treatments-xml/data/03/A8/87/03A887A3FF84FFBB1DE82EB7A00259C4.xml

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<mods:title>Meristotheca lysonensis sp. nov. (Solieriaceae, Rhodophyta), a new flattened species from Vietnamese waters</mods:title>
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<mods:namePart>Nguyen, Xuan-Vy</mods:namePart>
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<mods:affiliation>Institute of Oceanography, Viet Nam Academy of Science and Technology, 01 Cau Da, Nha Trang City, Viet Nam. &amp; Graduate University of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Viet Nam. &amp; nguyenxuanvi @ gmail. com; https: // orcid. org / 0000 - 0002 - 7260 - 5127</mods:affiliation>
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<mods:namePart>Nguyen, Xuan-Thuy</mods:namePart>
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<mods:affiliation>Institute of Oceanography, Viet Nam Academy of Science and Technology, 01 Cau Da, Nha Trang City, Viet Nam. &amp; xuanthuynguyen 1293 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9568 - 7690</mods:affiliation>
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<mods:namePart>Kittle Iii, Ronald P.</mods:namePart>
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<mods:affiliation>Department of Biology, University of Louisiana at Lafayette, 410 E. St. Mary Blvd., Lafayette, LA 70503, USA. &amp; ronaldkittleull @ gmail. com; https: // orcid. org / 0000 - 0002 - 9371 - 7152 &amp; Marine Science Department, University of Hawaii-Hilo 200 W. Kawili St. Hilo, HI 96720, USA. &amp; mcdermid @ hawaii. edu; https: // orcid. org / 0000 - 0002 - 7663 - 6545</mods:affiliation>
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<taxonomicName authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[136,412,724,751]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="4" pageNumber="141" phylum="Rhodophyta" rank="species" species="lysonensis">
<emphasis bold="true" box="[136,412,724,751]" italics="true" pageId="4" pageNumber="141">Meristotheca lysonensis</emphasis>
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X.-V. Nguyen, X.-T. Nguyen, Kittle &amp; McDermid 
<emphasis bold="true" box="[1008,1095,724,750]" italics="true" pageId="4" pageNumber="141">sp. nov.</emphasis>
(
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)
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<emphasis bold="true" box="[136,291,1958,1980]" pageId="4" pageNumber="141">FIGURES 3–6</emphasis>
. 
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<emphasis box="[303,534,1958,1979]" italics="true" pageId="4" pageNumber="141">Meristotheca lysonensis</emphasis>
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X.-V. Nguyen, X.-T. Nguyen, Kittle 
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McDermid, 
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. 3: Wet habit of a vegetative thallus, holotype specimen (ION-CS210215b), tetrasporangial plant with secondary holdfasts (black arrows). 4. A mature segment of the blade, with the rough surface by the presence of warty or short spinose projections (ION-CS210215b). 5. Dried specimen, isotype, a male plant (ION-CS210301b). 6. Dried specimen, isotype (ION-01011b), tetrasporangial plant.
</paragraph>
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<emphasis bold="true" box="[136,305,1283,1305]" pageId="5" pageNumber="142">FIGURES 7–10</emphasis>
. Vegetative structures of 
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<emphasis box="[559,793,1283,1304]" italics="true" pageId="5" pageNumber="142">Meristotheca lysonensis</emphasis>
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X.-V. Nguyen, X.-T. Nguyen, Kittle 
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McDermid, 
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. (IONCS210215b). 7. Cross section through the basal part showing the layers of outer layers (white arrow), inner layer cellular cortex (black arrow) and filamentous medulla (arrowhead). 8. Cross section near the apex of a blade. 9. Longitudinal section showing secondary pitconnections (arrowheads) between adjacent medullary cells. 10. Crosswise filaments (arrowhead) transversely arranged in the medullary region.
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: 
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15.3743 
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N
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109.1329 
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E
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, coll. X.- 
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. Nguyen &amp; X.- 
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- 
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b, tetrasporophyte, 
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). GenBank accession number for 
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and COI-5 
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: 
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11.5648 
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109.1269 
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E
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, 
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, coll. X.- 
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. 
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&amp; X. - 
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. 
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(
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- 
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b, tetrasporophyte, 
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). 
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accession number for 
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and COI-5 
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: 
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. 
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<collectingCountry box="[322,455,1676,1702]" name="Vietnam" pageId="5" pageNumber="142">VIETNAM</collectingCountry>
. 
<collectingRegion box="[465,606,1675,1702]" country="Vietnam" name="Quang Ngai" pageId="5" pageNumber="142">Quang Ngai</collectingRegion>
: 
<location LSID="urn:lsid:plazi:treatment:03A887A3FF84FFBB1DE82EB7A00259C4:8EDE606EFF85FFB91F082AEFA0785E9D" box="[616,774,1675,1702]" country="Vietnam" latitude="15.3743" longLatPrecision="7" longitude="109.1329" name="Ly Son Island" pageId="5" pageNumber="142" stateProvince="Quang Ngai">Ly Son Island</location>
; 
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15.3743 
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N
</geoCoordinate>
; 
<geoCoordinate box="[921,1056,1675,1702]" degrees="109.1329" direction="east" orientation="longitude" pageId="5" pageNumber="142" precision="5" value="109.1329">
109.1329 
<superScript attach="left" box="[1026,1033,1675,1689]" fontSize="6" pageId="5" pageNumber="142">°</superScript>
E
</geoCoordinate>
, 
<date box="[1068,1226,1675,1702]" pageId="5" pageNumber="142" value="2003-04-02">
<collectingDate box="[1068,1226,1675,1702]" pageId="5" pageNumber="142" value="2003-04-02">02 April 2003</collectingDate>
</date>
, coll. 
<collectionCode box="[1292,1316,1676,1702]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="5" pageNumber="142" type="Herbarium">H</collectionCode>
</materialsCitation>
.- 
<materialsCitation collectionCode="H, T" country="Vietnam" location="Nguyen" pageId="5" pageNumber="142" specimenCount="1" stateProvince="Quang Ngai">
D. 
<location LSID="urn:lsid:plazi:treatment:03A887A3FF84FFBB1DE82EB7A00259C4:8EDE606EFF85FFB918332AEFA6D25E9E" box="[1363,1452,1676,1702]" country="Vietnam" name="Nguyen" pageId="5" pageNumber="142" stateProvince="Quang Ngai">Nguyen</location>
&amp; 
<collectionCode box="[166,189,1712,1738]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="5" pageNumber="142" type="Herbarium">H</collectionCode>
.- 
<collectionCode box="[201,221,1712,1738]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="142">T</collectionCode>
</materialsCitation>
. 
<materialsCitation box="[233,1117,1711,1738]" collectionCode="L" country="Vietnam" location="GenBank" municipality="Pham" pageId="5" pageNumber="142" specimenCount="1" stateProvince="Quang Ngai">
<collectingMunicipality box="[233,297,1711,1737]" pageId="5" pageNumber="142">Pham</collectingMunicipality>
(
<emphasis bold="true" box="[314,367,1711,1737]" pageId="5" pageNumber="142">ION</emphasis>
-01011b, male plant, 
<figureCitation box="[608,677,1711,1737]" captionStart="FIGURES 3–6" captionStartId="4.[136,248,1958,1980]" captionTargetBox="[183,1404,775,1934]" captionTargetId="figure-212@4.[183,1404,775,1934]" captionTargetPageId="4" captionText="FIGURES 3–6. Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. 3: Wet habit of a vegetative thallus, holotype specimen (ION-CS210215b), tetrasporangial plant with secondary holdfasts (black arrows). 4. A mature segment of the blade, with the rough surface by the presence of warty or short spinose projections (ION-CS210215b). 5. Dried specimen, isotype, a male plant (ION-CS210301b). 6. Dried specimen, isotype (ION-01011b), tetrasporangial plant." figureDoi="http://doi.org/10.5281/zenodo.7380825" httpUri="https://zenodo.org/record/7380825/files/figure.png" pageId="5" pageNumber="142">Fig. 6</figureCitation>
). 
<location LSID="urn:lsid:plazi:treatment:03A887A3FF84FFBB1DE82EB7A00259C4:8EDE606EFF85FFB91FDB2ACCA05B5EF1" box="[699,805,1711,1737]" country="Vietnam" municipality="Pham" name="GenBank" pageId="5" pageNumber="142" stateProvince="Quang Ngai">GenBank</location>
accession number for 
<emphasis box="[1063,1100,1711,1737]" italics="true" pageId="5" pageNumber="142">rbc</emphasis>
<collectionCode box="[1101,1117,1712,1738]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="5" pageNumber="142" type="Herbarium">L</collectionCode>
</materialsCitation>
<materialsCitation accessionNumber="MZ770760 and MZ770757" collectionCode="P" country="Vietnam" pageId="5" pageNumber="142" specimenCount="1" stateProvince="Quang Ngai">
and COI-5 
<collectionCode box="[1248,1265,1711,1738]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="142" type="Herbarium">P</collectionCode>
: 
<accessionNumber isEnumeration="true" pageId="5" pageNumber="142">MZ770760 and MZ770757</accessionNumber>
</materialsCitation>
</paragraph>
<paragraph blockId="5.[136,1452,1495,2062]" box="[189,1398,1783,1810]" pageId="5" pageNumber="142">
<emphasis bold="true" box="[189,281,1783,1809]" pageId="5" pageNumber="142">Habitat</emphasis>
: Growing on dead corals in the mid-to lower intertidal zone where subjected to strong wave action.
</paragraph>
</subSubSection>
<subSubSection box="[189,1374,1819,1846]" pageId="5" pageNumber="142" type="etymology">
<paragraph blockId="5.[136,1452,1495,2062]" box="[189,1374,1819,1846]" pageId="5" pageNumber="142">
<emphasis bold="true" box="[189,318,1820,1846]" pageId="5" pageNumber="142">Etymology</emphasis>
: The specific epithet refers to Ly Son Island, the 
<typeStatus box="[871,919,1820,1846]" pageId="5" pageNumber="142">type</typeStatus>
locality of the new species in 
<collectingCountry box="[1258,1370,1819,1846]" name="Vietnam" pageId="5" pageNumber="142">Viet Nam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="144" pageId="5" pageNumber="142" type="description">
<paragraph blockId="5.[136,1452,1495,2062]" lastBlockId="6.[136,1452,158,653]" lastPageId="6" lastPageNumber="143" pageId="5" pageNumber="142">
<emphasis bold="true" box="[189,327,1855,1881]" pageId="5" pageNumber="142">Description</emphasis>
: Thalli prostrate or forming decumbent prostrate clumps; thallus attached to the substratum by many secondary holdfasts; thalli flattened, gelatinous in texture, deep rose-red to dark red, 
<quantity box="[1080,1190,1892,1918]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="142" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 cm</quantity>
tall with small papillae on the surface; divided into many broad segments in an irregularly dichotomous manner with rounded apices, becoming gradually narrower above and tapering below to a cuneate or sometimes cordate or heart-shaped base with a short slender stipe. Mature blades are 
<quantity box="[494,576,2000,2026]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="142" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
broad, margins entire at the beginning, later giving rise to irregularly branched obvious proliferations, broader or narrower intermixed; surface smooth, sometimes roughened by the presence of warty or short spinose processes (
<figureCitation box="[512,613,158,184]" captionStart="FIGURES 3–6" captionStartId="4.[136,248,1958,1980]" captionTargetBox="[183,1404,775,1934]" captionTargetId="figure-212@4.[183,1404,775,1934]" captionTargetPageId="4" captionText="FIGURES 3–6. Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. 3: Wet habit of a vegetative thallus, holotype specimen (ION-CS210215b), tetrasporangial plant with secondary holdfasts (black arrows). 4. A mature segment of the blade, with the rough surface by the presence of warty or short spinose projections (ION-CS210215b). 5. Dried specimen, isotype, a male plant (ION-CS210301b). 6. Dried specimen, isotype (ION-01011b), tetrasporangial plant." figureDoi="http://doi.org/10.5281/zenodo.7380825" httpUri="https://zenodo.org/record/7380825/files/figure.png" pageId="6" pageNumber="143">Figs 3, 4</figureCitation>
), pinkish or yellowish when dried (
<figureCitation box="[1008,1109,158,185]" captionStart="FIGURES 3–6" captionStartId="4.[136,248,1958,1980]" captionTargetBox="[183,1404,775,1934]" captionTargetId="figure-212@4.[183,1404,775,1934]" captionTargetPageId="4" captionText="FIGURES 3–6. Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. 3: Wet habit of a vegetative thallus, holotype specimen (ION-CS210215b), tetrasporangial plant with secondary holdfasts (black arrows). 4. A mature segment of the blade, with the rough surface by the presence of warty or short spinose projections (ION-CS210215b). 5. Dried specimen, isotype, a male plant (ION-CS210301b). 6. Dried specimen, isotype (ION-01011b), tetrasporangial plant." figureDoi="http://doi.org/10.5281/zenodo.7380825" httpUri="https://zenodo.org/record/7380825/files/figure.png" pageId="6" pageNumber="143">Figs 5, 6</figureCitation>
). Blades are 
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thick in the basal portions (
<figureCitation box="[384,454,194,220]" captionStart="FIGURES 7–10" captionStartId="5.[136,248,1283,1305]" captionTargetBox="[183,1404,190,1259]" captionTargetId="figure-18@5.[183,1404,190,1259]" captionTargetPageId="5" captionText="FIGURES 7–10. Vegetative structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b). 7. Cross section through the basal part showing the layers of outer layers (white arrow), inner layer cellular cortex (black arrow) and filamentous medulla (arrowhead). 8. Cross section near the apex of a blade. 9. Longitudinal section showing secondary pitconnections (arrowheads) between adjacent medullary cells. 10. Crosswise filaments (arrowhead) transversely arranged in the medullary region." figureDoi="http://doi.org/10.5281/zenodo.7380827" httpUri="https://zenodo.org/record/7380827/files/figure.png" pageId="6" pageNumber="143">Fig. 7</figureCitation>
), becoming progressively thinner upward, 320–450 µm thick below the apices (
<figureCitation box="[1367,1437,194,220]" captionStart="FIGURES 7–10" captionStartId="5.[136,248,1283,1305]" captionTargetBox="[183,1404,190,1259]" captionTargetId="figure-18@5.[183,1404,190,1259]" captionTargetPageId="5" captionText="FIGURES 7–10. Vegetative structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b). 7. Cross section through the basal part showing the layers of outer layers (white arrow), inner layer cellular cortex (black arrow) and filamentous medulla (arrowhead). 8. Cross section near the apex of a blade. 9. Longitudinal section showing secondary pitconnections (arrowheads) between adjacent medullary cells. 10. Crosswise filaments (arrowhead) transversely arranged in the medullary region." figureDoi="http://doi.org/10.5281/zenodo.7380827" httpUri="https://zenodo.org/record/7380827/files/figure.png" pageId="6" pageNumber="143">Fig. 8</figureCitation>
). Thalli are multiaxial and internally consist of a filamentous medulla. In traverse section, the outer cortex consists of 3– 4 layers of small, elongate cells, 12–20 µm in diameter, the inner cortex consists of 4–5 layers of bigger, ellipsoidal, or rounded cells, 40–140 µm in diameter. The medulla occupies about 20–30% of the blade thickness. Axial filaments run parallel to the longitudinal plane of the blade; adjacent axial filaments frequently linked by secondary pit-connections (
<figureCitation box="[144,214,374,400]" captionStart="FIGURES 7–10" captionStartId="5.[136,248,1283,1305]" captionTargetBox="[183,1404,190,1259]" captionTargetId="figure-18@5.[183,1404,190,1259]" captionTargetPageId="5" captionText="FIGURES 7–10. Vegetative structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b). 7. Cross section through the basal part showing the layers of outer layers (white arrow), inner layer cellular cortex (black arrow) and filamentous medulla (arrowhead). 8. Cross section near the apex of a blade. 9. Longitudinal section showing secondary pitconnections (arrowheads) between adjacent medullary cells. 10. Crosswise filaments (arrowhead) transversely arranged in the medullary region." figureDoi="http://doi.org/10.5281/zenodo.7380827" httpUri="https://zenodo.org/record/7380827/files/figure.png" pageId="6" pageNumber="143">Fig. 9</figureCitation>
). Crosswise filaments originate from inner cortical cells and traverse the blade to connect with inner cortical cells of the opposite side (
<figureCitation box="[428,512,410,436]" captionStart="FIGURES 7–10" captionStartId="5.[136,248,1283,1305]" captionTargetBox="[183,1404,190,1259]" captionTargetId="figure-18@5.[183,1404,190,1259]" captionTargetPageId="5" captionText="FIGURES 7–10. Vegetative structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b). 7. Cross section through the basal part showing the layers of outer layers (white arrow), inner layer cellular cortex (black arrow) and filamentous medulla (arrowhead). 8. Cross section near the apex of a blade. 9. Longitudinal section showing secondary pitconnections (arrowheads) between adjacent medullary cells. 10. Crosswise filaments (arrowhead) transversely arranged in the medullary region." figureDoi="http://doi.org/10.5281/zenodo.7380827" httpUri="https://zenodo.org/record/7380827/files/figure.png" pageId="6" pageNumber="143">Fig. 10</figureCitation>
). Tetrasporangia in scattered patches throughout the outer cortex on both dorsal and ventral sides of the entire blade except at tips and lowermost portions. Tetrasporangia are cut off from cortical cells located in the second layer of the outer cortex. In a few cases, tetrasporangial initials appear laterally, pit-connected to their parental cells (
<figureCitation box="[355,436,518,544]" captionStart="FIGURES 11–14" captionStartId="6.[136,248,1951,1973]" captionTargetBox="[229,1361,677,1927]" captionTargetId="figure-266@6.[227,1361,677,1927]" captionTargetPageId="6" captionText="FIGURES 11–14. Reproductive structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b) 11. Tetrasporangial initial laterally pit-connected (arrowhead) to its parental cell. 12. Tetrasporangium basally pit-connected (arrowhead) to the parental cells. 13. Zonately divided tetrasporangium (arrowhead) with a basal pit-connection with the parental cell. 14. Spermatangia (arrowhead) produced from the outermost cortical cells (ION-CS210301b)." figureDoi="http://doi.org/10.5281/zenodo.7380829" httpUri="https://zenodo.org/record/7380829/files/figure.png" pageId="6" pageNumber="143">Fig. 11</figureCitation>
). Tetrasporangial initials are basally attached to their supporting parental cortical cells (
<figureCitation captionStart="FIGURES 11–14" captionStartId="6.[136,248,1951,1973]" captionTargetBox="[229,1361,677,1927]" captionTargetId="figure-266@6.[227,1361,677,1927]" captionTargetPageId="6" captionText="FIGURES 11–14. Reproductive structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b) 11. Tetrasporangial initial laterally pit-connected (arrowhead) to its parental cell. 12. Tetrasporangium basally pit-connected (arrowhead) to the parental cells. 13. Zonately divided tetrasporangium (arrowhead) with a basal pit-connection with the parental cell. 14. Spermatangia (arrowhead) produced from the outermost cortical cells (ION-CS210301b)." figureDoi="http://doi.org/10.5281/zenodo.7380829" httpUri="https://zenodo.org/record/7380829/files/figure.png" pageId="6" pageNumber="143">Fig. 12</figureCitation>
, solid triangle), and basal pit-connections remain in mature tetrasporangia. Tetrasporangia zonately divided, 10–15 µm diameter and 40–45 µm long (
<figureCitation box="[517,600,590,616]" captionStart="FIGURES 11–14" captionStartId="6.[136,248,1951,1973]" captionTargetBox="[229,1361,677,1927]" captionTargetId="figure-266@6.[227,1361,677,1927]" captionTargetPageId="6" captionText="FIGURES 11–14. Reproductive structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b) 11. Tetrasporangial initial laterally pit-connected (arrowhead) to its parental cell. 12. Tetrasporangium basally pit-connected (arrowhead) to the parental cells. 13. Zonately divided tetrasporangium (arrowhead) with a basal pit-connection with the parental cell. 14. Spermatangia (arrowhead) produced from the outermost cortical cells (ION-CS210301b)." figureDoi="http://doi.org/10.5281/zenodo.7380829" httpUri="https://zenodo.org/record/7380829/files/figure.png" pageId="6" pageNumber="143">Fig. 13</figureCitation>
). Spermatangia produced from the outermost cortical cells (
<figureCitation box="[1266,1349,590,616]" captionStart="FIGURES 11–14" captionStartId="6.[136,248,1951,1973]" captionTargetBox="[229,1361,677,1927]" captionTargetId="figure-266@6.[227,1361,677,1927]" captionTargetPageId="6" captionText="FIGURES 11–14. Reproductive structures of Meristotheca lysonensis X.-V. Nguyen, X.-T. Nguyen, Kittle et McDermid, sp. nov. (ION- CS210215b) 11. Tetrasporangial initial laterally pit-connected (arrowhead) to its parental cell. 12. Tetrasporangium basally pit-connected (arrowhead) to the parental cells. 13. Zonately divided tetrasporangium (arrowhead) with a basal pit-connection with the parental cell. 14. Spermatangia (arrowhead) produced from the outermost cortical cells (ION-CS210301b)." figureDoi="http://doi.org/10.5281/zenodo.7380829" httpUri="https://zenodo.org/record/7380829/files/figure.png" pageId="6" pageNumber="143">Fig. 14</figureCitation>
). Female gametophytes were not observed.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7380829" ID-Zenodo-Dep="7380829" httpUri="https://zenodo.org/record/7380829/files/figure.png" pageId="6" pageNumber="143" startId="6.[136,248,1951,1973]" targetBox="[229,1361,677,1927]" targetPageId="6">
<paragraph blockId="6.[136,1452,1951,2081]" pageId="6" pageNumber="143">
<emphasis bold="true" box="[136,313,1951,1973]" pageId="6" pageNumber="143">FIGURES 11–14</emphasis>
. Reproductive structures of 
<taxonomicName authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[585,817,1951,1972]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="6" pageNumber="143" phylum="Rhodophyta" rank="species" species="lysonensis">
<emphasis box="[585,817,1951,1972]" italics="true" pageId="6" pageNumber="143">Meristotheca lysonensis</emphasis>
</taxonomicName>
X.-V. Nguyen, X.-T. Nguyen, Kittle 
<emphasis box="[1172,1189,1952,1973]" italics="true" pageId="6" pageNumber="143">et</emphasis>
McDermid, 
<emphasis box="[1313,1381,1952,1973]" italics="true" pageId="6" pageNumber="143">sp. nov</emphasis>
. (IONCS210215b) 11. Tetrasporangial initial laterally pit-connected (arrowhead) to its parental cell. 12. Tetrasporangium basally pit-connected (arrowhead) to the parental cells. 13. Zonately divided tetrasporangium (arrowhead) with a basal pit-connection with the parental cell. 14. Spermatangia (arrowhead) produced from the outermost cortical cells (ION-CS210301b).
</paragraph>
</caption>
<paragraph blockId="7.[136,1452,158,508]" pageId="7" pageNumber="144">
R 
<emphasis bold="true" box="[208,297,158,184]" pageId="7" pageNumber="144">emarks</emphasis>
:A morphological comparison of 
<taxonomicName authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[664,814,158,184]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="lysonensis">
<emphasis box="[664,814,158,184]" italics="true" pageId="7" pageNumber="144">M. lysonensis</emphasis>
</taxonomicName>
and other related species is given in 
<tableCitation box="[1211,1293,158,184]" captionStart="TABLE 1" captionStartId="7.[137,229,556,580]" captionText="TABLE 1. Comparative morphology for Meristotheca lysonensis and five other related flattened Meristotheca species (Rhodophyta). –: data not available." pageId="7" pageNumber="144">Table 1</tableCitation>
. 
<emphasis italics="true" pageId="7" pageNumber="144">
<taxonomicName authority="Nguyen &amp; Nguyen &amp; Kittle Iii, 2022" authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="lysonensis" status="sp. nov.">Meristotheca lysonensis</taxonomicName>
<taxonomicNameLabel box="[257,340,195,220]" pageId="7" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName authorityName="Okamura" authorityYear="1930" box="[484,595,195,220]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="coacta">
<emphasis box="[484,595,195,220]" italics="true" pageId="7" pageNumber="144">M. coacta</emphasis>
</taxonomicName>
based on 
<emphasis box="[703,860,194,220]" italics="true" pageId="7" pageNumber="144">M. lysonensis’</emphasis>
larger size and greater blade thickness, blades that do not bear proliferations along the margins, and usually smooth blade margins. 
<taxonomicName authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[1004,1157,230,256]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="lysonensis">
<emphasis box="[1004,1157,230,256]" italics="true" pageId="7" pageNumber="144">M. lysonensis</emphasis>
</taxonomicName>
clade habit is compressed to flattened. Pit-connections between tetrasporangial initials and the supporting parental cells change from lateral (young tetrasporangium) to basal (divided tetrasporangium). 
<taxonomicName authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[814,1083,302,328]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="lysonensis">
<emphasis box="[814,1083,302,328]" italics="true" pageId="7" pageNumber="144">Meristotheca lysonensis</emphasis>
</taxonomicName>
also differs from 
<taxonomicName authorityName="P. W. Gabrielson &amp; Kraft" authorityYear="1985" box="[1279,1451,302,328]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="procumbens">
<emphasis box="[1279,1451,302,328]" italics="true" pageId="7" pageNumber="144">M. procumbens</emphasis>
</taxonomicName>
in terms of thallus size and thickness, and the smaller medulla thickness in cross-sections. Position of pit-connections between tetrasporangial initials and the parental cells differ: basal position in 
<emphasis box="[1033,1286,374,400]" italics="true" pageId="7" pageNumber="144">
<taxonomicName authority="Nguyen &amp; Nguyen &amp; Kittle Iii, 2022" authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[1033,1188,374,400]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="lysonensis" status="sp. nov.">M. lysonensis</taxonomicName>
<taxonomicNameLabel box="[1198,1286,375,400]" pageId="7" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, unlike lateral position in 
<taxonomicName authorityName="P. W. Gabrielson &amp; Kraft" authorityYear="1985" box="[260,432,410,436]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="procumbens">
<emphasis box="[260,432,410,436]" italics="true" pageId="7" pageNumber="144">M. procumbens</emphasis>
</taxonomicName>
. This new species also differs from 
<taxonomicName box="[830,969,410,436]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="papulosa">
<emphasis box="[830,969,410,436]" italics="true" pageId="7" pageNumber="144">M. papulosa</emphasis>
</taxonomicName>
because 
<taxonomicName authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[1069,1220,410,436]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="lysonensis">
<emphasis box="[1069,1220,410,436]" italics="true" pageId="7" pageNumber="144">M. lysonensis</emphasis>
</taxonomicName>
has many secondary holdfasts compared to single or a few erect blades arising from a discoid holdfast in 
<taxonomicName box="[1140,1285,446,472]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="papulosa">
<emphasis box="[1140,1285,446,472]" italics="true" pageId="7" pageNumber="144">M. papulosa</emphasis>
</taxonomicName>
. There are no branchlets in 
<taxonomicName authorityName="Nguyen &amp; Nguyen &amp; Kittle Iii" authorityYear="2022" box="[287,439,482,508]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="lysonensis">
<emphasis box="[287,439,482,508]" italics="true" pageId="7" pageNumber="144">M. lysonensis</emphasis>
</taxonomicName>
whereas 
<taxonomicName box="[545,685,482,508]" class="Florideophyceae" family="Solieriaceae" genus="Meristotheca" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="144" phylum="Rhodophyta" rank="species" species="papulosa">
<emphasis box="[545,685,482,508]" italics="true" pageId="7" pageNumber="144">M. papulosa</emphasis>
</taxonomicName>
shows branchlets.
</paragraph>
</subSubSection>
</treatment>
</document>