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<document id="E0BB6693DE23D4954A01F8488940CDD9" ID-CLB-Dataset="26761" ID-DOI="10.11646/zootaxa.4394.1.2" ID-GBIF-Dataset="96807e95-89bd-4f9c-8e64-dd44cf0ca98a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3066097" ID-ZooBank="6AEE9169-0FC2-4728-A690-52FFA1707FC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558368195341" checkinUser="plazi" docAuthor="Heikkilä, Maria, Brown, John W., Baixeras, Joaquin, Mey, Wolfram &amp; Kozlov, Mikhail V." docDate="2018" docId="03B2FF08FFCE1402FF548289165FFF17" docLanguage="en" docName="zootaxa.4394.1.2.pdf.imf" docOrigin="Zootaxa 4394 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Antiquatortia Heikkilä &amp; Brown &amp; Baixeras &amp; Mey &amp; Kozlov 2018, gen. nov." docType="treatment" docVersion="10" lastPageNumber="49" masterDocId="FF8B8770FFC9140AFFC386241322FFC2" masterDocTitle="Re-examining the rare and the lost: a review of fossil Tortricidae (Lepidoptera)" masterLastPageNumber="60" masterPageNumber="41" pageNumber="48" updateTime="1698731243689" updateUser="ExternalLinkService">
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<mods:title id="62B24624445F2F65157BC642F16E0F67">Re-examining the rare and the lost: a review of fossil Tortricidae (Lepidoptera)</mods:title>
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<mods:namePart id="36FC5FB3DF1FE0051185F89EDC7551D1">Heikkilä, Maria</mods:namePart>
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<mods:namePart id="861F82CC05E9EB1A9C6A675E084AB0AA">Brown, John W.</mods:namePart>
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<mods:namePart id="F2BD2B10809FDD4945E5FD823314451A">Baixeras, Joaquin</mods:namePart>
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<mods:namePart id="F1E145F3D6AE889714AF4328985A3F3D">Mey, Wolfram</mods:namePart>
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<mods:namePart id="09A19D14A6C37A96C5D5CC0DCA564D79">Kozlov, Mikhail V.</mods:namePart>
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<mods:date id="56D603DE49D2D9DF3CF007E690646E1F">2018</mods:date>
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<treatment id="03B2FF08FFCE1402FF548289165FFF17" ID-GBIF-Taxon="157232478" LSID="urn:lsid:plazi:treatment:03B2FF08FFCE1402FF548289165FFF17" httpUri="http://treatment.plazi.org/id/03B2FF08FFCE1402FF548289165FFF17" lastPageId="8" lastPageNumber="49" pageId="7" pageNumber="48">
<subSubSection id="C3011D95FFCE140DFF5482891184FB05" box="[151,678,1197,1223]" pageId="7" pageNumber="48" type="nomenclature">
<paragraph id="8BA44E1EFFCE140DFF5482891184FB05" blockId="7.[151,678,1197,1223]" box="[151,678,1197,1223]" pageId="7" pageNumber="48">
<heading id="D0ECF972FFCE140DFF5482891184FB05" bold="true" box="[151,678,1197,1223]" fontSize="11" level="1" pageId="7" pageNumber="48" reason="1">
<emphasis id="B96F920CFFCE140DFF5482891184FB05" bold="true" box="[151,678,1197,1223]" pageId="7" pageNumber="48">
<taxonomicName id="4C1B359DFFCE140DFF5482891109FB05" authority="Brown &amp; Baixeras" authorityName="Heikkilä &amp; Brown &amp; Baixeras &amp; Mey &amp; Kozlov" authorityYear="2018" box="[151,555,1197,1223]" class="Insecta" family="Tortricidae" genus="Antiquatortia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96F920CFFCE140DFF5482891218FB05" bold="true" box="[151,314,1197,1223]" italics="true" pageId="7" pageNumber="48">Antiquatortia</emphasis>
Brown &amp; Baixeras
</taxonomicName>
,
<taxonomicNameLabel id="A25C2F77FFCE140DFDFB82891184FB05" box="[568,678,1197,1223]" pageId="7" pageNumber="48" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3011D95FFCE140DFF5482D11066FAC8" box="[151,836,1268,1290]" pageId="7" pageNumber="48" type="materials_examined">
<paragraph id="8BA44E1EFFCE140DFF5482D11066FAC8" blockId="7.[151,836,1268,1290]" box="[151,836,1268,1290]" pageId="7" pageNumber="48">
<typeStatus id="54A0F0BCFFCE140DFF5482D113EBFAC8" box="[151,201,1269,1290]" pageId="7" pageNumber="48">Type</typeStatus>
species:
<taxonomicName id="4C1B359DFFCE140DFEE582D111C9FAC8" authority="Brown &amp; Baixeras" authorityName="Heikkilä &amp; Brown &amp; Baixeras &amp; Mey &amp; Kozlov" authorityYear="2018" box="[294,747,1268,1290]" class="Insecta" family="Tortricidae" genus="Antiquatortia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="histuroides" status="sp. nov.">
<emphasis id="B96F920CFFCE140DFEE582D11107FAC8" box="[294,549,1268,1290]" italics="true" pageId="7" pageNumber="48">Antiquatortia histuroides</emphasis>
Brown &amp; Baixeras
</taxonomicName>
,
<taxonomicNameLabel id="A25C2F77FFCE140DFD3682D11066FAC8" box="[757,836,1269,1290]" pageId="7" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3011D95FFCE140DFF548312172BF926" pageId="7" pageNumber="48" type="description">
<paragraph id="8BA44E1EFFCE140DFF548312160FFA1D" blockId="7.[151,1437,1334,2016]" pageId="7" pageNumber="48">
<emphasis id="B96F920CFFCE140DFF548312120BFA8D" bold="true" box="[151,297,1334,1359]" pageId="7" pageNumber="48">Description.</emphasis>
Head: Vertex rough-scaled, frons with narrow scales. Antenna slender, slightly serrate, ca 0.4 times length of forewing, lacking conspicuous sensory setae, with two rows of scales per flagellomere. Ocelli present. Chaetosemata not visible. Labial palpus long, porrect; second segment ca 4.0 times length of first and third segment, expanded medially on dorsal side, third segment slender, rod-like, slightly tapering apically. Maxillary palpus not visible. Haustellum well developed, coiled, unscaled at base, with stout sensilla on distal half.
</paragraph>
<paragraph id="8BA44E1EFFCE140DFF0483CE1245F902" blockId="7.[151,1437,1334,2016]" pageId="7" pageNumber="48">
Thorax: Foreleg tibia with epiphysis; mid- and hindtibia with two pairs of spurs, the more medial spur longer; tarsus with apical spines on ventral side. Forewing discal cell length about 0.6 forewing length; all veins (Sc, R
<subScript id="179F4C5BFFCE140DFAA9803A1650F9EE" attach="left" box="[1386,1394,1566,1580]" fontSize="6" pageId="7" pageNumber="48">1</subScript>
-R
<subScript id="179F4C5BFFCE140DFA4E803A16B7F9EE" attach="left" box="[1421,1429,1566,1580]" fontSize="6" pageId="7" pageNumber="48">5</subScript>
, M
<subScript id="179F4C5BFFCE140DFF738060139AF990" attach="left" box="[176,184,1604,1618]" fontSize="6" pageId="7" pageNumber="48">1</subScript>
-M
<subScript id="179F4C5BFFCE140DFF19806013C0F990" attach="left" box="[218,226,1604,1618]" fontSize="6" pageId="7" pageNumber="48">3</subScript>
, CuA
<subScript id="179F4C5BFFCE140DFEE58060120CF990" attach="left" box="[294,302,1604,1618]" fontSize="6" pageId="7" pageNumber="48">1</subScript>
-CuA
<subScript id="179F4C5BFFCE140DFEAF80601256F990" attach="left" box="[364,372,1604,1618]" fontSize="6" pageId="7" pageNumber="48">2</subScript>
and 1A+2A) present and separate, R
<subScript id="179F4C5BFFCE140DFCD880601001F990" attach="left" box="[795,803,1604,1618]" fontSize="6" pageId="7" pageNumber="48">5</subScript>
extending to costa; neither chorda nor M-stem evident. Patches of upraised scales on the forewing. Hindwing with costa slightly sinuate, with a concave region preapically (
<figureCitation id="1320529BFFCE140DFF5C80A413C5F95B" box="[159,231,1664,1689]" captionStart="FIGURE 2" captionStartId="8.[151,250,1522,1544]" captionTargetBox="[154,1433,266,1499]" captionTargetId="figure@8.[153,1434,265,1500]" captionTargetPageId="8" captionText="FIGURE 2. Antiquatortia histuroides Brown &amp; Baixeras, gen. nov., sp. nov. AMNH DR8-43. Dominican amber. a. Ventral view. Length of forewing 7 mm. b. Wing venation c. Dorsal view of forewing with tufts of erect scales. d. Papillae anales. Photos: Mark Metz. Illustration: Joaquin Baixeras." httpUri="https://zenodo.org/record/3094741/files/figure.png" pageId="7" pageNumber="48">Figs 2</figureCitation>
ab); Sc+R
<subScript id="179F4C5BFFCE140DFEB280AB125BF95F" attach="left" box="[369,377,1679,1693]" fontSize="6" pageId="7" pageNumber="48">1</subScript>
and Rs approximate at base, M
<subScript id="179F4C5BFFCE140DFD2580AB11CCF95F" attach="left" box="[742,750,1679,1693]" fontSize="6" pageId="7" pageNumber="48">2</subScript>
and M
<subScript id="179F4C5BFFCE140DFC8280AB106BF95F" attach="left" box="[833,841,1679,1693]" fontSize="6" pageId="7" pageNumber="48">3</subScript>
apparently parallel; M
<subScript id="179F4C5BFFCE140DFB8D80AB1774F95F" attach="left" box="[1102,1110,1679,1693]" fontSize="6" pageId="7" pageNumber="48">3</subScript>
and CuA
<subScript id="179F4C5BFFCE140DFB0780AB17EEF95F" attach="left" box="[1220,1228,1679,1693]" fontSize="6" pageId="7" pageNumber="48">1</subScript>
stalked; frenulum with three acanthi.
</paragraph>
<paragraph id="8BA44E1EFFCE140DFF0480EF172BF926" blockId="7.[151,1437,1334,2016]" box="[199,1033,1739,1764]" pageId="7" pageNumber="48">Abdomen: Female genitalia with papillae anales weak, but apparently flat.</paragraph>
</subSubSection>
<subSubSection id="C3011D95FFCE140DFF0480CB12C6F81D" pageId="7" pageNumber="48" type="diagnosis">
<paragraph id="8BA44E1EFFCE140DFF0480CB12C6F81D" blockId="7.[151,1437,1334,2016]" pageId="7" pageNumber="48">
<emphasis id="B96F920CFFCE140DFF0480CB1263F8CA" bold="true" box="[199,321,1775,1800]" pageId="7" pageNumber="48">Diagnosis.</emphasis>
In wing venation
<taxonomicName id="4C1B359DFFCE140DFDD380D41185F8C5" authorityName="Heikkilä &amp; Brown &amp; Baixeras &amp; Mey &amp; Kozlov" authorityYear="2018" box="[528,679,1776,1799]" class="Insecta" family="Tortricidae" genus="Antiquatortia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B96F920CFFCE140DFDD380D41185F8C5" box="[528,679,1776,1799]" italics="true" pageId="7" pageNumber="48">Antiquatortia</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C1B359DFFCE140DFCFF80CB1086F8CA" authorityName="Poincar &amp; Brown" authorityYear="1993" box="[828,932,1775,1800]" class="Insecta" family="Tortricidae" genus="Polyvena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B96F920CFFCE140DFCFF80CB1086F8CA" box="[828,932,1775,1800]" italics="true" pageId="7" pageNumber="48">Polyvena</emphasis>
</taxonomicName>
, but in
<taxonomicName id="4C1B359DFFCE140DFC3F80CB1746F8CA" authorityName="Poincar &amp; Brown" authorityYear="1993" box="[1020,1124,1775,1800]" class="Insecta" family="Tortricidae" genus="Polyvena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B96F920CFFCE140DFC3F80CB1746F8CA" box="[1020,1124,1775,1800]" italics="true" pageId="7" pageNumber="48">Polyvena</emphasis>
</taxonomicName>
the chorda and the M-stem are present, whereas both are absent in
<taxonomicName id="4C1B359DFFCE140DFDB781301029F8E9" authorityName="Heikkilä &amp; Brown &amp; Baixeras &amp; Mey &amp; Kozlov" authorityYear="2018" box="[628,779,1812,1835]" class="Insecta" family="Tortricidae" genus="Antiquatortia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B96F920CFFCE140DFDB781301029F8E9" box="[628,779,1812,1835]" italics="true" pageId="7" pageNumber="48">Antiquatortia</emphasis>
</taxonomicName>
. Both specimens possess patches of what appear to be upraised scales on the forewing, a feature characteristic of both sexes of
<taxonomicName id="4C1B359DFFCE140DFC3C8113175FF892" box="[1023,1149,1847,1872]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="tribe" tribe="Polyorthini">Polyorthini</taxonomicName>
and
<taxonomicName id="4C1B359DFFCE140DFB7D81131608F892" box="[1214,1322,1847,1872]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="tribe" tribe="Tortricini">Tortricini</taxonomicName>
, although occurring sporadically elsewhere in
<taxonomicName id="4C1B359DFFCE140DFDF4817F107BF8B1" authority="(Horak 1998)" authorityName="Horak" authorityYear="1998" baseAuthorityName="Horak" baseAuthorityYear="1998" box="[567,857,1883,1908]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="family">
Tortricidae (
<bibRefCitation id="EF8A33EFFFCE140DFD00817F1073F8B1" author="Horak, M." box="[707,849,1883,1908]" pageId="7" pageNumber="48" pagination="199 - 215" refId="ref12685" refString="Horak, M. (1998) The Tortricoidea. In: Kristensen, N. P. (Ed.), Lepidoptera, moths and butterflies, 1 Evolution, systematics and biogeography. Handbuch der Zoologie / Handbook of Zoology 35. Walter de Gruyter, Berlin, New York, pp. 199 - 215." type="book chapter" year="1998">Horak 1998</bibRefCitation>
)
</taxonomicName>
. Because the forewing pattern is obscured in both fossils, no comparison of pattern can be made. In addition, because
<taxonomicName id="4C1B359DFFCE140DFC4D815B10D4F85A" authorityName="Poincar &amp; Brown" authorityYear="1993" box="[910,1014,1919,1944]" class="Insecta" family="Tortricidae" genus="Polyvena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B96F920CFFCE140DFC4D815B10D4F85A" box="[910,1014,1919,1944]" italics="true" pageId="7" pageNumber="48">Polyvena</emphasis>
</taxonomicName>
is a male and
<taxonomicName id="4C1B359DFFCE140DFB5481A4160CF855" authorityName="Heikkilä &amp; Brown &amp; Baixeras &amp; Mey &amp; Kozlov" authorityYear="2018" box="[1175,1326,1920,1943]" class="Insecta" family="Tortricidae" genus="Antiquatortia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B96F920CFFCE140DFB5481A4160CF855" box="[1175,1326,1920,1943]" italics="true" pageId="7" pageNumber="48">Antiquatortia</emphasis>
</taxonomicName>
a female, no comparisons regarding other external features are particularly relevant (i.e., number of acanthi in the frenulum, male secondary features, etc.)
</paragraph>
</subSubSection>
<subSubSection id="C3011D95FFC11402FF0486B3165FFF17" pageId="8" pageNumber="49" type="etymology">
<paragraph id="8BA44E1EFFC11402FF0486B316BEFF72" blockId="8.[151,1436,151,213]" box="[199,1436,151,176]" pageId="8" pageNumber="49">
<emphasis id="B96F920CFFC11402FF0486B3126CFF72" bold="true" box="[199,334,151,176]" pageId="8" pageNumber="49">Etymology.</emphasis>
The genus name is a Latinized derivative name freely inspired in the contraction of the Latin word
</paragraph>
<paragraph id="8BA44E1EFFC11402FF548698165FFF17" blockId="8.[151,1436,151,213]" box="[151,1405,188,213]" pageId="8" pageNumber="49">
“antiqua” (meaning old, ancient) and the
<typeStatus id="54A0F0BCFFC11402FDA5869911B4FF17" box="[614,662,189,213]" pageId="8" pageNumber="49">type</typeStatus>
genus of the family
<taxonomicName id="4C1B359DFFC11402FCBE869810DAFF17" baseAuthorityName="Horak" baseAuthorityYear="1998" box="[893,1016,188,213]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
<taxonomicName id="4C1B359DFFC11402FBC986991775FF16" box="[1034,1111,189,212]" class="Insecta" family="Tortricidae" genus="Tortrix" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B96F920CFFC11402FBC986991775FF16" box="[1034,1111,189,212]" italics="true" pageId="8" pageNumber="49">Tortrix</emphasis>
</taxonomicName>
”. It is feminine in gender.
</paragraph>
</subSubSection>
</treatment>
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