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<document id="9CC0A4D37A4D7BDFCB399D24962DCAE5" ID-CLB-Dataset="53324" ID-DOI="10.11646/phytotaxa.576.2.1" ID-GBIF-Dataset="1a9bd59f-7293-432f-b3c1-6df0bc659474" ID-ISSN="1179-3163" ID-Zenodo-Dep="7461280" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1671521402645" checkinUser="plazi" docAuthor="Hassan, Walaa A., Al-Shaye, Najla A., Alghamdi, Salma, Korany, Shereen M. &amp; Iamonico, Duilio" docDate="2022" docId="03C987964B4EFFAB06B902B854526096" docLanguage="en" docName="phytotaxa.576.2.1.pdf" docOrigin="Phytotaxa 576 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.576.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Amaranthus sparganicephalus Thell." docType="treatment" docVersion="7" lastPageNumber="145" masterDocId="FFF0FFEE4B47FFA106310226570A6226" masterDocTitle="Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia" masterLastPageNumber="157" masterPageNumber="135" pageNumber="144" updateTime="1699413697922" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="07F4F9F8D58B250E89C7CCE4CAE92AB9">Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia</mods:title>
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<mods:namePart id="BB7980D4EE1F257D971533C994F7D9E7">Hassan, Walaa A.</mods:namePart>
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<mods:namePart id="A0E762491C76E35B32A3D2DF9E989DA1">Al-Shaye, Najla A.</mods:namePart>
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<mods:affiliation id="DADCAC15C416E75D8C1CA22DFBED5558">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; naaalshaye @ pnu. edu. sa; https: // orcid. org / 0000 - 0002 - 0447 - 8613</mods:affiliation>
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<mods:namePart id="18C640F9023CEF2BE37B131FCBA8B4BF">Alghamdi, Salma</mods:namePart>
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<mods:affiliation id="20F4275D56F7B04CF281ED7416561B6E">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; saatalghamdi @ pnu. edu. sa; https: // orcid. org / 0000 - 0001 - 7716 - 2233</mods:affiliation>
<mods:nameIdentifier id="46F8BAB3EA6CD0D5545919AF1FA458CB" type="email">saatalghamdi@pnu.edu.sa</mods:nameIdentifier>
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<mods:namePart id="111DA5F11565A4F61AE96A2524D047F4">Korany, Shereen M.</mods:namePart>
<mods:nameIdentifier id="334EB4743FD14F5C8C729B8814F9D0A8" type="ORCID">0000-0002-9355-3952</mods:nameIdentifier>
<mods:affiliation id="6325486CF5B63DE055917977F4841E6C">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; smkorany @ pnu. edu. as; https: // orcid. org / 0000 - 0002 - 9355 - 3952</mods:affiliation>
<mods:nameIdentifier id="E7C3A8436C50FAB519D9F6113C448A56" type="email">smkorany@pnu.edu.as</mods:nameIdentifier>
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<mods:namePart id="CF2C4C82A34A59120D9AE4C59C2312AB">Iamonico, Duilio</mods:namePart>
<mods:nameIdentifier id="04518AA7D6A5E9F2E8F1B7928A6F6461" type="ORCID">0000-0001-5491-7568</mods:nameIdentifier>
<mods:affiliation id="04EFAB81E69A932CBA3420F714FC80D6">Ce. R. S. I. Te. S., University of Rome Sapienza, viale XXIV Maggio 7, 04100 - Latina, Italy &amp; duilio. iamonico @ uniroma 1. it; https: // orcid. org / 0000 - 0001 - 5491 - 7568 &amp; Authors that contributed equally to the work</mods:affiliation>
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<mods:title id="351603A4A1C970F63575FBC2CE907E7B">Phytotaxa</mods:title>
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<mods:date id="05A2AD2D88F98B69F09BB02500C1230A">2022</mods:date>
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<treatment id="03C987964B4EFFAB06B902B854526096" ID-DOI="http://doi.org/10.5281/zenodo.7461254" ID-GBIF-Taxon="206458914" ID-Zenodo-Dep="7461254" LSID="urn:lsid:plazi:treatment:03C987964B4EFFAB06B902B854526096" httpUri="http://treatment.plazi.org/id/03C987964B4EFFAB06B902B854526096" lastPageId="10" lastPageNumber="145" pageId="9" pageNumber="144">
<subSubSection id="C37A650B4B4EFFA806B902B853E8629E" box="[136,1250,158,185]" pageId="9" pageNumber="144" type="nomenclature">
<paragraph id="8BDF36804B4EFFA806B902B853E8629E" blockId="9.[136,1451,158,256]" box="[136,1250,158,185]" pageId="9" pageNumber="144">
<heading id="D09781EC4B4EFFA806B902B853E8629E" box="[136,1250,158,185]" fontSize="11" level="2" pageId="9" pageNumber="144" reason="4">
<emphasis id="B914EA924B4EFFA806B902B85797629E" bold="true" box="[136,157,158,184]" pageId="9" pageNumber="144">6.</emphasis>
<taxonomicName id="4C604D034B4EFFA8069502B8555B629E" authority="Thell." authorityName="Thell." box="[164,593,158,185]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="sparganicephalus">
<emphasis id="B914EA924B4EFFA8069502B85502629F" bold="true" box="[164,520,158,185]" italics="true" pageId="9" pageNumber="144">Amaranthus sparganicephalus</emphasis>
Thell.
</taxonomicName>
in Ascherson &amp; Graebner, Syn. Mitteleur. Fl. 5: 312. 1914
</heading>
</paragraph>
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<subSubSection id="C37A650B4B4EFFA806B902E0531562D9" pageId="9" pageNumber="144" type="materials_examined">
<paragraph id="8BDF36804B4EFFA806B902E0531562D9" blockId="9.[136,1451,158,256]" pageId="9" pageNumber="144">
<materialsCitation id="3B083CDD4B4EFFA806B902E0531162D9" ID-GBIF-Occurrence="3999584327" collectingDate="1985-09-19" collectorName="J. Qara &amp; A. G. Miller" country="Oman" elevation="880" location="Aqarnahawat" municipality="Type" pageId="9" pageNumber="144" specimenCount="2" stateProvince="Zufar" typeStatus="neotype">
<collectingMunicipality id="6BBBACFA4B4EFFA806B902E057B262FD" box="[136,184,198,219]" pageId="9" pageNumber="144">Type</collectingMunicipality>
(
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, here designated):—
<collectingCountry id="F37776104B4EFFA807EB02E0552C62FA" box="[474,550,198,220]" name="Oman" pageId="9" pageNumber="144">OMAN</collectingCountry>
,
<collectingRegion id="49A4F8624B4EFFA8040102E0557C62FA" box="[560,630,198,220]" country="Oman" name="Zufar" pageId="9" pageNumber="144">Dhofar</collectingRegion>
,
<collectorName id="269553564B4EFFA8044E02E055CF62FD" box="[639,709,198,219]" pageId="9" pageNumber="144">J. Qara</collectorName>
, nr.
<location id="8EBF605B4B4EFFA804DD02E0546462FA" LSID="urn:lsid:plazi:treatment:03C987964B4EFFAB06B902B854526096:8EBF605B4B4EFFA804DD02E0546462FA" box="[748,878,198,220]" country="Oman" municipality="Type" name="Aqarnahawat" pageId="9" pageNumber="144" stateProvince="Zufar">Aqarnahawat</location>
,
<taxonomicName id="4C604D034B4EFFA8054902E054B162FD" box="[888,955,198,219]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="B914EA924B4EFFA8054902E054B162FD" box="[888,955,198,219]" italics="true" pageId="9" pageNumber="144">Acacia</emphasis>
</taxonomicName>
hollow,
<elevation id="004DD1B34B4EFFA8023F02E0537462FA" box="[1038,1150,198,220]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="9" pageNumber="144" unit="m" value="880.0">
<quantity id="4C989B654B4EFFA8023F02E0534162FD" box="[1038,1099,198,219]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="9" pageNumber="144" unit="m" value="880.0">880 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDE10404B4EFFA802B802E053F262FA" box="[1161,1272,198,220]" pageId="9" pageNumber="144" value="1985-09-19">
<collectingDate id="EF9AE9A84B4EFFA802B802E053F262FA" box="[1161,1272,198,220]" pageId="9" pageNumber="144" value="1985-09-19">19.09.1985</collectingDate>
</date>
,
<emphasis id="B914EA924B4EFFA8033002E052A162FD" box="[1281,1451,198,219]" italics="true" pageId="9" pageNumber="144">
<collectorName id="269553564B4EFFA8033002E0527F62FD" box="[1281,1397,198,219]" pageId="9" pageNumber="144">A. G. Miller</collectorName>
7693
</emphasis>
(E00687024!, image of the neotype available at https://data.rbge.org.uk/herb/E00687024)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A650B4B4EFFAB06B9030054526096" lastPageId="10" lastPageNumber="145" pageId="9" pageNumber="144" type="description">
<paragraph id="8BDF36804B4EFFA806B90300548B605B" blockId="9.[136,1452,294,674]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA806B9030056326366" bold="true" box="[136,312,294,320]" pageId="9" pageNumber="144">Description:—</emphasis>
Herbs,
<quantity id="4C989B654B4EFFA807B8030156FB6367" box="[393,497,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="6.0" metricValueMin="0.7" pageId="9" pageNumber="144" unit="cm" value="3.35" valueMax="6.0" valueMin="0.7">0.76 cm</quantity>
tall, monoecious, annual (therophyte). Stems erect, glabrous (pubescent in the upper part), yellowish, simple or branched (branches sometimes decumbent). Leaves green to dark-green, ovate (1.55.5 × 0.63.0 cm), with entire margins, apex obtuse to retuse, mucronate, base cuneate, glabrous (short hairs on the nerves of the abaxial surface), long petioled (petiole up to 6.0 cm long). Synflorescences arranged in axillary spherical glomerules, ±
<quantity id="4C989B654B4EFFA807150392565C63E8" box="[292,342,436,462]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="144" unit="cm" value="1.0">1 cm</quantity>
in diameter, brown to dark-brown. Floral bracts ovate (0.30.5 × ca.
<quantity id="4C989B654B4EFFA8027C0392539663E8" box="[1101,1180,436,462]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="144" unit="cm" value="0.2">0.2 cm</quantity>
), about 1/2 times shorter than the perianth, acute to acuminate, mucronate. Staminate flowers with 3 tepals, green, ovate-lanceolate; stamens 3. Pistillate flowers with 3 tepals, green, ovate-lanceolate (1.01.3 × ca.
<quantity id="4C989B654B4EFFA805A803DC54FE6032" box="[921,1012,506,532]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="144" unit="mm" value="0.3">0.3 mm</quantity>
), with obtuse apex, median vein green; stigmas 2. Fruits arranged in stellate heads (divergent capsules), each one brown, as a double cone (2.753.25 × ca.
<quantity id="4C989B654B4EFFA806B9006657E8607C" box="[136,226,576,603]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="144" unit="mm" value="1.5">1.5 mm</quantity>
), longer than the perianth, the half base longitudinally sulcate, dehiscent with point of junction of lid and base cristate-crenulate. Seeds lenticular (
<quantity id="4C989B654B4EFFA80429004555AA6058" box="[536,672,611,638]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="9" pageNumber="144" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
in diameter), black.
</paragraph>
<paragraph id="8BDF36804B4EFFA8068C00AE55CA6087" blockId="9.[136,1452,294,674]" box="[189,704,647,674]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA8068C00AE56716084" bold="true" box="[189,379,648,674]" pageId="9" pageNumber="144">Iconography:—</emphasis>
<bibRefCitation id="EFF14B714B4EFFA8074A00A155696084" author="Townsend, C. C." box="[379,611,647,674]" pageId="9" pageNumber="144" pagination="1 - 136" refId="ref18438" refString="Townsend, C. C. (1985) Amaranthaceae. In: Polhill, R. M. (Ed.) Flora of Tropical East Africa. A. A. Balkema, Rotterdam, pp. 1 - 136." type="book chapter" year="1985">Townsend (1985: 33</bibRefCitation>
,
<figureCitation id="135B2A054B4EFFA8045E00A155B86087" box="[623,690,647,673]" captionStart="FIGURE 6" captionStartId="9.[136,229,1478,1500]" captionTargetId="figure-232@9.[265,1323,699,1454]" captionTargetPageId="9" captionText="FIGURE 6. Map of Amaranthus sparaganicephalus." figureDoi="http://doi.org/10.5281/zenodo.7465630" httpUri="https://zenodo.org/record/7465630/files/figure.png" pageId="9" pageNumber="144">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF1F66084B4EFFA806B907E0558D67FD" ID-DOI="http://doi.org/10.5281/zenodo.7465630" ID-Zenodo-Dep="7465630" box="[136,647,1478,1500]" httpUri="https://zenodo.org/record/7465630/files/figure.png" pageId="9" pageNumber="144" startId="9.[136,229,1478,1500]" targetBox="[265,1323,699,1454]" targetPageId="9">
<paragraph id="8BDF36804B4EFFA806B907E0558D67FD" blockId="9.[136,647,1478,1500]" box="[136,647,1478,1500]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA806B907E057F767FD" bold="true" box="[136,253,1478,1500]" pageId="9" pageNumber="144">FIGURE 6.</emphasis>
Map of
<taxonomicName id="4C604D034B4EFFA8077E07E0558867FD" box="[335,642,1478,1499]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="sparaganicephalus">
<emphasis id="B914EA924B4EFFA8077E07E0558867FD" box="[335,642,1478,1499]" italics="true" pageId="9" pageNumber="144">Amaranthus sparaganicephalus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BDF36804B4EFFA8068C042455FE643A" blockId="9.[136,1452,1538,2086]" box="[189,756,1538,1565]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA8068C04245655643A" bold="true" box="[189,351,1538,1564]" pageId="9" pageNumber="144">Phenology:—</emphasis>
Flowering time from march to april.
</paragraph>
<paragraph id="8BDF36804B4EFFA8068C040354AA6419" blockId="9.[136,1452,1538,2086]" box="[189,928,1573,1600]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA8068C040356EF6419" bold="true" box="[189,485,1573,1599]" pageId="9" pageNumber="144">Habitat and elevation:—</emphasis>
Uncultivated land, around
<quantity id="4C989B654B4EFFA805210403546F6466" box="[784,869,1573,1600]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="144" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BDF36804B4EFFA8068C046155A16444" blockId="9.[136,1452,1538,2086]" box="[189,683,1607,1634]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA8068C046156E26444" bold="true" box="[189,488,1607,1634]" pageId="9" pageNumber="144">Chromosome number:—</emphasis>
Not still counted.
</paragraph>
<paragraph id="8BDF36804B4EFFA8068C044C52676481" blockId="9.[136,1452,1538,2086]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA8068C044C566A64A2" bold="true" box="[189,352,1642,1668]" pageId="9" pageNumber="144">Chorology:—</emphasis>
Species native to eastern tropical Africa (
<collectingCountry id="F37776104B4EFFA8057F044C54BA64A2" box="[846,944,1642,1668]" name="Djibouti" pageId="9" pageNumber="144">Djibouti</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA805F1044C530564A2" box="[960,1039,1642,1668]" name="Eritrea" pageId="9" pageNumber="144">Eritrea</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA80211044C538864A2" box="[1056,1154,1642,1668]" name="Ethiopia" pageId="9" pageNumber="144">Ethiopia</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA802A3044D53D464A3" box="[1170,1246,1643,1669]" name="Kenya" pageId="9" pageNumber="144">Kenya</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA802DE044C524464A2" box="[1263,1358,1642,1668]" name="Somalia" pageId="9" pageNumber="144">Somalia</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA8036F044C52AD64A2" box="[1374,1447,1642,1668]" name="Sudan" pageId="9" pageNumber="144">Sudan</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA806B904AB57FB6481" box="[136,241,1677,1703]" name="Tanzania" pageId="9" pageNumber="144">Tanzania</collectingCountry>
), Arabian Peninsula (
<collectingCountry id="F37776104B4EFFA807EE04AB557F6481" box="[479,629,1677,1703]" name="Saudi Arabia" pageId="9" pageNumber="144">Saudi Arabia</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA8044E04A855DA648E" box="[639,720,1678,1704]" name="Yemen" pageId="9" pageNumber="144">Yemen</collectingCountry>
,
<collectingCountry id="F37776104B4EFFA804EA04AB542E6481" box="[731,804,1677,1703]" name="Oman" pageId="9" pageNumber="144">Oman</collectingCountry>
), and
<collectingCountry id="F37776104B4EFFA8055604AB54B76481" box="[871,957,1677,1703]" name="Yemen" pageId="9" pageNumber="144">Socotra</collectingCountry>
(
<bibRefCitation id="EFF14B714B4EFFA805FB04AB53616481" author="POWO" box="[970,1131,1677,1703]" pageId="9" pageNumber="144" refId="ref17700" refString="POWO (2022 b) Plants of the World Online. Amaranthus sparaganicephalus Thell. Available from: https: // powo. science. kew. org / taxon / urn: lsid: ipni. org: names: 59735 - 1 (accessed 29 March 2022)" type="url" year="2022">POWO 2022b</bibRefCitation>
and literature therein).
</paragraph>
<paragraph id="8BDF36804B4EFFA8068C048955ED64EC" blockId="9.[136,1452,1538,2086]" box="[189,743,1711,1738]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA8068C0489557864EF" bold="true" box="[189,626,1711,1737]" pageId="9" pageNumber="144">
Occurrence in
<collectingCountry id="F37776104B4EFFA8075F0489550664EF" box="[366,524,1711,1737]" name="Saudi Arabia" pageId="9" pageNumber="144">Saudi Arabia</collectingCountry>
(
<figureCitation id="135B2A054B4EFFA8042A0489556864EF" box="[539,610,1711,1737]" captionStart="FIGURE 6" captionStartId="9.[136,229,1478,1500]" captionTargetId="figure-232@9.[265,1323,699,1454]" captionTargetPageId="9" captionText="FIGURE 6. Map of Amaranthus sparaganicephalus." figureDoi="http://doi.org/10.5281/zenodo.7465630" httpUri="https://zenodo.org/record/7465630/files/figure.png" pageId="9" pageNumber="144">Fig. 6</figureCitation>
):
</emphasis>
<collectingRegion id="49A4F8624B4EFFA804BE049655E864EC" box="[655,738,1712,1738]" country="Saudi Arabia" name="Ar Riyad" pageId="9" pageNumber="144">Riyadh</collectingRegion>
.
</paragraph>
<paragraph id="8BDF36804B4EFFA8068C04F4559C6A00" blockId="9.[136,1452,1538,2086]" pageId="9" pageNumber="144">
<emphasis id="B914EA924B4EFFA8068C04F4545464CA" bold="true" box="[189,862,1746,1773]" pageId="9" pageNumber="144">
Typification of the name
<taxonomicName id="4C604D034B4EFFA807D504F4543364CB" box="[484,825,1746,1773]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="sprganicephalus">
<emphasis id="B914EA924B4EFFA807D504F4543364CB" bold="true" box="[484,825,1746,1773]" italics="true" pageId="9" pageNumber="144">Amaranthus sprganicephalus</emphasis>
</taxonomicName>
:—
</emphasis>
<taxonomicName id="4C604D034B4EFFA8056F04F453B964CA" box="[862,1203,1746,1772]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="sparganicephalus">
<emphasis id="B914EA924B4EFFA8056F04F453B964CA" box="[862,1203,1746,1772]" italics="true" pageId="9" pageNumber="144">Amaranthus sparganicephalus</emphasis>
</taxonomicName>
was validly published by
<bibRefCitation id="EFF14B714B4EFFA8069904D356876529" author="Thellung, A." box="[168,397,1781,1807]" pageId="9" pageNumber="144" pagination="225 - 356" refId="ref18248" refString="Thellung, A. (1914) Amaranthus L. In: Ascherson, P. &amp; Graebner, P. (Eds.) Synopsis der Mitteleuropaischen Flora, vol. 5. Verlag Von Gebruder Borntraeger. Leipzig, Germany, pp. 225 - 356." type="book chapter" year="1914">Thellung (1914: 312</bibRefCitation>
, a note) by a short diagnosis (in german) in which the following two characters were highlighted: the shape of the fruiting glomerules (which resembles those of the members belonging to the genus
<taxonomicName id="4C604D034B4EFFA802CA053E52896514" box="[1275,1411,1816,1842]" class="Liliopsida" family="Typhaceae" genus="Sparganium" kingdom="Plantae" order="Poales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="B914EA924B4EFFA802CA053E52896514" box="[1275,1411,1816,1842]" italics="true" pageId="9" pageNumber="144">Sparganium</emphasis>
</taxonomicName>
L.; the specific epithet “
<taxonomicName id="4C604D034B4EFFA80742051C55336572" box="[371,569,1850,1876]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="sparganicephalus">
<emphasis id="B914EA924B4EFFA80742051C55336572" box="[371,569,1850,1876]" italics="true" pageId="9" pageNumber="144">sparganicephalus</emphasis>
</taxonomicName>
” derives from this feature) and that of each fruit (“polyedrisch abgeflachten” = “polyhedral flattened”); a morphologic comparison with
<taxonomicName id="4C604D034B4EFFA8050D057B533D655E" authority="Lam." authorityName="Lam." box="[828,1079,1885,1912]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="angustifolius">
<emphasis id="B914EA924B4EFFA8050D057B54FB6551" box="[828,1009,1885,1911]" italics="true" pageId="9" pageNumber="144">A. angustifolius</emphasis>
Lam.
</taxonomicName>
(currently
<taxonomicName id="4C604D034B4EFFA8028D057B560B65BF" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="subSpecies" species="graecizans" subSpecies="graecizans">
<emphasis id="B914EA924B4EFFA8028D057B52536551" box="[1212,1369,1885,1911]" italics="true" pageId="9" pageNumber="144">A. graecizans</emphasis>
subsp.
<emphasis id="B914EA924B4EFFA806B905A6560B65BF" box="[136,257,1920,1945]" italics="true" pageId="9" pageNumber="144">graecizans</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C604D034B4EFFA8077205A6554D65BC" authority="Benth." authorityName="Benth." box="[323,583,1920,1946]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="macrocarpus">
<emphasis id="B914EA924B4EFFA8077205A656FF65BF" box="[323,501,1920,1946]" italics="true" pageId="9" pageNumber="144">A. macrocarpus</emphasis>
Benth.
</taxonomicName>
was also given. Finally the provenance (“tropischen Africa ... und Arabiens” = tropical Africa and
<collectingCountry id="F37776104B4EFFA807B605845514659A" box="[391,542,1954,1980]" name="Saudi Arabia" pageId="9" pageNumber="144">Saudi Arabia</collectingCountry>
) and
<specimenCount id="9D66FD094B4EFFA8045505845416659A" box="[612,796,1954,1980]" pageId="9" pageNumber="144" type="generic">three specimens</specimenCount>
(“
<collectingCountry id="F37776104B4EFFA80509058454B7659A" box="[824,957,1954,1980]" name="Ethiopia" pageId="9" pageNumber="144">Abessinien</collectingCountry>
: Massaua: Hildebrandt n. 716!; ostafrican. Grabenrand, 1904, Merken! ... “Chedrasch et Chedolia, Ehrenberg!”) were reported. According to the Art. 9.6 of ICN, these three citations are
<typeStatus id="54DB88224B4EFFA807A705CE56F16A24" box="[406,507,2024,2050]" pageId="9" pageNumber="144" type="syntype">syntypes</typeStatus>
, original material for the name
<taxonomicName id="4C604D034B4EFFA8056805CE53346A24" box="[857,1086,2024,2050]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="144" phylum="Tracheophyta" rank="species" species="sparganicephalus">
<emphasis id="B914EA924B4EFFA8056805CE53346A24" box="[857,1086,2024,2050]" italics="true" pageId="9" pageNumber="144">A. sparganicephalus</emphasis>
</taxonomicName>
(Art. 9.4 of ICN), and useful for the lectotypification purpose (Art. 9.3 of ICN).
</paragraph>
<paragraph id="8BDF36804B4DFFAB068C02B85560604E" blockId="10.[136,1452,158,689]" pageId="10" pageNumber="145">
<bibRefCitation id="EFF14B714B4DFFAB068C02B856B5629F" author="Verdcourt, B." box="[189,447,158,185]" pageId="10" pageNumber="145" pagination="252" refId="ref18758" refString="Verdcourt, B. (1967) Tropical African Plants: XXVIII. AMARANTHACEAE. Amaranthus sparganiocephalus. Kew Bulletin 21 (2): 252. https: // doi. org / 10.2307 / 4108530" type="journal article" year="1967">Verdcourt (1967: 252)</bibRefCitation>
listed
<specimenCount id="9D66FD094B4DFFAB042002B855A4629E" box="[529,686,158,184]" pageId="10" pageNumber="145" type="generic">16 specimens</specimenCount>
of
<taxonomicName id="4C604D034B4DFFAB04EA02B8533F629E" box="[731,1077,158,184]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="145" phylum="Tracheophyta" rank="species" species="sparganicephalus">
<emphasis id="B914EA924B4DFFAB04EA02B8533F629E" box="[731,1077,158,184]" italics="true" pageId="10" pageNumber="145">Amaranthus sparganicephalus</emphasis>
</taxonomicName>
adding “
<typeStatus id="54DB88224B4DFFAB029302B853F7629E" box="[1186,1277,158,184]" pageId="10" pageNumber="145" type="syntype">syntype</typeStatus>
” after three of them, i.e. “ARABIA. Chedrasch and Chedolia, Ehrennberg (B)”, “
<collectingCountry id="F37776104B4DFFAB054E02E454F162FA" box="[895,1019,194,220]" name="Eritrea" pageId="10" pageNumber="145">ERITREA</collectingCountry>
. Massawa, Hildebrandt 716 (B)”, and “
<collectingCountry id="F37776104B4DFFAB06A302C0562D6326" box="[146,295,230,256]" name="Tanzania" pageId="10" pageNumber="145">TANZANIA</collectingCountry>
. Grabenland, 1904, Merker (B)”. These
<specimenCount id="9D66FD094B4DFFAB04D002C0549E6326" box="[737,916,230,256]" pageId="10" pageNumber="145" type="generic">three specimens</specimenCount>
was also cited (as “Types”) by
<bibRefCitation id="EFF14B714B4DFFAB02DB02C057A76302" author="Townsend, C. C." pageId="10" pageNumber="145" pagination="1 - 136" refId="ref18438" refString="Townsend, C. C. (1985) Amaranthaceae. In: Polhill, R. M. (Ed.) Flora of Tropical East Africa. A. A. Balkema, Rotterdam, pp. 1 - 136." type="book chapter" year="1985">Townsend (1985: 32)</bibRefCitation>
in his treatment of
<taxonomicName id="4C604D034B4DFFAB07A6032C554F6302" box="[407,581,266,292]" class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="145" phylum="Tracheophyta" rank="family">Amaranthaceae</taxonomicName>
for the
<emphasis id="B914EA924B4DFFAB0496032C54F16302" box="[679,1019,266,292]" italics="true" pageId="10" pageNumber="145">Flora of Tropical East Africa</emphasis>
. According to
<emphasis id="B914EA924B4DFFAB029A032C52466302" box="[1195,1356,266,292]" italics="true" pageId="10" pageNumber="145">Shenzen Code</emphasis>
, neither
<bibRefCitation id="EFF14B714B4DFFAB06B903085688636F" author="Verdcourt, B." box="[136,386,302,329]" pageId="10" pageNumber="145" pagination="252" refId="ref18758" refString="Verdcourt, B. (1967) Tropical African Plants: XXVIII. AMARANTHACEAE. Amaranthus sparganiocephalus. Kew Bulletin 21 (2): 252. https: // doi. org / 10.2307 / 4108530" type="journal article" year="1967">Verdcourt (1967: 252)</bibRefCitation>
nor
<bibRefCitation id="EFF14B714B4DFFAB0787030855AC636E" author="Townsend, C. C." box="[438,678,302,329]" pageId="10" pageNumber="145" pagination="1 - 136" refId="ref18438" refString="Townsend, C. C. (1985) Amaranthaceae. In: Polhill, R. M. (Ed.) Flora of Tropical East Africa. A. A. Balkema, Rotterdam, pp. 1 - 136." type="book chapter" year="1985">Townsend (1985: 32)</bibRefCitation>
proposed a correct typification, since they just re-listed the
<typeStatus id="54DB88224B4DFFAB0378030952A1636F" box="[1353,1451,303,329]" pageId="10" pageNumber="145" type="syntype">syntypes</typeStatus>
which was originally reported by
<bibRefCitation id="EFF14B714B4DFFAB043B037455F4634A" author="Thellung, A." box="[522,766,338,364]" pageId="10" pageNumber="145" pagination="225 - 356" refId="ref18248" refString="Thellung, A. (1914) Amaranthus L. In: Ascherson, P. &amp; Graebner, P. (Eds.) Synopsis der Mitteleuropaischen Flora, vol. 5. Verlag Von Gebruder Borntraeger. Leipzig, Germany, pp. 225 - 356." type="book chapter" year="1914">Thellung (1914: 312)</bibRefCitation>
in the protologue. As a consequence, a lectoptypification is necessary. Note moreover that both Verdecourt and Townsend reported after the
<typeStatus id="54DB88224B4DFFAB02350351536C63B7" box="[1028,1126,375,401]" pageId="10" pageNumber="145" type="syntype">syntypes</typeStatus>
the symbol “†” which would indicate that the specimen was are no longer exsisting. R. Vogt (pers. comm.) informed one of us (DI) that no original material for
<taxonomicName id="4C604D034B4DFFAB0726039956F463FE" box="[279,510,446,473]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="145" phylum="Tracheophyta" rank="species" species="sparganicephalus">
<emphasis id="B914EA924B4DFFAB0726039956F463FE" box="[279,510,446,473]" italics="true" pageId="10" pageNumber="145">A. sparganicephalus</emphasis>
</taxonomicName>
is preserved at B being probably lost/destroyed during the II World War. Lacking material useful for the lectotypification purpose (Arts. 9.3 and 9.4 of ICN), a neotyipfication is required under the Art. 9.8 of ICN. We here propose to designate, as
<typeStatus id="54DB88224B4DFFAB044A002155DE6007" box="[635,724,519,545]" pageId="10" pageNumber="145" type="neotype">neotype</typeStatus>
of the name
<taxonomicName id="4C604D034B4DFFAB05520021534D6006" box="[867,1095,518,545]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="145" phylum="Tracheophyta" rank="species" species="sparganicephalus">
<emphasis id="B914EA924B4DFFAB05520021534D6006" box="[867,1095,518,545]" italics="true" pageId="10" pageNumber="145">A. sparganicephalus</emphasis>
</taxonomicName>
, a well preserved specimen at E (barcode E00687024) collected in Oman in 1985 which matches the Thellungs description and the current application of the name (see e.g.,
<bibRefCitation id="EFF14B714B4DFFAB074C00685554604E" author="Townsend, C. C." box="[381,606,590,617]" pageId="10" pageNumber="145" pagination="1 - 136" refId="ref18438" refString="Townsend, C. C. (1985) Amaranthaceae. In: Polhill, R. M. (Ed.) Flora of Tropical East Africa. A. A. Balkema, Rotterdam, pp. 1 - 136." type="book chapter" year="1985">Townsend 1985: 32</bibRefCitation>
).
</paragraph>
<paragraph id="8BDF36804B4DFFAB068C005454526096" blockId="10.[136,1452,158,689]" pageId="10" pageNumber="145">
<emphasis id="B914EA924B4DFFAB068C005456FA60AA" bold="true" box="[189,496,626,652]" pageId="10" pageNumber="145">Specimina visa selecta:—</emphasis>
<materialsCitation id="3B083CDD4B4DFFAB07C00054545E6096" ID-GBIF-Occurrence="3999584306" collectingDate="1995-04-07" country="Saudi Arabia" location="Raidah Village" pageId="10" pageNumber="145" specimenCount="1">
<collectingCountry id="F37776104B4DFFAB07C0005455B760AA" box="[497,701,626,652]" name="Saudi Arabia" pageId="10" pageNumber="145">SAUDI ARABIA</collectingCountry>
.
<location id="8EBF605B4B4DFFAB04FB0054547E60AA" LSID="urn:lsid:plazi:treatment:03C987964B4EFFAB06B902B854526096:8EBF605B4B4DFFAB04FB0054547E60AA" box="[714,884,626,652]" country="Saudi Arabia" name="Raidah Village" pageId="10" pageNumber="145">Raidah Village</location>
near base of scarp.
<locationDeviation id="9A6D2FA24B4DFFAB026E0054525460AA" box="[1119,1374,626,653]" location="Abha" pageId="10" pageNumber="145">
<quantity id="4C989B654B4DFFAB026E005453A360AA" box="[1119,1193,626,653]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="145" unit="km" value="25.0">25 km</quantity>
NNW of Abha
</locationDeviation>
, waste ground in villane,
<date id="FFDE10404B4DFFAB076400B056FE6097" box="[341,500,662,689]" pageId="10" pageNumber="145" value="1995-04-07">
<collectingDate id="EF9AE9A84B4DFFAB076400B056FE6097" box="[341,500,662,689]" pageId="10" pageNumber="145" value="1995-04-07">07 April 1995</collectingDate>
</date>
,
<emphasis id="B914EA924B4DFFAB07CE00B055A66096" box="[511,684,662,688]" italics="true" pageId="10" pageNumber="145">Collinette 9337</emphasis>
(E00121397!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>