise,
|
||
<bibRefCitation id="EFE7EFEC0A03947CFF6A6AF5FEAEB488" author="Ziesenhenne FC" box="[160,374,177,202]" pageId="13" pageNumber="378" pagination="185 - 201" refId="ref11224" refString="Ziesenhenne FC. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation fo Scientific Research, editor. Essays in the Natural Sciences in honor of Captain Allan Hancock. Los Angeles: University of Southern California Press; p. 185 - 201. doi: 10.5962 / bhl. title. 6088." type="book chapter" year="1955">Ziesenhenne (1955</bibRefCitation>
|
||
, 194) mentioned that
|
||
<taxonomicName id="4C76E99E0A03947CFDBB6AF5FCD6B488" authorityName="Ziesenhenne" authorityYear="1955" box="[625,782,177,202]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A03947CFDBB6AF5FCD6B488" box="[625,782,177,202]" italics="true" pageId="13" pageNumber="378">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
showed one colour combination
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF0A6AA3FC59B575" blockId="13.[192,1124,231,311]" pageId="13" pageNumber="378">In which the disk is brown to olive and the arms greenish, with the arms banded distally with white... Some specimens have broken upper arm plates.. .. The majority of the specimens of this color phase have concealed, or partially concealed, radial shields.</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF6A6B10FBEFB52F" blockId="13.[160,1156,340,1058]" box="[160,1079,340,365]" pageId="13" pageNumber="378">
|
||
All of the above are now considered diagnostic characters of
|
||
<taxonomicName id="4C76E99E0A03947CFC9F6B10FC00B52F" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[853,984,340,365]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFC9F6B10FC00B52F" box="[853,984,340,365]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFC156B10FBEFB52F" box="[991,1079,340,365]" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF0A6B32FD73B67F" blockId="13.[160,1156,340,1058]" pageId="13" pageNumber="378">
|
||
Moreover,
|
||
<taxonomicName id="4C76E99E0A03947CFEF76B32FE06B5CD" authorityName="Ziesenhenne" authorityYear="1955" box="[317,478,374,399]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A03947CFEF76B32FE06B5CD" box="[317,478,374,399]" italics="true" pageId="13" pageNumber="378">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C76E99E0A03947CFDEA6B32FD70B5CD" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[544,680,374,399]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFDEA6B32FD70B5CD" box="[544,680,374,399]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFD796B32FCD7B5CD" box="[691,783,374,399]" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differ in their distributions; the former is widely distributed from southern
|
||
<collectingCountry id="F361D28D0A03947CFD586BDDFD19B5F0" box="[658,705,409,434]" name="United States of America" pageId="13" pageNumber="378">USA</collectingCountry>
|
||
to
|
||
<collectingCountry id="F361D28D0A03947CFD206BDDFCC5B5F0" box="[746,797,409,434]" name="Peru" pageId="13" pageNumber="378">Peru</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE7EFEC0A03947CFCE66BDDFBAEB5F0" author="Granja-Fernandez R & Pineda-Enriquez T & Solis-Marin FA & Laguarda-Figueras A" box="[812,1142,409,434]" pageId="13" pageNumber="378" pagination="11 - 41" refId="ref9695" refString="Granja-Fernandez R, Pineda-Enriquez T, Solis-Marin FA, Laguarda-Figueras A. 2020. Ophioderma hendleri sp. nov. (Echinodermata: Ophiuroidea: Ophiodermatidae) and its congeners from the Eastern Pacific. Eur J Taxon. 729: 11 - 41. doi: 10.5852 / ejt. 2020.729.1187" type="journal article" year="2020">Granja-Fernández et al. 2020</bibRefCitation>
|
||
), while
|
||
<taxonomicName id="4C76E99E0A03947CFF2D6BF8FEB5B597" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[231,365,444,469]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFF2D6BF8FEB5B597" box="[231,365,444,469]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFEBD6BF8FE09B597" box="[375,465,444,469]" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is restricted to the northern Mexican Pacific. In the Gulf of California both species appear to be the most common and abundant
|
||
<taxonomicName id="4C76E99E0A03947CFC7A6B9AFBE0B5B5" authorityName="Muller and Troschel" authorityYear="1840" box="[944,1080,478,503]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9024E0F0A03947CFC7A6B9AFBE0B5B5" box="[944,1080,478,503]" italics="true" pageId="13" pageNumber="378">Ophioderma</emphasis>
|
||
</taxonomicName>
|
||
, along with
|
||
<taxonomicName id="4C76E99E0A03947CFF1D6845FE93B658" box="[215,331,513,538]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="hendleri">
|
||
<emphasis id="B9024E0F0A03947CFF1D6845FE93B658" box="[215,331,513,538]" italics="true" pageId="13" pageNumber="378">O. hendleri</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C76E99E0A03947CFE0F6845FD9FB658" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[453,583,513,538]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFE0F6845FD9FB658" box="[453,583,513,538]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFD866845FD79B658" box="[588,673,513,538]" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is most often found in shallower waters (i.e. intertidal and tide pools; RGF, KJHG pers. obs).
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF0A6802FC74B79F" blockId="13.[160,1156,340,1058]" pageId="13" pageNumber="378">
|
||
The new species was also misidentified as
|
||
<taxonomicName id="4C76E99E0A03947CFD726802FCD6B61D" authorityName="Lyman" authorityYear="1860" box="[696,782,582,607]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFD726802FCD6B61D" box="[696,782,582,607]" italics="true" pageId="13" pageNumber="378">O. teres</emphasis>
|
||
</taxonomicName>
|
||
in the scientific collections and literature (
|
||
<bibRefCitation id="EFE7EFEC0A03947CFED9682DFEA6B6C0" author="Ives JE" box="[275,382,617,642]" pageId="13" pageNumber="378" pagination="76 - 77" refId="ref9999" refString="Ives JE. 1889. Variation in Ophiura panamensis and Ophiura teres. Proc Acad Nat Sci Phila. 41: 76 - 77." type="journal article" year="1889">Ives 1889</bibRefCitation>
|
||
). Since the
|
||
<typeStatus id="54CD2CBF0A03947CFDCB682DFDBDB6C0" box="[513,613,617,642]" pageId="13" pageNumber="378" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C76E99E0A03947CFD40682DFD04B6C0" authorityName="Lyman" authorityYear="1860" box="[650,732,617,642]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFD40682DFD04B6C0" box="[650,732,617,642]" italics="true" pageId="13" pageNumber="378">O. teres</emphasis>
|
||
</taxonomicName>
|
||
is lost, we based our comparisons on the original description of the species, as well as on a specimen identified by Lyman as
|
||
<taxonomicName id="4C76E99E0A03947CFF6A68EAFF2DB685" authorityName="Lyman" authorityYear="1860" box="[160,245,686,711]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFF6A68EAFF2DB685" box="[160,245,686,711]" italics="true" pageId="13" pageNumber="378">O. teres</emphasis>
|
||
</taxonomicName>
|
||
(MCZ OPH-112). Although this specimen is not a topotype (i.e. it is not from
|
||
<collectingCountry id="F361D28D0A03947CFF6A6895FF25B6A8" box="[160,253,721,746]" name="Panama" pageId="13" pageNumber="378">Panama</collectingCountry>
|
||
, the type locality of
|
||
<taxonomicName id="4C76E99E0A03947CFE3C6895FD94B6A8" authorityName="Lyman" authorityYear="1860" box="[502,588,721,746]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFE3C6895FD94B6A8" box="[502,588,721,746]" italics="true" pageId="13" pageNumber="378">O. teres</emphasis>
|
||
</taxonomicName>
|
||
, but from Acapulco,
|
||
<collectingCountry id="F361D28D0A03947CFC836895FC45B6A8" box="[841,925,721,746]" name="Mexico" pageId="13" pageNumber="378">Mexico</collectingCountry>
|
||
), it agrees with the description of
|
||
<taxonomicName id="4C76E99E0A03947CFEF568B0FE57B74F" authorityName="Lyman" authorityYear="1860" box="[319,399,756,781]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFEF568B0FE57B74F" box="[319,399,756,781]" italics="true" pageId="13" pageNumber="378">O. teres</emphasis>
|
||
</taxonomicName>
|
||
and was included as additional material in Lyman’s first descriptions of the species (
|
||
<bibRefCitation id="EFE7EFEC0A03947CFE826952FE0BB76D" author="Lyman T" box="[328,467,790,815]" pageId="13" pageNumber="378" pagination="252 - 262" refId="ref10267" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proc Boston Soc Nat Hist. 7: 193 - 204, 252 - 262. doi: 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman 1860</bibRefCitation>
|
||
, 257, 1865, 38).
|
||
<taxonomicName id="4C76E99E0A03947CFD426952FC92B76D" baseAuthorityName="Lyman" baseAuthorityYear="1860" box="[648,842,790,815]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFD426952FC92B76D" box="[648,842,790,815]" italics="true" pageId="13" pageNumber="378">Ophioderma teres</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C76E99E0A03947CFC106952FB85B76D" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[986,1117,790,815]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFC106952FB85B76D" box="[986,1117,790,815]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFBAE6952FF09B710" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in having DAPs divided into multiple pieces (up to five pieces), rounded arms, and a speckled colour pattern on disc and arms (
|
||
<figureCitation id="134D8E980A03947CFD5A6918FD36B737" box="[656,750,860,885]" captionStart="Figure 4" captionStartId="12.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@12.[163,1153,142,1597]" captionTargetPageId="12" captionText="Figure 4. Some morphological characters of Eastern Pacific Ophioderma that differ from those of Ophioderma occultum sp. nov. Ophioderma hendleri (holotype: ICML-UNAM 18315, DD = 14.8 mm): (a) close-up of a jaw; (b) dorsal arm. Ophioderma panamense (holotype: NHMD-107679, DD = 21.9 mm): (c) portion of dorsal disc and arm. Ophioderma pentacanthum (paratype: MCZ OPH-4519," figureDoi="http://doi.org/10.5281/zenodo.6758484" httpUri="https://zenodo.org/record/6758484/files/figure.png" pageId="13" pageNumber="378">Figure 4</figureCitation>
|
||
(j);
|
||
<tableCitation id="C6F4A7A60A03947CFCC76918FC87B737" box="[781,863,860,885]" captionStart="Table 1" captionStartId="8.[126,182,439,462]" captionTargetPageId="8" captionText="Table 1. Main characters that distinguish Eastern Pacific Ophioderma species." httpUri="http://table.plazi.org/id/DF09C2950A069479FFB46BF3FCA2B58C" pageId="13" pageNumber="378" tableUuid="DF09C2950A069479FFB46BF3FCA2B58C">Table 1</tableCitation>
|
||
). Confusions between the two species may arise from the fact that both present divided DAPs, as well as a misinterpretation of
|
||
<taxonomicName id="4C76E99E0A03947CFE6969E5FE2FB7F8" baseAuthorityName="Lyman" baseAuthorityYear="1860" box="[419,503,929,954]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFE6969E5FE2FB7F8" box="[419,503,929,954]" italics="true" pageId="13" pageNumber="378">O. teres</emphasis>
|
||
</taxonomicName>
|
||
speckles on the arms (
|
||
<figureCitation id="134D8E980A03947CFCCC69E5FCBEB7F8" box="[774,870,929,954]" captionStart="Figure 4" captionStartId="12.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@12.[163,1153,142,1597]" captionTargetPageId="12" captionText="Figure 4. Some morphological characters of Eastern Pacific Ophioderma that differ from those of Ophioderma occultum sp. nov. Ophioderma hendleri (holotype: ICML-UNAM 18315, DD = 14.8 mm): (a) close-up of a jaw; (b) dorsal arm. Ophioderma panamense (holotype: NHMD-107679, DD = 21.9 mm): (c) portion of dorsal disc and arm. Ophioderma pentacanthum (paratype: MCZ OPH-4519," figureDoi="http://doi.org/10.5281/zenodo.6758484" httpUri="https://zenodo.org/record/6758484/files/figure.png" pageId="13" pageNumber="378">Figure 4</figureCitation>
|
||
(j)) as the white blotches present in the median and distal arm sections of
|
||
<taxonomicName id="4C76E99E0A03947CFD006980FC96B79F" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[714,846,964,989]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFD006980FC96B79F" box="[714,846,964,989]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFC9F6980FC74B79F" box="[853,940,964,989]" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF0A69A2FCD6B060" blockId="13.[160,1156,340,1058]" pageId="13" pageNumber="378">
|
||
Differences among
|
||
<taxonomicName id="4C76E99E0A03947CFE5769A2FDF9B7BD" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[413,545,998,1023]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A03947CFE5769A2FDF9B7BD" box="[413,545,998,1023]" italics="true" pageId="13" pageNumber="378">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A03947CFDE269A2FD58B7BD" box="[552,640,998,1023]" pageId="13" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and all its congeners from the Eastern Pacific are summarised and illustrated in
|
||
<tableCitation id="C6F4A7A60A03947CFDEA6E4DFDABB060" box="[544,627,1033,1058]" captionStart="Table 1" captionStartId="8.[126,182,439,462]" captionTargetPageId="8" captionText="Table 1. Main characters that distinguish Eastern Pacific Ophioderma species." httpUri="http://table.plazi.org/id/DF09C2950A069479FFB46BF3FCA2B58C" pageId="13" pageNumber="378" tableUuid="DF09C2950A069479FFB46BF3FCA2B58C">Table 1</tableCitation>
|
||
and
|
||
<figureCitation id="134D8E980A03947CFD666E4DFCD0B060" box="[684,776,1033,1058]" captionStart="Figure 4" captionStartId="12.[160,225,1629,1652]" captionTargetBox="[164,1152,143,1596]" captionTargetId="figure-3@12.[163,1153,142,1597]" captionTargetPageId="12" captionText="Figure 4. Some morphological characters of Eastern Pacific Ophioderma that differ from those of Ophioderma occultum sp. nov. Ophioderma hendleri (holotype: ICML-UNAM 18315, DD = 14.8 mm): (a) close-up of a jaw; (b) dorsal arm. Ophioderma panamense (holotype: NHMD-107679, DD = 21.9 mm): (c) portion of dorsal disc and arm. Ophioderma pentacanthum (paratype: MCZ OPH-4519," figureDoi="http://doi.org/10.5281/zenodo.6758484" httpUri="https://zenodo.org/record/6758484/files/figure.png" pageId="13" pageNumber="378">Figure 4</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF6A6E09FDE1B025" blockId="13.[160,1156,1101,1370]" box="[160,569,1101,1127]" pageId="13" pageNumber="378">
|
||
<emphasis id="B9024E0F0A03947CFF6A6E09FDE1B025" bold="true" box="[160,569,1101,1127]" italics="true" pageId="13" pageNumber="378">Genetic delimitation of the species</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF6A6E35FD3CB118" blockId="13.[160,1156,1101,1370]" pageId="13" pageNumber="378">COI sequences obtained in this study had a total length of 680 bp; however, they were cut to align them with those retrieved from BOLD and GenBank, resulting in the analysis of 426 bp. Of these, 305 (71.60%) sites were conserved and 121 (28.40%) were variable, all parsimony informative. On the other hand, 16S sequences varied from 489 to 500 bp, and the alignment presented several indels. The latter were removed, and 454 bp were analysed. In this case, 328 (74.25%) sites were conserved and 126 (27.75%) were variable, of which 123 (27.09%) were parsimony informative.</paragraph>
|
||
<paragraph id="8BC9921D0A03947CFF6A6F86FBA8B2E4" blockId="13.[160,1157,1474,1702]" pageId="13" pageNumber="378">
|
||
DD =
|
||
<quantity id="4C8E3FF80A03947CFF2B6F86FEE5B19B" box="[225,317,1474,1497]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="13" pageNumber="378" unit="mm" value="21.1">21.1 mm</quantity>
|
||
): (d) close-up of a jaw; (e) dorsal arm.
|
||
<taxonomicName id="4C76E99E0A03947CFD1B6F86FC6BB19B" box="[721,947,1474,1497]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="peruanum">
|
||
<emphasis id="B9024E0F0A03947CFD1B6F86FC6BB19B" box="[721,947,1474,1497]" italics="true" pageId="13" pageNumber="378">Ophioderma peruanum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54CD2CBF0A03947CFC096F86FBC6B19B" box="[963,1054,1474,1497]" pageId="13" pageNumber="378" type="holotype">holotype</typeStatus>
|
||
: CZA-363, DD =
|
||
<quantity id="4C8E3FF80A03947CFF176FA4FEE0B1B5" box="[221,312,1504,1527]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="13" pageNumber="378" unit="mm" value="32.5">32.5 mm</quantity>
|
||
): (f) portion of dorsal disc and arm.
|
||
<materialsCitation id="3B1E98400A03947CFD6E6FA4FB59B256" ID-GBIF-Occurrence="3819603303" collectionCode="ICML-UNAM" pageId="13" pageNumber="378" specimenCode="ICML-UNAM 3.24" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="4C76E99E0A03947CFD6E6FA4FC40B1B5" authorityName="Caso" authorityYear="1986" box="[676,920,1504,1527]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="sodipallaresi">
|
||
<emphasis id="B9024E0F0A03947CFD6E6FA4FC40B1B5" box="[676,920,1504,1527]" italics="true" pageId="13" pageNumber="378">Ophioderma sodipallaresi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54CD2CBF0A03947CFC6C6FA4FBD9B1B5" box="[934,1025,1504,1527]" pageId="13" pageNumber="378" type="holotype">holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DBD03A660A03947CFBC66FA4FF16B256" collectionCode="ICML-UNAM" pageId="13" pageNumber="378">ICML-UNAM 3.24</specimenCode>
|
||
.3, DD =
|
||
<quantity id="4C8E3FF80A03947CFEE86FB9FEA5B256" box="[290,381,1533,1556]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="13" pageNumber="378" unit="mm" value="19.1">19.1 mm</quantity>
|
||
): (g) portion of dorsal disc and arm; (h) close-up of a jaw; (i) distal genital slit
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1E98400A03947CFF6A6C5EFF11B20C" ID-GBIF-Occurrence="3819603304" collectionCode="MCZ" pageId="13" pageNumber="378" specimenCode="OPH-112" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="4C76E99E0A03947CFF6A6C5EFE93B273" baseAuthorityName="Lyman" baseAuthorityYear="1860" box="[160,331,1562,1585]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A03947CFF6A6C5EFE93B273" box="[160,331,1562,1585]" italics="true" pageId="13" pageNumber="378">Ophioderma teres</emphasis>
|
||
</taxonomicName>
|
||
(non-type specimen:
|
||
<collectionCode id="ED670AD80A03947CFDEC6C5EFD8AB273" box="[550,594,1562,1585]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="378" type="Museum">MCZ</collectionCode>
|
||
<specimenCode id="DBD03A660A03947CFD906C5EFD6CB273" box="[602,692,1562,1585]" collectionCode="OPH-" pageId="13" pageNumber="378">OPH-112</specimenCode>
|
||
, DD =
|
||
<quantity id="4C8E3FF80A03947CFD316C5EFC8EB273" box="[763,854,1562,1585]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="13" pageNumber="378" unit="mm" value="24.8">24.8 mm</quantity>
|
||
): (j) portion of dorsal disc and arm
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1E98400A03947CFF186C73FF11B22E" ID-GBIF-Occurrence="3819603307" collectionCode="MCZ" pageId="13" pageNumber="378" specimenCode="OPH-114" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="4C76E99E0A03947CFF186C73FE17B20C" authorityName="H. L. Clark" authorityYear="1940" box="[210,463,1591,1614]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="subSpecies" species="teres" subSpecies="unicolor">
|
||
<emphasis id="B9024E0F0A03947CFF186C73FE17B20C" box="[210,463,1591,1614]" italics="true" pageId="13" pageNumber="378">Ophioderma teres unicolor</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54CD2CBF0A03947CFE116C73FDEEB20C" box="[475,566,1591,1614]" pageId="13" pageNumber="378" type="holotype">holotype</typeStatus>
|
||
:
|
||
<collectionCode id="ED670AD80A03947CFD8A6C73FDB4B20C" box="[576,620,1591,1614]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="378" type="Museum">MCZ</collectionCode>
|
||
<specimenCode id="DBD03A660A03947CFDB96C73FD15B20C" box="[627,717,1591,1614]" collectionCode="OPH-" pageId="13" pageNumber="378">OPH-114</specimenCode>
|
||
, DD =
|
||
<quantity id="4C8E3FF80A03947CFCC56C73FC80B20C" box="[783,856,1591,1614]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="378" unit="mm" value="30.0">30 mm</quantity>
|
||
): (k) portion of dorsal disc and arm
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1E98400A03947CFF1B6C11FEAAB2CB" ID-GBIF-Occurrence="3819603306" collectionCode="LACM" pageId="13" pageNumber="378" specimenCode="LACM 90, 391.1" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="4C76E99E0A03947CFF1B6C11FE78B22E" authorityName="Hendler" authorityYear="1996" box="[209,416,1621,1644]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="vansyoci">
|
||
<emphasis id="B9024E0F0A03947CFF1B6C11FE78B22E" box="[209,416,1621,1644]" italics="true" pageId="13" pageNumber="378">Ophioderma vansyoci</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54CD2CBF0A03947CFE606C11FDDDB22E" box="[426,517,1621,1644]" pageId="13" pageNumber="378" type="holotype">holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DBD03A660A03947CFDC46C11FDBEB22E" box="[526,614,1621,1644]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="13" pageNumber="378" type="Museum">LACM 90</specimenCode>
|
||
–
|
||
<specimenCode id="DBD03A660A03947CFDBB6C11FD71B22E" box="[625,681,1621,1644]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="13" pageNumber="378" type="Museum">391.1</specimenCode>
|
||
, DD =
|
||
<quantity id="4C8E3FF80A03947CFD206C11FCE9B22E" box="[746,817,1621,1644]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="378" unit="mm" value="31.0">31 mm</quantity>
|
||
): (l) portion of dorsal disc and arm; (m) close-up of a jaw
|
||
</materialsCitation>
|
||
.
|
||
<taxonomicName id="4C76E99E0A03947CFEB16C36FDBDB2CB" authorityName="Lutken" authorityYear="1856" box="[379,613,1650,1673]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="13" pageNumber="378" phylum="Echinodermata" rank="species" species="variegatum">
|
||
<emphasis id="B9024E0F0A03947CFEB16C36FDBDB2CB" box="[379,613,1650,1673]" italics="true" pageId="13" pageNumber="378">Ophioderma variegatum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54CD2CBF0A03947CFDBB6C36FD09B2CB" box="[625,721,1650,1673]" pageId="13" pageNumber="378" type="lectotype">lectotype</typeStatus>
|
||
: NHMD-107680, DD =
|
||
<quantity id="4C8E3FF80A03947CFC7A6C36FBD2B2CB" box="[944,1034,1650,1673]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="13" pageNumber="378" unit="mm" value="12.4">12.4 mm</quantity>
|
||
): (n) portion of dorsal disc and arm; (o) close-up of a jaw. See the Supplemental material for more information.
|
||
</paragraph>
|
||
<paragraph id="8BC9921D0A00947FFF0A6ACAFCB2B640" blockId="14.[160,1157,142,514]" pageId="14" pageNumber="379">
|
||
The topology of the trees was consistent between the two genes (COI, 16S) and the two methods (ML, NJ). The main clade grouped the species of
|
||
<taxonomicName id="4C76E99E0A00947FFCA26AF5FC28B488" authorityName="Muller and Troschel" authorityYear="1840" box="[872,1008,177,202]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9024E0F0A00947FFCA26AF5FC28B488" box="[872,1008,177,202]" italics="true" pageId="14" pageNumber="379">Ophioderma</emphasis>
|
||
</taxonomicName>
|
||
, followed by a subclade grouping
|
||
<taxonomicName id="4C76E99E0A00947FFE5D6A90FDD7B4AF" baseAuthorityName="Lyman" baseAuthorityYear="1860" box="[407,527,212,237]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" isUncertain="true" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B9024E0F0A00947FFE5D6A90FE71B4AF" box="[407,425,212,237]" italics="true" pageId="14" pageNumber="379">O</emphasis>
|
||
. cf.
|
||
<emphasis id="B9024E0F0A00947FFE166A90FDD7B4AF" box="[476,527,212,237]" italics="true" pageId="14" pageNumber="379">teres</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9024E0F0A00947FFDD36A90FCF9B4AF" box="[537,801,212,237]" italics="true" pageId="14" pageNumber="379">
|
||
unicolor,
|
||
<taxonomicName id="4C76E99E0A00947FFD4B6A90FCF9B4AF" authorityName="Ziesenhenne" authorityYear="1955" box="[641,801,212,237]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="panamense">O. panamense</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C76E99E0A00947FFC956A90FC3EB4AF" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[863,998,212,237]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A00947FFC956A90FC3EB4AF" box="[863,998,212,237]" italics="true" pageId="14" pageNumber="379">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A00947FFC3A6A90FB94B4AF" box="[1008,1100,212,237]" pageId="14" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and another subclade grouping the two latter species; both presented high bootstrap values (81–100). Sequences of
|
||
<taxonomicName id="4C76E99E0A00947FFE786B5DFDE0B570" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[434,568,281,306]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A00947FFE786B5DFDE0B570" box="[434,568,281,306]" italics="true" pageId="14" pageNumber="379">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A00947FFD8B6B5DFD43B570" box="[577,667,281,306]" pageId="14" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
formed a well-supported clade (bootstrap values: 93/100) for both molecular markers with ML and NJ, separating it from the rest of the species. Particularly for COI, BOLD sequences included in the analysis were grouped with
|
||
<taxonomicName id="4C76E99E0A00947FFF116BC5FEB9B5D8" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[219,353,385,410]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A00947FFF116BC5FEB9B5D8" box="[219,353,385,410]" italics="true" pageId="14" pageNumber="379">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A00947FFEA06BC5FE1EB5D8" box="[362,454,385,410]" pageId="14" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, which confirmed the hypothesis that those specimens were
|
||
<taxonomicName id="4C76E99E0A00947FFF6A6BE0FEF4B5FF" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[160,300,420,445]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A00947FFF6A6BE0FEF4B5FF" box="[160,300,420,445]" italics="true" pageId="14" pageNumber="379">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A00947FFEF36BE0FE4FB5FF" box="[313,407,420,445]" pageId="14" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and not
|
||
<taxonomicName id="4C76E99E0A00947FFDC56BE0FD6CB5FF" authorityName="Ziesenhenne" authorityYear="1955" box="[527,692,420,445]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A00947FFDC56BE0FD6CB5FF" box="[527,692,420,445]" italics="true" pageId="14" pageNumber="379">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
, as suspected based on photographs. According to the analysed data, the sister taxon of
|
||
<taxonomicName id="4C76E99E0A00947FFD296B82FCB0B59D" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[739,872,454,479]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A00947FFD296B82FCB0B59D" box="[739,872,454,479]" italics="true" pageId="14" pageNumber="379">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A00947FFCA56B82FC18B59D" box="[879,960,454,479]" pageId="14" pageNumber="379" rank="species">sp. nov</taxonomicNameLabel>
|
||
is
|
||
<taxonomicName id="4C76E99E0A00947FFC2A6B82FBA6B59D" authorityName="Ziesenhenne" authorityYear="1955" box="[992,1150,454,479]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A00947FFC2A6B82FBA6B59D" box="[992,1150,454,479]" italics="true" pageId="14" pageNumber="379">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
, whereas
|
||
<taxonomicName id="4C76E99E0A00947FFECF6BADFEA3B640" box="[261,379,489,514]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="hendleri">
|
||
<emphasis id="B9024E0F0A00947FFECF6BADFEA3B640" box="[261,379,489,514]" italics="true" pageId="14" pageNumber="379">O. hendleri</emphasis>
|
||
</taxonomicName>
|
||
is the most distantly related to it (
|
||
<figureCitation id="134D8E980A00947FFCCB6BADFC86B640" box="[769,862,489,514]" captionStart="Figure 5" captionStartId="14.[160,225,1609,1632]" captionTargetBox="[165,1150,590,1575]" captionTargetId="figure-160@14.[163,1153,588,1577]" captionTargetPageId="14" captionText="Figure 5. Maximum likelihood tree of the species analysed in this study. (a) COI; (b) 16S. Values at branches indicate bootstrap values (maximum likelihood/neighbour-joining); asterisks indicate specimens of Ophioderma occultum sp. nov. previously identified as Ophioderma panamense. Sequences obtained in this study are indicated in bold. See the Supplemental material for more information." figureDoi="http://doi.org/10.5281/zenodo.6758488" httpUri="https://zenodo.org/record/6758488/files/figure.png" pageId="14" pageNumber="379">Figure 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF09C2950A00947FFF6A6C0DFBB2B2FA" ID-DOI="http://doi.org/10.5281/zenodo.6758488" ID-Zenodo-Dep="6758488" httpUri="https://zenodo.org/record/6758488/files/figure.png" pageId="14" pageNumber="379" startId="14.[160,225,1609,1632]" targetBox="[165,1150,590,1575]" targetPageId="14">
|
||
<paragraph id="8BC9921D0A00947FFF6A6C0DFBB2B2FA" blockId="14.[160,1156,1609,1720]" pageId="14" pageNumber="379">
|
||
<emphasis id="B9024E0F0A00947FFF6A6C0DFF24B222" bold="true" box="[160,252,1609,1632]" pageId="14" pageNumber="379">Figure 5.</emphasis>
|
||
Maximum likelihood tree of the species analysed in this study. (a) COI; (b) 16S. Values at branches indicate bootstrap values (maximum likelihood/neighbour-joining); asterisks indicate specimens of
|
||
<taxonomicName id="4C76E99E0A00947FFF306CC7FE09B2D8" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[250,465,1667,1690]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A00947FFF306CC7FE09B2D8" box="[250,465,1667,1690]" italics="true" pageId="14" pageNumber="379">Ophioderma occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A00947FFE136CC7FDF0B2D8" box="[473,552,1667,1690]" pageId="14" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
|
||
previously identified as
|
||
<taxonomicName id="4C76E99E0A00947FFCD56CC7FBD5B2D8" authorityName="Ziesenhenne" authorityYear="1955" box="[799,1037,1667,1690]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="14" pageNumber="379" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A00947FFCD56CC7FBD5B2D8" box="[799,1037,1667,1690]" italics="true" pageId="14" pageNumber="379">Ophioderma panamense</emphasis>
|
||
</taxonomicName>
|
||
. Sequences obtained in this study are indicated in bold. See the Supplemental material for more information.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9921D0A01947EFF0A6ACAFD7EB54D" blockId="15.[160,1156,142,271]" pageId="15" pageNumber="380">
|
||
The estimated mean genetic distances among species were 8.76–19.95% for COI, and 3.16–22.69% for 16S. The lowest genetic distances were recorded between
|
||
<taxonomicName id="4C76E99E0A01947EFC176AF5FB87B488" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[989,1119,177,202]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="15" pageNumber="380" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A01947EFC176AF5FB87B488" box="[989,1119,177,202]" italics="true" pageId="15" pageNumber="380">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A01947EFBAE6AF5FF09B4AF" pageId="15" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C76E99E0A01947EFECD6A90FE7BB4AF" authorityName="Ziesenhenne" authorityYear="1955" box="[263,419,212,237]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="15" pageNumber="380" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A01947EFECD6A90FE7BB4AF" box="[263,419,212,237]" italics="true" pageId="15" pageNumber="380">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
, and the greatest between
|
||
<taxonomicName id="4C76E99E0A01947EFD1C6A90FC81B4AF" authority="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso, 2022" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Rodríguez-Troncoso" authorityYear="2022" box="[726,857,212,237]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="15" pageNumber="380" phylum="Echinodermata" rank="species" species="occultum" status="sp. nov.">
|
||
<emphasis id="B9024E0F0A01947EFD1C6A90FC81B4AF" box="[726,857,212,237]" italics="true" pageId="15" pageNumber="380">O. occultum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231F3740A01947EFC956A90FC6EB4AF" box="[863,950,212,237]" pageId="15" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C76E99E0A01947EFC266A90FB5BB4AF" authority="(COI)" baseAuthorityName="COI" box="[1004,1155,212,237]" class="Ophiuroidea" family="Ophiopezidae" genus="Ophiopeza" kingdom="Animalia" order="Ophiacanthida" pageId="15" pageNumber="380" phylum="Echinodermata" rank="species" species="fallax">
|
||
<emphasis id="B9024E0F0A01947EFC266A90FB9DB4AF" box="[1004,1093,212,237]" italics="true" pageId="15" pageNumber="380">O. fallax</emphasis>
|
||
(COI)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C76E99E0A01947EFF186AB2FEB7B54D" authorityName="Ziesenhenne" authorityYear="1955" box="[210,367,246,271]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="15" pageNumber="380" phylum="Echinodermata" rank="species" species="panamense">
|
||
<emphasis id="B9024E0F0A01947EFF186AB2FEB7B54D" box="[210,367,246,271]" italics="true" pageId="15" pageNumber="380">O. panamense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C76E99E0A01947EFE636AB2FDDBB54D" box="[425,515,246,271]" class="Ophiuroidea" family="Ophiopezidae" genus="Ophiopeza" kingdom="Animalia" order="Ophiacanthida" pageId="15" pageNumber="380" phylum="Echinodermata" rank="species" species="fallax">
|
||
<emphasis id="B9024E0F0A01947EFE636AB2FDDBB54D" box="[425,515,246,271]" italics="true" pageId="15" pageNumber="380">O. fallax</emphasis>
|
||
</taxonomicName>
|
||
(16S;
|
||
<tableCitation id="C6F4A7A60A01947EFD8D6AB2FD42B54D" box="[583,666,246,271]" captionStart="Table 2" captionStartId="15.[160,216,1534,1557]" captionTargetPageId="15" captionText="Table 2. Mean genetic distances (%) among the analysed species (COI, lower left; 16S, upper right)." httpUri="http://table.plazi.org/id/DF09C2950A01947EFF6A6FBAFBA4B257" pageId="15" pageNumber="380" tableUuid="DF09C2950A01947EFF6A6FBAFBA4B257">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |