808 lines
143 KiB
XML
808 lines
143 KiB
XML
<document id="C6473EE71A0AB459109BC149C6B6056B" ID-CLB-Dataset="55753" ID-DOI="10.11646/zootaxa.5115.3.5" ID-GBIF-Dataset="a11ef934-b343-4214-bb58-8a8e478df7fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="6358394" ID-ZooBank="CCDE4450-C609-4640-AE94-F49F95CCCD1C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647346345120" checkinUser="plazi" docAuthor="Komai, Tomoyuki & Koreeda, Reo" docDate="2022" docId="03C9878CFFBF4B4D8BA1FCE5FEF5243E" docLanguage="en" docName="zootaxa.5115.3.5.pdf" docOrigin="Zootaxa 5115 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Richalpheus laeuadigitus Komai & Koreeda 2022, n. sp." docType="treatment" docUuid="234F1113-5EBD-40FC-9FD8-39E93A1BFF2E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="407" masterDocId="FFF0FFF4FFBE4B478B36FF98FFE12067" masterDocTitle="A new species of the infaunal alpheid shrimp genus Richalpheus Anker & Jeng 2006 (Decapoda: Caridea) from Kagoshima, Japan" masterLastPageNumber="408" masterPageNumber="397" pageNumber="398" updateTime="1701998973334" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="123A4D3AAA08DF27A75C173C0FD16DB2">
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<mods:title id="FC3BAC5F8A8532338CDCC05646E699E3">A new species of the infaunal alpheid shrimp genus Richalpheus Anker & Jeng 2006 (Decapoda: Caridea) from Kagoshima, Japan</mods:title>
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<mods:name id="A7E9DE13644B64CB282C667C0C576A2E" type="personal">
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<mods:roleTerm id="2A64D28E12F166612DA53133C05F9CB8">Author</mods:roleTerm>
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<mods:namePart id="2B92680359EF39C4E500D96B9BE55AA7">Komai, Tomoyuki</mods:namePart>
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<mods:affiliation id="69647E273DE76BF7D0EE24252128333B">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba, 260 - 8682 Japan</mods:affiliation>
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<mods:namePart id="0A755A631CFAE046BD8D7F39CAFA6EDF">Koreeda, Reo</mods:namePart>
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<mods:nameIdentifier id="54192689D7DD8E2DDBA45922CAEF4E6A" type="ORCID">0000-0002-6977-3357</mods:nameIdentifier>
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<mods:affiliation id="4955B024B2BD6841EA37110115D6775A">Graduate School of Agriculture, Forestry and Fisheries, Kagoshima University, 1 - 21 - 24, Korimoto, Kagoshima, 890 - 0065 Japan k 4920583 @ kadai. jp; https: // orcid. org / 0000 - 0002 - 6977 - 3357</mods:affiliation>
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<mods:nameIdentifier id="9966053A428F7A093EB074BDC4A5BC4E" type="email">k4920583@kadai.jp</mods:nameIdentifier>
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<mods:title id="2AE51CF6D377B1DB474AE11A2FF938D5">Zootaxa</mods:title>
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<mods:part id="046A3E46B44ABC5C4BDE9CE1A55290CE">
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<mods:date id="7552FD58015A5F9851FD3B32E2008CE9">2022</mods:date>
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<mods:number id="03B5DD133E19543EE8295DAF72704A9C">2022-03-15</mods:number>
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<treatment id="03C9878CFFBF4B4D8BA1FCE5FEF5243E" ID-DOI="http://doi.org/10.5281/zenodo.6511708" ID-GBIF-Taxon="194951999" ID-Zenodo-Dep="6511708" LSID="urn:lsid:zoobank.org:act:234F1113-5EBD-40FC-9FD8-39E93A1BFF2E" httpUri="http://treatment.plazi.org/id/03C9878CFFBF4B4D8BA1FCE5FEF5243E" lastPageId="10" lastPageNumber="407" pageId="1" pageNumber="398">
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<subSubSection id="C37A6511FFBF4B468BA1FCE5FDF523FF" box="[151,532,893,920]" pageId="1" pageNumber="398" type="nomenclature">
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<paragraph id="8BDF369AFFBF4B468BA1FCE5FDF523FF" blockId="1.[151,773,893,993]" box="[151,532,893,920]" pageId="1" pageNumber="398">
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<heading id="D09781F6FFBF4B468BA1FCE5FDF523FF" bold="true" box="[151,532,893,920]" fontSize="11" level="1" pageId="1" pageNumber="398" reason="1">
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<emphasis id="B914EA88FFBF4B468BA1FCE5FDF523FF" bold="true" box="[151,532,893,920]" italics="true" pageId="1" pageNumber="398">
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<taxonomicName id="4C604D19FFBF4B468BA1FCE5FE2B23FF" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[151,458,893,920]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">Richalpheus laeuadigitus</taxonomicName>
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<taxonomicNameLabel id="A22757F3FFBF4B468AE4FCE6FDF523FF" box="[466,532,894,920]" pageId="1" pageNumber="398" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C37A6511FFBF4B468BA1FC3AFCE423DB" box="[151,773,929,956]" pageId="1" pageNumber="398" type="vernacular_names">
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<paragraph id="8BDF369AFFBF4B468BA1FC3AFCE423DB" blockId="1.[151,773,893,993]" box="[151,773,929,956]" pageId="1" pageNumber="398">[New Japanese name: Usubeni-ana-yadori-teppou-ebi]]</paragraph>
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</subSubSection>
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<subSubSection id="C37A6511FFBF4B468BA1FC5EFEA12386" box="[151,320,966,993]" pageId="1" pageNumber="398" type="description">
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<paragraph id="8BDF369AFFBF4B468BA1FC5EFEA12386" blockId="1.[151,773,893,993]" box="[151,320,966,993]" pageId="1" pageNumber="398">
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(
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<figureCitation id="135B2A1FFFBF4B468BA9FC5EFEEA2386" box="[159,267,966,993]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1929,1953]" captionStartId-1="4.[151,250,1812,1836]" captionStartId-2="5.[151,250,1943,1967]" captionStartId-3="6.[151,250,1943,1967]" captionStartId-4="7.[151,250,1715,1739]" captionTargetBox-0="[151,1435,181,1905]" captionTargetBox-1="[228,1358,190,1778]" captionTargetBox-2="[204,1389,191,1910]" captionTargetBox-3="[198,1385,197,1904]" captionTargetBox-4="[158,1426,190,1680]" captionTargetId-0="figure-17@3.[151,1436,181,1905]" captionTargetId-1="figure-30@4.[223,1365,181,1788]" captionTargetId-2="figure-17@5.[193,1395,181,1919]" captionTargetId-3="figure-30@6.[183,1404,181,1919]" captionTargetId-4="figure-17@7.[151,1436,181,1691]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Richalpheus laeuadigitus n. sp., habitus in lateral view, showing colouration in fresh condition.A, holotype, female (cl 10.5 mm), KAUM-AT 1005; B, paratype, female (cl 7.4 mm), KAUM-AT 902; C, paratype, female (cl 4.5 mm), KAUM-AT 1352." captionText-1="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." captionText-2="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." captionText-3="FIGURE 4. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, major (left) cheliped, lateral view; B, same, mesial view; C, same, merus, dorsal view; E, same, chela, flexor view; F, same, extensor view." captionText-4="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi-0="http://doi.org/10.5281/zenodo.6358396" figureDoi-1="http://doi.org/10.5281/zenodo.6358398" figureDoi-2="http://doi.org/10.5281/zenodo.6358402" figureDoi-3="http://doi.org/10.5281/zenodo.6358404" figureDoi-4="http://doi.org/10.5281/zenodo.6358406" httpUri-0="https://zenodo.org/record/6358396/files/figure.png" httpUri-1="https://zenodo.org/record/6358398/files/figure.png" httpUri-2="https://zenodo.org/record/6358402/files/figure.png" httpUri-3="https://zenodo.org/record/6358404/files/figure.png" httpUri-4="https://zenodo.org/record/6358406/files/figure.png" pageId="1" pageNumber="398">Figs. 1–5</figureCitation>
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,
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<figureCitation id="135B2A1FFFBF4B468A21FC5EFED92386" box="[279,312,966,993]" captionStart="FIGURE 6" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[165,1423,181,1851]" captionTargetId="figure-30@8.[165,1423,181,1852]" captionTargetPageId="8" captionText="FIGURE 6. A, tidal flat of Kiire, Kagoshima Prefecture, where specimens of Richalpheus laeuadigitus n. sp. were collected; B, catches by yabby pump at Kiire, Kagoshima Prefecture: 1, Richalpheus laeuadigitus n. sp.; 2, Balanoglossus misakiensis Kuwano, 1902; 3, Neotrypaea harmandi (Bouvier, 1901) or N. japonica (Ortmann, 1891); 4, Paratrypaea bouvieri (Nobili, 1904); 5, Pinnixa balanoglossana Sakai, 1934." figureDoi="http://doi.org/10.5281/zenodo.6358410" httpUri="https://zenodo.org/record/6358410/files/figure.png" pageId="1" pageNumber="398">6B</figureCitation>
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)
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</paragraph>
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<subSubSection id="C37A6511FFBF4B468BA1FB95FC90250A" pageId="1" pageNumber="398" type="materials_examined">
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<emphasis id="B914EA88FFBF4B468BA1FB95FE602440" bold="true" box="[151,385,1037,1063]" pageId="1" pageNumber="398">Material examined.</emphasis>
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<materialsCitation id="3B083CC7FFBF4B468ABFFB96FB10242B" ID-GBIF-Occurrence="3758808306" collectingDate="2020-05-09" collectionCode="R" country="Japan" elevation="2" latitude="31.372168" location="Kiire" longLatPrecision="12" longitude="130.54167" municipality="Kyushu" pageId="1" pageNumber="398" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
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<typeStatus id="54DB8838FFBF4B468ABFFB96FE14244F" box="[393,501,1038,1064]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
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. KAUM-AT-1005, female (cl
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),
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,
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<collectingRegion id="49A4F878FFBF4B468F3BFB96FAEB244F" box="[1037,1290,1038,1064]" country="Japan" name="Kagoshima" pageId="1" pageNumber="398">Kagoshima Prefecture</collectingRegion>
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,
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<collectingMunicipality id="6BBBACE0FFBF4B468E20FB96FA91244F" box="[1302,1392,1038,1064]" pageId="1" pageNumber="398">Kyushu</collectingMunicipality>
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,
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<collectingCountry id="F377760AFFBF4B468E4AFB96FF23242B" name="Japan" pageId="1" pageNumber="398">Japan</collectingCountry>
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,
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,
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<geoCoordinate id="EE54505DFFBF4B468A69FBAAFE10242A" box="[351,497,1074,1101]" degrees="130" direction="east" minutes="32.50" orientation="longitude" pageId="1" pageNumber="398" precision="9" value="130.54167">130°32.50’E</geoCoordinate>
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,
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<quantity id="4C989B7FFFBF4B468ACAFBAAFDDD242B" box="[508,572,1074,1100]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="398" unit="m" value="0.2">
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<elevation id="004DD1A9FFBF4B468ACAFBAAFDDD242B" box="[508,572,1074,1100]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="398" unit="m" value="0.2">0.2 m</elevation>
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</quantity>
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deep at low tide,
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<date id="FFDE105AFFBF4B468835FBAAFC6C242B" box="[771,909,1074,1100]" pageId="1" pageNumber="398" value="2020-05-09">
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<collectingDate id="EF9AE9B2FFBF4B468835FBAAFC6C242B" box="[771,909,1074,1100]" pageId="1" pageNumber="398" value="2020-05-09">9 May 2020</collectingDate>
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</date>
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, yabby pump, coll.
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<collectionCode id="ED71AE5FFFBF4B468F5FFBAAFB9E242B" box="[1129,1151,1074,1100]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="398">R</collectionCode>
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. Koreeda.
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</materialsCitation>
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</paragraph>
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<materialsCitation id="3B083CC7FFBF4B468BF1FBCEFB912417" ID-GBIF-Occurrence="3758808312" box="[199,1136,1110,1136]" collectingDate="2020-05-09" collectionCode="R" country="Japan" elevation="2" latitude="31.372168" location="Collection" longLatPrecision="12" longitude="130.54167" municipality="Kyushu" pageId="1" pageNumber="398" specimenCode="KAUM-AT 902" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paratype">
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<typeStatus id="54DB8838FFBF4B468BF1FBCEFED92417" box="[199,312,1110,1136]" pageId="1" pageNumber="398" type="paratype">Paratypes</typeStatus>
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.
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<location id="8EBF6041FFBF4B468A74FBCEFE562417" LSID="urn:lsid:plazi:treatment:03C9878CFFBF4B4D8BA1FCE5FEF5243E:8EBF6041FFBF4B468A74FBCEFE562417" box="[322,439,1110,1136]" name="Collection" pageId="1" pageNumber="398">Collection</location>
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||
data same as holotype:
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<specimenCode id="DBC69EE1FFBF4B4689F4FBCEFC962417" box="[706,887,1110,1136]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 902</specimenCode>
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,
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<specimenCount id="9D66FD13FFBF4B4688B4FBCEFC022417" box="[898,995,1110,1136]" pageId="1" pageNumber="398" type="female">1 female</specimenCount>
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(cl
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)
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</materialsCitation>
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;
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<materialsCitation id="3B083CC7FFBF4B468F4AFBCEFEE424F3" ID-GBIF-Occurrence="3758808308" collectingDate="2020-05-09" collectionCode="R" country="Japan" elevation="2" isTailEnumeration="true" latitude="31.372168" location="Collection" longLatPrecision="12" longitude="130.54167" municipality="Kyushu" pageId="1" pageNumber="398" specimenCode="KAUM-AT 920" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paratype">
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<specimenCode id="DBC69EE1FFBF4B468F4AFBCEFAD12417" box="[1148,1328,1110,1136]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 920</specimenCode>
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,
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<specimenCount id="9D66FD13FFBF4B468E0DFBCEFA7D2417" box="[1339,1436,1110,1136]" pageId="1" pageNumber="398" type="female">1 female</specimenCount>
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(
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<quantity id="4C989B7FFFBF4B468BA9FBE2FF1C24F3" box="[159,253,1146,1172]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="398" unit="mm" value="8.2">8.2 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B083CC7FFBF4B468A25FBE2FD2E24F3" ID-GBIF-Occurrence="3758808302" box="[275,719,1146,1172]" collectionCode="KAUM-AT" country="Japan" pageId="1" pageNumber="398" specimenCode="KAUM-AT 921" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
<specimenCode id="DBC69EE1FFBF4B468A25FBE2FE2824F3" box="[275,457,1146,1172]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 921</specimenCode>
|
||
,
|
||
<specimenCount id="9D66FD13FFBF4B468AE1FBE2FDDA24F3" box="[471,571,1146,1172]" pageId="1" pageNumber="398" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="4C989B7FFFBF4B46895DFBE2FD2924F3" box="[619,712,1146,1172]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="398" unit="mm" value="8.2">8.2 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B083CC7FFBF4B4689EBFBE2FF2E24DF" ID-GBIF-Occurrence="3758808309" accessionNumber="LC683777" collectionCode="LC" country="Japan" pageId="1" pageNumber="398" specimenCode="KAUM-AT 922" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
DNA voucher (
|
||
<accessionNumber id="9433AB79FFBF4B4688BBFBE2FBED24F2" box="[909,1036,1146,1173]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC683777" pageId="1" pageNumber="398">
|
||
<collectionCode id="ED71AE5FFFBF4B4688BBFBE2FC5224F2" box="[909,947,1146,1173]" pageId="1" pageNumber="398">LC</collectionCode>
|
||
683777
|
||
</accessionNumber>
|
||
);
|
||
<specimenCode id="DBC69EE1FFBF4B468F17FBE2FB3924F3" box="[1057,1240,1146,1172]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 922</specimenCode>
|
||
,
|
||
<specimenCount id="9D66FD13FFBF4B468FD3FBE2FAA824F3" box="[1253,1353,1146,1172]" pageId="1" pageNumber="398" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="4C989B7FFFBF4B468E4FFBE2FF2624DF" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="398" unit="mm" value="5.3">5.3 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B083CC7FFBF4B468BEDFB06FD9224DF" ID-GBIF-Occurrence="3758808303" accessionNumber="LC683777" box="[219,627,1182,1209]" collectionCode="LC" country="Japan" isTailEnumeration="true" pageId="1" pageNumber="398" specimenCode="KAUM-AT 923" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
<specimenCode id="DBC69EE1FFBF4B468BEDFB06FE6E24DF" box="[219,399,1182,1208]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 923</specimenCode>
|
||
,
|
||
<specimenCount id="9D66FD13FFBF4B468AACFB06FE0424DF" box="[410,485,1182,1208]" pageId="1" pageNumber="398" type="male">1 male</specimenCount>
|
||
(cl
|
||
<quantity id="4C989B7FFFBF4B468927FB06FD8B24DF" box="[529,618,1182,1209]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="398" unit="mm" value="5.7">5.7 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B083CC7FFBF4B46894BFB06FBC224DF" ID-GBIF-Occurrence="3758808301" box="[637,1059,1182,1208]" collectionCode="KAUM-AT" country="Japan" pageId="1" pageNumber="398" specimenCode="KAUM-AT 1202" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
<specimenCode id="DBC69EE1FFBF4B46894BFB06FCA124DF" box="[637,832,1182,1208]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 1202</specimenCode>
|
||
,
|
||
<specimenCount id="9D66FD13FFBF4B46887CFB06FC7424DF" box="[842,917,1182,1208]" pageId="1" pageNumber="398" type="male">1 male</specimenCount>
|
||
(cl
|
||
<quantity id="4C989B7FFFBF4B4688F7FB06FBFA24DF" box="[961,1051,1182,1208]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="1" pageNumber="398" unit="mm" value="6.9">6.9 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B083CC7FFBF4B468F18FB06FB7124BB" ID-GBIF-Occurrence="3758808310" collectingDate="2020-06-21" collectionCode="KAUM-AT" collectorName="Naoto Shimizu" country="Japan" latitude="31.3725" location="Kiire" longLatPrecision="12" longitude="130.543" pageId="1" pageNumber="398" specimenCode="KAUM-AT 1203" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
<location id="8EBF6041FFBF4B468F18FB06FB8B24DF" LSID="urn:lsid:plazi:treatment:03C9878CFFBF4B4D8BA1FCE5FEF5243E:8EBF6041FFBF4B468F18FB06FB8B24DF" box="[1070,1130,1182,1208]" country="Japan" latitude="31.3725" longLatPrecision="12" longitude="130.543" name="Kiire" pageId="1" pageNumber="398" stateProvince="Kagoshima">Kiire</location>
|
||
,
|
||
<geoCoordinate id="EE54505DFFBF4B468F43FB06FB1D24DE" box="[1141,1276,1182,1209]" degrees="31" direction="north" minutes="22.35" orientation="latitude" pageId="1" pageNumber="398" precision="9" value="31.3725">31°22.35’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54505DFFBF4B468E31FB06FA7924DE" box="[1287,1432,1182,1209]" degrees="130" direction="east" minutes="32.58" orientation="longitude" pageId="1" pageNumber="398" precision="9" value="130.543">130°32.58’E</geoCoordinate>
|
||
,
|
||
<date id="FFDE105AFFBF4B468BA1FB5AFED324BB" box="[151,306,1218,1244]" pageId="1" pageNumber="398" value="2020-06-21">
|
||
<collectingDate id="EF9AE9B2FFBF4B468BA1FB5AFED324BB" box="[151,306,1218,1244]" pageId="1" pageNumber="398" value="2020-06-21">21 June 2020</collectingDate>
|
||
</date>
|
||
, yabby pump, coll.
|
||
<collectorName id="2695534CFFBF4B468925FB5AFD2124BB" box="[531,704,1218,1244]" pageId="1" pageNumber="398">Naoto Shimizu</collectorName>
|
||
;
|
||
<specimenCode id="DBC69EE1FFBF4B4689FBFB5AFC7124BB" box="[717,912,1218,1244]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 1203</specimenCode>
|
||
,
|
||
<specimenCount id="9D66FD13FFBF4B4688ABFB5AFC1E24BB" box="[925,1023,1218,1244]" pageId="1" pageNumber="398" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="4C989B7FFFBF4B468F1BFB5AFB6924BB" box="[1069,1160,1218,1244]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="398" unit="mm" value="7.3">7.3 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B083CC7FFBF4B468FAAFB5AFE292567" ID-GBIF-Occurrence="3758808305" collectingDate="2020-06-21" collectionCode="KAUM-AT" collectorName="Naoto Shimizu" country="Japan" latitude="31.3725" location="Kiire" longLatPrecision="12" longitude="130.543" pageId="1" pageNumber="398" specimenCode="KAUM-AT 1490" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
same data.
|
||
<specimenCode id="DBC69EE1FFBF4B468E2AFB5AFF332567" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 1490</specimenCode>
|
||
,
|
||
<specimenCount id="9D66FD13FFBF4B468BEAFB7EFEDC2567" box="[220,317,1254,1280]" pageId="1" pageNumber="398" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="4C989B7FFFBF4B468A51FB7EFE212567" box="[359,448,1254,1280]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="1" pageNumber="398" unit="mm" value="6.9">6.9 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B083CC7FFBF4B468AE4FB7EFAEC2543" ID-GBIF-Occurrence="3758808311" collectingDate="2021-08-22" collectionCode="KAUM-AT" collectorName="Naoto Shimizu." country="Japan" county="Matsuzaki River" latitude="31.438667" location="Tarumizu" longLatPrecision="12" longitude="130.73517" municipality="Shinjo" pageId="1" pageNumber="398" specimenCode="KAUM-AT 1352" specimenCount="1" specimenCount-female="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
river mouth of
|
||
<collectingCounty id="62BE4E16FFBF4B46894EFB7EFCD22567" box="[632,819,1254,1280]" pageId="1" pageNumber="398">Matsuzaki River</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BBBACE0FFBF4B468808FB7EFC6B2567" box="[830,906,1254,1280]" pageId="1" pageNumber="398">Shinjo</collectingMunicipality>
|
||
,
|
||
<location id="8EBF6041FFBF4B4688A5FB7EFBE02567" LSID="urn:lsid:plazi:treatment:03C9878CFFBF4B4D8BA1FCE5FEF5243E:8EBF6041FFBF4B4688A5FB7EFBE02567" box="[915,1025,1254,1280]" country="Japan" county="Matsuzaki River" latitude="31.438667" longLatPrecision="12" longitude="130.73517" municipality="Shinjo" name="Tarumizu" pageId="1" pageNumber="398" stateProvince="Kagoshima">Tarumizu</location>
|
||
,
|
||
<collectingRegion id="49A4F878FFBF4B468F3DFB7EFAE62567" box="[1035,1287,1254,1280]" country="Japan" name="Kagoshima" pageId="1" pageNumber="398">Kagoshima Prefecture</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE54505DFFBF4B468E27FB7EFA762567" box="[1297,1431,1254,1280]" degrees="31" direction="north" minutes="26.32" orientation="latitude" pageId="1" pageNumber="398" precision="9" value="31.438667">31°26.32’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54505DFFBF4B468BA1FA92FEC92543" box="[151,296,1290,1316]" degrees="130" direction="east" minutes="44.11" orientation="longitude" pageId="1" pageNumber="398" precision="9" value="130.73517">130°44.11’E</geoCoordinate>
|
||
, tidal flat,
|
||
<date id="FFDE105AFFBF4B468A96FA92FDB42543" box="[416,597,1290,1316]" pageId="1" pageNumber="398" value="2021-08-22">
|
||
<collectingDate id="EF9AE9B2FFBF4B468A96FA92FDB42543" box="[416,597,1290,1316]" pageId="1" pageNumber="398" value="2021-08-22">22 August 2021</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="2695534CFFBF4B4689AEFA92FCA82543" box="[664,841,1290,1316]" pageId="1" pageNumber="398">Naoto Shimizu.</collectorName>
|
||
<specimenCode id="DBC69EE1FFBF4B468879FA92FBF32542" box="[847,1042,1290,1317]" collectionCode="KAUM-AT" pageId="1" pageNumber="398">KAUM-AT 1352</specimenCode>
|
||
,
|
||
<specimenCount id="9D66FD13FFBF4B468F28FA92FB9E2543" box="[1054,1151,1290,1316]" pageId="1" pageNumber="398" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="4C989B7FFFBF4B468F9DFA92FAE42543" box="[1195,1285,1290,1317]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="398" unit="mm" value="4.5">4.5 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B083CC7FFBF4B468E2FFA92FE25250B" ID-GBIF-Occurrence="3758808307" collectingDate="2021-04-01" collectionCode="R" country="Japan" county="Ogawa River" latitude="31.218334" location="Kimotsuki" longLatPrecision="12" longitude="130.757" municipality="Minamiohsumi" pageId="1" pageNumber="398" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
river mouth of
|
||
<collectingCounty id="62BE4E16FFBF4B468B81FAB6FEB1252F" box="[183,336,1326,1352]" pageId="1" pageNumber="398">Ogawa River</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BBBACE0FFBF4B468A6BFAB6FDEA252F" box="[349,523,1326,1352]" pageId="1" pageNumber="398">Minamiohsumi</collectingMunicipality>
|
||
,
|
||
<location id="8EBF6041FFBF4B46892EFAB6FD70252F" LSID="urn:lsid:plazi:treatment:03C9878CFFBF4B4D8BA1FCE5FEF5243E:8EBF6041FFBF4B46892EFAB6FD70252F" box="[536,657,1326,1352]" country="Japan" county="Ogawa River" latitude="31.218334" longLatPrecision="12" longitude="130.757" municipality="Minamiohsumi" name="Kimotsuki" pageId="1" pageNumber="398" stateProvince="Kagoshima">Kimotsuki</location>
|
||
,
|
||
<collectingRegion id="49A4F878FFBF4B4689A8FAB6FC7D252F" box="[670,924,1326,1352]" country="Japan" name="Kagoshima" pageId="1" pageNumber="398">Kagoshima Prefecture</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE54505DFFBF4B46889FFAB6FBD1252F" box="[937,1072,1326,1352]" degrees="31" direction="north" minutes="13.10" orientation="latitude" pageId="1" pageNumber="398" precision="9" value="31.218334">31°13.10’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54505DFFBF4B468F0BFAB6FB2F252E" box="[1085,1230,1326,1353]" degrees="130" direction="east" minutes="45.42" orientation="longitude" pageId="1" pageNumber="398" precision="9" value="130.757">130°45.42’E</geoCoordinate>
|
||
, tidal flat,
|
||
<date id="FFDE105AFFBF4B468E7AFAB6FF33250B" pageId="1" pageNumber="398" value="2021-04-01">
|
||
<collectingDate id="EF9AE9B2FFBF4B468E7AFAB6FF33250B" pageId="1" pageNumber="398" value="2021-04-01">1 April 2021</collectingDate>
|
||
</date>
|
||
, yabby pump, coll.
|
||
<collectionCode id="ED71AE5FFFBF4B468A98FACAFE25250B" box="[430,452,1362,1388]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="398">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B083CC7FFBF4B468AF9FACAFC8C250A" ID-GBIF-Occurrence="3758808304" accessionNumber="LC683778" box="[463,877,1362,1389]" collectionCode="LC" country="Japan" location="Koreeda" pageId="1" pageNumber="398" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
|
||
<location id="8EBF6041FFBF4B468AF9FACAFDD0250B" LSID="urn:lsid:plazi:treatment:03C9878CFFBF4B4D8BA1FCE5FEF5243E:8EBF6041FFBF4B468AF9FACAFDD0250B" box="[463,561,1362,1388]" country="Japan" name="Koreeda" pageId="1" pageNumber="398" stateProvince="Kagoshima">Koreeda</location>
|
||
, DNA voucher (
|
||
<accessionNumber id="9433AB79FFBF4B4689D1FACAFC87250A" box="[743,870,1362,1389]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC683778" pageId="1" pageNumber="398">
|
||
<collectionCode id="ED71AE5FFFBF4B4689D1FACAFCEC250A" box="[743,781,1362,1389]" pageId="1" pageNumber="398">LC</collectionCode>
|
||
683778
|
||
</accessionNumber>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A6511FFBF4B468BF1FAEDFB3D2647" pageId="1" pageNumber="398" type="diagnosis">
|
||
<paragraph id="8BDF369AFFBF4B468BF1FAEDFB3D2647" blockId="1.[151,1437,1037,2036]" pageId="1" pageNumber="398">
|
||
<emphasis id="B914EA88FFBF4B468BF1FAEDFEA325E8" bold="true" box="[199,322,1397,1423]" pageId="1" pageNumber="398">Diagnosis.</emphasis>
|
||
Telson (
|
||
<figureCitation id="135B2A1FFFBF4B468A90FAEEFDE125F7" box="[422,512,1398,1424]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="1" pageNumber="398">Fig. 2D</figureCitation>
|
||
) 1.9–2.0 times as long as wide. Antennular peduncle (
|
||
<figureCitation id="135B2A1FFFBF4B468F45FAEEFB0B25F7" box="[1139,1258,1398,1424]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="1" pageNumber="398">Fig. 2E, F</figureCitation>
|
||
) elongate, with article 2 more than twice as long as wide, article 1 with ventromesial carina bearing acute tooth; tip of antennular stylocerite reaching 0.6 of peduncular article 1. Major chela (4A–D; 5B) with dactylus relatively stout, with bulgelike ridge on occlusal margin; pollex with longitudinal groove in proximal 0.7 of occlusal margin. Posterior flap of uropodal exopod (
|
||
<figureCitation id="135B2A1FFFBF4B468A55F99EFE532647" box="[355,434,1542,1568]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="1" pageNumber="398">Fig. 2J</figureCitation>
|
||
) less developed, with distolateral margin truncate or shallowly concave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A6511FFBF4B4E8BF1F9B1FCE22476" lastPageId="9" lastPageNumber="406" pageId="1" pageNumber="398" type="description">
|
||
<paragraph id="8BDF369AFFBF4B468BF1F9B1FB672623" blockId="1.[151,1437,1037,2036]" box="[199,1158,1577,1604]" pageId="1" pageNumber="398">
|
||
<emphasis id="B914EA88FFBF4B468BF1F9B1FEB82624" bold="true" box="[199,345,1577,1603]" pageId="1" pageNumber="398">Description.</emphasis>
|
||
<emphasis id="B914EA88FFBF4B468A56F9B2FDF92623" box="[352,536,1578,1604]" italics="true" pageId="1" pageNumber="398">
|
||
<typeStatus id="54DB8838FFBF4B468A56F9B2FE262623" box="[352,455,1578,1604]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
|
||
female
|
||
</emphasis>
|
||
. Body (
|
||
<figureCitation id="135B2A1FFFBF4B468944F9B2FD212623" box="[626,704,1578,1604]" captionStart="FIGURE 1" captionStartId="3.[151,250,1929,1953]" captionTargetBox="[151,1435,181,1905]" captionTargetId="figure-17@3.[151,1436,181,1905]" captionTargetPageId="3" captionText="FIGURE 1. Richalpheus laeuadigitus n. sp., habitus in lateral view, showing colouration in fresh condition.A, holotype, female (cl 10.5 mm), KAUM-AT 1005; B, paratype, female (cl 7.4 mm), KAUM-AT 902; C, paratype, female (cl 4.5 mm), KAUM-AT 1352." figureDoi="http://doi.org/10.5281/zenodo.6358396" httpUri="https://zenodo.org/record/6358396/files/figure.png" pageId="1" pageNumber="398">Figs. 1</figureCitation>
|
||
;
|
||
<figureCitation id="135B2A1FFFBF4B4689F8F9B2FCF02623" box="[718,785,1578,1604]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="1" pageNumber="398">2A, B</figureCitation>
|
||
) somewhat compressed laterally.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBF4B468BF1F9D6FBCD26B3" blockId="1.[151,1437,1037,2036]" pageId="1" pageNumber="398">
|
||
Carapace (
|
||
<figureCitation id="135B2A1FFFBF4B468A09F9D6FE37260F" box="[319,470,1614,1640]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="1" pageNumber="398">Fig. 2A, E, F</figureCitation>
|
||
) smooth, becoming deep posteriorly, subequal in length to anterior 3 pleomeres combined; frontal margin broadly rounded, deflexed in lateral view, without clearly defined rostrum or orbital teeth; anterolateral margin rounded, extending as far as frontal margin; ventrolateral area of branchiostegite membranous, with rounded notch at base of pereopod 1 (cheliped); cardiac notch deep, slit-like.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBF4B468BF1F946FCBF2793" blockId="1.[151,1437,1037,2036]" pageId="1" pageNumber="398">
|
||
Pleon (
|
||
<figureCitation id="135B2A1FFFBF4B468A20F946FE6D269F" box="[278,396,1758,1784]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="1" pageNumber="398">Fig. 2B, C</figureCitation>
|
||
) elongate, pleomeres 1 and 2 combined subequal in length to pleomeres 3–5 combined, pleomere 2 longest; pleuron 2 slightly asymmetrical with posterior margin more produced than anterior margin; pleura 4 and 5 rounded posteroventrally. Pleomere 6 with clearly demarcated posteroventral plate. Telson (
|
||
<figureCitation id="135B2A1FFFBF4B468FE4F8BEFAC82727" box="[1234,1321,1830,1856]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="1" pageNumber="398">Fig. 2D</figureCitation>
|
||
) widest in proximal fifth, 1.9 times as long as wide, tapering to rounded posterior margin; dorsal surface with 2 pairs of spiniform setae inserted in deep pits, each spiniform seta slightly more than 0.1 times of telson length, first pair closer to lateral margin, located at 0.3 of telson length, second pair at some more distantly from lateral margin, located at 0.6 of telson length; posterior margin with 2 pairs of spiniform setae at posterolateral angles, lateral pair minute, mesial pair 0.2 times of telson length; anal tubercles feebly developed.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1FF0FFC3A20B2" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Eyes (
|
||
<figureCitation id="135B2A1FFFBC4B458A3BFF0FFE6120D6" box="[269,384,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="2" pageNumber="399">Fig. 2E, F</figureCitation>
|
||
) completely concealed in dorsal and lateral views, partly visible in frontal view; anteromesial margin of eyestalks rounded; corneas relatively small, but well pigmented.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1FF47FAD121CA" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Antennular peduncles (
|
||
<figureCitation id="135B2A1FFFBC4B458AFBFF47FDA1209E" box="[461,576,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="2" pageNumber="399">Fig. 2E, F</figureCitation>
|
||
) elongate, 0.6 times as long as carapace, moderately stout, slightly flattened dorsoventrally, overreaching distal margin of antennal scaphocerite by length of peduncular article 3. Article 1 dorsomesial margin bearing short row of 5 (left) or 6 (right) minute spiniform setae in proximal half; stylocerite pressed against peduncle, tip free-standing, acutely pointed, reaching beyond 0.6 of article 1; ventromesial carina with sharp tooth directed anteriorly. Article 2 more than twice as long as wide, subequal in length to visible part of article 1. Article 3 about half-length of article 2. Lateral flagellum with shorter ramus carrying tufts of aesthetascs.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1FE2FFC22225A" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Antennal peduncle (
|
||
<figureCitation id="135B2A1FFFBC4B458A9FFE2FFDFB21B6" box="[425,538,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="2" pageNumber="399">Fig. 2E, F</figureCitation>
|
||
) with basicerite moderately stout, with relatively large ventrolateral distal spine. Scaphocerite ovate with lateral margin gently convex, 0.3 times as long as carapace, 1.9 times as long as wide, bearing small blunt distolateral tooth reaching beyond distal margin of subtruncate blade. Carpocerite reaching well beyond scaphocerite, but not reaching distal end of antennular peduncle.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1FDDFFDE02206" blockId="2.[151,1437,151,1977]" box="[199,513,583,609]" pageId="2" pageNumber="399">Epistomal sclerite unarmed.</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1FDF3FCD2233A" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Mouthparts typical for genus on external observation. Maxilliped 3 endopod (
|
||
<figureCitation id="135B2A1FFFBC4B458F06FDF3FB4922E2" box="[1072,1192,619,645]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="2" pageNumber="399">Fig. 3A, B</figureCitation>
|
||
) falling slightly short of distal margin of article 2 of antennular peduncle; coxal lateral plate (
|
||
<figureCitation id="135B2A1FFFBC4B4588F2FD17FBFC22CE" box="[964,1053,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="2" pageNumber="399">Fig. 5A</figureCitation>
|
||
) developed into thick, distally tapering projection; antepenultimate article (basis-ischium-merus fused) strongly flattened, gently sinuous in dorsal view; penultimate and ultimate article much more slender than antepenultimate article, ultimate article gradually tapering to subacute apex, 1.3 times as long as penultimate article; mesial face of penultimate article sparsely setose, that of ultimate article having several transverse tracts of grooming setae; exopod (
|
||
<figureCitation id="135B2A1FFFBC4B458F11FC87FB7D235E" box="[1063,1180,799,825]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="2" pageNumber="399">Fig. 3A, B</figureCitation>
|
||
) long, but not reaching distal margin of antepenultimate article; arthrobranch large.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1FCFFFB992532" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Chelipeds (pereopods 1) greatly unequal and dissimilar. Major (left) cheliped (
|
||
<figureCitation id="135B2A1FFFBC4B458F05FCFFFB5B23E6" box="[1075,1210,871,897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[198,1385,197,1904]" captionTargetId="figure-30@6.[183,1404,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, major (left) cheliped, lateral view; B, same, mesial view; C, same, merus, dorsal view; E, same, chela, flexor view; F, same, extensor view." figureDoi="http://doi.org/10.5281/zenodo.6358404" httpUri="https://zenodo.org/record/6358404/files/figure.png" pageId="2" pageNumber="399">Figs. 4A–D</figureCitation>
|
||
;
|
||
<figureCitation id="135B2A1FFFBC4B458FF2FCFEFB0723E6" box="[1220,1254,870,897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="2" pageNumber="399">5B</figureCitation>
|
||
) elongate, chela folded against merus when not in use, overreaching distal margin of scaphocerite by full length of chela when extended. Basis with subtriangular, plate-like projection on distolateral margin. Ischium very short, unarmed, articulation to merus strongly oblique. Merus slender, widened distally in dorsal view, fairly depressed dorsoventrally, gently arcuate in lateral view; ventrolateral margin ending in rounded distal tooth; dorsal surface rounded, distally upturned; ventral surface shallowly excavate. Carpus very short, cup-shaped; dorsal surface strongly constricted near base. Chela stout, subequal in length to carapace, 3 times as long as wide. Palm 1.8 times as long as wide; dorsal face generally smooth, rounded transversely, with deep transverse constriction near articulation with dactylus; ventral face with shallow, broad longitudinal groove along midline, fitting ventral face of merus when chela carried folded; fingers somewhat twisted, gaping when closed, tips touching, but not strongly crossing. Pollex strongly deflexed at distal to base, distal part strongly hooked, terminating in blunt apex; occlusal margin of dactylus longitudinally grooved, flanked by generally convex ridges. Dactylus noticeably curved, terminating in blunt apex; upper surface carinate along extensor margin; occlusal margin with bulge-like ridge along proximal 0.7, terminating distally in blunt point, fitting to groove on pollex, distal 0.3 of occlusal margin unarmed.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1FAC7FED72612" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Minor (right) cheliped (
|
||
<figureCitation id="135B2A1FFFBC4B458AEEFAC7FDDF251E" box="[472,574,1375,1401]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="2" pageNumber="399">Figs. 3C</figureCitation>
|
||
;
|
||
<figureCitation id="135B2A1FFFBC4B45897AFAC6FD72251E" box="[588,659,1374,1401]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="2" pageNumber="399">5C, D</figureCitation>
|
||
) slender, chela folded against merus when not in use, overreaching distal margin of scaphocerite by length of dactylus. Ischium unarmed, short, articulation to merus strongly oblique. Merus dorsoventrally flattened, only slightly arcuate, unarmed. Carpus very short, cup-shaped. Chela slender, 0.6 times as long as carapace, slightly longer than merus; palm smooth, ventral (flexor) surface somewhat inflated; fingers about 1.6 times longer than palm, slender, tips crossing. Pollex gently bowed, distal part strongly hooked, terminating in acute tip; occlusal margin with row of minute, barb-like teeth in proximal half, unarmed in distal half. Dactylus gently curved, terminating in acute tip; occlusal margin armed with row of minute, barb-like teeth also in proximal half.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1F9E7FBF62762" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Pereopod 2 (
|
||
<figureCitation id="135B2A1FFFBC4B458A6DF9E7FE5426FE" box="[347,437,1663,1689]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="2" pageNumber="399">Fig. 3D</figureCitation>
|
||
) overreaching distal end of antennal carpocerite by length of chela. Ischium about 0.7 length of merus, unarmed. Merus subequal on length to carpus. Carpus divided into 4 segments, segment length ratio approximately equal to 4.5: 1.2: 1.0: 2.0. Chela 0.4 times as long as carpus, tapering distally, with brush-like tuft of setae on distal half of palm to pollex. Dactylus 0.7 times as long as palm.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFBC4B458BF1F897FB2127DE" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="399">
|
||
Pereopod 3 (
|
||
<figureCitation id="135B2A1FFFBC4B458A62F897FE46274E" box="[340,423,1807,1833]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="2" pageNumber="399">Fig. 3E</figureCitation>
|
||
) strongly compressed. Ischium short, unarmed. Merus with convex dorsal and ventral margins, 3.5 times as long as greatest width. Carpus short, less than half-length of merus, with prominent distoventral spiniform seta. Propodus narrowing distally, armed with 2 widely spaced, strong spiniform setae on flexor margin; flexor distal angle with 2 unequal spiniform setae (mesial seta stronger than lateral seta). Dactylus (
|
||
<figureCitation id="135B2A1FFFBC4B458E24F8E3FA8627F2" box="[1298,1383,1914,1941]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="2" pageNumber="399">Fig. 5F</figureCitation>
|
||
) 0.5 length of propodus, slightly curved mesially, subspatulate with subacute tip; flexor surface flat.
|
||
</paragraph>
|
||
<caption id="DF1F6612FFBD4B448BA1F811FF30278E" ID-DOI="http://doi.org/10.5281/zenodo.6358396" ID-Zenodo-Dep="6358396" httpUri="https://zenodo.org/record/6358396/files/figure.png" pageId="3" pageNumber="400" startId="3.[151,250,1929,1953]" targetBox="[151,1435,181,1905]" targetPageId="3">
|
||
<paragraph id="8BDF369AFFBD4B448BA1F811FF30278E" blockId="3.[151,1436,1929,2025]" pageId="3" pageNumber="400">
|
||
<emphasis id="B914EA88FFBD4B448BA1F811FEF327C6" bold="true" box="[151,274,1929,1953]" pageId="3" pageNumber="400">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C604D19FFBD4B448A21F811FDF627C6" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[279,535,1929,1953]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFBD4B448A21F811FDF627C6" box="[279,535,1929,1953]" italics="true" pageId="3" pageNumber="400">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFBD4B44892AF811FDB527C6" bold="true" box="[540,596,1929,1953]" pageId="3" pageNumber="400">
|
||
<taxonomicNameLabel id="A22757F3FFBD4B44892AF811FDB527C6" box="[540,596,1929,1953]" pageId="3" pageNumber="400" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus in lateral view, showing colouration in fresh condition. A, holotype, female (cl 10.5 mm), KAUM-AT 1005; B, paratype, female (cl 7.4 mm), KAUM-AT 902; C, paratype, female (cl 4.5 mm), KAUM-AT 1352.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F6612FFBA4B438BA1F88CFC352787" ID-DOI="http://doi.org/10.5281/zenodo.6358398" ID-Zenodo-Dep="6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="4" pageNumber="401" startId="4.[151,250,1812,1836]" targetBox="[228,1358,190,1778]" targetPageId="4">
|
||
<paragraph id="8BDF369AFFBA4B438BA1F88CFC352787" blockId="4.[151,1437,1812,2016]" pageId="4" pageNumber="401">
|
||
<emphasis id="B914EA88FFBA4B438BA1F88CFEF5274B" bold="true" box="[151,276,1812,1836]" pageId="4" pageNumber="401">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C604D19FFBA4B438A2AF88CFDFF274B" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[284,542,1812,1836]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFBA4B438A2AF88CFDFF274B" box="[284,542,1812,1836]" italics="true" pageId="4" pageNumber="401">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFBA4B438910F88CFD80274B" bold="true" box="[550,609,1812,1836]" pageId="4" pageNumber="401">
|
||
<taxonomicNameLabel id="A22757F3FFBA4B438910F88CFD80274B" box="[550,609,1812,1836]" pageId="4" pageNumber="401" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F6612FFBB4B428BA1F80FFEA7279F" ID-DOI="http://doi.org/10.5281/zenodo.6358402" ID-Zenodo-Dep="6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="5" pageNumber="402" startId="5.[151,250,1943,1967]" targetBox="[204,1389,191,1910]" targetPageId="5">
|
||
<paragraph id="8BDF369AFFBB4B428BA1F80FFEA7279F" blockId="5.[151,1436,1943,2040]" pageId="5" pageNumber="402">
|
||
<emphasis id="B914EA88FFBB4B428BA1F80FFEF327C8" bold="true" box="[151,274,1943,1967]" pageId="5" pageNumber="402">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C604D19FFBB4B428A21F800FDF627D7" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[279,535,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="402" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFBB4B428A21F800FDF627D7" box="[279,535,1944,1968]" italics="true" pageId="5" pageNumber="402">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFBB4B42892AF800FDB527D7" bold="true" box="[540,596,1944,1968]" pageId="5" pageNumber="402">
|
||
<taxonomicNameLabel id="A22757F3FFBB4B42892AF800FDB527D7" box="[540,596,1944,1968]" pageId="5" pageNumber="402" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 10.5 mm), KAUM-AT 1005. A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F6612FFB84B418BA1F80FFB1527B3" ID-DOI="http://doi.org/10.5281/zenodo.6358404" ID-Zenodo-Dep="6358404" httpUri="https://zenodo.org/record/6358404/files/figure.png" pageId="6" pageNumber="403" startId="6.[151,250,1943,1967]" targetBox="[198,1385,197,1904]" targetPageId="6">
|
||
<paragraph id="8BDF369AFFB84B418BA1F80FFB1527B3" blockId="6.[151,1436,1943,2004]" pageId="6" pageNumber="403">
|
||
<emphasis id="B914EA88FFB84B418BA1F80FFEF227C8" bold="true" box="[151,275,1943,1967]" pageId="6" pageNumber="403">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C604D19FFB84B418A2CF800FDFB27D7" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[282,538,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB84B418A2CF800FDFB27D7" box="[282,538,1944,1968]" italics="true" pageId="6" pageNumber="403">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB84B418917F800FDBB27D7" bold="true" box="[545,602,1944,1968]" pageId="6" pageNumber="403">
|
||
<taxonomicNameLabel id="A22757F3FFB84B418917F800FDBB27D7" box="[545,602,1944,1968]" pageId="6" pageNumber="403" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 10.5 mm), KAUM-AT 1005. A, major (left) cheliped, lateral view; B, same, mesial view; C, same, merus, dorsal view; E, same, chela, flexor view; F, same, extensor view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F6612FFB94B408BA1F92BFB0227A0" ID-DOI="http://doi.org/10.5281/zenodo.6358406" ID-Zenodo-Dep="6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="7" pageNumber="404" startId="7.[151,250,1715,1739]" targetBox="[158,1426,190,1680]" targetPageId="7">
|
||
<paragraph id="8BDF369AFFB94B408BA1F92BFB0227A0" blockId="7.[151,1437,1715,1991]" pageId="7" pageNumber="404">
|
||
<emphasis id="B914EA88FFB94B408BA1F92BFEF326AC" bold="true" box="[151,274,1715,1739]" pageId="7" pageNumber="404">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C604D19FFB94B408A21F92BFDF626AC" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[279,535,1715,1739]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB94B408A21F92BFDF626AC" box="[279,535,1715,1739]" italics="true" pageId="7" pageNumber="404">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB94B40892BF92BFDB426AC" bold="true" box="[541,597,1715,1739]" pageId="7" pageNumber="404">
|
||
<taxonomicNameLabel id="A22757F3FFB94B40892BF92BFDB426AC" box="[541,597,1715,1739]" pageId="7" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923. A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted). Abbreviations: CLP = coxal lateral plate; EXP = exopod.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F6612FFB64B4F8BA1F8CBFDCA27BF" ID-DOI="http://doi.org/10.5281/zenodo.6358410" ID-Zenodo-Dep="6358410" httpUri="https://zenodo.org/record/6358410/files/figure.png" pageId="8" pageNumber="405" startId="8.[151,250,1875,1899]" targetBox="[165,1423,181,1851]" targetPageId="8">
|
||
<paragraph id="8BDF369AFFB64B4F8BA1F8CBFDCA27BF" blockId="8.[151,1437,1875,2008]" pageId="8" pageNumber="405">
|
||
<emphasis id="B914EA88FFB64B4F8BA1F8CBFEF3270B" bold="true" box="[151,274,1875,1900]" pageId="8" pageNumber="405">FIGURE 6.</emphasis>
|
||
A, tidal flat of Kiire, Kagoshima Prefecture, where specimens of
|
||
<taxonomicName id="4C604D19FFB64B4F8895F8CCFB42270B" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[931,1187,1876,1900]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB64B4F8895F8CCFB42270B" box="[931,1187,1876,1900]" italics="true" pageId="8" pageNumber="405">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB64B4F8F9FF8CCFB00270B" bold="true" box="[1193,1249,1876,1900]" pageId="8" pageNumber="405">
|
||
<taxonomicNameLabel id="A22757F3FFB64B4F8F9FF8CCFB00270B" box="[1193,1249,1876,1900]" pageId="8" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were collected; B, catches by yabby pump at Kiire, Kagoshima Prefecture: 1,
|
||
<taxonomicName id="4C604D19FFB64B4F89C2F8E0FC1427F7" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[756,1013,1912,1936]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB64B4F89C2F8E0FC1427F7" box="[756,1013,1912,1936]" italics="true" pageId="8" pageNumber="405">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB64B4F88CAF8E0FBD727F7" bold="true" box="[1020,1078,1912,1936]" pageId="8" pageNumber="405">
|
||
<taxonomicNameLabel id="A22757F3FFB64B4F88CAF8E0FBD727F7" box="[1020,1078,1912,1936]" pageId="8" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 2,
|
||
<taxonomicName id="4C604D19FFB64B4F8F68F8E0FEF127D3" authority="Kuwano, 1902" authorityName="Kuwano" authorityYear="1902" class="Enteropneusta" family="Ptychoderidae" genus="Balanoglossus" kingdom="Animalia" pageId="8" pageNumber="405" phylum="Hemichordata" rank="species" species="misakiensis">
|
||
<emphasis id="B914EA88FFB64B4F8F68F8E0FA8F27F7" box="[1118,1390,1912,1936]" italics="true" pageId="8" pageNumber="405">Balanoglossus misakiensis</emphasis>
|
||
<bibRefCitation id="EFF14B6BFFB64B4F8E43F8E0FEF127D3" author="Kuwano, H." pageId="8" pageNumber="405" pagination="77 - 84" refId="ref5593" refString="Kuwano, H. (1902) On a new Enteropneust from Misaki, Balanoglossus misakiensis n. sp. Annotationes Zoologicae Japonenses, 4, 77 - 84." type="journal article" year="1902">Kuwano, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
; 3,
|
||
<taxonomicName id="4C604D19FFB64B4F8A05F804FD5627D3" authority="(Bouvier, 1901)" baseAuthorityName="Bouvier" baseAuthorityYear="1901" box="[307,695,1948,1972]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="harmandi">
|
||
<emphasis id="B914EA88FFB64B4F8A05F804FDF327D3" box="[307,530,1948,1972]" italics="true" pageId="8" pageNumber="405">Neotrypaea harmandi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF14B6BFFB64B4F8928F804FD5027D3" author="Bouvier, E. L." box="[542,689,1948,1972]" pageId="8" pageNumber="405" pagination="332 - 334" refId="ref5268" refString="Bouvier, E. L. (1901) Sur quelques crustaces du Japon, offerts au museum par M. le Dr. Harmand. Bulletin du Museum National d'Histoire Naturelle, Paris, 7, 332 - 334. https: // doi. org / 10.5962 / bhl. part. 1625" type="journal article" year="1901">Bouvier, 1901</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C604D19FFB64B4F89EFF804FBE027D3" authority="(Ortmann, 1891)" baseAuthorityName="Ortmann" baseAuthorityYear="1891" box="[729,1025,1948,1972]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="B914EA88FFB64B4F89EFF804FCAE27D4" box="[729,847,1948,1971]" italics="true" pageId="8" pageNumber="405">N. japonica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF14B6BFFB64B4F886DF804FC1827D3" author="Ortmann, A. E." box="[859,1017,1948,1972]" pageId="8" pageNumber="405" pagination="1 - 58" refId="ref5710" refString="Ortmann, A. E. (1891) Die Decapoden-Krebse des Strassburger Museums mit besonderer Berucksichtigung der von Herrn Dr. Doderlein dei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen III. Theil. Die Abtheilungen der Reptantia Boas: Homaridae, Loricata und Thalassinidea. Zoologische Jahrbucher. Abteilung fur Systematik, 6, 1 - 58. https: // doi. org / 10.5962 / bhl. part. 26455" type="journal article" year="1891">Ortmann, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; 4,
|
||
<taxonomicName id="4C604D19FFB64B4F8F12F804FA7827D3" authority="(Nobili, 1904)" baseAuthorityName="Nobili" baseAuthorityYear="1904" box="[1060,1433,1948,1972]" class="Malacostraca" family="Callianassidae" genus="Paratrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="bouvieri">
|
||
<emphasis id="B914EA88FFB64B4F8F12F804FB1F27D3" box="[1060,1278,1948,1972]" italics="true" pageId="8" pageNumber="405">Paratrypaea bouvieri</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF14B6BFFB64B4F8E3AF804FA7027D3" author="Nobili, G." box="[1292,1425,1948,1972]" pageId="8" pageNumber="405" pagination="230 - 238" refId="ref5673" refString="Nobili, G. (1904) Diagnoses preliminaires de vingt-huit especes nouvelles de Stomatopodes et Decapodes Macroures de la Mer Rouge. Bulletin du Museum d'Histoire Naturelle de Paris, 10, 230 - 238." type="journal article" year="1904">Nobili, 1904</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; 5,
|
||
<taxonomicName id="4C604D19FFB64B4F8B87F858FDC627BF" authority="Sakai, 1934" authorityName="Sakai" authorityYear="1934" box="[177,551,1984,2008]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="balanoglossana">
|
||
<emphasis id="B914EA88FFB64B4F8B87F858FE4727BF" box="[177,422,1984,2008]" italics="true" pageId="8" pageNumber="405">Pinnixa balanoglossana</emphasis>
|
||
<bibRefCitation id="EFF14B6BFFB64B4F8A9AF858FDC627BF" author="Sakai, T." box="[428,551,1984,2008]" pageId="8" pageNumber="405" pagination="37 - 43" refId="ref5884" refString="Sakai, T. (1934) Species of the genus Pinnixa (Pinnotherid crab) found in the Far East. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2, 37 - 43." type="journal article" year="1934">Sakai, 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FF0FFBCF20D6" blockId="9.[151,1437,150,2013]" box="[199,1070,150,177]" pageId="9" pageNumber="406">
|
||
Pereopod 4 (
|
||
<figureCitation id="135B2A1FFFB74B4E8A63FF0FFE4B20D6" box="[341,426,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="9" pageNumber="406">Fig. 3F</figureCitation>
|
||
;
|
||
<figureCitation id="135B2A1FFFB74B4E8A80FF0EFE3820D6" box="[438,473,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">5G</figureCitation>
|
||
) generally similar to pereopod 3, but slightly shorter.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FF23FB99217A" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">
|
||
Pereopod 5 (
|
||
<figureCitation id="135B2A1FFFB74B4E8A61FF23FE4E20B2" box="[343,431,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="9" pageNumber="406">Fig. 3G</figureCitation>
|
||
) shorter and more slender than pereopods 3 and 4. Propodus 1.3 times as long as carpus, with grooming apparatus consisting of 4 transverse rows of stiff setae in distal 0.2, otherwise unarmed. Dactylus (
|
||
<figureCitation id="135B2A1FFFB74B4E8B96FE9BFF16217A" box="[160,247,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">Fig. 5H</figureCitation>
|
||
) short, about 0.2 times as long as propodus, subconical, with flat flexor surface.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FEBFFACA2102" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">Gill formula including 5 pleurobranchs above pereopods 1–5, 1 arthrobranch above maxilliped 3, 2 lobe-like epipods (maxillipeds 1 and 2), no strap-like epipods on maxilliped 3 and pereopods 1–5, no setobranchs.</paragraph>
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FEF7FE672192" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">
|
||
Pleopod protopods (
|
||
<figureCitation id="135B2A1FFFB74B4E8A9DFEF7FDF321EE" box="[427,530,366,393]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">Fig. 5I, J</figureCitation>
|
||
) each with thick ovigerous setae on mesial margin. Pleopod 1 (
|
||
<figureCitation id="135B2A1FFFB74B4E8FE8FEF7FACA21EE" box="[1246,1323,366,393]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">Fig. 5I</figureCitation>
|
||
) endopod small, about half-length of exopod, tapering distally, bearing numerous setae distally. Pleopod 2 (
|
||
<figureCitation id="135B2A1FFFB74B4E8FEBFE0BFACD21CA" box="[1245,1324,402,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">Fig. 5J</figureCitation>
|
||
) endopod subequal in length to exopod, with long, slender appendix interna (about half length of endopod) arising at proximal 0.4 of mesial margin.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FE67FDF122CE" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">
|
||
Uropod (
|
||
<figureCitation id="135B2A1FFFB74B4E8A18FE67FE9F227E" box="[302,382,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="9" pageNumber="406">Fig. 2J</figureCitation>
|
||
) with protopod having broadly rounded lateral lobe and 1 small medial spine dorsally at articulation to endopod. Exopod with posterior flap very short, posterior margin roundly truncate, without incision or concavity; lateral margin slightly convex; diaeresis with feebly marked, broadly triangular tooth just lateral to posterolateral spiniform seta, and two much larger triangular teeth defined by similarly deep incisions. Endopod slightly longer than exopod, oval.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FD2BFE4923C2" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">
|
||
<typeStatus id="54DB8838FFB74B4E8BF1FD2BFED922AA" box="[199,312,691,717]" pageId="9" pageNumber="406" type="paratype">Paratypes</typeStatus>
|
||
. Generally similar to
|
||
<typeStatus id="54DB8838FFB74B4E891BFD2BFD7222AA" box="[557,659,691,717]" pageId="9" pageNumber="406" type="holotype">holotype</typeStatus>
|
||
. Telson 1.9–2.0 times as long as wide. Antennular peduncle article 1 with row of 5–6 minute spiniform setae on proximal half of dorsomesial margin. Antennal scaphocerite with distolateral tooth varying from sharply triangular (
|
||
<figureCitation id="135B2A1FFFB74B4E89AEFD63FD112372" box="[664,752,762,789]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">Fig. 5K</figureCitation>
|
||
) to blunt as in
|
||
<typeStatus id="54DB8838FFB74B4E88AAFD63FBE32372" box="[924,1026,763,789]" pageId="9" pageNumber="406" type="holotype">holotype</typeStatus>
|
||
. Male pleopod 1 with endopod (
|
||
<figureCitation id="135B2A1FFFB74B4E8E59FD63FF56235E" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">Fig. 5L</figureCitation>
|
||
) small (length less than half of exopod), bearing sparse setae. Male pleopod 2 with appendices masculina and interna (
|
||
<figureCitation id="135B2A1FFFB74B4E8BC2FCDBFEAE233A" box="[244,335,834,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">Fig. 5M</figureCitation>
|
||
) arising proximal to mid-length of endopod mesial margin; appendix masculina rod-like, distinctly shorter than appendix interna, bearing 1 stiff apical seta. Posterolateral margin of uropodal exopod distal flap truncate or slightly concave.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FC36FCE22476" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">
|
||
<emphasis id="B914EA88FFB74B4E8BF1FC36FE4C23AF" bold="true" box="[199,429,942,968]" pageId="9" pageNumber="406">Colouration in life.</emphasis>
|
||
Body and appendages (
|
||
<figureCitation id="135B2A1FFFB74B4E89F6FC37FCDB23AE" box="[704,826,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1929,1953]" captionTargetBox="[151,1435,181,1905]" captionTargetId="figure-17@3.[151,1436,181,1905]" captionTargetPageId="3" captionText="FIGURE 1. Richalpheus laeuadigitus n. sp., habitus in lateral view, showing colouration in fresh condition.A, holotype, female (cl 10.5 mm), KAUM-AT 1005; B, paratype, female (cl 7.4 mm), KAUM-AT 902; C, paratype, female (cl 4.5 mm), KAUM-AT 1352." figureDoi="http://doi.org/10.5281/zenodo.6358396" httpUri="https://zenodo.org/record/6358396/files/figure.png" pageId="9" pageNumber="406">Fig. 1A–C</figureCitation>
|
||
) generally pale pink or pale orange semi-transparent, with numerous scattered red chromatophores on carapace, pleon, telson, antennae and major cheliped, occasionally forming intense tinge; pereopods 2–5 semi-transparent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A6511FFB74B4E8BF1FB82FCFD243E" pageId="9" pageNumber="406" type="distribution">
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FB82FCFD243E" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">
|
||
<emphasis id="B914EA88FFB74B4E8BF1FB82FE802453" bold="true" box="[199,353,1050,1076]" pageId="9" pageNumber="406">Distribution.</emphasis>
|
||
Presently known only from three locations in
|
||
<collectingRegion id="49A4F878FFB74B4E8844FB83FB912452" box="[882,1136,1051,1077]" country="Japan" name="Kagoshima" pageId="9" pageNumber="406">Kagoshima Prefecture</collectingRegion>
|
||
, Kyushu,
|
||
<collectingCountry id="F377760AFFB74B4E8FD4FB83FAC42452" box="[1250,1317,1051,1077]" name="Japan" pageId="9" pageNumber="406">Japan</collectingCountry>
|
||
, i.e., Kiire (
|
||
<typeStatus id="54DB8838FFB74B4E8BA9FBA7FF30243E" box="[159,209,1087,1113]" pageId="9" pageNumber="406">type</typeStatus>
|
||
locality), Tarumizu and Kimotsuki; lower intertidal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A6511FFB74B4E8BF1FBFBFB2926FE" pageId="9" pageNumber="406" type="biology_ecology">
|
||
<paragraph id="8BDF369AFFB74B4E8BF1FBFBFB2926FE" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="406">
|
||
<emphasis id="B914EA88FFB74B4E8BF1FBFBFECD241A" bold="true" box="[199,300,1123,1149]" pageId="9" pageNumber="406">Ecology.</emphasis>
|
||
Eight specimens from Kiire, including the
|
||
<typeStatus id="54DB8838FFB74B4E8824FBFBFC95241A" box="[786,884,1123,1149]" pageId="9" pageNumber="406" type="holotype">holotype</typeStatus>
|
||
female, were all collected from tidal flat consisting of soft mud, where there is a spring of fresh-water; scattered burrow openings were seen on the bottom surface (
|
||
<figureCitation id="135B2A1FFFB74B4E8B96FB33FF1A24A1" box="[160,251,1195,1222]" captionStart="FIGURE 6" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[165,1423,181,1851]" captionTargetId="figure-30@8.[165,1423,181,1852]" captionTargetPageId="8" captionText="FIGURE 6. A, tidal flat of Kiire, Kagoshima Prefecture, where specimens of Richalpheus laeuadigitus n. sp. were collected; B, catches by yabby pump at Kiire, Kagoshima Prefecture: 1, Richalpheus laeuadigitus n. sp.; 2, Balanoglossus misakiensis Kuwano, 1902; 3, Neotrypaea harmandi (Bouvier, 1901) or N. japonica (Ortmann, 1891); 4, Paratrypaea bouvieri (Nobili, 1904); 5, Pinnixa balanoglossana Sakai, 1934." figureDoi="http://doi.org/10.5281/zenodo.6358410" httpUri="https://zenodo.org/record/6358410/files/figure.png" pageId="9" pageNumber="406">Fig. 6A</figureCitation>
|
||
). Sampling made on
|
||
<date id="FFDE105AFFB74B4E8ADBFB33FD9624A2" box="[493,631,1195,1221]" pageId="9" pageNumber="406" value="2020-05-09">9 May 2020</date>
|
||
with the yabby pump yielded individuals of a large ptychoderid worm
|
||
<taxonomicName id="4C604D19FFB74B4E8BA1FB57FD92248E" authority="Kuwano, 1902" authorityName="Kuwano" authorityYear="1902" box="[151,627,1231,1257]" class="Enteropneusta" family="Ptychoderidae" genus="Balanoglossus" kingdom="Animalia" pageId="9" pageNumber="406" phylum="Hemichordata" rank="species" species="misakiensis">
|
||
<emphasis id="B914EA88FFB74B4E8BA1FB57FE22248E" box="[151,451,1231,1257]" italics="true" pageId="9" pageNumber="406">Balanoglossus misakiensis</emphasis>
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8AFDFB57FD92248E" author="Kuwano, H." box="[459,627,1231,1257]" pageId="9" pageNumber="406" pagination="77 - 84" refId="ref5593" refString="Kuwano, H. (1902) On a new Enteropneust from Misaki, Balanoglossus misakiensis n. sp. Annotationes Zoologicae Japonenses, 4, 77 - 84." type="journal article" year="1902">Kuwano, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
, along with six individuals of the present new species (KAUM-AT 902, 920–923, and 1005), individuals of three callianassid ghost shrimp
|
||
<taxonomicName id="4C604D19FFB74B4E88A1FB6BFAAC256A" authority="(Bouvier, 1901)" baseAuthorityName="Bouvier" baseAuthorityYear="1901" box="[919,1357,1267,1293]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="harmandi">
|
||
<emphasis id="B914EA88FFB74B4E88A1FB6BFB6E256A" box="[919,1167,1267,1293]" italics="true" pageId="9" pageNumber="406">Neotrypaea harmandi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8F96FB6BFAA4256A" author="Bouvier, E. L." box="[1184,1349,1267,1293]" pageId="9" pageNumber="406" pagination="332 - 334" refId="ref5268" refString="Bouvier, E. L. (1901) Sur quelques crustaces du Japon, offerts au museum par M. le Dr. Harmand. Bulletin du Museum National d'Histoire Naturelle, Paris, 7, 332 - 334. https: // doi. org / 10.5962 / bhl. part. 1625" type="journal article" year="1901">Bouvier, 1901</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C604D19FFB74B4E8E6DFB6BFE4A2556" authority="(Ortmann, 1891)" baseAuthorityName="Ortmann" baseAuthorityYear="1891" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="B914EA88FFB74B4E8E6DFB6BFF022557" italics="true" pageId="9" pageNumber="406">N. japonica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8BC2FA8FFE432556" author="Ortmann, A. E." box="[244,418,1303,1329]" pageId="9" pageNumber="406" pagination="1 - 58" refId="ref5710" refString="Ortmann, A. E. (1891) Die Decapoden-Krebse des Strassburger Museums mit besonderer Berucksichtigung der von Herrn Dr. Doderlein dei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen III. Theil. Die Abtheilungen der Reptantia Boas: Homaridae, Loricata und Thalassinidea. Zoologische Jahrbucher. Abteilung fur Systematik, 6, 1 - 58. https: // doi. org / 10.5962 / bhl. part. 26455" type="journal article" year="1891">Ortmann, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C604D19FFB74B4E8AD3FA8FFC662556" authority="(Nobili, 1904)" baseAuthorityName="Nobili" baseAuthorityYear="1904" box="[485,903,1303,1329]" class="Malacostraca" family="Callianassidae" genus="Paratrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="bouvieri">
|
||
<emphasis id="B914EA88FFB74B4E8AD3FA8FFD382556" box="[485,729,1303,1329]" italics="true" pageId="9" pageNumber="406">Paratrypaea bouvieri</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF14B6BFFB74B4E89DCFA8FFC9E2556" author="Nobili, G." box="[746,895,1303,1329]" pageId="9" pageNumber="406" pagination="230 - 238" refId="ref5673" refString="Nobili, G. (1904) Diagnoses preliminaires de vingt-huit especes nouvelles de Stomatopodes et Decapodes Macroures de la Mer Rouge. Bulletin du Museum d'Histoire Naturelle de Paris, 10, 230 - 238." type="journal article" year="1904">Nobili, 1904</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and individuals of a pinnotherid crab
|
||
<taxonomicName id="4C604D19FFB74B4E8E73FA8FFE362532" authority="Sakai, 1934" authorityName="Sakai" authorityYear="1934" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="balanoglossana">
|
||
<emphasis id="B914EA88FFB74B4E8E73FA8FFEA92532" italics="true" pageId="9" pageNumber="406">Pinnixa balanoglossana</emphasis>
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8A79FAA3FE362532" author="Sakai, T." box="[335,471,1339,1365]" pageId="9" pageNumber="406" pagination="37 - 43" refId="ref5884" refString="Sakai, T. (1934) Species of the genus Pinnixa (Pinnotherid crab) found in the Far East. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2, 37 - 43." type="journal article" year="1934">Sakai, 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
, from those burrow openings (
|
||
<figureCitation id="135B2A1FFFB74B4E881BFAA3FC672531" box="[813,902,1339,1366]" captionStart="FIGURE 6" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[165,1423,181,1851]" captionTargetId="figure-30@8.[165,1423,181,1852]" captionTargetPageId="8" captionText="FIGURE 6. A, tidal flat of Kiire, Kagoshima Prefecture, where specimens of Richalpheus laeuadigitus n. sp. were collected; B, catches by yabby pump at Kiire, Kagoshima Prefecture: 1, Richalpheus laeuadigitus n. sp.; 2, Balanoglossus misakiensis Kuwano, 1902; 3, Neotrypaea harmandi (Bouvier, 1901) or N. japonica (Ortmann, 1891); 4, Paratrypaea bouvieri (Nobili, 1904); 5, Pinnixa balanoglossana Sakai, 1934." figureDoi="http://doi.org/10.5281/zenodo.6358410" httpUri="https://zenodo.org/record/6358410/files/figure.png" pageId="9" pageNumber="406">Fig. 6B</figureCitation>
|
||
). The other
|
||
<specimenCount id="9D66FD13FFB74B4E8F3AFAA3FB532532" box="[1036,1202,1339,1365]" pageId="9" pageNumber="406" type="generic">two specimens</specimenCount>
|
||
collected at Kiire by another occasion (
|
||
<date id="FFDE105AFFB74B4E8A50FAC7FDE3251E" box="[358,514,1375,1401]" pageId="9" pageNumber="406" value="2020-06-21">21 June 2020</date>
|
||
; KAUM-AT 1202, 1203) were extracted from sediments together with specimens of
|
||
<taxonomicName id="4C604D19FFB74B4E8B80FA1BFDBF25FA" authority="de Haan, 1844" authorityName="de Haan" authorityYear="1844" box="[182,606,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="brevicristatus">
|
||
<emphasis id="B914EA88FFB74B4E8B80FA1BFE5025FA" box="[182,433,1411,1437]" italics="true" pageId="9" pageNumber="406">Alpheus brevicristatus</emphasis>
|
||
de Haan, 1844
|
||
</taxonomicName>
|
||
, but the collection field was also inhabited by
|
||
<taxonomicName id="4C604D19FFB74B4E8F5FFA1BFA7425FA" authorityName="Kuwano" authorityYear="1902" box="[1129,1429,1411,1437]" class="Enteropneusta" family="Ptychoderidae" genus="Balanoglossus" kingdom="Animalia" pageId="9" pageNumber="406" phylum="Hemichordata" rank="species" species="misakiensis">
|
||
<emphasis id="B914EA88FFB74B4E8F5FFA1BFA7425FA" box="[1129,1429,1411,1437]" italics="true" pageId="9" pageNumber="406">Balanoglossus misakiensis</emphasis>
|
||
</taxonomicName>
|
||
. It is possible that the new species is commensal with one of those infaunal animals except for
|
||
<taxonomicName id="4C604D19FFB74B4E8F8EFA3FFF2D2583" authorityName="Sakai" authorityYear="1934" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="balanoglossana">
|
||
<emphasis id="B914EA88FFB74B4E8F8EFA3FFF2D2583" italics="true" pageId="9" pageNumber="406">Pinnixa balanoglossana</emphasis>
|
||
</taxonomicName>
|
||
that is commensal with
|
||
<taxonomicName id="4C604D19FFB74B4E8AEBFA53FD9F2582" authorityName="Kuwano" authorityYear="1902" box="[477,638,1483,1509]" class="Enteropneusta" family="Ptychoderidae" genus="Balanoglossus" kingdom="Animalia" pageId="9" pageNumber="406" phylum="Hemichordata" rank="species" species="misakiensis">
|
||
<emphasis id="B914EA88FFB74B4E8AEBFA53FD9F2582" box="[477,638,1483,1509]" italics="true" pageId="9" pageNumber="406">B. misakiensis</emphasis>
|
||
</taxonomicName>
|
||
(cf. T.
|
||
<bibRefCitation id="EFF14B6BFFB74B4E89F9FA53FCAE2582" author="Sakai, T." box="[719,847,1483,1509]" pageId="9" pageNumber="406" pagination="37 - 43" refId="ref5884" refString="Sakai, T. (1934) Species of the genus Pinnixa (Pinnotherid crab) found in the Far East. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2, 37 - 43." type="journal article" year="1934">Sakai 1934</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF14B6BFFB74B4E886CFA53FC742582" author="Sakai, T." box="[858,917,1483,1509]" pageId="9" pageNumber="406" refId="ref5923" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, xxix + 773 pp. [English text] + 461 pp. (Japanese text) + 251 pp. (plate volume)]" type="book" year="1976">1976</bibRefCitation>
|
||
; K.
|
||
<bibRefCitation id="EFF14B6BFFB74B4E88F5FA53FBA52582" author="Sakai, K." box="[963,1092,1483,1509]" pageId="9" pageNumber="406" pagination="1 - 6" refId="ref5839" refString="Sakai, K. (2002) Ecological notes on Pinnixa balanoglossana Sakai (Pinnotheridae) inhabited the coast of Noto Peninsula, the Sea of Japan. Report of the Noto Marine Center, 8, 1 - 6. [in Japanese]" type="journal article" year="2002">Sakai 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8F79FA53FA712582" author="Shimetsugu, M. & Kimura, S." box="[1103,1424,1483,1509]" pageId="9" pageNumber="406" pagination="207 - 210" refId="ref6040" refString="Shimetsugu, M. & Kimura, S. (2018) Records of Pinnixa balanoglossana Sakai, 1934 (Decapoda, Brachyura, Pinnotheridae) from Mie Prefecture, Japan. Nankiseibutu, 60 (2), 207 - 210. [in Japanese]" type="journal article" year="2018">Shimetsugu & Kimura 2018</bibRefCitation>
|
||
), though it is not easy to specify at present what is the host of
|
||
<taxonomicName id="4C604D19FFB74B4E8802FA77FC3D266E" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[820,988,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB74B4E8802FA77FC3D266E" box="[820,988,1519,1545]" italics="true" pageId="9" pageNumber="406">R. laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB74B4E88D5FA77FBC0266E" bold="true" box="[995,1057,1519,1545]" pageId="9" pageNumber="406">
|
||
<taxonomicNameLabel id="A22757F3FFB74B4E88D5FA77FBC0266E" box="[995,1057,1519,1545]" pageId="9" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The specimens from a tidal flat at the river mouth of Matsuzaki River, Tarumizu (KAUM-AT 1490) and from a tidal flat at the river mouth of Ogawa River, Kimotsuki (KAUM-AT 1352) were extracted from mud substrate also by using yabby pump, but no other invertebrates were found. Other species of
|
||
<taxonomicName id="4C604D19FFB74B4E8969F9C3FCA02612" box="[607,833,1627,1653]" pageId="9" pageNumber="406">
|
||
<emphasis id="B914EA88FFB74B4E8969F9C3FD082612" box="[607,745,1627,1653]" italics="true" pageId="9" pageNumber="406">Richalpheus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are also supposed to be associated with other infaunal invertebrates, possibly callianassid ghost shrimp, but not positively identified yet (
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8F07F9E7FB5A26FE" author="Anker, A." box="[1073,1211,1663,1689]" pageId="9" pageNumber="406" pagination="59 - 68" refId="ref5038" refString="Anker, A. (2011) Description of a third species in the alpheid shrimp genus Richalpheus Anker & Jeng, 2006 (Crustacea, Decapoda). Zootaxa, 3059 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 3059.1.4" type="journal article" year="2011">Anker 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A6511FFB74B4D8BF1F93BFF11238A" lastPageId="10" lastPageNumber="407" pageId="9" pageNumber="406" type="discussion">
|
||
<paragraph id="8BDF369AFFB74B4D8BF1F93BFED02125" blockId="9.[151,1437,150,2013]" lastBlockId="10.[151,1437,151,1113]" lastPageId="10" lastPageNumber="407" pageId="9" pageNumber="406">
|
||
<emphasis id="B914EA88FFB74B4E8BF1F93BFEDA26DA" bold="true" box="[199,315,1699,1725]" pageId="9" pageNumber="406">Remarks.</emphasis>
|
||
The configuration of fingers of the major cheliped provides diagnostic characters for species differentiation in
|
||
<taxonomicName id="4C604D19FFB74B4E8A08F95FFD8D2686" authority="(Anker 2011)" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[318,620,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B914EA88FFB74B4E8A08F95FFE292686" box="[318,456,1735,1761]" italics="true" pageId="9" pageNumber="406">Richalpheus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8AEEF95FFD852686" author="Anker, A." box="[472,612,1735,1761]" pageId="9" pageNumber="406" pagination="59 - 68" refId="ref5038" refString="Anker, A. (2011) Description of a third species in the alpheid shrimp genus Richalpheus Anker & Jeng, 2006 (Crustacea, Decapoda). Zootaxa, 3059 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 3059.1.4" type="journal article" year="2011">Anker 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. In this regard, the present new species is most similar to
|
||
<taxonomicName id="4C604D19FFB74B4E8FC8F95FFA972686" authorityName="Anker & Jeng" authorityYear="2006" box="[1278,1398,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB74B4E8FC8F95FFA972686" box="[1278,1398,1735,1761]" italics="true" pageId="9" pageNumber="406">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C604D19FFB74B4E8BA1F973FEA02762" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[151,321,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="laeuadigitus" status="n. sp. and r.">
|
||
<emphasis id="B914EA88FFB74B4E8BA1F973FEA02762" box="[151,321,1771,1797]" italics="true" pageId="9" pageNumber="406">R. laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22757F3FFB74B4E8A7CF973FE3A2763" box="[330,475,1771,1797]" pageId="9" pageNumber="406" rank="subSpecies">
|
||
<emphasis id="B914EA88FFB74B4E8A7CF973FE6B2762" bold="true" box="[330,394,1771,1797]" pageId="9" pageNumber="406">n. sp.</emphasis>
|
||
and
|
||
<taxonomicName id="4C604D19FFB74B4E8AF5F973FE3A2763" authorityName="Anker & Jeng" authorityYear="2006" box="[451,475,1771,1796]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB74B4E8AF5F973FE3A2763" box="[451,475,1771,1796]" italics="true" pageId="9" pageNumber="406">R.</emphasis>
|
||
</taxonomicName>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B914EA88FFB74B4E8AD5F973FDDB2762" box="[483,570,1771,1797]" italics="true" pageId="9" pageNumber="406">palmeri</emphasis>
|
||
, the dactylus has a broad bulge-like ridge, occupying proximal half or more of the dactylar occlusal margin and fitting into a deep longitudinal groove on the pollex (
|
||
<figureCitation id="135B2A1FFFB74B4E8FB5F897FAE9274E" box="[1155,1288,1807,1833]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[198,1385,197,1904]" captionTargetId="figure-30@6.[183,1404,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, major (left) cheliped, lateral view; B, same, mesial view; C, same, merus, dorsal view; E, same, chela, flexor view; F, same, extensor view." figureDoi="http://doi.org/10.5281/zenodo.6358404" httpUri="https://zenodo.org/record/6358404/files/figure.png" pageId="9" pageNumber="406">Figs. 4E, D</figureCitation>
|
||
,
|
||
<figureCitation id="135B2A1FFFB74B4E8E22F896FAD6274E" box="[1300,1335,1806,1833]" captionStart="FIGURE 5" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[158,1426,190,1680]" captionTargetId="figure-17@7.[151,1436,181,1691]" captionTargetPageId="7" captionText="FIGURE 5. Richalpheus laeuadigitus n. sp. A–J, holotype, female (cl 10.5 mm), KAUM-AT 1005; K–M, paratype, male (cl 5.7 mm), KAUM-AT 923.A, coxal lateral plate of right maxilliped 3, obliquely lateral view; B, pollex and dactylus occlusal margin of left major cheliped, showing groove on pollex; C, chela of left minor cheliped (pereopod 1), flexor view; D, same, pollex and dactylus; E, distal part of dactylus of left minor cheliped, mesial view, showing details of armature on occlusal margin; F, G, distal part of propodus and dactylus of pereopods 3 and 4, flexor view; H, distal part of propodus and dactylus of pereopods 5, lateral view; I, L, left pleopod 1, ventral view (marginal setae on rami omitted); J, left pleopod 2, ventral view (marginal setae on rami omitted); K, distal part of left scaphocerite, dorsal view (setae omitted); M, endopod of left pleopod 2 with appendices interna and masculina, dorsal view (setae omitted).Abbreviations: CLP = coxal lateral plate; EXP = exopod." figureDoi="http://doi.org/10.5281/zenodo.6358406" httpUri="https://zenodo.org/record/6358406/files/figure.png" pageId="9" pageNumber="406">5B</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8BA1F8ABFE9B272A" author="Anker, A. & Jeng, M. S." box="[151,378,1843,1869]" pageId="9" pageNumber="406" pagination="379 - 391" refId="ref5197" refString="Anker, A. & Jeng, M. S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutiere, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379 - 391. https: // doi. org / 10.1651 / S- 2647.1" type="journal article" year="2006">Anker & Jeng 2006</bibRefCitation>
|
||
: fig. 4). In
|
||
<taxonomicName id="4C604D19FFB74B4E8ACEF8ABFD70272A" authorityName="Anker" authorityYear="2011" box="[504,657,1843,1869]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="alpheoides">
|
||
<emphasis id="B914EA88FFB74B4E8ACEF8ABFD70272A" box="[504,657,1843,1869]" italics="true" pageId="9" pageNumber="406">R. alpheoides</emphasis>
|
||
</taxonomicName>
|
||
, the dactylus has a well-defined, distally excavated tooth, situated in the proximal half of the dactylar occlusal margin, and fitting into a deep rounded fossa on the pollex (cf.
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8E60F8CFFF3327F2" author="Anker, A." pageId="9" pageNumber="406" pagination="59 - 68" refId="ref5038" refString="Anker, A. (2011) Description of a third species in the alpheid shrimp genus Richalpheus Anker & Jeng, 2006 (Crustacea, Decapoda). Zootaxa, 3059 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 3059.1.4" type="journal article" year="2011">Anker 2011</bibRefCitation>
|
||
: fig. 2D–F). In
|
||
<taxonomicName id="4C604D19FFB74B4E8AB7F8E3FDFF27F2" authorityName="Anker & Dworschak" authorityYear="2007" box="[385,542,1915,1941]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="dahabensis">
|
||
<emphasis id="B914EA88FFB74B4E8AB7F8E3FDFF27F2" box="[385,542,1915,1941]" italics="true" pageId="9" pageNumber="406">R. dahabensis</emphasis>
|
||
</taxonomicName>
|
||
, the dactylus has neither a bulge nor a tooth, whereas the pollex has a proximal groove (
|
||
<bibRefCitation id="EFF14B6BFFB74B4E8BC5F807FDC027DE" author="Anker, A. & Dworschak, P. C." box="[243,545,1951,1977]" pageId="9" pageNumber="406" pagination="2331 - 2340" refId="ref5138" refString="Anker, A. & Dworschak, P. C. (2007) Description of a new species of Richalpheus Anker and Jeng, 2006 (Crustacea: Decapoda: Alpheidae) from the Red Sea. Journal of Natural History, 41, 2331 - 2340. https: // doi. org / 10.1080 / 10503300701579943" type="journal article" year="2007">Anker & Dworschak 2007</bibRefCitation>
|
||
: fig. 3A).
|
||
<taxonomicName id="4C604D19FFB74B4E89A4F807FC4C27DE" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[658,941,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB74B4E89A4F807FC4C27DE" box="[658,941,1951,1977]" italics="true" pageId="9" pageNumber="406">Richalpheus laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB74B4E8883F807FC1227DE" bold="true" box="[949,1011,1951,1977]" pageId="9" pageNumber="406">
|
||
<taxonomicNameLabel id="A22757F3FFB74B4E8883F807FC1227DE" box="[949,1011,1951,1977]" pageId="9" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C604D19FFB74B4E8FB1F807FB1C27DE" authorityName="Anker & Jeng" authorityYear="2006" box="[1159,1277,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB74B4E8FB1F807FB1C27DE" box="[1159,1277,1951,1977]" italics="true" pageId="9" pageNumber="406">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
in the following particulars: the telson is relatively wider in the new species than in
|
||
<taxonomicName id="4C604D19FFB74B4E8880F85BFBCD27BA" authorityName="Anker & Jeng" authorityYear="2006" box="[950,1068,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB74B4E8880F85BFBCD27BA" box="[950,1068,1987,2013]" italics="true" pageId="9" pageNumber="406">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
(twice or less than twice as long as wide versus twice or more;
|
||
<figureCitation id="135B2A1FFFB44B4D8ADDFF0FFDA020D6" box="[491,577,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="10" pageNumber="407">Fig. 2D</figureCitation>
|
||
versus
|
||
<bibRefCitation id="EFF14B6BFFB44B4D89A3FF0FFC9420D6" author="Anker, A. & Jeng, M. S." box="[661,885,151,177]" pageId="10" pageNumber="407" pagination="379 - 391" refId="ref5197" refString="Anker, A. & Jeng, M. S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutiere, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379 - 391. https: // doi. org / 10.1651 / S- 2647.1" type="journal article" year="2006">Anker & Jeng 2006</bibRefCitation>
|
||
: fig. 2k); the antennular stylocerite reaches to the 0.6 of the basal peduncular article in
|
||
<taxonomicName id="4C604D19FFB44B4D890BFF23FD0620B2" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[573,743,187,213]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB44B4D890BFF23FD0620B2" box="[573,743,187,213]" italics="true" pageId="10" pageNumber="407">R. laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB44B4D89D9FF23FCCE20B2" bold="true" box="[751,815,187,213]" pageId="10" pageNumber="407">
|
||
<taxonomicNameLabel id="A22757F3FFB44B4D89D9FF23FCCE20B2" box="[751,815,187,213]" pageId="10" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="135B2A1FFFB44B4D8876FF23FC7720B2" box="[832,918,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="10" pageNumber="407">Fig. 2F</figureCitation>
|
||
), rather than reaching to the
|
||
<quantity id="4C989B7FFFB44B4D8FEAFF23FAFC20B2" box="[1244,1309,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" pageId="10" pageNumber="407" unit="in" value="0.8">0.8 in</quantity>
|
||
<taxonomicName id="4C604D19FFB44B4D8E13FF23FA7D20B2" authorityName="Anker & Jeng" authorityYear="2006" box="[1317,1436,187,213]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB44B4D8E13FF23FA7D20B2" box="[1317,1436,187,213]" italics="true" pageId="10" pageNumber="407">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EFF14B6BFFB44B4D8BF2FF47FE49209E" author="Anker, A. & Jeng, M. S." box="[196,424,223,249]" pageId="10" pageNumber="407" pagination="379 - 391" refId="ref5197" refString="Anker, A. & Jeng, M. S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutiere, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379 - 391. https: // doi. org / 10.1651 / S- 2647.1" type="journal article" year="2006">Anker & Jeng 2006</bibRefCitation>
|
||
: fig. 2c); the uropodal diaeresis has two similarly deep incisions in
|
||
<taxonomicName id="4C604D19FFB44B4D8F9FFF47FAB2209E" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[1193,1363,223,249]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB44B4D8F9FFF47FAB2209E" box="[1193,1363,223,249]" italics="true" pageId="10" pageNumber="407">R. laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB44B4D8E6AFF47FA7D209E" bold="true" box="[1372,1436,223,249]" pageId="10" pageNumber="407">
|
||
<taxonomicNameLabel id="A22757F3FFB44B4D8E6AFF47FA7D209E" box="[1372,1436,223,249]" pageId="10" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="135B2A1FFFB44B4D8B96FE9BFF11217A" box="[160,240,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1812,1836]" captionTargetBox="[228,1358,190,1778]" captionTargetId="figure-30@4.[223,1365,181,1788]" captionTargetPageId="4" captionText="FIGURE 2. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005. A, carapace and cephalic appendages, left lateral view; B, pleon, telson and pleonal appendages, left lateral view; C, pleomere 6, left lateral view; D, telson, dorsal view; E, anterior part of carapace and cephalic appendages, lateral view; F, same, dorsal view (marginal setae on left scaphocerite omitted); G, proximomesial margins of articles 1 of antennular peduncles, dorsal view, showing detail of armature; H, ventromesial ridge on article 1 of antennular peduncle, lateral view; I, distal part of left scaphocerite, dorsal view (setae omitted); J, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6358398" httpUri="https://zenodo.org/record/6358398/files/figure.png" pageId="10" pageNumber="407">Fig. 2J</figureCitation>
|
||
), whereas it has a very shallow lateral incision and a deep mesial incision in
|
||
<taxonomicName id="4C604D19FFB44B4D8F62FE9BFB2B217A" authorityName="Anker & Jeng" authorityYear="2006" box="[1108,1226,259,285]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB44B4D8F62FE9BFB2B217A" box="[1108,1226,259,285]" italics="true" pageId="10" pageNumber="407">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EFF14B6BFFB44B4D8FC8FE9BFF322126" author="Anker, A. & Jeng, M. S." pageId="10" pageNumber="407" pagination="379 - 391" refId="ref5197" refString="Anker, A. & Jeng, M. S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutiere, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379 - 391. https: // doi. org / 10.1651 / S- 2647.1" type="journal article" year="2006">Anker & Jeng 2006</bibRefCitation>
|
||
: fig. 2j).
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB44B4D8BF1FED3FF24227E" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="407">
|
||
Other than the characters of the major cheliped fingers,
|
||
<taxonomicName id="4C604D19FFB44B4D880DFED3FC052102" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[827,996,331,357]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB44B4D880DFED3FC052102" box="[827,996,331,357]" italics="true" pageId="10" pageNumber="407">R. laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB44B4D88DAFED3FBCA2102" bold="true" box="[1004,1067,331,357]" pageId="10" pageNumber="407">
|
||
<taxonomicNameLabel id="A22757F3FFB44B4D88DAFED3FBCA2102" box="[1004,1067,331,357]" pageId="10" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is very similar to
|
||
<taxonomicName id="4C604D19FFB44B4D8FCEFED3FA742102" authorityName="Anker & Dworschak" authorityYear="2007" box="[1272,1429,331,357]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="dahabensis">
|
||
<emphasis id="B914EA88FFB44B4D8FCEFED3FA742102" box="[1272,1429,331,357]" italics="true" pageId="10" pageNumber="407">R. dahabensis</emphasis>
|
||
</taxonomicName>
|
||
. Nevertheless, the following subtle differences might be diagnostic for discrimination between the two taxa: the telson is relatively wider in
|
||
<taxonomicName id="4C604D19FFB44B4D8A98FE0BFDB721CA" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[430,598,403,429]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB44B4D8A98FE0BFDB721CA" box="[430,598,403,429]" italics="true" pageId="10" pageNumber="407">R. laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB44B4D896AFE0BFD7B21CA" bold="true" box="[604,666,403,429]" pageId="10" pageNumber="407">
|
||
<taxonomicNameLabel id="A22757F3FFB44B4D896AFE0BFD7B21CA" box="[604,666,403,429]" pageId="10" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName id="4C604D19FFB44B4D89C4FE0BFC6F21CA" authorityName="Anker & Dworschak" authorityYear="2007" box="[754,910,403,429]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="dahabensis">
|
||
<emphasis id="B914EA88FFB44B4D89C4FE0BFC6F21CA" box="[754,910,403,429]" italics="true" pageId="10" pageNumber="407">R. dahabensis</emphasis>
|
||
</taxonomicName>
|
||
(1.9–2.0 times as long as wide versus 2.3 times as long); the exopod of the maxilliped 3 does not reach the distal margin of the endopod antepenultimate article in
|
||
<taxonomicName id="4C604D19FFB44B4D8BA1FE43FEA12192" authority="Komai & Koreeda, 2022" authorityName="Komai & Koreeda" authorityYear="2022" box="[151,320,475,501]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="laeuadigitus" status="sp. nov.">
|
||
<emphasis id="B914EA88FFB44B4D8BA1FE43FEA12192" box="[151,320,475,501]" italics="true" pageId="10" pageNumber="407">R. laeuadigitus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914EA88FFB44B4D8A7EFE43FE672192" bold="true" box="[328,390,475,501]" pageId="10" pageNumber="407">
|
||
<taxonomicNameLabel id="A22757F3FFB44B4D8A7EFE43FE672192" box="[328,390,475,501]" pageId="10" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="135B2A1FFFB44B4D8AA0FE43FDEE2192" box="[406,527,475,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[204,1389,191,1910]" captionTargetId="figure-17@5.[193,1395,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Richalpheus laeuadigitus n. sp., holotype, female (cl 10.5 mm), KAUM-AT 1005.A, left maxilliped 3, lateral view; B, same, dorsal view (coxal lateral plate broken off); C, minor (right) cheliped (pereopod 1), lateral view; D–G, left pereopods 2–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6358402" httpUri="https://zenodo.org/record/6358402/files/figure.png" pageId="10" pageNumber="407">Fig. 3A, B</figureCitation>
|
||
), rather than overreaching it in
|
||
<taxonomicName id="4C604D19FFB44B4D8846FE43FBEC2192" authorityName="Anker & Dworschak" authorityYear="2007" box="[880,1037,475,501]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="dahabensis">
|
||
<emphasis id="B914EA88FFB44B4D8846FE43FBEC2192" box="[880,1037,475,501]" italics="true" pageId="10" pageNumber="407">R. dahabensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EFF14B6BFFB44B4D8F09FE43FA8A2192" author="Anker, A. & Dworschak, P. C." box="[1087,1387,475,501]" pageId="10" pageNumber="407" pagination="2331 - 2340" refId="ref5138" refString="Anker, A. & Dworschak, P. C. (2007) Description of a new species of Richalpheus Anker and Jeng, 2006 (Crustacea: Decapoda: Alpheidae) from the Red Sea. Journal of Natural History, 41, 2331 - 2340. https: // doi. org / 10.1080 / 10503300701579943" type="journal article" year="2007">Anker & Dworschak 2007</bibRefCitation>
|
||
: fig. 2F).
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB44B4D8BF1FDBBFDA422AA" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="407">
|
||
<taxonomicName id="4C604D19FFB44B4D8BF1FDBBFE33225A" authorityName="Anker" authorityYear="2011" box="[199,466,547,573]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="alpheoides">
|
||
<emphasis id="B914EA88FFB44B4D8BF1FDBBFE33225A" box="[199,466,547,573]" italics="true" pageId="10" pageNumber="407">Richalpheus alpheoides</emphasis>
|
||
</taxonomicName>
|
||
further differs from the new species,
|
||
<taxonomicName id="4C604D19FFB44B4D8840FDBBFBF2225A" authorityName="Anker & Dworschak" authorityYear="2007" box="[886,1043,547,573]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="dahabensis">
|
||
<emphasis id="B914EA88FFB44B4D8840FDBBFBF2225A" box="[886,1043,547,573]" italics="true" pageId="10" pageNumber="407">R. dahabensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C604D19FFB44B4D8F7CFDBBFB21225A" authorityName="Anker & Jeng" authorityYear="2006" box="[1098,1216,547,573]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB44B4D8F7CFDBBFB21225A" box="[1098,1216,547,573]" italics="true" pageId="10" pageNumber="407">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
in the less elongate antennular peduncle, the more distally placed tip of the antennular stylocerite, which reaches nearly the distal margin of the peduncular article 1 (
|
||
<bibRefCitation id="EFF14B6BFFB44B4D8ACBFDF3FD6922E2" author="Anker, A." box="[509,648,619,645]" pageId="10" pageNumber="407" pagination="59 - 68" refId="ref5038" refString="Anker, A. (2011) Description of a third species in the alpheid shrimp genus Richalpheus Anker & Jeng, 2006 (Crustacea, Decapoda). Zootaxa, 3059 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 3059.1.4" type="journal article" year="2011">Anker 2011</bibRefCitation>
|
||
: fig. 1A), the more deeply incised distolateral margin of the uropodal exopod (
|
||
<bibRefCitation id="EFF14B6BFFB44B4D8BCCFD17FE6722CE" author="Anker, A." box="[250,390,655,681]" pageId="10" pageNumber="407" pagination="59 - 68" refId="ref5038" refString="Anker, A. (2011) Description of a third species in the alpheid shrimp genus Richalpheus Anker & Jeng, 2006 (Crustacea, Decapoda). Zootaxa, 3059 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 3059.1.4" type="journal article" year="2011">Anker 2011</bibRefCitation>
|
||
: fig. 1N), and the ventromesial carina of the article 1 of the antennular peduncle forming a blunt convexity (
|
||
<bibRefCitation id="EFF14B6BFFB44B4D8A63FD2BFE3F22AA" author="Anker, A." box="[341,478,691,717]" pageId="10" pageNumber="407" pagination="59 - 68" refId="ref5038" refString="Anker, A. (2011) Description of a third species in the alpheid shrimp genus Richalpheus Anker & Jeng, 2006 (Crustacea, Decapoda). Zootaxa, 3059 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 3059.1.4" type="journal article" year="2011">Anker 2011</bibRefCitation>
|
||
: fig. 1E).
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB44B4D8BF1FD4FFECE233A" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="407">
|
||
16S sequences (502 bp) were obtained from the
|
||
<specimenCount id="9D66FD13FFB44B4D89EDFD4FFC972296" box="[731,886,727,753]" pageId="10" pageNumber="407" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
of the new species (KAUM-AT 902, female from Kiire, and KAUM-AT 1352, female from Kimotsuki). They are very similar for one another with only one base pair difference. The genetic divergence between
|
||
<taxonomicName id="4C604D19FFB44B4D89B5FC87FD18235E" authorityName="Anker & Jeng" authorityYear="2006" box="[643,761,799,825]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB44B4D89B5FC87FD18235E" box="[643,761,799,825]" italics="true" pageId="10" pageNumber="407">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
is 11.8–12.1%, well supporting that the two taxa are specifically distinct.
|
||
</paragraph>
|
||
<paragraph id="8BDF369AFFB44B4D8BF1FCFFFF11238A" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="407">
|
||
The present new species is the first representative of
|
||
<taxonomicName id="4C604D19FFB44B4D882BFCFFFC4623E6" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[797,935,871,897]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B914EA88FFB44B4D882BFCFFFC4623E6" box="[797,935,871,897]" italics="true" pageId="10" pageNumber="407">Richalpheus</emphasis>
|
||
</taxonomicName>
|
||
to be recorded from
|
||
<collectingCountry id="F377760AFFB44B4D8FA0FCFFFB3723E6" box="[1174,1238,871,897]" name="Japan" pageId="10" pageNumber="407">Japan</collectingCountry>
|
||
with certainty. In Japan's red list of marine creatures (
|
||
<bibRefCitation id="EFF14B6BFFB44B4D891FFC13FB6423C2" author="Ministry of Environment & Government of Japan" box="[553,1157,907,933]" pageId="10" pageNumber="407" refId="ref5623" refString="Ministry of Environment, Government of Japan (2017) Crustacea. In: Japan's Red List of Marine Creatures. Available from: https: // www. env. go. jp / press / files / jp / 106405. pdf (accessed 21 February 2022)" type="book" year="2017">
|
||
Ministry of Environment, Government of
|
||
<collectingCountry id="F377760AFFB44B4D8F35FC13FBA223C2" box="[1027,1091,907,933]" name="Japan" pageId="10" pageNumber="407">Japan</collectingCountry>
|
||
2017
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C604D19FFB44B4D8FAFFC13FAEE23C2" authorityName="Anker & Jeng" authorityYear="2006" box="[1177,1295,907,933]" class="Malacostraca" family="Alpheidae" genus="Richalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="palmeri">
|
||
<emphasis id="B914EA88FFB44B4D8FAFFC13FAEE23C2" box="[1177,1295,907,933]" italics="true" pageId="10" pageNumber="407">R. palmeri</emphasis>
|
||
</taxonomicName>
|
||
is listed and ranked as DD (data deficient), but no information on the occurrence is provided or no voucher specimens could be located.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A6511FFB44B4D8BF1FC6FFEF5243E" pageId="10" pageNumber="407" type="etymology">
|
||
<paragraph id="8BDF369AFFB44B4D8BF1FC6FFEF5243E" blockId="10.[151,1437,151,1113]" pageId="10" pageNumber="407">
|
||
<emphasis id="B914EA88FFB44B4D8BF1FC6FFEAF2476" bold="true" box="[199,334,1015,1041]" pageId="10" pageNumber="407">Etymology.</emphasis>
|
||
From the combination of the Latin words,
|
||
<emphasis id="B914EA88FFB44B4D880AFC6FFC9B2476" box="[828,890,1015,1041]" italics="true" pageId="10" pageNumber="407">laeua</emphasis>
|
||
(= bulge) and
|
||
<emphasis id="B914EA88FFB44B4D8F10FC6FFB932476" box="[1062,1138,1015,1041]" italics="true" pageId="10" pageNumber="407">digitus</emphasis>
|
||
(= finger), in reference to the bulge-like ridge on the occlusal margin of the major cheliped dactylus in the new species. Used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |