243 lines
30 KiB
XML
243 lines
30 KiB
XML
<document id="E5D2119F12B22780E33F15C7FF32C91D" ID-DOI="10.11646/zootaxa.5002.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5122655" ID-ZooBank="5DC2A66A-3F04-42D0-8A8C-F0686054E556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626888359662" checkinUser="plazi" docAuthor="Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott" docDate="2021" docId="03A887D05F6CFFB9FF3EEE3AFF1A4DB5" docLanguage="en" docName="zootaxa.5002.1.1.pdf" docOrigin="Zootaxa 5002 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomosvaryella nimroozensis Motamedinia & Skevington, sp. nov." docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="FF91FFA85F42FF89FFA9EB42FFAF4F30" masterDocTitle="Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species" masterLastPageNumber="103" masterPageNumber="1" pageNumber="47" updateTime="1699012478263" updateUser="plazi">
|
||
<mods:mods id="07181405A7A0B7572B1744D0898D32E5" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="851D13632F6E18E99324BE286EDCF467">
|
||
<mods:title id="19AEF762997F76A975A344E39D020664">Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="EA2875D6CD4654F0A3A3E0CD41579A72" type="personal">
|
||
<mods:role id="C26BC8E6589C151B746379190F5FC96A">
|
||
<mods:roleTerm id="7CAE6EF33991E9713EBA8C30D5E21C2C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A53B578252D983E4F21E19F3892C87D3">Motamedinia, Behnam</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="805E46A8233B3A02D03E60204664AAE1" type="personal">
|
||
<mods:role id="581126B64984909EBB509AAC993A9504">
|
||
<mods:roleTerm id="5E30763037EF2212288E94467A6C2770">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7F3DFE503B0E9A1E8972755412CD27FD">Skevington, Jeffrey H.</mods:namePart>
|
||
<mods:nameIdentifier id="38FA8488A19312C1E9B7EA134BD5757C" type="ORCID">0000-0002-1445-9870</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="E77437DD6395759593336AC79BEB67F2" type="email">jhskevington@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="022D310A2121A587C4062B0549EF4BB0" type="personal">
|
||
<mods:role id="728D7AB3B1411CF8E6B0F79198AE126F">
|
||
<mods:roleTerm id="83CDBD30C3C55C9B4719145FE848C9F8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="EBF1D7A54971FFA5F295FEABE0B823F0">Kelso, Scott</mods:namePart>
|
||
<mods:nameIdentifier id="149D66776F618A194C5ED13926015ECA" type="ORCID">0000-0001-5559-9477</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="19C07556FBC60E50E335EB28EBB02C0F" type="email">scott.kelso@canada.ca</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="EB796165C35596F7400063FDFAB8C469">text</mods:typeOfResource>
|
||
<mods:relatedItem id="9FA63E39BFF6DB35B89EC9BD61D93629" type="host">
|
||
<mods:titleInfo id="46F0E6B90AA422B0CD57A043160D1F33">
|
||
<mods:title id="886B41269B7C678D3A931B2F3DBF6F8C">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="DDFF23B83C6EB4C2B3BD9610BEB385EF">
|
||
<mods:date id="8B33B5A78C1877442F8460A9EBF0F429">2021</mods:date>
|
||
<mods:detail id="4F181EA047B4F8E4C8DF8F7DDE147D22" type="pubDate">
|
||
<mods:number id="6170D4D5542483F67B7AA2F215D26ACB">2021-07-16</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="8246E045277FBF066D8301C085962E66" type="volume">
|
||
<mods:number id="C28DF7E745BCA8340BF07585048759CF">5002</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="339473C97C4B3813F9BB12997A6164C7" type="issue">
|
||
<mods:number id="DA48AD5D69BD1C03B5A73695C8E55D62">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="1AA48D7CA7D32877E593932194CA971F" unit="page">
|
||
<mods:start id="4EC4B5D69F911BA5696B8CEE9586A842">1</mods:start>
|
||
<mods:end id="6477E9E2A82A096EB33191030046897C">103</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="2B4C198192DC51AA64C964BBF7EC7803">journal article</mods:classification>
|
||
<mods:identifier id="746DB5DC2F29520BDF65DF12A8F94E8A" type="DOI">10.11646/zootaxa.5002.1.1</mods:identifier>
|
||
<mods:identifier id="44CDC73DA3EC1D3D43C4EC32DDB165DB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="217690EFB33661105DB3364763D5B1F8" type="Zenodo-Dep">5122655</mods:identifier>
|
||
<mods:identifier id="ED006E51200B4DB9186D512292D5BCEF" type="ZooBank">5DC2A66A-3F04-42D0-8A8C-F0686054E556</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A887D05F6CFFB9FF3EEE3AFF1A4DB5" ID-DOI="http://doi.org/10.5281/zenodo.5122587" ID-Zenodo-Dep="5122587" LSID="urn:lsid:plazi:treatment:03A887D05F6CFFB9FF3EEE3AFF1A4DB5" httpUri="http://treatment.plazi.org/id/03A887D05F6CFFB9FF3EEE3AFF1A4DB5" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
|
||
<subSubSection id="C31B654D5F6CFFA7FF3EEE3AFC1A4AA3" box="[151,949,1400,1427]" pageId="46" pageNumber="47" type="nomenclature">
|
||
<paragraph id="8BBE36C65F6CFFA7FF3EEE3AFC1A4AA3" blockId="46.[151,949,1400,1463]" box="[151,949,1400,1427]" pageId="46" pageNumber="47">
|
||
<heading id="D0F681AA5F6CFFA7FF3EEE3AFC1A4AA3" bold="true" box="[151,949,1400,1427]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="3">
|
||
<emphasis id="B975EAD45F6CFFA7FF3EEE3AFC1A4AA3" bold="true" box="[151,949,1400,1427]" pageId="46" pageNumber="47">
|
||
<taxonomicName id="4C014D455F6CFFA7FF3EEE3AFCE04AA2" ID-CoL="9HKN7" authority="Motamedinia & Skevington" authorityName="Motamedinia & Skevington" box="[151,847,1400,1427]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="nimroozensis" status="sp. nov.">
|
||
<emphasis id="B975EAD45F6CFFA7FF3EEE3AFE424AA2" bold="true" box="[151,493,1400,1427]" italics="true" pageId="46" pageNumber="47">Tomosvaryella nimroozensis</emphasis>
|
||
Motamedinia & Skevington
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657AF5F6CFFA7FCFEEE3BFC1A4AA3" box="[855,949,1401,1427]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F6CFFA7FF3EEEDEFE0D4A86" box="[151,418,1436,1463]" pageId="46" pageNumber="47" type="description">
|
||
<paragraph id="8BBE36C65F6CFFA7FF3EEEDEFE0D4A86" blockId="46.[151,949,1400,1463]" box="[151,418,1436,1463]" pageId="46" pageNumber="47">
|
||
<figureCitation id="133A2A435F6CFFA7FF3EEEDEFEB24A87" box="[151,285,1436,1463]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="46" pageNumber="47">Figs 25A–F</figureCitation>
|
||
,
|
||
<figureCitation id="133A2A435F6CFFA7FE81EEDEFEE94A87" box="[296,326,1436,1463]" captionStart="FIGURE 56" captionStartId="88.[151,250,1562,1586]" captionTargetBox="[151,1436,896,1538]" captionTargetId="figure-53@88.[151,1436,896,1538]" captionTargetPageId="88" captionText="FIGURE 56. Distribution of Tomosvaryella inopinata, T. minuscula, T. nimroozensis sp. nov., T. soziana sp. nov. in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122783" httpUri="https://zenodo.org/record/5122783/files/figure.png" pageId="46" pageNumber="47">56</figureCitation>
|
||
,
|
||
<figureCitation id="133A2A435F6CFFA7FEFBEEDEFE0D4A86" box="[338,418,1436,1462]" captionStart="FIGURE 68" captionStartId="96.[151,250,1518,1542]" captionTargetBox="[151,1436,182,1494]" captionTargetId="figure-31@96.[151,1436,181,1494]" captionTargetPageId="96" captionText="FIGURE 68. Male habitus of Tomosvaryella species A–B) T. nimroozensis sp. nov., holotype (JSS51819, CNC) A) in dorsal view B) in lateral view, C–D) T. oshidae sp. nov., holotype (JSS51818, CNC) C) in dorsal view D) in lateral view, E–F) T. osteodes sp. nov., holotype (CNCD175298, CNC) E) in dorsal view F) in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122809" httpUri="https://zenodo.org/record/5122809/files/figure.png" pageId="46" pageNumber="47">68A–B</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F6CFFA7FF3EEEA6FCB44977" pageId="46" pageNumber="47" type="diagnosis">
|
||
<paragraph id="8BBE36C65F6CFFA7FF3EEEA6FCB44977" blockId="46.[151,1437,1508,2039]" pageId="46" pageNumber="47">
|
||
<emphasis id="B975EAD45F6CFFA7FF3EEEA6FEBB4ACE" bold="true" box="[151,276,1508,1534]" pageId="46" pageNumber="47">Diagnosis:</emphasis>
|
||
This species can be recognized by the asymmetrical surstyli in dorsal view, broadened in basal, left surstylus knobbed apically, right surstylus narrowed in apical half (
|
||
<figureCitation id="133A2A435F6CFFA7FCF4ED4AFC6A4913" box="[861,965,1544,1571]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="46" pageNumber="47">Fig. 25A</figureCitation>
|
||
); gonopods equal in height in ventral view (
|
||
<figureCitation id="133A2A435F6CFFA7FF09ED6EFEA84977" box="[160,263,1580,1607]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="46" pageNumber="47">Fig. 25B</figureCitation>
|
||
); small phallic guide in lateral view (
|
||
<figureCitation id="133A2A435F6CFFA7FD0FED6EFCA04977" box="[678,783,1580,1607]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="46" pageNumber="47">Fig. 25D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F6CFFB9FF6EED12FB1D4EB9" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
|
||
<paragraph id="8BBE36C65F6CFFB9FF6EED12FB1D4EB9" blockId="46.[151,1437,1508,2039]" lastBlockId="48.[151,1437,151,645]" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
|
||
<emphasis id="B975EAD45F6CFFA7FF6EED12FEFE495A" bold="true" box="[199,337,1616,1642]" pageId="46" pageNumber="47">Description</emphasis>
|
||
:
|
||
<emphasis id="B975EAD45F6CFFA7FEC9ED12FE1B495A" bold="true" box="[352,436,1616,1642]" pageId="46" pageNumber="47">MALE</emphasis>
|
||
: Body length (excluding antennae):
|
||
<quantity id="4CF99B235F6CFFA7FCE7ED13FC08495B" box="[846,935,1617,1643]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="46" pageNumber="47" unit="mm" value="3.9">3.9 mm</quantity>
|
||
.
|
||
<emphasis id="B975EAD45F6CFFA7FC1BED12FC5F495B" bold="true" box="[946,1008,1616,1643]" italics="true" pageId="46" pageNumber="47">Head</emphasis>
|
||
. Scape, pedicel, flagellum, arista dark brown. Pedicel gray pollinose in apical half, with three upper setae and 1-2 lower setae. Flagellum gray pollinose and tapering (LF:WF = 3.0); arista dark, with thickened base. Eyes meeting for a distance of 6‒8 facets. Frons dark silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose with scattered short black setae.
|
||
<emphasis id="B975EAD45F6CFFA7FF77EDA2FE9449CA" box="[222,315,1760,1787]" italics="true" pageId="46" pageNumber="47">
|
||
<emphasis id="B975EAD45F6CFFA7FF77EDA2FE9B49CB" bold="true" box="[222,308,1760,1787]" italics="true" pageId="46" pageNumber="47">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Postpronotal lobe light yellow, gray pollinose with 4-6 yellowish setae along the upper margin (up to
|
||
<quantity id="4CF99B235F6CFFA7FF1CEC47FEB2482F" box="[181,285,1797,1823]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="mm" value="0.02">0.02 mm</quantity>
|
||
). Prescutum, scutum and scutellum black. Scutum with two uniseriate rows of intra-alar setae. Scutellum with 14–16 thin short setae along lateral margin (up to
|
||
<quantity id="4CF99B235F6CFFA7FC95EC6BFC0A4873" box="[828,933,1833,1859]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="mm" value="0.02">0.02 mm</quantity>
|
||
). Subscutellum black, gray pollinose. Pleura black but dark brown on pteropleuron, hypopleuron and sternopleuron.
|
||
<emphasis id="B975EAD45F6CFFA7FC1BEC0EFC404856" bold="true" box="[946,1007,1868,1894]" italics="true" pageId="46" pageNumber="47">Wing</emphasis>
|
||
. Length:
|
||
<quantity id="4CF99B235F6CFFA7FBF3EC0EFB6E4857" box="[1114,1217,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="46" pageNumber="47" unit="mm" value="3.85">3.85 mm</quantity>
|
||
. LW:MWW = 5.13. Wing almost entirely covered with microtrichia. Cross-vein r-m reaches dm at the middle. M
|
||
<subScript id="178534835F6CFFA7FB18ECC1FB1648A1" attach="left" box="[1201,1209,1923,1937]" fontSize="6" pageId="46" pageNumber="47">1</subScript>
|
||
strongly undulating in middle. Halter length:
|
||
<quantity id="4CF99B235F6CFFA7FE1CECD7FDA0489F" box="[437,527,1941,1967]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="46" pageNumber="47" unit="mm" value="0.4">0.4 mm</quantity>
|
||
. Whitish, narrowly brown at base.
|
||
<emphasis id="B975EAD45F6CFFA7FC33ECD7FC7F489F" bold="true" box="[922,976,1941,1967]" italics="true" pageId="46" pageNumber="47">Legs</emphasis>
|
||
. Coxae dark brown, gray pollinose. Mid coxa with four dark anterior setae. Trochanters brown, smooth. Femora dark with light brown apices, gray pollinose. Mid and hind femora bearing two rows of small spines in apical half and with one wrinkled indentation at the base. Tibiae brown but dark at middle, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with 1–2 wrinkled indentations in middle. Tarsi light brown with scattered brown setae at anterior margin, basitarsi longer than other tarsomeres. Pulvilli yellow. Claws light brown with black tips.
|
||
<emphasis id="B975EAD45F6DFFA6FB20ECC9FB514895" box="[1161,1278,1931,1958]" italics="true" pageId="47" pageNumber="48">
|
||
<emphasis id="B975EAD45F6DFFA6FB20ECC9FB584896" bold="true" box="[1161,1271,1931,1958]" italics="true" pageId="47" pageNumber="48">Abdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
Ground color dark. Tergite 1 densely gray pollinose, with 8–10 dark lateral setae (up to
|
||
<quantity id="4CF99B235F6DFFA6FC4DECF2FBE348FA" box="[996,1100,1968,1994]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="47" pageNumber="48" unit="mm" value="0.11">0.11 mm</quantity>
|
||
). Tergites 2–5 gray pollinose, with scattered brown setae. Sternites light brown laterally and dark brown centrally, gray pollinose.
|
||
<emphasis id="B975EAD45F6DFFA6FA83EC91FA3348DD" box="[1322,1436,2003,2030]" italics="true" pageId="47" pageNumber="48">
|
||
<emphasis id="B975EAD45F6DFFA6FA83EC91FA3A48DE" bold="true" box="[1322,1429,2003,2030]" italics="true" pageId="47" pageNumber="48">Genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.5). Surstyli asymmetrical, elongated (
|
||
<figureCitation id="133A2A435F72FFB9FD04EBF9FCBA4FE5" box="[685,789,186,213]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="48" pageNumber="49">Fig. 25A</figureCitation>
|
||
). Both surstyli broad at base, base of left surstylus triangular-shaped, base of right surstylus square-shaped, left surstylus entirely constricted at middle and knobbed apically, right surstylus narrowed in apical half, left surstylus higher than right one (
|
||
<figureCitation id="133A2A435F72FFB9FC5EEA41FBCD4E2D" box="[1015,1122,258,285]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="48" pageNumber="49">Fig. 25A</figureCitation>
|
||
). Genital capsule in ventral view: gonopods wide, equal in height (
|
||
<figureCitation id="133A2A435F72FFB9FDE4EA65FD1A4E71" box="[589,693,294,321]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="48" pageNumber="49">Fig. 25B</figureCitation>
|
||
). Subepandrial sclerite small (
|
||
<figureCitation id="133A2A435F72FFB9FBA1EA65FBDF4E71" box="[1032,1136,294,321]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="48" pageNumber="49">Fig. 25B</figureCitation>
|
||
). Genital capsule in lateral view: both surstyli bent in middle towards sternite (
|
||
<figureCitation id="133A2A435F72FFB9FD70EA09FCF24E55" box="[729,861,330,357]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="48" pageNumber="49">Fig. 25E–F</figureCitation>
|
||
). Phallic guide short, phallus with three ejaculatory ducts (
|
||
<figureCitation id="133A2A435F72FFB9FF4BEA2DFEE54EB9" box="[226,330,366,393]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="48" pageNumber="49">Fig. 25D</figureCitation>
|
||
). Ejaculatory apodeme rather spade-shaped (
|
||
<figureCitation id="133A2A435F72FFB9FCE9EA2DFC084EB9" box="[832,935,366,393]" captionStart="FIGURE 25" captionStartId="47.[151,250,1753,1777]" captionTargetBox="[159,1421,184,1727]" captionTargetId="figure-21@47.[151,1436,179,1727]" captionTargetPageId="47" captionText="FIGURE 25. Male genitalia of Tomosvaryella nimroozensis sp. nov., holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="48" pageNumber="49">Fig. 25C</figureCitation>
|
||
).
|
||
<emphasis id="B975EAD45F72FFB9FC13EA2CFB934EB8" bold="true" box="[954,1084,366,392]" pageId="48" pageNumber="49">FEMALE:</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7E664E5F6DFFA6FF3EED9BFC204825" ID-DOI="http://doi.org/10.5281/zenodo.5122723" ID-Zenodo-Dep="5122723" httpUri="https://zenodo.org/record/5122723/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1753,1777]" targetBox="[159,1421,184,1727]" targetPageId="47">
|
||
<paragraph id="8BBE36C65F6DFFA6FF3EED9BFC204825" blockId="47.[151,1436,1753,1813]" pageId="47" pageNumber="48">
|
||
<emphasis id="B975EAD45F6DFFA6FF3EED9BFE8D49C1" bold="true" box="[151,290,1753,1777]" pageId="47" pageNumber="48">FIGURE 25.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C014D455F6DFFA6FE4BED9BFCAB49C1" box="[482,772,1753,1777]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="nimroozensis" status="sp. nov.">
|
||
<emphasis id="B975EAD45F6DFFA6FE4BED9BFCAB49C1" box="[482,772,1753,1777]" italics="true" pageId="47" pageNumber="48">Tomosvaryella nimroozensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B975EAD45F6DFFA6FCA5ED9BFCF149C1" bold="true" box="[780,862,1753,1777]" pageId="47" pageNumber="48">
|
||
<taxonomicNameLabel id="A24657AF5F6DFFA6FCA5ED9BFCF149C1" box="[780,862,1753,1777]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (JSS51819, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D) phallus in lateral view E, F) lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31B654D5F72FFB9FF6EEAD1FD0B4EC5" pageId="48" pageNumber="49" type="etymology">
|
||
<paragraph id="8BBE36C65F72FFB9FF6EEAD1FB644E9D" blockId="48.[151,1437,151,645]" box="[199,1227,403,429]" pageId="48" pageNumber="49">
|
||
<emphasis id="B975EAD45F72FFB9FF6EEAD1FEFD4E9D" bold="true" box="[199,338,403,429]" pageId="48" pageNumber="49">Etymology:</emphasis>
|
||
The specific epithet refers to Nimrooz, from where the
|
||
<typeStatus id="54BA88645F72FFB9FC6AEAD1FB8A4E9D" box="[963,1061,403,429]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
was collected.
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F72FFB9FF6EEAF4FD0B4EC5" blockId="48.[151,1437,151,645]" pageId="48" pageNumber="49">
|
||
<emphasis id="B975EAD45F72FFB9FF6EEAF4FE104EE0" bold="true" box="[199,447,438,464]" pageId="48" pageNumber="49">Specimen examined:</emphasis>
|
||
<materialsCitation id="3B693C9B5F72FFB9FE6EEAF4FD0B4EC5" accessionNumber="JSS51819" collectingDate="2016-04-07" collectionCode="CNC" collectorName="Sistan & Baluchestan & M. Enaiatnia" country="Iran" latitude="31.05" location="Zabol" longLatPrecision="1218" longitude="61.366665" pageId="48" pageNumber="49" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B975EAD45F72FFB9FE6EEAF4FDC04EE0" bold="true" box="[455,623,438,464]" pageId="48" pageNumber="49">
|
||
<typeStatus id="54BA88645F72FFB9FE6EEAF4FDC54EE0" box="[455,618,438,464]" pageId="48" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectingCountry id="F31676565F72FFB9FDDEEAF5FD6F4EE1" box="[631,704,439,465]" name="Iran" pageId="48" pageNumber="49">IRAN</collectingCountry>
|
||
:
|
||
<collectorName id="26F453105F72FFB9FD65EAF5FCBF4EE1" box="[716,784,439,465]" pageId="48" pageNumber="49">Sistan</collectorName>
|
||
&
|
||
<collectorName id="26F453105F72FFB9FC9EEAF5FC6F4EE1" box="[823,960,439,465]" pageId="48" pageNumber="49">Baluchestan</collectorName>
|
||
:
|
||
<location id="8EDE601D5F72FFB9FC71EAF5FBB34EE1" LSID="urn:lsid:plazi:treatment:03A887D05F6CFFB9FF3EEE3AFF1A4DB5:8EDE601D5F72FFB9FC71EAF5FBB34EE1" box="[984,1052,439,465]" country="Iran" latitude="31.05" longLatPrecision="1218" longitude="61.366665" name="Zabol" pageId="48" pageNumber="49">Zabol</location>
|
||
,
|
||
<geoCoordinate id="EE3550015F72FFB9FB81EAF5FB244EE1" box="[1064,1163,439,465]" degrees="31" direction="north" minutes="03" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="31.05">31°03’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3550015F72FFB9FB31EAF5FB574EE1" box="[1176,1272,439,465]" degrees="61" direction="east" minutes="22" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="61.366665">61°22’E</geoCoordinate>
|
||
,
|
||
<date id="FFBF10065F72FFB9FAACEAF5FAC04EE1" box="[1285,1391,439,465]" pageId="48" pageNumber="49" value="2016-04-07">
|
||
<collectingDate id="EFFBE9EE5F72FFB9FAACEAF5FAC04EE1" box="[1285,1391,439,465]" pageId="48" pageNumber="49" value="2016-04-07">7.iv.2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F453105F72FFB9FAD5EAF5FEAB4EC5" pageId="48" pageNumber="49">M. Enaiatnia</collectorName>
|
||
,
|
||
<collectingMethod id="52404ED15F72FFB9FEA6EA99FE324EC5" box="[271,413,475,501]" pageId="48" pageNumber="49">Malaise trap</collectingMethod>
|
||
, JSS51819 (
|
||
<specimenCount id="9D07FD4F5F72FFB9FD80EA99FDE24EC5" box="[553,589,475,501]" pageId="48" pageNumber="49" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED10AE035F72FFB9FDF0EA99FD384EC5" box="[601,663,475,501]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="48" pageNumber="49">CNC</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F72FFB9FF6EEABCFF1A4DB5" pageId="48" pageNumber="49" type="distribution">
|
||
<paragraph id="8BBE36C65F72FFB9FF6EEABCFDA84D29" blockId="48.[151,1437,151,645]" box="[199,519,510,537]" pageId="48" pageNumber="49">
|
||
<emphasis id="B975EAD45F72FFB9FF6EEABCFECC4D28" bold="true" box="[199,355,510,536]" pageId="48" pageNumber="49">Distribution:</emphasis>
|
||
<collectingCountry id="F31676565F72FFB9FEC3EABDFE384D29" box="[362,407,511,537]" name="Iran" pageId="48" pageNumber="49">Iran</collectingCountry>
|
||
(
|
||
<figureCitation id="133A2A435F72FFB9FE0FEABDFE544D29" box="[422,507,510,537]" captionStart="FIGURE 56" captionStartId="88.[151,250,1562,1586]" captionTargetBox="[151,1436,896,1538]" captionTargetId="figure-53@88.[151,1436,896,1538]" captionTargetPageId="88" captionText="FIGURE 56. Distribution of Tomosvaryella inopinata, T. minuscula, T. nimroozensis sp. nov., T. soziana sp. nov. in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122783" httpUri="https://zenodo.org/record/5122783/files/figure.png" pageId="48" pageNumber="49">Fig. 56</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F72FFB9FF6EE961FF1A4DB5" blockId="48.[151,1437,151,645]" pageId="48" pageNumber="49">
|
||
<emphasis id="B975EAD45F72FFB9FF6EE961FEBC4D0D" bold="true" box="[199,275,547,573]" pageId="48" pageNumber="49">Notes:</emphasis>
|
||
<taxonomicName id="4C014D455F72FFB9FEB5E961FE634D0C" box="[284,460,547,572]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="nimroozensis" status="sp. nov.">
|
||
<emphasis id="B975EAD45F72FFB9FEB5E961FE634D0C" box="[284,460,547,572]" italics="true" pageId="48" pageNumber="49">T. nimroozensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657AF5F72FFB9FE7DE961FD9F4D0D" box="[468,560,547,573]" pageId="48" pageNumber="49" rank="species">
|
||
<emphasis id="B975EAD45F72FFB9FE7DE961FD854D0D" bold="true" box="[468,554,547,573]" pageId="48" pageNumber="49">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is not genetically similar to any other known
|
||
<taxonomicName id="4C014D455F72FFB9FB97E961FAEC4D0D" box="[1086,1347,547,573]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B975EAD45F72FFB9FB97E961FB4D4D0D" box="[1086,1250,547,573]" italics="true" pageId="48" pageNumber="49">Tomosvaryella</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
<taxonomicName id="4C014D455F72FFB9FAE2E961FE2A4D51" baseAuthorityName="Becker" baseAuthorityYear="1900" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pilosiventris">
|
||
<emphasis id="B975EAD45F72FFB9FAE2E961FE2A4D51" italics="true" pageId="48" pageNumber="49">Tomosvaryella pilosiventris</emphasis>
|
||
</taxonomicName>
|
||
is closest to
|
||
<taxonomicName id="4C014D455F72FFB9FDB5E905FD634D50" box="[540,716,583,608]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="nimroozensis" status="sp. nov.">
|
||
<emphasis id="B975EAD45F72FFB9FDB5E905FD634D50" box="[540,716,583,608]" italics="true" pageId="48" pageNumber="49">T. nimroozensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657AF5F72FFB9FD7DE905FC9F4D51" box="[724,816,583,609]" pageId="48" pageNumber="49" rank="species">
|
||
<emphasis id="B975EAD45F72FFB9FD7DE905FC854D51" bold="true" box="[724,810,583,609]" pageId="48" pageNumber="49">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
with 12.2% pairwise difference (Supplementary
|
||
<tableCitation id="C683037D5F72FFB9FAF7E905FF0B4DB5" captionStart="TABLE 1" captionStartId="5.[151,239,810,836]" captionText="TABLE 1. Cytochrome c oxidase subunit I (COI) and internal transcribed spacer (ITS) primers." pageId="48" pageNumber="49">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |