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<document id="D03C58ACC0BD1EAD73BAF3D17EBFC7EA" ID-DOI="10.11646/zootaxa.3986.4.4" ID-GBIF-Dataset="e2fa9510-b1da-46f6-9387-2b5ed055d63f" ID-ISSN="1175-5326" ID-Zenodo-Dep="239739" ID-ZooBank="D8055FFB-5890-4273-8230-F899EA0D8478" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461264358145" checkinUser="plazi" docAuthor="Bukejs, Andris, Alekseev, Vitalii I. &amp; Jäch, Manfred A." docDate="2015" docId="03A887D3DD2E582C1D99CD7636FBF922" docLanguage="en" docName="zt03986p460.pdf" docOrigin="Zootaxa 3986 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterlimnius Hinton 1935" docType="treatment" docVersion="7" lastPageNumber="453" masterDocId="FF91FFABDD2F582D1D0EC97C3536FFCF" masterDocTitle="The riffle beetles (Coleoptera: Elmidae) of the Eocene Baltic amber: Heterelmis groehni sp. nov. and Heterlimnius samlandicus (Bollow, 1940) comb. nov." masterLastPageNumber="460" masterPageNumber="452" pageNumber="453" updateTime="1698610159723" updateUser="plazi">
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<mods:title id="C1588CA3D958F16A53052B0169F20C29">The riffle beetles (Coleoptera: Elmidae) of the Eocene Baltic amber: Heterelmis groehni sp. nov. and Heterlimnius samlandicus (Bollow, 1940) comb. nov.</mods:title>
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<mods:namePart id="DE2AB15EDE53F764AB8BFACB4135A636">Bukejs, Andris</mods:namePart>
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<mods:namePart id="DF05E5C845DB7534B0577BAAF10578AE">Alekseev, Vitalii I.</mods:namePart>
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<mods:namePart id="A4466B61BE2F9405742FC358FFA84CBF">Jäch, Manfred A.</mods:namePart>
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<mods:title id="654A0716364A570362C7FDF70477AB1A">Zootaxa</mods:title>
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<mods:date id="D571F906C47D97825754A5398E9FDEBB">2015</mods:date>
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<mods:number id="F75B6EEB17D9E6D2CC42D3D6477A2C86">3986</mods:number>
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<mods:identifier id="2965FA2B3373BAFBA3E86CC120C2367F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A887D3DD2E582C1D99CD7636FBF922" ID-DOI="http://doi.org/10.5281/zenodo.6114208" ID-GBIF-Taxon="119637411" ID-Zenodo-Dep="6114208" LSID="urn:lsid:plazi:treatment:03A887D3DD2E582C1D99CD7636FBF922" httpUri="http://treatment.plazi.org/id/03A887D3DD2E582C1D99CD7636FBF922" lastPageNumber="453" pageId="1" pageNumber="453">
<subSubSection id="C31B654EDD2E582C1D99CD763723FBEB" box="[151,533,1034,1060]" pageId="1" pageNumber="453" type="nomenclature">
<paragraph id="8BBE36C5DD2E582C1D99CD763723FBEB" blockId="1.[151,533,1034,1060]" box="[151,533,1034,1060]" pageId="1" pageNumber="453">
<heading id="D0F681A9DD2E582C1D99CD763723FBEB" bold="true" box="[151,533,1034,1060]" fontSize="11" level="1" pageId="1" pageNumber="453" reason="1">
<treatmentCitationGroup id="AB1111EBDD2E582C1D99CD763723FBEB" box="[151,533,1034,1060]" pageId="1" pageNumber="453">
<taxonomicName id="4C014D46DD2E582C1D99CD763723FBEB" ID-CoL="4W94" authority="Hinton 1935: 178" authorityName="Hinton" authorityPageNumber="178" authorityYear="1935" box="[151,533,1034,1060]" class="Insecta" family="Elmidae" genus="Heterlimnius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1D99CD763723FBEB" bold="true" box="[151,533,1034,1060]" pageId="1" pageNumber="453">
<emphasis id="B975EAD7DD2E582C1D99CD763401FBEB" bold="true" box="[151,311,1034,1060]" italics="true" pageId="1" pageNumber="453">Heterlimnius</emphasis>
<treatmentCitation id="0AA010D4DD2E582C1C30CD763723FBEB" author="Hinton" box="[318,533,1034,1060]" page="178" pageId="1" pageNumber="453" year="1935">
<bibRefCitation id="EF904B34DD2E582C1C30CD7634EAFBEB" author="Hinton" box="[318,476,1034,1060]" pageId="1" pageNumber="453" refString="Hinton, H. E. (1935) Notes on the Dryopoidea (Col.). Stylops, 4 (8), 169 - 179." type="journal article" year="1935">Hinton 1935</bibRefCitation>
: 178
</treatmentCitation>
</emphasis>
</taxonomicName>
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</heading>
</paragraph>
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<subSubSection id="C31B654EDD2E582C1D99CD2D36FBF922" pageId="1" pageNumber="453" type="reference_group">
<paragraph id="8BBE36C5DD2E582C1D99CD2D3198FBA8" blockId="1.[151,1198,1105,1127]" box="[151,1198,1105,1127]" pageId="1" pageNumber="453">
<treatmentCitationGroup id="AB1111EBDD2E582C1D99CD2D3198FBA8" box="[151,1198,1105,1127]" pageId="1" pageNumber="453">
<taxonomicName id="4C014D46DD2E582C1D99CD2D34CCFBA8" authority="Bollow 1940: 117" authorityName="Bollow" authorityPageNumber="117" authorityYear="1940" box="[151,506,1105,1127]" class="Insecta" family="Dryopidae" genus="Palaeoriohelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1D99CD2D3408FBA9" box="[151,318,1105,1126]" italics="true" pageId="1" pageNumber="453">Palaeoriohelmis</emphasis>
<treatmentCitation id="0AA010D4DD2E582C1C4ACD2D34CCFBA8" author="Bollow" box="[324,506,1105,1127]" page="117" pageId="1" pageNumber="453" year="1940">
<bibRefCitation id="EF904B34DD2E582C1C4ACD2D34FDFBA8" author="Bollow" box="[324,459,1105,1127]" pageId="1" pageNumber="453" refString="Bollow, H. (1940) Die erste Helminidae (Col, Dryop.) aus Bernstein. Mitteilungen der Munchner Entomologischen Gesellschaft, 30, 117 - 119." type="journal article" year="1940">Bollow 1940</bibRefCitation>
: 117
</treatmentCitation>
</taxonomicName>
<emphasis id="B975EAD7DD2E582C1F0FCD2D3768FBA9" bold="true" box="[513,606,1105,1126]" pageId="1" pageNumber="453">syn. nov.</emphasis>
(
<typeStatus id="54BA8867DD2E582C1F65CD2D37AEFBA8" box="[619,664,1105,1127]" pageId="1" pageNumber="453">type</typeStatus>
species:
<taxonomicName id="4C014D46DD2E582C1FFBCD2D319EFBA8" authority="Bollow, 1940" authorityName="Bollow" authorityYear="1940" box="[757,1192,1105,1127]" class="Insecta" family="Dryopidae" genus="Palaeoriohelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="species" species="samlandica">
<emphasis id="B975EAD7DD2E582C1FFBCD2D312EFBA9" box="[757,1048,1105,1126]" italics="true" pageId="1" pageNumber="453">Palaeoriohelmis samlandica</emphasis>
<bibRefCitation id="EF904B34DD2E582C1910CD2D319EFBA8" author="Bollow" box="[1054,1192,1105,1127]" pageId="1" pageNumber="453" refString="Bollow, H. (1940) Die erste Helminidae (Col, Dryop.) aus Bernstein. Mitteilungen der Munchner Entomologischen Gesellschaft, 30, 117 - 119." type="journal article" year="1940">Bollow, 1940</bibRefCitation>
</taxonomicName>
)
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</paragraph>
<paragraph id="8BBE36C5DD2E582C1D99CDE836DFFAF2" blockId="1.[151,1436,1172,1773]" pageId="1" pageNumber="453">
The genus
<taxonomicName id="4C014D46DD2E582C1C19CDE834E6FB63" box="[279,464,1172,1196]" class="Insecta" family="Dryopidae" genus="Palaeoriohelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1C19CDE834E6FB63" box="[279,464,1172,1196]" italics="true" pageId="1" pageNumber="453">Palaeoriohelmis</emphasis>
</taxonomicName>
is herewith synonymized with the genus
<taxonomicName id="4C014D46DD2E582C1EBDCDE83173FB63" ID-CoL="4W94" box="[947,1093,1172,1196]" class="Insecta" family="Elmidae" genus="Heterlimnius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1EBDCDE83173FB63" box="[947,1093,1172,1196]" italics="true" pageId="1" pageNumber="453">Heterlimnius</emphasis>
</taxonomicName>
on the basis of the following morphological characteristics: (1) subovate body with plastron mainly on ventral surface; (2) filiform 11- segmented antennae with symmetrical antennomere 11; (3) pronotum with basal sublateral longitudinal grooves and with very shallow basal transverse depression; (4) prosternum produced anteriorly; (5) elytra with uniform, slightly convex, not carinate intervals; (6) tarsal claws simple, curved, long.
</paragraph>
<paragraph id="8BBE36C5DD2E582C1DC9CC3B3461FA20" blockId="1.[151,1436,1172,1773]" pageId="1" pageNumber="453">
<bibRefCitation id="EF904B34DD2E582C1DC9CC3B345AFAAF" author="Bollow" box="[199,364,1351,1376]" pageId="1" pageNumber="453" refString="Bollow, H. (1940) Die erste Helminidae (Col, Dryop.) aus Bernstein. Mitteilungen der Munchner Entomologischen Gesellschaft, 30, 117 - 119." type="journal article" year="1940">Bollow (1940)</bibRefCitation>
compared
<taxonomicName id="4C014D46DD2E582C1CE4CC343795FAAF" box="[490,675,1352,1376]" class="Insecta" family="Dryopidae" genus="Palaeoriohelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1CE4CC343795FAAF" box="[490,675,1352,1376]" italics="true" pageId="1" pageNumber="453">Palaeoriohelmis</emphasis>
</taxonomicName>
with two extant West Palearctic genera:
<taxonomicName id="4C014D46DD2E582C197DCC34304DFAAF" authority="Reitter, 1883" authorityName="Reitter" authorityYear="1883" box="[1139,1403,1351,1376]" class="Insecta" family="Dryopidae" genus="Lathelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C197DCC3431D7FAAF" box="[1139,1249,1352,1376]" italics="true" pageId="1" pageNumber="453">Lathelmis</emphasis>
Reitter, 1883
</taxonomicName>
[=
<taxonomicName id="4C014D46DD2E582C1D99CC1134BDFA4B" ID-CoL="62YJ4" authority="Illiger, 1802" authorityName="Illiger" authorityYear="1802" box="[151,395,1388,1413]" class="Insecta" family="Elmidae" genus="Limnius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1D99CC1135C7FA4B" box="[151,241,1389,1412]" italics="true" pageId="1" pageNumber="453">Limnius</emphasis>
Illiger, 1802
</taxonomicName>
] and
<taxonomicName id="4C014D46DD2E582C1CC0CC10362BFA4B" ID-CoL="643QN" authority="Mulsant &amp; Rey, 1872" authorityName="Mulsant &amp; Rey" authorityYear="1872" box="[462,797,1388,1413]" class="Insecta" family="Elmidae" genus="Riolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1CC0CC103720FA4B" box="[462,534,1388,1412]" italics="true" pageId="1" pageNumber="453">Riolus</emphasis>
Mulsant &amp; Rey, 1872
</taxonomicName>
, stating that the new genus was related with
<taxonomicName id="4C014D46DD2E582C1820CC103040FA4B" ID-CoL="643QN" box="[1326,1398,1388,1412]" class="Insecta" family="Elmidae" genus="Riolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1820CC103040FA4B" box="[1326,1398,1388,1412]" italics="true" pageId="1" pageNumber="453">Riolus</emphasis>
</taxonomicName>
on account of the lacking “eingeschnittenen Dorsalstreifen des Halsschildes” [grooves of the pronotum]. However, examination of the
<typeStatus id="54BA8867DD2E582C1C70CCC834D6FA02" box="[382,480,1460,1485]" pageId="1" pageNumber="453" type="holotype">holotype</typeStatus>
revealed that a pair of basal sublateral longitudinal grooves is indeed present in
<taxonomicName id="4C014D46DD2E582C1D99CCA43466FA3F" box="[151,336,1496,1520]" class="Insecta" family="Dryopidae" genus="Palaeoriohelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1D99CCA43466FA3F" box="[151,336,1496,1520]" italics="true" pageId="1" pageNumber="453">Palaeoriohelmis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBE36C5DD2E582C1DC9CC8136FBF922" blockId="1.[151,1436,1172,1773]" pageId="1" pageNumber="453">
It should be mentioned here that
<taxonomicName id="4C014D46DD2E582C1F3ACC8037F0F9DB" ID-CoL="4W94" box="[564,710,1532,1556]" class="Insecta" family="Elmidae" genus="Heterlimnius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1F3ACC8037F0F9DB" box="[564,710,1532,1556]" italics="true" pageId="1" pageNumber="453">Heterlimnius</emphasis>
</taxonomicName>
cannot be distinguished reliably from
<taxonomicName id="4C014D46DD2E582C197DCC8031CCF9DC" ID-CoL="63FPZ" box="[1139,1274,1532,1555]" class="Insecta" family="Elmidae" genus="Optioservus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C197DCC8031CCF9DC" box="[1139,1274,1532,1555]" italics="true" pageId="1" pageNumber="453">Optioservus</emphasis>
</taxonomicName>
. These genera are very similar in general morphology and distribution (both occur in the East Palearctic, Nearctic and northern Oriental Regions), and members of both genera have been historically confused with one another. At present, 14 species are recorded in
<taxonomicName id="4C014D46DD2E582C1C90CF143706F94F" ID-CoL="4W94" box="[414,560,1640,1664]" class="Insecta" family="Elmidae" genus="Heterlimnius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1C90CF143706F94F" box="[414,560,1640,1664]" italics="true" pageId="1" pageNumber="453">Heterlimnius</emphasis>
</taxonomicName>
, and
<quantity id="4CF99B20DD2E582C1F63CF143790F94F" box="[621,678,1639,1664]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="1" pageNumber="453" unit="in" value="19.0">19 in</quantity>
<taxonomicName id="4C014D46DD2E582C1FA3CF143602F9B0" ID-CoL="63FPZ" box="[685,820,1640,1663]" class="Insecta" family="Elmidae" genus="Optioservus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD7DD2E582C1FA3CF143602F9B0" box="[685,820,1640,1663]" italics="true" pageId="1" pageNumber="453">Optioservus</emphasis>
</taxonomicName>
. The only possibly reliable character to distinguish the two genera is provided by the ventral sclerites of the larval meso- and metathorax (
<bibRefCitation id="EF904B34DD2E582C1954CFF031C2F96B" author="Kamite" box="[1114,1268,1676,1701]" pageId="1" pageNumber="453" refString="Kamite, Y. (2013) Revision of the genus Optioservus Sanderson, part 1: Generic redescription and revision of the Nearctic species (O. fastiditus and O. quadrimaculatus species groups) (Coleoptera: Elmidae). Koleopterologische Rundschau, 83, 133 - 164." type="journal article" year="2013">Kamite 2013</bibRefCitation>
), but larvae of several species are still unknown. Quite possibly, these two genera will have to be synonymized following comprehensive molecular studies, which have not been carried out so far.
</paragraph>
</subSubSection>
</treatment>
</document>
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