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<document id="3BBDFE77597898A2442DA1DCF2879A18" ID-CLB-Dataset="21325" ID-DOI="10.11646/zootaxa.4759.4.12" ID-GBIF-Dataset="146289cf-b286-4e54-9430-0418d9039312" ID-ISSN="1175-5326" ID-Zenodo-Dep="3974746" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1596728105938" checkinUser="felipe" docAuthor="Dufresnes, Christophe &amp; Crochet, Pierre-André" docDate="2020" docId="03A6FF5BFFE9B83F0DDB31B5FEDF35D1" docLanguage="en" docName="zootaxa.4759.4.12.pdf.imf" docOrigin="Zootaxa 4759 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hyla" docType="treatment" docVersion="9" lastPageNumber="598" masterDocId="FF9F8723FFE9B83E0D1C3422FFE23757" masterDocTitle="The valid nomen for the tree frog (genus Hyla) of Tunisia and Eastern Algeria" masterLastPageNumber="599" masterPageNumber="597" pageNumber="597" updateTime="1698844160887" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="085C01608FD4CFE0B98233B3AF14B1F8">The valid nomen for the tree frog (genus Hyla) of Tunisia and Eastern Algeria</mods:title>
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<mods:namePart id="8F9F5B64A1466EF6223AA31BC2E1EF2A">Dufresnes, Christophe</mods:namePart>
<mods:affiliation id="BBA8D4244E7D26B6EDB6D6D30CB49B30">LASER team, College of Biology and the Environment, Nanjing Forestry University, Nanjing, China</mods:affiliation>
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<mods:namePart id="33A8111934B1A071940489799202DE50">Crochet, Pierre-André</mods:namePart>
<mods:affiliation id="767693B5CEEFF05B03624CE815E9CF52">CEFE, CNRS, Montpellier, France</mods:affiliation>
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<mods:date id="B6F8F443A1F10BA1057D5B6C3CE4858D">2020</mods:date>
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<treatment id="03A6FF5BFFE9B83F0DDB31B5FEDF35D1" ID-DOI="http://doi.org/10.5281/zenodo.4323858" ID-GBIF-Taxon="166240037" ID-Zenodo-Dep="4323858" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1" httpUri="http://treatment.plazi.org/id/03A6FF5BFFE9B83F0DDB31B5FEDF35D1" lastPageId="1" lastPageNumber="598" pageId="0" pageNumber="597">
<subSubSection id="C3151DC6FFE9B83E0DDB31B5FF1232F8" box="[199,240,1431,1455]" pageId="0" pageNumber="597" type="nomenclature">
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<heading id="D0F8F921FFE9B83E0DDB31B5FF1232F8" box="[199,240,1431,1455]" level="2" pageId="0" pageNumber="597">
<taxonomicName id="4C0F35CEFFE9B83E0DDB31B5FF1232F8" ID-CoL="4ZYM" authority="Laurenti, 1768" box="[199,240,1431,1455]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="0" pageNumber="597" phylum="Chordata" rank="genus">
<location id="8ED01896FFE9B83E0DDB31B5FF1232F8" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E0DDB31B5FF1232F8" box="[199,240,1431,1455]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="The" pageId="0" pageNumber="597">The</location>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3151DC6FFE9B83F0DEB31B5FEDF35D1" lastPageId="1" lastPageNumber="598" pageId="0" pageNumber="597" type="discussion">
<paragraph id="8BB04E4DFFE9B83E0DEB31B5FD7430D4" blockId="0.[151,1437,603,2032]" pageId="0" pageNumber="597">
first condition to be met for a nomen to be valid according to the
<location id="8ED01896FFE9B83E0EB031B5FC0632F8" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E0EB031B5FC0632F8" box="[940,996,1431,1455]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="Code" pageId="0" pageNumber="597">Code</location>
is availability.
<location id="8ED01896FFE9B83E099931B5FB5132F8" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E099931B5FB5132F8" box="[1157,1203,1431,1455]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="Any" pageId="0" pageNumber="597">Any</location>
species-group nomen proposed after 1999 is only available when included in a published work (as defined by Art. 8) that explicitly designates a name-bearing type (Art. 16.4).
<location id="8ED01896FFE9B83E0CEB31FDFD9732A0" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E0CEB31FDFD9732A0" box="[503,629,1503,1527]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="Regrettably" pageId="0" pageNumber="597">Regrettably</location>
, the description of
<taxonomicName id="4C0F35CEFFE9B83E0E5D31FDFC3D32A0" authorityName="Escoriza, Ben Hassine" authorityYear="2019" box="[833,991,1503,1527]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="0" pageNumber="597" phylum="Chordata" rank="species" species="numidica">
<emphasis id="B97B925FFFE9B83E0E5D31FDFC3D32A0" box="[833,991,1503,1527]" italics="true" pageId="0" pageNumber="597">Hyla numidica</emphasis>
</taxonomicName>
fails to specify any type specimen.
<location id="8ED01896FFE9B83E084331C2FA9732A0" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E084331C2FA9732A0" box="[1375,1397,1504,1527]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="In" pageId="0" pageNumber="597">In</location>
the corresponding paragraph (“Type specimen”, p.
<quantity id="4CF7E3A8FFE9B83E0F933221FD30314C" box="[655,722,1539,1563]" metricMagnitude="0" metricUnit="m" metricValue="4.826" pageId="0" pageNumber="597" unit="in" value="190.0">190 in</quantity>
<bibRefCitation id="EF9E33BCFFE9B83E0FC43221FBF8314C" author="Escoriza, D. &amp; Ben Hassine, J." box="[728,1050,1539,1563]" pageId="0" pageNumber="597" refId="ref1558" refString="Escoriza, D. &amp; Ben Hassine, J. (2019) Amphibians of North Africa. 1 st Edition. Elsevier, London, 335 pp. https: // doi. org / 10.1016 / C 2017 - 0 - 03447 - X" type="book" year="2019">Escoriza &amp; Ben Hassine 2019</bibRefCitation>
), the authors only mention the name of a collection and a type locality: “
<taxonomicName id="4C0F35CEFFE9B83E0F033205FD5C3168" authority="Dufresnes &amp; Crochet, 2020" authorityName="Dufresnes &amp; Crochet" authorityYear="2020" box="[543,702,1575,1599]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="0" pageNumber="597" phylum="Chordata" rank="species" species="numidica" status="sp. nov.">
<emphasis id="B97B925FFFE9B83E0F033205FD5C3168" box="[543,702,1575,1599]" italics="true" pageId="0" pageNumber="597">Hyla numidica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2482F24FFE9B83E0FDB320AFCFD3168" box="[711,799,1576,1599]" pageId="0" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
: MCNB (Museu de Ciències Naturals de Barcelona). Type locality: Nefza (
<collectingCountry id="F3180EDDFFE9B83E0C5D3269FE7A3134" box="[321,408,1611,1635]" name="Tunisia" pageId="0" pageNumber="597">Tunisia</collectingCountry>
)”.
<location id="8ED01896FFE9B83E0CAF326EFE363134" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E0CAF326EFE363134" box="[435,468,1612,1635]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="No" pageId="0" pageNumber="597">No</location>
indication about the specimens that should be considered as the types of
<taxonomicName id="4C0F35CEFFE9B83E09C93269FA933134" authorityName="Dufresnes &amp; Crochet" authorityYear="2020" box="[1237,1393,1611,1635]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="0" pageNumber="597" phylum="Chordata" rank="species" species="numidica">
<emphasis id="B97B925FFFE9B83E09C93269FA933134" box="[1237,1393,1611,1635]" italics="true" pageId="0" pageNumber="597">Hyla numidica</emphasis>
</taxonomicName>
can be found elsewhere in their work.
<location id="8ED01896FFE9B83E0F1F324DFD8D31D0" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E0F1F324DFD8D31D0" box="[515,623,1647,1671]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="Therefore" pageId="0" pageNumber="597">Therefore</location>
, the conditions of Art. 16.4 (“
<location id="8ED01896FFE9B83E0EB5324DFC0E31D0" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E0EB5324DFC0E31D0" box="[937,1004,1647,1671]" country="Tunisia" county="ZooBank" municipality="Official Register of Zoological Nomenclature" name="Every" pageId="0" pageNumber="597">Every</location>
new specific and subspecific name published after 1999, …, must be accompanied in the original publication by the explicit fixation of a
<typeStatus id="54B4F0EFFFE9B83E09AD32B1FAF331FC" box="[1201,1297,1683,1707]" pageId="0" pageNumber="597" type="holotype">holotype</typeStatus>
, or
<typeStatus id="54B4F0EFFFE9B83E082432B6FA7A31FB" box="[1336,1432,1684,1708]" pageId="0" pageNumber="597" type="syntype">syntypes</typeStatus>
, …”) are not met, as the mere designation of a collection is clearly insufficient to be considered as an explicit fixation of name-bearing types. A registration in the
<collectingMunicipality id="6BD4D437FFE9B83E0F6D32F9FBB831A4" box="[625,1114,1755,1779]" pageId="0" pageNumber="597">Official Register of Zoological Nomenclature</collectingMunicipality>
(
<collectingCounty id="62D136C1FFE9B83E097432F9FB2D31A4" box="[1128,1231,1755,1779]" pageId="0" pageNumber="597">ZooBank</collectingCounty>
) made
<emphasis id="B97B925FFFE9B83E080232FEFA7E31A4" box="[1310,1436,1756,1779]" italics="true" pageId="0" pageNumber="597">a posteriori</emphasis>
(on
<date id="FFB1688DFFE9B83E0DDE32DDFE703040" box="[194,402,1788,1815]" pageId="0" pageNumber="597" value="2019-09-09">
<collectingDate id="EFF59165FFE9B83E0DDE32DDFE703040" box="[194,402,1788,1815]" pageId="0" pageNumber="597" value="2019-09-09">
September 9
<superScript id="7C7AE305FFE9B83E0C5432DEFEB6305C" attach="left" box="[328,340,1788,1803]" fontSize="6" pageId="0" pageNumber="597">th</superScript>
2019
</collectingDate>
</date>
) and mentioning the specimen MZB 2019-0924 as
<typeStatus id="54B4F0EFFFE9B83E0EA732DDFBF93040" box="[955,1051,1791,1815]" pageId="0" pageNumber="597" type="holotype">holotype</typeStatus>
, does not fulfil the requirements for availability either, since the designation needs to be done in the work itself—a ZooBank registration is not a published work in the sense of the Code. As a consequence,
<taxonomicName id="4C0F35CEFFE9B83E0FA93365FB6D3008" authority="Escoriza, Ben Hassine, 2019" authorityName="Escoriza, Ben Hassine" authorityYear="2019" box="[693,1167,1863,1887]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="0" pageNumber="597" phylum="Chordata" rank="species" species="numidica">
<emphasis id="B97B925FFFE9B83E0FA93365FCB13008" box="[693,851,1863,1887]" italics="true" pageId="0" pageNumber="597">Hyla numidica</emphasis>
<bibRefCitation id="EF9E33BCFFE9B83E0E473365FB6D3008" author="Escoriza, D. &amp; Ben Hassine, J." box="[859,1167,1863,1887]" pageId="0" pageNumber="597" refId="ref1558" refString="Escoriza, D. &amp; Ben Hassine, J. (2019) Amphibians of North Africa. 1 st Edition. Elsevier, London, 335 pp. https: // doi. org / 10.1016 / C 2017 - 0 - 03447 - X" type="book" year="2019">Escoriza, Ben Hassine, 2019</bibRefCitation>
</taxonomicName>
is an unavailable nomen that cannot be used in taxonomic nomenclature.
</paragraph>
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In contrast, the description of
<taxonomicName id="4C0F35CEFFE9B83E0F1E33ADFD3D30F0" authorityName="Dufresnes, Beddek, Skorinov, Fumagalli, Perrin, Crochet, Litvinchuk" authorityYear="2019" box="[514,735,1935,1959]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="0" pageNumber="597" phylum="Chordata" rank="species" species="carthaginiensis">
<emphasis id="B97B925FFFE9B83E0F1E33ADFD3D30F0" box="[514,735,1935,1959]" italics="true" pageId="0" pageNumber="597">Hyla carthaginiensis</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EF9E33BCFFE9B83E0E7633ADFBB430F0" author="Dufresnes, C. &amp; Beddek, M. &amp; Skorinov, D. V. &amp; Fumagalli, L. &amp; Perrin, N. &amp; Crochet, P. A. &amp; Litvinchuk, S. N." box="[874,1110,1935,1959]" pageId="0" pageNumber="597" pagination="291 - 299" refId="ref1466" refString="Dufresnes, C., Beddek, M., Skorinov, D. V., Fumagalli, L., Perrin, N., Crochet, P. A. &amp; Litvinchuk, S. N. (2019) Diversification and speciation in tree frogs from the Maghreb (Hyla meridionalis sensu lato), with description of a new African endemic. Molecular Phylogenetics and Evolution, 134, 291 - 299. https: // doi. org / 10.1016 / j. ympev. 2019.02.009" type="journal article" year="2019">
Dufresnes
<emphasis id="B97B925FFFE9B83E0EC733B2FBF230F0" box="[987,1040,1935,1959]" italics="true" pageId="0" pageNumber="597">et al.</emphasis>
(2019
</bibRefCitation>
, p. 296-298) specifically states the type specimen (
<typeStatus id="54B4F0EFFFE9B83E0C7A3391FE27309C" box="[358,453,1971,1995]" pageId="0" pageNumber="597" type="holotype">holotype</typeStatus>
ZISP 13722, portrayed in their Fig. 6) and the type locality (
<location id="8ED01896FFE9B83E09533391FB70309C" LSID="urn:lsid:plazi:treatment:03A6FF5BFFE9B83F0DDB31B5FEDF35D1:8ED01896FFE9B83E09533391FB70309C" box="[1103,1170,1971,1995]" country="Liberia" municipality="Featured" name="Qusur" pageId="0" pageNumber="597">Qusur</location>
<collectingCountry id="F3180EDDFFE9B83E09853391FB0B309C" box="[1177,1257,1971,1995]" name="Liberia" pageId="0" pageNumber="597">Liberia</collectingCountry>
,
<collectingCountry id="F3180EDDFFE9B83E09EF3391FAA4309C" box="[1267,1350,1971,1995]" name="Tunisia" pageId="0" pageNumber="597">Tunisia</collectingCountry>
), whose identity was confirmed by genome-wide data.
<collectingMunicipality id="6BD4D437FFE9B83E0F9533F5FD0430B8" box="[649,742,2007,2031]" pageId="0" pageNumber="597">Featured</collectingMunicipality>
in the
<date id="FFB1688DFFE9B83E0E2D33FAFC7C30B8" box="[817,926,2007,2032]" pageId="0" pageNumber="597" value="2019-05">
<collectingDate id="EFF59165FFE9B83E0E2D33FAFC7C30B8" box="[817,926,2007,2032]" pageId="0" pageNumber="597" value="2019-05">May 2019</collectingDate>
</date>
issue of
<collectorName id="26FA2B9BFFE9B83E0EE333F5FAE830B8" box="[1023,1290,2007,2031]" pageId="0" pageNumber="597">Molecular Phylogenetics</collectorName>
&amp;
<collectorName id="26FA2B9BFFE9B83E083033F5FA7A30B8" box="[1324,1432,2007,2031]" pageId="0" pageNumber="597">Evolution</collectorName>
,
<bibRefCitation id="EF9E33BCFFE8B83F0D8B34B8FE7237E5" author="Dufresnes, C. &amp; Beddek, M. &amp; Skorinov, D. V. &amp; Fumagalli, L. &amp; Perrin, N. &amp; Crochet, P. A. &amp; Litvinchuk, S. N." box="[151,400,154,178]" pageId="1" pageNumber="598" pagination="291 - 299" refId="ref1466" refString="Dufresnes, C., Beddek, M., Skorinov, D. V., Fumagalli, L., Perrin, N., Crochet, P. A. &amp; Litvinchuk, S. N. (2019) Diversification and speciation in tree frogs from the Maghreb (Hyla meridionalis sensu lato), with description of a new African endemic. Molecular Phylogenetics and Evolution, 134, 291 - 299. https: // doi. org / 10.1016 / j. ympev. 2019.02.009" type="journal article" year="2019">
Dufresnes
<emphasis id="B97B925FFFE8B83F0C1734B9FEA037E5" box="[267,322,154,178]" italics="true" pageId="1" pageNumber="598">et al.</emphasis>
(2019)
</bibRefCitation>
was published online by
<collectorName id="26FA2B9BFFE8B83F0FBA34B8FCE437E5" box="[678,774,154,178]" pageId="1" pageNumber="598">February</collectorName>
15
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(time-stamped p. 291).
<collectorName id="26FA2B9BFFE8B83F092534B8FB8537E5" box="[1081,1127,154,178]" pageId="1" pageNumber="598">And</collectorName>
because the publication was registered in
<collectorName id="26FA2B9BFFE8B83F0C34349CFE693781" box="[296,395,190,214]" pageId="1" pageNumber="598">ZooBank</collectorName>
beforehand (p. 296), the nomenclatural act it contains became active at this date already—see
<collectorName id="26FA2B9BFFE8B83F0D8B34C0FF0337AD" box="[151,225,226,250]" pageId="1" pageNumber="598">Article</collectorName>
8.5 of the
<collectorName id="26FA2B9BFFE8B83F0C4E34C0FE6837AD" box="[338,394,226,250]" pageId="1" pageNumber="598">Code</collectorName>
regarding online publication.
<collectorName id="26FA2B9BFFE8B83F0FD634C0FD1137AD" box="[714,755,226,250]" pageId="1" pageNumber="598">The</collectorName>
nomen
<taxonomicName id="4C0F35CEFFE8B83F0E5434C0FBC437AD" authorityName="Dufresnes, Beddek, Skorinov, Fumagalli, Perrin, Crochet, Litvinchuk" authorityYear="2019" box="[840,1062,226,250]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="carthaginiensis">
<emphasis id="B97B925FFFE8B83F0E5434C0FBC437AD" box="[840,1062,226,250]" italics="true" pageId="1" pageNumber="598">Hyla carthaginiensis</emphasis>
</taxonomicName>
is thus available, and as detailed in
<bibRefCitation id="EF9E33BCFFE8B83F0D8B3524FE703649" author="Dufresnes, C. &amp; Beddek, M. &amp; Skorinov, D. V. &amp; Fumagalli, L. &amp; Perrin, N. &amp; Crochet, P. A. &amp; Litvinchuk, S. N." box="[151,402,262,286]" pageId="1" pageNumber="598" pagination="291 - 299" refId="ref1466" refString="Dufresnes, C., Beddek, M., Skorinov, D. V., Fumagalli, L., Perrin, N., Crochet, P. A. &amp; Litvinchuk, S. N. (2019) Diversification and speciation in tree frogs from the Maghreb (Hyla meridionalis sensu lato), with description of a new African endemic. Molecular Phylogenetics and Evolution, 134, 291 - 299. https: // doi. org / 10.1016 / j. ympev. 2019.02.009" type="journal article" year="2019">
Dufresnes
<emphasis id="B97B925FFFE8B83F0C173525FEA03649" box="[267,322,262,286]" italics="true" pageId="1" pageNumber="598">et al.</emphasis>
(2019)
</bibRefCitation>
, all other nomina proposed for
<collectorName id="26FA2B9BFFE8B83F0FFA3524FC633649" box="[742,897,262,286]" pageId="1" pageNumber="598">Mediterranean</collectorName>
tree frogs correspond to the North-
<collectorName id="26FA2B9BFFE8B83F081B3524FABB3649" box="[1287,1369,262,286]" pageId="1" pageNumber="598">African</collectorName>
/
<collectorName id="26FA2B9BFFE8B83F087D3524FF2B3616" pageId="1" pageNumber="598">European</collectorName>
species (
<taxonomicName id="4C0F35CEFFE8B83F0C313509FE363615" authorityName="Boettger" authorityYear="1874" box="[301,468,298,322]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="B97B925FFFE8B83F0C313509FE363615" box="[301,468,298,322]" italics="true" pageId="1" pageNumber="598">H. meridionalis</emphasis>
</taxonomicName>
).
<figureCitation id="133452C8FFE8B83F0CF63508FDA63615" box="[490,580,298,322]" captionStart="FIGURE 1" captionStartId="1.[151,244,1606,1628]" captionTargetBox="[265,1318,673,1565]" captionTargetId="figure@1.[268,1313,673,1564]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Hyla meridionalis (yellow) and Hyla carthaginiensis (red). For each, the type localities of proposed nomina are shown by stars (large: oldest available; small: junior synonym and/or unavailable), with their order of priority listed in the inset. The exact origin of the Hyla africana holotype was not documented, but historical DNA barcoding indicated genetic similarities with pupulations from Morocco and the introduced ranges, corresponding to H. meridionalis (Dufresnes et al. 2019). Photo credits: CD (Hyla meridionalis from France) and Spartak Litvinchuk (Hyla carthaginiensis from Tunisia)." figureDoi="http://doi.org/10.5281/zenodo.3974750" httpUri="https://zenodo.org/record/3974750/files/figure.png" pageId="1" pageNumber="598">Figure 1</figureCitation>
provides a visual summary.
</paragraph>
<paragraph id="8BB04E4DFFE8B83F0DDB356CFF3F36A1" blockId="1.[151,1437,151,646]" pageId="1" pageNumber="598">
Therefore, the only name available for the Tunisian/Numidian species is
<taxonomicName id="4C0F35CEFFE8B83F0EC6356CFB5B3631" authorityName="Dufresnes, Beddek, Skorinov, Fumagalli, Perrin, Crochet, Litvinchuk" authorityYear="2019" box="[986,1209,334,358]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="carthaginiensis">
<emphasis id="B97B925FFFE8B83F0EC6356CFB5B3631" box="[986,1209,334,358]" italics="true" pageId="1" pageNumber="598">Hyla carthaginiensis</emphasis>
</taxonomicName>
. Note that even with a valid description,
<taxonomicName id="4C0F35CEFFE8B83F0C6E3550FDF136DD" authorityName="Escoriza, Ben Hassine" authorityYear="2019" box="[370,531,370,394]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="numidica">
<emphasis id="B97B925FFFE8B83F0C6E3550FDF136DD" box="[370,531,370,394]" italics="true" pageId="1" pageNumber="598">Hyla numidica</emphasis>
</taxonomicName>
would have been a junior synonym: according to the publishers website (https:// www.elsevier.com/books/amphibians-of-north-africa/escoriza/978-0-12-815476-2),
<bibRefCitation id="EF9E33BCFFE8B83F090535B4FA8936F9" author="Escoriza, D. &amp; Ben Hassine, J." box="[1049,1387,406,430]" pageId="1" pageNumber="598" refId="ref1558" refString="Escoriza, D. &amp; Ben Hassine, J. (2019) Amphibians of North Africa. 1 st Edition. Elsevier, London, 335 pp. https: // doi. org / 10.1016 / C 2017 - 0 - 03447 - X" type="book" year="2019">Escoriza &amp; Ben Hassine (2019)</bibRefCitation>
was released on
<date id="FFB1688DFFE8B83F0C0A3598FE333685" box="[278,465,442,466]" pageId="1" pageNumber="598" value="2019-03-19">March 19th 2019</date>
, hence a few weeks later than the online publication of
<taxonomicName id="4C0F35CEFFE8B83F093C3598FB1D3685" authorityName="Dufresnes, Beddek, Skorinov, Fumagalli, Perrin, Crochet, Litvinchuk" authorityYear="2019" box="[1056,1279,442,466]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="carthaginiensis">
<emphasis id="B97B925FFFE8B83F093C3598FB1D3685" box="[1056,1279,442,466]" italics="true" pageId="1" pageNumber="598">Hyla carthaginiensis</emphasis>
</taxonomicName>
(
<date id="FFB1688DFFE8B83F08113598FF3336A1" pageId="1" pageNumber="598" value="2019-02-15">
February 15
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2019
</date>
).
</paragraph>
<paragraph id="8BB04E4DFFE8B83F0DDB3620FEDF35D1" blockId="1.[151,1437,151,646]" pageId="1" pageNumber="598">
In conclusion, the two tree frog species that inhabit North Africa are
<taxonomicName id="4C0F35CEFFE8B83F0ED53620FB6C354D" authorityName="Boettger" authorityYear="1874" box="[969,1166,514,538]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="B97B925FFFE8B83F0ED53620FB6C354D" box="[969,1166,514,538]" italics="true" pageId="1" pageNumber="598">Hyla meridionalis</emphasis>
</taxonomicName>
, which is widespread in
<collectingCountry id="F3180EDDFFE8B83F0D8B3604FF18356A" box="[151,250,550,573]" name="Morocco" pageId="1" pageNumber="598">Morocco</collectingCountry>
,
<collectingCountry id="F3180EDDFFE8B83F0C183604FEB53569" box="[260,343,550,574]" name="Algeria" pageId="1" pageNumber="598">Algeria</collectingCountry>
, as well as in introduced European and insular ranges; and
<taxonomicName id="4C0F35CEFFE8B83F0ECC3604FB4D3569" authorityName="Dufresnes, Beddek, Skorinov, Fumagalli, Perrin, Crochet, Litvinchuk" authorityYear="2019" box="[976,1199,550,574]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="carthaginiensis">
<emphasis id="B97B925FFFE8B83F0ECC3604FB4D3569" box="[976,1199,550,574]" italics="true" pageId="1" pageNumber="598">Hyla carthaginiensis</emphasis>
</taxonomicName>
, which is restricted to
<collectingCountry id="F3180EDDFFE8B83F0D8B3668FF043535" box="[151,230,586,610]" name="Tunisia" pageId="1" pageNumber="598">Tunisia</collectingCountry>
and eastern
<collectingCountry id="F3180EDDFFE8B83F0C763668FE583535" box="[362,442,586,610]" name="Algeria" pageId="1" pageNumber="598">Algeria</collectingCountry>
(
<bibRefCitation id="EF9E33BCFFE8B83F0CDB3668FD523535" author="Dufresnes, C. &amp; Beddek, M. &amp; Skorinov, D. V. &amp; Fumagalli, L. &amp; Perrin, N. &amp; Crochet, P. A. &amp; Litvinchuk, S. N." box="[455,688,586,610]" pageId="1" pageNumber="598" pagination="291 - 299" refId="ref1466" refString="Dufresnes, C., Beddek, M., Skorinov, D. V., Fumagalli, L., Perrin, N., Crochet, P. A. &amp; Litvinchuk, S. N. (2019) Diversification and speciation in tree frogs from the Maghreb (Hyla meridionalis sensu lato), with description of a new African endemic. Molecular Phylogenetics and Evolution, 134, 291 - 299. https: // doi. org / 10.1016 / j. ympev. 2019.02.009" type="journal article" year="2019">
Dufresnes
<emphasis id="B97B925FFFE8B83F0F273669FD923535" box="[571,624,586,610]" italics="true" pageId="1" pageNumber="598">et al.</emphasis>
2019
</bibRefCitation>
).
<taxonomicName id="4C0F35CEFFE8B83F0FDD3668FCBC3535" authorityName="Escoriza, Ben Hassine" authorityYear="2019" box="[705,862,586,610]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="598" phylum="Chordata" rank="species" species="numidica">
<emphasis id="B97B925FFFE8B83F0FDD3668FCBC3535" box="[705,862,586,610]" italics="true" pageId="1" pageNumber="598">Hyla numidica</emphasis>
</taxonomicName>
is an unavailable name (and is also younger) and thus cannot be used.
</paragraph>
</subSubSection>
</treatment>
</document>