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<document id="67B8ED28350B8274D54A211ECFFF6B57" ID-CLB-Dataset="28525" ID-DOI="10.11646/zootaxa.4559.1.1" ID-GBIF-Dataset="bd04f2d1-6de5-43f6-9fca-16a5482835fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="2585011" ID-ZooBank="B5A75104-8C94-462B-A5BD-BB0F28D5C9A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1550795223334" checkinUser="plazi" docAuthor="Packer, Laurence &amp; Dumesh, Sheila" docDate="2019" docId="03C987C3FFEEFF8071E6FAFCFCD4F9E3" docLanguage="en" docName="zootaxa.4559.1.1.pdf" docOrigin="Zootaxa 4559 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chilicola (Oroediscelis) calchaqui Packer &amp; Dumesh 2019, new species" docType="treatment" docUuid="D71B6883-6A86-4529-A03A-60694BE04D61" docUuidSource="ZooBank" docVersion="8" lastPageNumber="26" masterDocId="FFF0FFBBFFF6FF997171FFA3FFA5FF81" masterDocTitle="Fifteen new species of Chilicola (Oroediscelis) (Hymenoptera: Colletidae: Xeromelissinae) with illustrated keys to the males and females of the subgenus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="25" updateTime="1698699904863" updateUser="ExternalLinkService">
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<mods:title id="1D796EFF8CDEB399729782C841A249BB">Fifteen new species of Chilicola (Oroediscelis) (Hymenoptera: Colletidae: Xeromelissinae) with illustrated keys to the males and females of the subgenus</mods:title>
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<mods:namePart id="34BEFC316B15FDEE15A0609543A46A9C">Packer, Laurence</mods:namePart>
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<mods:namePart id="CBAAC3F9012DCFB0ADC0285D2E4C5A63">Dumesh, Sheila</mods:namePart>
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<treatment id="03C987C3FFEEFF8071E6FAFCFCD4F9E3" ID-DOI="http://doi.org/10.5281/zenodo.5945213" ID-GBIF-Taxon="154007299" ID-Zenodo-Dep="5945213" LSID="urn:lsid:zoobank.org:act:D71B6883-6A86-4529-A03A-60694BE04D61" httpUri="http://treatment.plazi.org/id/03C987C3FFEEFF8071E6FAFCFCD4F9E3" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="C37A655EFFEEFF8171E6FAFCFC65FA1A" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BDF36D5FFEEFF8171E6FAFCFC70FAFB" blockId="24.[151,981,1375,1471]" box="[151,981,1375,1402]" pageId="24" pageNumber="25">
<heading id="D09781B9FFEEFF8171E6FAFCFC70FAFB" bold="true" box="[151,981,1375,1402]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B914EAC7FFEEFF8171E6FAFCFC70FAFB" bold="true" box="[151,981,1375,1402]" pageId="24" pageNumber="25">
<taxonomicName id="4C604D56FFEEFF8171E6FAFCFC9DFAFB" authority="Packer and Dumesh" authorityName="Packer &amp; Dumesh" authorityYear="2019" box="[151,824,1375,1402]" class="Insecta" family="Colletidae" genus="Chilicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Oroediscelis">
<emphasis id="B914EAC7FFEEFF8171E6FAFCFEA3FAF8" bold="true" box="[151,262,1375,1401]" italics="true" pageId="24" pageNumber="25">Chilicola</emphasis>
(
<emphasis id="B914EAC7FFEEFF817067FAFCFE0FFAF8" bold="true" box="[278,426,1375,1401]" italics="true" pageId="24" pageNumber="25">Oroediscelis</emphasis>
)
<emphasis id="B914EAC7FFEEFF8170CDFAFCFD96FAF8" bold="true" box="[444,563,1375,1401]" italics="true" pageId="24" pageNumber="25">calchaqui</emphasis>
Packer and Dumesh
</taxonomicName>
,
<taxonomicNameLabel id="A22757BCFFEEFF817234FAC3FC70FAFB" box="[837,981,1376,1402]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
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urn:lsid:zoobank.org:act:
<uuid id="FFC60C00FFEEFF8170D8FA21FC65FA1A" box="[425,960,1410,1435]" pageId="24" pageNumber="25">D71B6883-6A86-4529-A03A-60694BE04D61</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C37A655EFFEEFF8171E6FA05FE3BFA3E" box="[151,414,1446,1471]" pageId="24" pageNumber="25" type="description">
<paragraph id="8BDF36D5FFEEFF8171E6FA05FE3BFA3E" blockId="24.[151,981,1375,1471]" box="[151,414,1446,1471]" pageId="24" pageNumber="25">
(
<figureCitation id="135B2A50FFEEFF8171EEFA05FE82FA3E" box="[159,295,1446,1471]" captionStart="FIGURES 6869" captionStartId="25.[151,264,1074,1097]" captionTargetBox="[151,1436,445,1054]" captionTargetId="figure@25.[151,1436,445,1054]" captionTargetPageId="25" captionText="FIGURES 6869. Chilicola calchaqui. 68. female lateral habitus (holotype); 69. female face (holotype)." httpUri="https://zenodo.org/record/2585035/files/figure.png" pageId="24" pageNumber="25">Figs. 6869</figureCitation>
,
<figureCitation id="135B2A50FFEEFF817043FA04FEFBFA3E" box="[306,350,1447,1471]" captionStart="FIGURES 129130" captionStartId="41.[151,264,681,704]" captionTargetBox="[161,1418,187,657]" captionTargetId="figure@41.[151,1436,187,658]" captionTargetPageId="41" captionText="FIGURES 129130. Head, frontal view, to show relative length: width. 129. L~1.5XW (C. calchaqui holotype); 130. L&lt;1.35W (C. carPenteri paratype)." httpUri="https://zenodo.org/record/2585097/files/figure.png" pageId="24" pageNumber="25">129</figureCitation>
,
<figureCitation id="135B2A50FFEEFF81701BFA04FE30FA3E" box="[362,405,1447,1471]" captionStart="FIGURES 131132" captionStartId="41.[151,264,1270,1293]" captionTargetBox="[151,1436,776,1245]" captionTargetId="figure@41.[151,1436,774,1248]" captionTargetPageId="41" captionText="FIGURES 131132. Heads, side view to show relative length:width of malar space. 131. L clearly&gt; W (C. calchaqui holotype); 132. L&lt;W (C. carPenteri paratype)." httpUri="https://zenodo.org/record/2585099/files/figure.png" pageId="24" pageNumber="25">131</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37A655EFFEEFF8171E6FA4DFEEEF917" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BDF36D5FFEEFF8171E6FA4DFEEEF917" blockId="24.[151,1437,1518,2011]" pageId="24" pageNumber="25">
<emphasis id="B914EAC7FFEEFF8171E6FA4DFEB4F986" bold="true" box="[151,273,1518,1543]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
The relatively extreme length of the head is diagnostic for the female of this species, it is almost 1.5X longer than wide (99:68). Several species of the subgenus have heads that are 1.31.35X longer than wide, but not approaching the extreme relative length of this species. In addition to its extremely long head,
<taxonomicName id="4C604D56FFEEFF8175CCF994FAE9F9CE" authorityName="Packer &amp; Dumesh" authorityYear="2019" box="[1213,1356,1590,1615]" class="Insecta" family="Colletidae" genus="Chilicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="calchaqui">
<emphasis id="B914EAC7FFEEFF8175CCF994FAE9F9CE" box="[1213,1356,1590,1615]" italics="true" pageId="24" pageNumber="25">C. calchaqui</emphasis>
</taxonomicName>
differs from the other longer headed species in the very straight, almost unbranched and uninterrupted striae of the metapostnotum.
</paragraph>
</subSubSection>
<subSubSection id="C37A655EFFEEFF8071B6F901FEDAFAE7" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="8BDF36D5FFEEFF8171B6F901FBDAF93B" blockId="24.[151,1437,1518,2011]" box="[199,1151,1698,1723]" pageId="24" pageNumber="25">
<emphasis id="B914EAC7FFEEFF8171B6F901FEFCF93A" bold="true" box="[199,345,1698,1723]" pageId="24" pageNumber="25">Description.</emphasis>
<emphasis id="B914EAC7FFEEFF81702EF901FE1EF93A" box="[351,443,1698,1723]" italics="true" pageId="24" pageNumber="25">Female.</emphasis>
Length
<quantity id="4C989B30FFEEFF817366F901FDD5F93B" box="[535,624,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="24" pageNumber="25" unit="mm" value="5.5">5.5 mm</quantity>
; forewing length
<quantity id="4C989B30FFEEFF817244F900FC22F93A" box="[821,903,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="24" pageNumber="25" unit="mm" value="4.4">4.4mm</quantity>
, head width
<quantity id="4C989B30FFEEFF817565F900FBDEF93B" box="[1044,1147,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="24" pageNumber="25" unit="mm" value="1.38">1.38 mm</quantity>
.
</paragraph>
<paragraph id="8BDF36D5FFEEFF8171B6F965FAD4F812" blockId="24.[151,1437,1518,2011]" pageId="24" pageNumber="25">
<emphasis id="B914EAC7FFEEFF8171B6F965FEA1F95E" box="[199,260,1734,1759]" italics="true" pageId="24" pageNumber="25">Head</emphasis>
: almost 1.5 X as long as wide, 99:68; entirely black. Clypeus imbricate, punctures irregular in size and spacing, i=13 d; with weak longitudinal depression for basal 2/3; supraclypeal area punctures close in size to the smaller ones of clypeus; lower paraocular area punctures i=12 d; frontal area densely punctate, i=0.51d; facial fovea weakly imbricate, no less densely punctate than adjacent frontal area; vertexal area behind compound eye with punctures in longitudinal depressions; vertexal area behind ocellar triangle sparsely punctate, i=15 d. Malar space longer than wide (14:11). Longest hairs on face above antennal sockets, &lt;2 MOD; genal beard &lt;3 MOD.
</paragraph>
<paragraph id="8BDF36D5FFEEFF8071B6F83CFC49FE09" blockId="24.[151,1437,1518,2011]" lastBlockId="25.[151,1437,151,392]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<taxonomicName id="4C604D56FFEEFF8171B6F83CFE1BF837" authority=": pronotum" authorityName="Pronotum" box="[199,446,1951,1974]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAC7FFEEFF8171B6F83CFE9AF837" box="[199,319,1951,1974]" italics="true" pageId="24" pageNumber="25">Mesosoma</emphasis>
: pronotum
</taxonomicName>
with fine, dense punctures, i~d; mesoscutum somewhat dull; punctures distinct, i~d, somewhat sparser on disc, larger towards rear; mesoscutellum more weakly imbricate, punctures less regular, i=0.53 d; metanotum dull, punctures i=13 d; mesepisternum with scattered minute punctures among more abundant larger ones, i=13 d, sparser above and anterior to scrobe; metepisternum ruguloso-punctate above, coarsely imbricate below. Mesoscutum lacking long hairs, &lt;0,5 MOD; mesepisternum with longest hairs &lt;2.3 MOD. Stigma almost as long as marginal cell on costal margin (36:38); apex of stigma more gradually narrowing than in other species; distal stigmal perpendicular past middle of second submarginal cell. Metapostnotum striate, striae long, parallel, straight, rarely interrupted or branched; lateral surface of propodeum distinctly but shallowly punctate above, i=23 d, punctures increasingly shallow and obscure below.
</paragraph>
<caption id="DF1F665DFFEFFF8071E6FB91FB1BFBC8" box="[151,1214,1074,1097]" httpUri="https://zenodo.org/record/2585035/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,264,1074,1097]" targetBox="[151,1436,445,1054]" targetPageId="25">
<paragraph id="8BDF36D5FFEFFF8071E6FB91FB1BFBC8" blockId="25.[151,1214,1074,1097]" box="[151,1214,1074,1097]" pageId="25" pageNumber="26">
<emphasis id="B914EAC7FFEFFF8071E6FB91FEF1FBC9" bold="true" box="[151,340,1074,1097]" pageId="25" pageNumber="26">FIGURES 6869.</emphasis>
<taxonomicName id="4C604D56FFEFFF80702BFB90FD87FBC8" authorityName="Packer &amp; Dumesh" authorityYear="2019" box="[346,546,1075,1097]" class="Insecta" family="Colletidae" genus="Chilicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAC7FFEFFF80702BFB90FD87FBC8" box="[346,546,1075,1097]" italics="true" pageId="25" pageNumber="26">Chilicola calchaqui</emphasis>
</taxonomicName>
. 68. female lateral habitus (holotype); 69. female face (holotype).
</paragraph>
</caption>
<paragraph id="8BDF36D5FFEFFF8071B6FB38FEE9FAC2" blockId="25.[151,1437,1178,1635]" pageId="25" pageNumber="26">
<emphasis id="B914EAC7FFEFFF8071B6FB38FE99FB33" box="[199,316,1179,1202]" italics="true" pageId="25" pageNumber="26">Metasoma</emphasis>
: Terga shiny, shallowly imbricate; T1 finely punctate, i=13 d on disc, apical impressed area mostly impunctate; T2 punctures larger and more evenly spaced around basal 1/3, i~d; finely and irregularly punctate posteriorly, i=18d, only apical half of impressed area impunctate, translucent; T3 more evenly punctate, i=12 d except more sparsely punctate towards apex, apical half of impressed area impunctate (remaining terga telescoped and not visible).
</paragraph>
<paragraph id="8BDF36D5FFEFFF8071B6FAEEFEDAFAE7" blockId="25.[151,1437,1178,1635]" box="[199,383,1357,1382]" pageId="25" pageNumber="26">
<emphasis id="B914EAC7FFEFFF8071B6FAEEFEA4FAE7" box="[199,257,1357,1382]" italics="true" pageId="25" pageNumber="26">Male</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37A655EFFEFFF8071B6FAD1FD8AFA2F" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BDF36D5FFEFFF8071B6FAD1FD8AFA2F" blockId="25.[151,1437,1178,1635]" pageId="25" pageNumber="26">
<emphasis id="B914EAC7FFEFFF8071B6FAD1FE32FA0A" bold="true" box="[199,407,1394,1419]" pageId="25" pageNumber="26">Material studied.</emphasis>
<materialsCitation id="3B083C88FFEFFF8070D1FAD1FD8AFA2F" ID-GBIF-Occurrence="2005761106" collectingDate="1986-02" collectionCode="AMNH" collectorName="M. Fritz" country="Argentina" elevation="2400" location="Tafi" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Tucuman" typeStatus="holotype">
<typeStatus id="54DB8877FFEFFF8070D1FAD1FDA2FA0A" box="[416,519,1394,1419]" pageId="25" pageNumber="26" type="holotype">
<emphasis id="B914EAC7FFEFFF8070D1FAD1FDA2FA0A" box="[416,519,1394,1419]" italics="true" pageId="25" pageNumber="26">Holotype</emphasis>
</typeStatus>
female:
<collectingCountry id="F3777645FFEFFF80731CFAD1FCB6FA0B" box="[621,787,1394,1418]" name="Argentina" pageId="25" pageNumber="26">ARGENTINA</collectingCountry>
,
<collectingRegion id="49A4F837FFEFFF807251FAD0FC22FA0B" box="[800,903,1395,1418]" country="Argentina" name="Tucuman" pageId="25" pageNumber="26">
<emphasis id="B914EAC7FFEFFF807251FAD0FC22FA0B" box="[800,903,1395,1418]" italics="true" pageId="25" pageNumber="26">Tucumán</emphasis>
</collectingRegion>
,
<location id="8EBF600EFFEFFF8072E6FAD1FC62FA0A" LSID="urn:lsid:plazi:treatment:03C987C3FFEEFF8071E6FAFCFCD4F9E3:8EBF600EFFEFFF8072E6FAD1FC62FA0A" box="[919,967,1394,1419]" country="Argentina" name="Tafi" pageId="25" pageNumber="26" stateProvince="Tucuman">Tafi</location>
,
<elevation id="004DD1E6FFEFFF8072A5FAD1FB80FA0B" box="[980,1061,1394,1418]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="26" unit="m" value="2400.0">
<quantity id="4C989B30FFEFFF8072A5FAD1FB80FA0B" box="[980,1061,1394,1418]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="26" unit="m" value="2400.0">2400m</quantity>
</elevation>
,
<collectingDate id="EF9AE9FDFFEFFF807542FAD1FB21FA0A" box="[1075,1156,1394,1419]" pageId="25" pageNumber="26" value="1986-02">
<date id="FFDE1015FFEFFF807542FAD1FB21FA0A" box="[1075,1156,1394,1419]" pageId="25" pageNumber="26" value="1986-02">ii.1986</date>
</collectingDate>
,
<collectorName id="26955303FFEFFF8075E0FAD0FB55FA0A" box="[1169,1264,1394,1419]" pageId="25" pageNumber="26">M. Fritz</collectorName>
[
<collectionCode id="ED71AE10FFEFFF80758EFAD1FAFEFA0B" box="[1279,1371,1394,1418]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
] [See
<figureCitation id="135B2A50FFEFFF8071E6FA36FE92FA2F" box="[151,311,1429,1454]" captionStart="FIGURES 184185" captionStartId="53.[151,264,966,989]" captionTargetBox="[151,1436,185,950]" captionTargetId="figure@53.[151,1436,185,950]" captionTargetPageId="53" captionText="FIGURES 184185. Distribution maps of newly described Chilicola (Oroediscelis) species. 184. Large scale map of South America showing distribution of new species; 185. Small scale map of northern Argentina to show distribution within the country." httpUri="https://zenodo.org/record/2585153/files/figure.png" pageId="25" pageNumber="26">Figs. 184185</figureCitation>
for distribution map].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A655EFFEFFF8071B6FA19FC90FA77" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BDF36D5FFEFFF8071B6FA19FC90FA77" blockId="25.[151,1437,1178,1635]" pageId="25" pageNumber="26">
<emphasis id="B914EAC7FFEFFF8071B6FA19FEEBFA52" bold="true" box="[199,334,1466,1491]" pageId="25" pageNumber="26">Etymology.</emphasis>
The species is named after the indigenous Calchaqui people that used to inhabit the region of northwestern
<collectingCountry id="F3777645FFEFFF807041FA7EFE3AFA77" box="[304,415,1501,1526]" name="Argentina" pageId="25" pageNumber="26">Argentina</collectingCountry>
from whence this species is known.
</paragraph>
</subSubSection>
<subSubSection id="C37A655EFFEFFF8071B6F9A1FCD4F9E3" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BDF36D5FFEFFF8071B6F9A1FCD4F9E3" blockId="25.[151,1437,1178,1635]" pageId="25" pageNumber="26">
<emphasis id="B914EAC7FFEFFF8071B6F9A1FEEAF99A" bold="true" box="[199,335,1538,1563]" pageId="25" pageNumber="26">Comments.</emphasis>
While the male is unknown, it is expected to be readily identifiable through having a similarly elongate head. The very long head of this species is a convergent feature that has evolved independently at least five times elsewhere in the Xeromelissinae (
<bibRefCitation id="EFF14B24FFEFFF8073F6F9E9FCB8F9E2" author="Packer, L." box="[647,797,1610,1635]" pageId="25" pageNumber="26" pagination="84 - 91" refId="ref28934" refString="Packer, L. (2005) A new species of Geodiscelis (Hymenoptera: Colletidae: Xeromelissinae) from the Atacama desert of Chile. Journal of HymenoPtera Research, 14, 84 - 91." type="journal article" year="2005">Packer, 2005</bibRefCitation>
;
<bibRefCitation id="EFF14B24FFEFFF807259F9E9FCC0F9E3" author="Packer, L." box="[808,869,1610,1634]" pageId="25" pageNumber="26" pagination="72 - 96" refId="ref28971" refString="Packer, L. (2008) Phylogeny and classification of the Xeromelissinae (Hymenoptera: Apoidea, Colletidae) with special emphasis on the genus Chilicola. Systematic Entomology, 33, 72 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.2007.00398. x" type="journal article" year="2008">2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>