276 lines
34 KiB
XML
276 lines
34 KiB
XML
<document id="5EBE213C63D3AA91CD9F8D470451FA58" ID-CLB-Dataset="28123" ID-DOI="10.11646/zootaxa.4568.3.2" ID-GBIF-Dataset="d40eb95b-5a98-418a-b607-7c2cbec7c897" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601607" ID-ZooBank="8FD0AC89-424E-4CAC-92A5-A5600A481140" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553172834442" checkinUser="plazi" docAuthor="Kozłowska, Anna, Bates, Denis, Zalasiewicz, Jan & Radzevičius, Sigitas" docDate="2019" docId="03A687FCFFFC98568FA2D45BF8FAFC04" docLanguage="en" docName="zootaxa.4568.3.2.pdf" docOrigin="Zootaxa 4568 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gothograptus obtectus , Kozlowska-Dawidziuk 1990" docType="treatment" docVersion="8" lastPageNumber="462" masterDocId="FF9FFF84FFEA984D8F35D37CFD67FFA1" masterDocTitle="Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania" masterLastPageNumber="469" masterPageNumber="435" pageNumber="457" updateTime="1698703580205" updateUser="ExternalLinkService">
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<mods:title id="D9EDA0991A160178319994A6CC136A15">Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania</mods:title>
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<mods:namePart id="EA0820F5878DEC492D34174AFE4EB92D">Kozłowska, Anna</mods:namePart>
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<mods:namePart id="0F95E81AA2B3B4BA1DD3106DE58353D2">Bates, Denis</mods:namePart>
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<mods:namePart id="9ECC22A782E30C27A009467ED1287A5B">Zalasiewicz, Jan</mods:namePart>
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<mods:namePart id="F641FA0DAE212056E5F433616E42FEF8">Radzevičius, Sigitas</mods:namePart>
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<treatment id="03A687FCFFFC98568FA2D45BF8FAFC04" ID-DOI="http://doi.org/10.5281/zenodo.5935886" ID-GBIF-Taxon="154474984" ID-Zenodo-Dep="5935886" LSID="urn:lsid:plazi:treatment:03A687FCFFFC98568FA2D45BF8FAFC04" httpUri="http://treatment.plazi.org/id/03A687FCFFFC98568FA2D45BF8FAFC04" lastPageId="27" lastPageNumber="462" pageId="22" pageNumber="457">
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<subSubSection id="C3156561FFFC985B8FA2D45BFE75F8E3" box="[151,786,1831,1858]" pageId="22" pageNumber="457" type="nomenclature">
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<paragraph id="8BB036EAFFFC985B8FA2D45BFE75F8E3" blockId="22.[151,786,1831,1891]" box="[151,786,1831,1858]" pageId="22" pageNumber="457">
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<heading id="D0F88186FFFC985B8FA2D45BFE75F8E3" bold="true" box="[151,786,1831,1858]" fontSize="11" level="1" pageId="22" pageNumber="457" reason="1">
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<taxonomicName id="4C0F4D69FFFC985B8FA2D45BFE75F8E3" authority="Kozlowska-Dawidziuk, 1990" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[151,786,1831,1858]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="22" pageNumber="457" phylum="Hemichordata" rank="species" species="obtectus">
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<emphasis id="B97BEAF8FFFC985B8FA2D45BFE75F8E3" bold="true" box="[151,786,1831,1858]" pageId="22" pageNumber="457">
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<emphasis id="B97BEAF8FFFC985B8FA2D45BFCC0F8E0" bold="true" box="[151,423,1831,1857]" italics="true" pageId="22" pageNumber="457">Gothograptus obtectus</emphasis>
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<bibRefCitation id="EF9E4B1BFFFC985B8E9AD454FE75F8E3" author="Kozlowska-Dawidziuk, A." box="[431,786,1832,1858]" pageId="22" pageNumber="457" pagination="191 - 209" refId="ref11319" refString="Kozlowska-Dawidziuk, A. (1990) The genus Gothograptus (Graptolithina) from the Wenlock of Poland. Acta Palaeontologica Polonica, 35, 191 - 209." type="journal article" year="1990">Kozłowska-Dawidziuk, 1990</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156561FFFC985B8FA2D436FC73F8C2" box="[151,276,1866,1891]" pageId="22" pageNumber="457" type="description">
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<paragraph id="8BB036EAFFFC985B8FA2D436FC73F8C2" blockId="22.[151,786,1831,1891]" box="[151,276,1866,1891]" pageId="22" pageNumber="457">
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<figureCitation id="13342A6FFFFC985B8FA2D436FC73F8C2" box="[151,276,1866,1891]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStart-2="FIGURE 21" captionStartId-0="25.[151,250,1930,1952]" captionStartId-1="26.[151,250,1588,1610]" captionStartId-2="28.[151,250,1852,1874]" captionTargetBox-0="[176,1412,193,1916]" captionTargetBox-1="[170,1420,193,1578]" captionTargetBox-2="[263,1329,193,1852]" captionTargetId-0="figure@25.[176,1412,193,1916]" captionTargetId-1="figure@26.[170,1420,193,1578]" captionTargetId-2="figure@28.[263,1329,193,1834]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="28" captionText-0="FIGURE 19. Mature tubarium with appendix of Gothograptus obtectus Kozłowska-Dawidziuk, 1990 from the Bartoszyce IG- 1 core, depth 1677.5 m, lundgreni Biozone, Poland, ZPAL G.55/48. A. enlargement of distal part of specimen. B. enlargement of proximal end. C. whole specimen, reverse view. Abbreviations: lat. ap.—lateral apertural." captionText-1="FIGURE 20. Tubaria of Gothograptus obtectus Kozłowska-Dawidziuk, 1990, Zawada 1 core, depth 1533.3–1540.0 m, Poland, upper part of lundgreni Biozone. A. tubarium, ZPAL G.XIII/23. B. gerontic tubarium, holotype, ZPAL G.XIII/23. Abbreviations: conn.—connecting, v.—ventral." captionText-2="FIGURE 21. Mature tubaria of Gothograptus obtectus Kozłowska-Dawidziuk, 1990 from Lithuania, Šiupyliai-69 core, depth 1012.0 m, lundgreni Biozone. A, B, E. finite tubarium with no proximal end, ventro-lateral view, VU RET-19; A, whole specimen; B, enlargement of last thecae; E. enlargement of thecal hood. C–D. proximal part of tubarium, obverse view, VU RET-20; C. enlargement of medial part; D. whole specimen; F–H. fragment of tubarium, lateral-ventral view, VU RET-21; F. whole specimen; G. enlargement of proximal part; H. enlargement of distal part. Abbreviations: genic.—geniculum, lat. ap.— lateral apertural, m–v.—mid-ventral." httpUri-0="https://zenodo.org/record/2601657/files/figure.png" httpUri-1="https://zenodo.org/record/2601659/files/figure.png" httpUri-2="https://zenodo.org/record/2601661/files/figure.png" pageId="22" pageNumber="457">Figs 19–21</figureCitation>
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</paragraph>
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</subSubSection>
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<footnote id="E8142AE4FFFC985B8FA2D4EEFFF3F847" pageId="22" pageNumber="457">
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<paragraph id="8BB036EAFFFC985B8FA2D4EEFFF3F847" blockId="22.[151,1436,1938,2022]" pageId="22" pageNumber="457">
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1990.
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<taxonomicName id="4C0F4D69FFFC985B8FE3D4EEFE3CF809" authority=", Kozlowska-Dawidziuk" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[214,859,1938,1960]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="22" pageNumber="457" phylum="Hemichordata" rank="species" species="obtectus" subGenus="Gothograptus">
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<emphasis id="B97BEAF8FFFC985B8FE3D4EEFC04F809" box="[214,355,1938,1960]" italics="true" pageId="22" pageNumber="457">Gothograptus</emphasis>
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(
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<emphasis id="B97BEAF8FFFC985B8E47D4EEFC99F809" box="[370,510,1938,1960]" italics="true" pageId="22" pageNumber="457">Gothograptus</emphasis>
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)
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<emphasis id="B97BEAF8FFFC985B8D38D4EEFF06F809" box="[525,609,1938,1960]" italics="true" pageId="22" pageNumber="457">obtectus</emphasis>
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, Kozłowska-Dawidziuk
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</taxonomicName>
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; pp. 200–202; pl. 24: 1; pl. 25: 3; pl. 26: 1, 3, 4; fig. 5A– D.?2010
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<taxonomicName id="4C0F4D69FFFC985B8FEED4ACFCAFF847" box="[219,456,2000,2022]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="22" pageNumber="457" phylum="Hemichordata" rank="species" species="osgaleae">
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<emphasis id="B97BEAF8FFFC985B8FEED4ACFCAFF847" box="[219,456,2000,2022]" italics="true" pageId="22" pageNumber="457">Gothograptus osgaleae</emphasis>
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</taxonomicName>
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; Maletz, fig. 2A, B.
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</paragraph>
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</footnote>
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<caption id="DF706662FFFD985A8FA2D5ADF9D5F8E4" httpUri="https://zenodo.org/record/2601653/files/figure.png" pageId="23" pageNumber="458" startId="23.[151,250,1745,1767]" targetBox="[150,1436,195,1745]" targetPageId="23">
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<paragraph id="8BB036EAFFFD985A8FA2D5ADF9D5F8E4" blockId="23.[151,1436,1745,1861]" pageId="23" pageNumber="458">
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<emphasis id="B97BEAF8FFFD985A8FA2D5ADFC43F946" bold="true" box="[151,292,1745,1767]" pageId="23" pageNumber="458">FIGURE 17.</emphasis>
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Mature tubaria of
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<taxonomicName id="4C0F4D69FFFD985A8EC1D5ADF949F946" authority="Kozlowska-Dawidziuk, 1990" authorityName="Kozlowska-Dawidziuk" authorityYear="1990" box="[500,1070,1745,1767]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="23" pageNumber="458" phylum="Hemichordata" rank="species" species="kozlowskii">
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<emphasis id="B97BEAF8FFFD985A8EC1D5ADFF91F946" box="[500,758,1745,1767]" italics="true" pageId="23" pageNumber="458">Gothograptus kozlowskii</emphasis>
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Kozłowska-Dawidziuk, 1990
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</taxonomicName>
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from the Šiupyliai-69 core, depth 1009.1 m,
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<emphasis id="B97BEAF8FFFD985A8E30D58CFC00F8A7" box="[261,359,1776,1798]" italics="true" pageId="23" pageNumber="458">lundgreni</emphasis>
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Biozone, Lithuania. A–E. mature tubarium with seven pairs of thecae, VU RET-17; A. whole specimen, ventro-lateral view; B. enlargement of distal part of tubarium; C. enlargement of medial part of tubarium; D, E. orifice of th3
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<superScript id="7C7A9BA2FFFD985A8ABBD472F8F2F8BB" attach="none" box="[1422,1429,1806,1818]" fontSize="5" pageId="23" pageNumber="458">1</superScript>
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. F. fragment of mature tubarium showing overgrown proximal ventral orifice, ventral view, VU RET-18.
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</paragraph>
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</caption>
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<subSubSection id="C3156561FFFD985A8FA2D410F9A1F853" pageId="23" pageNumber="458" type="diagnosis">
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<paragraph id="8BB036EAFFFD985A8FA2D410F9A1F853" blockId="23.[151,1436,1900,2034]" pageId="23" pageNumber="458">
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<emphasis id="B97BEAF8FFFD985A8FA2D410FCE5F824" bold="true" box="[151,386,1900,1925]" pageId="23" pageNumber="458">Emended diagnosis</emphasis>
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. Tubarium usually with eight pairs of thecae and appendix. Genicular structures forming reticulated veil, covering thecal orifice and theca. Veil ended by a thicker list located above the lip of the previous theca. Mid-ventral lists are well-developed in proximal and medial thecae while becoming thinner or not developed in distal thecae. It differs from other
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<taxonomicName id="4C0F4D69FFFD985A8D07D4A5FFA9F850" authorityName=", Frech" authorityYear="1897" box="[562,718,2009,2033]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="23" pageNumber="458" phylum="Hemichordata" rank="genus">
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<emphasis id="B97BEAF8FFFD985A8D07D4A5FFA9F850" box="[562,718,2009,2033]" italics="true" pageId="23" pageNumber="458">Gothograptus</emphasis>
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</taxonomicName>
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species in the extreme development of veils.
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</paragraph>
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</subSubSection>
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<caption id="DF706662FFF298558FA2D4EEFFB7F849" httpUri="https://zenodo.org/record/2601655/files/figure.png" pageId="24" pageNumber="459" startId="24.[151,250,1938,1960]" targetBox="[216,1376,193,1926]" targetPageId="24">
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<paragraph id="8BB036EAFFF298558FA2D4EEFFB7F849" blockId="24.[151,1436,1938,2024]" pageId="24" pageNumber="459">
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<emphasis id="B97BEAF8FFF298558FA2D4EEFC78F809" bold="true" box="[151,287,1938,1960]" pageId="24" pageNumber="459">FIGURE 18.</emphasis>
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Details of mature tubarium of
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<taxonomicName id="4C0F4D69FFF298558D62D4EEF9E5F809" authority="Kozlowska-Dawidziuk, 1990" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[599,1154,1938,1960]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="24" pageNumber="459" phylum="Hemichordata" rank="species" species="kozlowskii">
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<emphasis id="B97BEAF8FFF298558D62D4EEFE34F809" box="[599,851,1938,1960]" italics="true" pageId="24" pageNumber="459">Gothograptus kozlowskii</emphasis>
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Kozłowska-Dawidziuk, 1990
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</taxonomicName>
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<emphasis id="B97BEAF8FFF298558BBDD4EEF982F809" bold="true" box="[1160,1253,1938,1960]" pageId="24" pageNumber="459">holotype</emphasis>
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, ZPAL G.XIII/43, Baltic erratic boulder 147 from Jarosławiec, Poland. A. ventral view; B. lateral view, reverse side of tubarium; C. proximal end, outside view; D. orifice of th5
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<superScript id="7C7A9BA2FFF298558EFDD4B3FCA8F87A" attach="right" box="[456,463,1999,2011]" fontSize="5" pageId="24" pageNumber="459">1</superScript>
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; E. opening of appendix.
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</paragraph>
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</caption>
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<caption id="DF706662FFF398548FA2D4F6F927F87C" httpUri="https://zenodo.org/record/2601657/files/figure.png" pageId="25" pageNumber="460" startId="25.[151,250,1930,1952]" targetBox="[176,1412,193,1916]" targetPageId="25">
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<paragraph id="8BB036EAFFF398548FA2D4F6F927F87C" blockId="25.[151,1435,1930,2013]" pageId="25" pageNumber="460">
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<emphasis id="B97BEAF8FFF398548FA2D4F6FC78F801" bold="true" box="[151,287,1930,1952]" pageId="25" pageNumber="460">FIGURE 19.</emphasis>
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Mature tubarium with appendix of
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<taxonomicName id="4C0F4D69FFF398548DBCD4F6F9F8F801" authority="Kozlowska-Dawidziuk, 1990" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[649,1183,1930,1952]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="25" pageNumber="460" phylum="Hemichordata" rank="species" species="obtectus">
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<emphasis id="B97BEAF8FFF398548DBCD4F6FE17F801" box="[649,880,1930,1952]" italics="true" pageId="25" pageNumber="460">Gothograptus obtectus</emphasis>
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Kozłowska-Dawidziuk, 1990
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</taxonomicName>
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from the Bartoszyce IG- 1 core, depth 1677.5 m,
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<emphasis id="B97BEAF8FFF398548EA5D4D5FC96F81E" box="[400,497,1961,1983]" italics="true" pageId="25" pageNumber="460">lundgreni</emphasis>
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Biozone, Poland, ZPAL G.55/48. A. enlargement of distal part of specimen. B. enlargement of proximal end. C. whole specimen, reverse view. Abbreviations: lat. ap.—lateral apertural.
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</paragraph>
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</caption>
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<caption id="DF706662FFF098578FA2D548FF1FF929" httpUri="https://zenodo.org/record/2601659/files/figure.png" pageId="26" pageNumber="461" startId="26.[151,250,1588,1610]" targetBox="[170,1420,193,1578]" targetPageId="26">
|
||
<paragraph id="8BB036EAFFF098578FA2D548FF1FF929" blockId="26.[151,1437,1588,1672]" pageId="26" pageNumber="461">
|
||
<emphasis id="B97BEAF8FFF098578FA2D548FC78F9EB" bold="true" box="[151,287,1588,1610]" pageId="26" pageNumber="461">FIGURE 20.</emphasis>
|
||
Tubaria of
|
||
<taxonomicName id="4C0F4D69FFF098578EA6D548FECDF9EB" authority="Kozlowska-Dawidziuk, 1990" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[403,938,1588,1610]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="26" pageNumber="461" phylum="Hemichordata" rank="species" species="obtectus">
|
||
<emphasis id="B97BEAF8FFF098578EA6D548FF1EF9EB" box="[403,633,1588,1610]" italics="true" pageId="26" pageNumber="461">Gothograptus obtectus</emphasis>
|
||
Kozłowska-Dawidziuk, 1990
|
||
</taxonomicName>
|
||
, Zawada 1 core, depth 1533.3–1540.0 m, Poland, upper part of
|
||
<emphasis id="B97BEAF8FFF098578E08D52FFCF8F9C8" box="[317,415,1619,1641]" italics="true" pageId="26" pageNumber="461">lundgreni</emphasis>
|
||
Biozone. A. tubarium, ZPAL G.XIII/23. B. gerontic tubarium,
|
||
<emphasis id="B97BEAF8FFF098578B41D52FF9B6F9C8" bold="true" box="[1140,1233,1619,1641]" pageId="26" pageNumber="461">holotype</emphasis>
|
||
, ZPAL G.XIII/23. Abbreviations: conn.—connecting, v.—ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3156561FFF098578FF2D5C8F9EBF8FC" pageId="26" pageNumber="461" type="materials_examined">
|
||
<paragraph id="8BB036EAFFF098578FF2D5C8F9EBF8FC" blockId="26.[151,1437,1716,2032]" pageId="26" pageNumber="461">
|
||
<emphasis id="B97BEAF8FFF098578FF2D5C8FC51F96C" bold="true" box="[199,310,1716,1741]" pageId="26" pageNumber="461">Material.</emphasis>
|
||
Material described herein comes from
|
||
<collectingCountry id="F318767AFFF098578DC4D5C8FE58F96C" box="[753,831,1716,1741]" name="Poland" pageId="26" pageNumber="461">Poland</collectingCountry>
|
||
and
|
||
<collectingCountry id="F318767AFFF098578C43D5C8FE82F96C" box="[886,997,1716,1741]" name="Lithuania" pageId="26" pageNumber="461">Lithuania</collectingCountry>
|
||
. Material from
|
||
<collectingCountry id="F318767AFFF098578BA1D5C8F981F96C" box="[1172,1254,1716,1741]" name="Poland" pageId="26" pageNumber="461">Poland</collectingCountry>
|
||
: Bartoszyce IG- 1 core, upper part of
|
||
<emphasis id="B97BEAF8FFF098578EB1D5A4FC97F951" box="[388,496,1752,1776]" italics="true" pageId="26" pageNumber="461">lundgreni</emphasis>
|
||
Biozone, from 1679.3–
|
||
<quantity id="4CF79B0FFFF098578DC8D5A4FE0EF951" box="[765,873,1752,1777]" metricMagnitude="3" metricUnit="m" metricValue="1.6774999999999998" pageId="26" pageNumber="461" unit="m" value="1677.5">1677.5 m</quantity>
|
||
, 20 mature and young specimens, Zawada 1 core, depth 1533.3–1540.0 m, upper part of
|
||
<emphasis id="B97BEAF8FFF098578D7DD580FFD3F8B5" box="[584,692,1788,1812]" italics="true" pageId="26" pageNumber="461">lundgreni</emphasis>
|
||
Biozone, several specimens. Material from
|
||
<collectingCountry id="F318767AFFF098578B91D580F868F8B4" box="[1188,1295,1788,1813]" name="Lithuania" pageId="26" pageNumber="461">Lithuania</collectingCountry>
|
||
comes from the upper part of the
|
||
<emphasis id="B97BEAF8FFF098578E4AD45CFC8CF899" box="[383,491,1824,1848]" italics="true" pageId="26" pageNumber="461">lundgreni</emphasis>
|
||
Biozone, from two cores: Šiupyliai-69, depth 1012.0 m and Likenai-396 core, depth 577.0 m. It consists of ten mature and 20 young specimens and some fragments of tubaria.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156561FFF098568FF2D414F9F5FD6C" lastPageId="27" lastPageNumber="462" pageId="26" pageNumber="461" type="description">
|
||
<paragraph id="8BB036EAFFF098568FF2D414F8F2FEC5" blockId="26.[151,1437,1716,2032]" lastBlockId="27.[151,1437,151,933]" lastPageId="27" lastPageNumber="462" pageId="26" pageNumber="461">
|
||
<emphasis id="B97BEAF8FFF098578FF2D414FC36F820" bold="true" box="[199,337,1896,1921]" pageId="26" pageNumber="461">Description</emphasis>
|
||
. The best preserved mature tubarium from
|
||
<collectingCountry id="F318767AFFF098578C01D414FEE2F820" box="[820,901,1896,1921]" name="Poland" pageId="26" pageNumber="461">Poland</collectingCountry>
|
||
, with seven pairs of thecae, is
|
||
<quantity id="4CF79B0FFFF098578BE3D414F84CF821" box="[1238,1323,1896,1921]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="26" pageNumber="461" unit="mm" value="9.6">9.6 mm</quantity>
|
||
long. The tubarium is narrower up to the third pair of thecae, about
|
||
<quantity id="4CF79B0FFFF098578C10D4F1FE00F805" box="[805,871,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="461" unit="mm" value="1.0">1 mm</quantity>
|
||
wide, and becomes much wider from the level of th4
|
||
<superScript id="7C7A9BA2FFF098578F8ED4CCFDA4F81F" attach="left" box="[187,195,1968,1982]" fontSize="6" pageId="26" pageNumber="461">1</superScript>
|
||
up to th6
|
||
<superScript id="7C7A9BA2FFF098578E19D4CCFC53F81F" attach="left" box="[300,308,1968,1982]" fontSize="6" pageId="26" pageNumber="461">2</superScript>
|
||
and reaches
|
||
<quantity id="4CF79B0FFFF098578EF2D4C8FF47F86A" box="[455,544,1972,1996]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="26" pageNumber="461" unit="mm" value="1.3">1.3 mm</quantity>
|
||
, and then the width decreases close to the appendix. The appendix is about
|
||
<quantity id="4CF79B0FFFF098578A4CD4C8FDA4F84E" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="26" pageNumber="461" unit="mm" value="1.4">1.4 mm</quantity>
|
||
long. In mature specimens, the first pair of thecae does not have any genicular structures, whereas the next thecae have typical reticulated veils covering the thecal openings. Veils grow proximally from the geniculum; they are connected to pleural lists and lateral apertural rods. A thick, distinctive transverse list ends every veil (
|
||
<figureCitation id="13342A6FFFF198568A5AD3C0FDADFF59" captionStart="FIGURE 19" captionStartId="25.[151,250,1930,1952]" captionTargetBox="[176,1412,193,1916]" captionTargetId="figure@25.[176,1412,193,1916]" captionTargetPageId="25" captionText="FIGURE 19. Mature tubarium with appendix of Gothograptus obtectus Kozłowska-Dawidziuk, 1990 from the Bartoszyce IG- 1 core, depth 1677.5 m, lundgreni Biozone, Poland, ZPAL G.55/48. A. enlargement of distal part of specimen. B. enlargement of proximal end. C. whole specimen, reverse view. Abbreviations: lat. ap.—lateral apertural." httpUri="https://zenodo.org/record/2601657/files/figure.png" pageId="27" pageNumber="462">Fig. 19A</figureCitation>
|
||
). The veils are longer in distal thecae, ending just above the genicular list of the previous theca. Some thicker transverse lists may be developed in the middle part of the veil. The lips are clearly visible among the meshwork of reticulum. The genicular lists become very thin in distal thecae and difficult to identify. Mid-ventral lists are very thin in distal thecae; in the last thecae it is difficult to distinguish them from other lists of the tubarium (
|
||
<figureCitation id="13342A6FFFF198568A2AD230F8EFFEC5" box="[1311,1416,332,357]" captionStart="FIGURE 20" captionStartId="26.[151,250,1588,1610]" captionTargetBox="[170,1420,193,1578]" captionTargetId="figure@26.[170,1420,193,1578]" captionTargetPageId="26" captionText="FIGURE 20. Tubaria of Gothograptus obtectus Kozłowska-Dawidziuk, 1990, Zawada 1 core, depth 1533.3–1540.0 m, Poland, upper part of lundgreni Biozone. A. tubarium, ZPAL G.XIII/23. B. gerontic tubarium, holotype, ZPAL G.XIII/23. Abbreviations: conn.—connecting, v.—ventral." httpUri="https://zenodo.org/record/2601659/files/figure.png" pageId="27" pageNumber="462">Fig. 20A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF198568FF2D213FF35FDC1" blockId="27.[151,1437,151,933]" pageId="27" pageNumber="462">
|
||
Specimens of
|
||
<taxonomicName id="4C0F4D69FFF198568E46D20CFF1EFE29" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[371,633,368,392]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="27" pageNumber="462" phylum="Hemichordata" rank="species" species="obtectus">
|
||
<emphasis id="B97BEAF8FFF198568E46D20CFF1EFE29" box="[371,633,368,392]" italics="true" pageId="27" pageNumber="462">Gothograptus obtectus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F318767AFFF198568DFFD213FE51FE29" box="[714,822,367,392]" name="Lithuania" pageId="27" pageNumber="462">Lithuania</collectingCountry>
|
||
are well preserved; three-dimensional, representing different stages of astogenetic growth. There is no complete mature tubarium; they are slightly damaged proximally or distally. The longest specimen, with no proximal end, is
|
||
<quantity id="4CF79B0FFFF198568CF6D2CBF97CFE6E" box="[963,1051,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="27" pageNumber="462" unit="mm" value="6.4">6.4 mm</quantity>
|
||
long and has nine pairs of thecae and a fragment of appendix. The specimen is slightly damaged laterally which makes it thinner (
|
||
<figureCitation id="13342A6FFFF198568BC5D2A0F83CFE55" box="[1264,1371,476,501]" captionStart="FIGURE 21" captionStartId="28.[151,250,1852,1874]" captionTargetBox="[263,1329,193,1852]" captionTargetId="figure@28.[263,1329,193,1834]" captionTargetPageId="28" captionText="FIGURE 21. Mature tubaria of Gothograptus obtectus Kozłowska-Dawidziuk, 1990 from Lithuania, Šiupyliai-69 core, depth 1012.0 m, lundgreni Biozone. A, B, E. finite tubarium with no proximal end, ventro-lateral view, VU RET-19; A, whole specimen; B, enlargement of last thecae; E. enlargement of thecal hood. C–D. proximal part of tubarium, obverse view, VU RET-20; C. enlargement of medial part; D. whole specimen; F–H. fragment of tubarium, lateral-ventral view, VU RET-21; F. whole specimen; G. enlargement of proximal part; H. enlargement of distal part. Abbreviations: genic.—geniculum, lat. ap.— lateral apertural, m–v.—mid-ventral." httpUri="https://zenodo.org/record/2601661/files/figure.png" pageId="27" pageNumber="462">Fig. 21A</figureCitation>
|
||
). The measurements of tubaria from
|
||
<collectingCountry id="F318767AFFF198568ED8D283FF3EFDB9" box="[493,601,511,536]" name="Lithuania" pageId="27" pageNumber="462">Lithuania</collectingCountry>
|
||
are similar to those of the population from
|
||
<collectingCountry id="F318767AFFF198568B08D283F9E9FDB9" box="[1085,1166,511,536]" name="Poland" pageId="27" pageNumber="462">Poland</collectingCountry>
|
||
. They are about
|
||
<quantity id="4CF79B0FFFF198568A72D17CF8FBFDB6" box="[1351,1436,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="27" pageNumber="462" unit="mm" value="0.7">0.7 mm</quantity>
|
||
wide at the level of the first pair of thecae and slightly wider up to the fourth or fifth pairs of thecae and then become about
|
||
<quantity id="4CF79B0FFFF198568E0FD134FCF7FDFE" box="[314,400,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="462" unit="mm" value="1.1">1.1 mm</quantity>
|
||
wide (
|
||
<figureCitation id="13342A6FFFF198568EE8D13BFF22FDC1" box="[477,581,583,608]" captionStart="FIGURE 21" captionStartId="28.[151,250,1852,1874]" captionTargetBox="[263,1329,193,1852]" captionTargetId="figure@28.[263,1329,193,1834]" captionTargetPageId="28" captionText="FIGURE 21. Mature tubaria of Gothograptus obtectus Kozłowska-Dawidziuk, 1990 from Lithuania, Šiupyliai-69 core, depth 1012.0 m, lundgreni Biozone. A, B, E. finite tubarium with no proximal end, ventro-lateral view, VU RET-19; A, whole specimen; B, enlargement of last thecae; E. enlargement of thecal hood. C–D. proximal part of tubarium, obverse view, VU RET-20; C. enlargement of medial part; D. whole specimen; F–H. fragment of tubarium, lateral-ventral view, VU RET-21; F. whole specimen; G. enlargement of proximal part; H. enlargement of distal part. Abbreviations: genic.—geniculum, lat. ap.— lateral apertural, m–v.—mid-ventral." httpUri="https://zenodo.org/record/2601661/files/figure.png" pageId="27" pageNumber="462">Fig. 21D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF198568FF2D111F9F5FD6C" blockId="27.[151,1437,151,933]" pageId="27" pageNumber="462">
|
||
Mature specimens of
|
||
<taxonomicName id="4C0F4D69FFF198568EF6D110FF25FD25" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[451,578,620,644]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="27" pageNumber="462" phylum="Hemichordata" rank="species" species="obtectus">
|
||
<emphasis id="B97BEAF8FFF198568EF6D110FF25FD25" box="[451,578,620,644]" italics="true" pageId="27" pageNumber="462">G. obtectus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F318767AFFF198568DB8D110FF9FFD24" box="[653,760,620,645]" name="Lithuania" pageId="27" pageNumber="462">Lithuania</collectingCountry>
|
||
show some diversity in genicular process development in proximal thecae. In some specimens the second pair of thecae has remnants of the incomplete rim of the genicular process; the next thecae are covered by genicular reticulated veils with typical thicker lists.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156561FFF198568FF2D1ABF8FAFC04" pageId="27" pageNumber="462" type="discussion">
|
||
<paragraph id="8BB036EAFFF198568FF2D1ABFE70FC99" blockId="27.[151,1437,151,933]" pageId="27" pageNumber="462">
|
||
<emphasis id="B97BEAF8FFF198568FF2D1ABFC53FD51" bold="true" box="[199,308,727,752]" pageId="27" pageNumber="462">Remarks</emphasis>
|
||
. Previously,
|
||
<taxonomicName id="4C0F4D69FFF198568EF0D1A4FFA3FD51" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[453,708,728,752]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="27" pageNumber="462" phylum="Hemichordata" rank="species" species="obtectus">
|
||
<emphasis id="B97BEAF8FFF198568EF0D1A4FFA3FD51" box="[453,708,728,752]" italics="true" pageId="27" pageNumber="462">Gothograptus obtectus</emphasis>
|
||
</taxonomicName>
|
||
was known only from the Zawada 1 core of
|
||
<collectingCountry id="F318767AFFF198568B83D1ABF863FD51" box="[1206,1284,727,752]" name="Poland" pageId="27" pageNumber="462">Poland</collectingCountry>
|
||
(Kozłowska-
|
||
<bibRefCitation id="EF9E4B1BFFF198568FA2D180FC30FCB5" author="Kozlowska-Dawidziuk, A." box="[151,343,764,789]" pageId="27" pageNumber="462" pagination="191 - 209" refId="ref11319" refString="Kozlowska-Dawidziuk, A. (1990) The genus Gothograptus (Graptolithina) from the Wenlock of Poland. Acta Palaeontologica Polonica, 35, 191 - 209." type="journal article" year="1990">Dawidziuk 1990</bibRefCitation>
|
||
), where it occurred in the upper part of the
|
||
<emphasis id="B97BEAF8FFF198568C66D180FED8FCB5" box="[851,959,764,788]" italics="true" pageId="27" pageNumber="462">lundgreni</emphasis>
|
||
Biozone. Our new data indicate that it is present in a similar biostratigraphic interval in
|
||
<collectingCountry id="F318767AFFF198568D91D063FE74FC99" box="[676,787,799,824]" name="Lithuania" pageId="27" pageNumber="462">Lithuania</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF198568FF2D038F825FC21" blockId="27.[151,1437,151,933]" pageId="27" pageNumber="462">
|
||
The
|
||
<taxonomicName id="4C0F4D69FFF198568FC8D038FC1BFCFD" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[253,380,836,860]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="27" pageNumber="462" phylum="Hemichordata" rank="species" species="obtectus">
|
||
<emphasis id="B97BEAF8FFF198568FC8D038FC1BFCFD" box="[253,380,836,860]" italics="true" pageId="27" pageNumber="462">G. obtectus</emphasis>
|
||
</taxonomicName>
|
||
tubaria are narrower in the proximal part; from the level of the fourth pair of thecae they becomes much wider. This is caused by a layer outside of the distal thecae, formed by the veils (
|
||
<figureCitation id="13342A6FFFF198568BF8D01BF851FC21" box="[1229,1334,871,896]" captionStart="FIGURE 21" captionStartId="28.[151,250,1852,1874]" captionTargetBox="[263,1329,193,1852]" captionTargetId="figure@28.[263,1329,193,1834]" captionTargetPageId="28" captionText="FIGURE 21. Mature tubaria of Gothograptus obtectus Kozłowska-Dawidziuk, 1990 from Lithuania, Šiupyliai-69 core, depth 1012.0 m, lundgreni Biozone. A, B, E. finite tubarium with no proximal end, ventro-lateral view, VU RET-19; A, whole specimen; B, enlargement of last thecae; E. enlargement of thecal hood. C–D. proximal part of tubarium, obverse view, VU RET-20; C. enlargement of medial part; D. whole specimen; F–H. fragment of tubarium, lateral-ventral view, VU RET-21; F. whole specimen; G. enlargement of proximal part; H. enlargement of distal part. Abbreviations: genic.—geniculum, lat. ap.— lateral apertural, m–v.—mid-ventral." httpUri="https://zenodo.org/record/2601661/files/figure.png" pageId="27" pageNumber="462">Fig. 21D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF198568FF2D0F0F8FAFC04" blockId="27.[151,1437,151,933]" box="[199,1437,908,933]" pageId="27" pageNumber="462">
|
||
The transverse lists of the
|
||
<taxonomicName id="4C0F4D69FFF198568ED9D0F0FF0FFC05" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[492,616,908,932]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="27" pageNumber="462" phylum="Hemichordata" rank="species" species="obtectus">
|
||
<emphasis id="B97BEAF8FFF198568ED9D0F0FF0FFC05" box="[492,616,908,932]" italics="true" pageId="27" pageNumber="462">G. obtectus</emphasis>
|
||
</taxonomicName>
|
||
veil may represent successive edges of the growing veil during astogeny.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |