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<document ID-DOI="10.11646/zootaxa.5239.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7634923" ID-ZooBank="7792E6A2-188A-4F92-81E7-04D6C7D247F7" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1676281994289" checkinUser="plazi" docAuthor="Arnscheid, Wilfried R. &amp; Sobczyk, Thomas" docDate="2023" docId="03A687FB9327DB712E83FE4C4CBBF8CF" docLanguage="en" docName="zootaxa.5239.3.3.pdf" docOrigin="Zootaxa 5239 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oiketicoides Heylaerts 1881" docType="treatment" docVersion="2" lastPageNumber="377" masterDocId="FF9FFF839323DB752E14FF964F25FF94" masterDocTitle="Taxonomic review of the Oiketicoides species (Lepidoptera: Psychidae: Oiketicinae Acanthopsychini) from Anatolia, the Middle East and Central Asia" masterLastPageNumber="394" masterPageNumber="373" pageNumber="377" updateTime="1676282755008" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Taxonomic review of the Oiketicoides species (Lepidoptera: Psychidae: Oiketicinae Acanthopsychini) from Anatolia, the Middle East and Central Asia</mods:title>
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<mods:namePart>Arnscheid, Wilfried R.</mods:namePart>
<mods:affiliation>Im Ostholz 58, 44879 Bochum, Germany.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sobczyk, Thomas</mods:namePart>
<mods:affiliation>Diesterwegstrasse 28, 02977 Hoyerswerda, Germany.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-10</mods:number>
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<mods:number>5239</mods:number>
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<mods:number>3</mods:number>
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<subSubSection box="[151,588,474,501]" pageId="4" pageNumber="377" type="nomenclature">
<paragraph blockId="4.[151,588,474,501]" box="[151,588,474,501]" pageId="4" pageNumber="377">
<heading bold="true" box="[151,588,474,501]" fontSize="11" level="1" pageId="4" pageNumber="377" reason="1">
<emphasis bold="true" box="[151,588,474,501]" pageId="4" pageNumber="377">
Genus
<taxonomicName authority="Heylaerts, 1881" authorityName="Heylaerts" authorityYear="1881" box="[238,588,474,501]" class="Insecta" family="Psychidae" genus="Oiketicoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[238,386,474,501]" italics="true" pageId="4" pageNumber="377">Oiketicoides</emphasis>
<bibRefCitation author="Heylaerts, F. J. M." box="[394,588,474,501]" pageId="4" pageNumber="377" pagination="29 - 73" refId="ref14424" refString="Heylaerts, F. J. M. (1881) Essai d'une monographie des Psychides de la faune europeene. Bulletin &amp; Annales de la Societe Entomologique de Belgique, 25, 29 - 73." type="journal article" year="1881">Heylaerts, 1881</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,1251,543,567]" pageId="4" pageNumber="377" type="reference_group">
<paragraph blockId="4.[151,1437,543,874]" box="[151,1251,543,567]" pageId="4" pageNumber="377">
<taxonomicName authority="Heylaerts, 1881" authorityName="Heylaerts" authorityYear="1881" box="[151,624,543,567]" class="Insecta" family="Psychidae" genus="Acanthopsyche" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="subGenus" subGenus="Oiketicoides">
<emphasis box="[151,455,543,567]" italics="true" pageId="4" pageNumber="377">Acanthopsyche (Oiketicoides)</emphasis>
<bibRefCitation author="Heylaerts, F. J. M." box="[461,624,543,567]" pageId="4" pageNumber="377" pagination="29 - 73" refId="ref14424" refString="Heylaerts, F. J. M. (1881) Essai d'une monographie des Psychides de la faune europeene. Bulletin &amp; Annales de la Societe Entomologique de Belgique, 25, 29 - 73." type="journal article" year="1881">Heylaerts, 1881</bibRefCitation>
</taxonomicName>
; 66.
<emphasis box="[672,1173,543,567]" italics="true" pageId="4" pageNumber="377">
Annales de la Société entomologique de
<collectingCountry box="[1083,1173,543,567]" name="Belgium" pageId="4" pageNumber="377">Belgique</collectingCountry>
</emphasis>
25: 66.
</paragraph>
</subSubSection>
<subSubSection box="[151,1199,574,598]" pageId="4" pageNumber="377" type="materials_examined">
<paragraph blockId="4.[151,1437,543,874]" box="[151,1199,574,598]" pageId="4" pageNumber="377">
<typeStatus box="[151,201,574,598]" pageId="4" pageNumber="377">Type</typeStatus>
species:
<taxonomicName authority="Lederer, 1858: 142" authorityName="Lederer" authorityPageNumber="142" authorityYear="1858" box="[294,668,574,598]" class="Insecta" family="Psychidae" genus="Psyche" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="species" species="inquinata">
<emphasis box="[294,469,574,598]" italics="true" pageId="4" pageNumber="377">Psyche inquinata</emphasis>
<bibRefCitation author="Lederer, J." box="[476,668,574,598]" pageId="4" pageNumber="377" pagination="135 - 143" refId="ref14711" refString="Lederer, J. (1858) Noch einige syrische Schmetterlinge. Wiener Entomologische Monatsschrift, 2 (5), 135 - 143." type="journal article" year="1858">Lederer, 1858: 142</bibRefCitation>
</taxonomicName>
by subsequent designation by
<bibRefCitation author="Hampson, G. F." box="[980,1195,574,598]" pageId="4" pageNumber="377" refId="ref14385" refString="Hampson, G. F. (1893) The Fauna of British India including Ceylon and Burma. Moths. Vol. 1. Taylor and Francis, London, xxiii + 527 pp. [1892]" type="book" year="1893">Hampson, 1893: 293</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[151,909,604,629]" pageId="4" pageNumber="377" type="reference_group">
<paragraph blockId="4.[151,1437,543,874]" box="[151,909,604,629]" pageId="4" pageNumber="377">
<treatmentCitationGroup box="[151,909,604,629]" pageId="4" pageNumber="377">
<treatmentCitation author="Strand, E." box="[151,464,604,629]" page="355" pageId="4" pageNumber="377" year="1912">
<taxonomicName ID-CoL="9253W" authority="Strand, 1912: 355" authorityName="Strand" authorityPageNumber="355" authorityYear="1912" box="[151,464,604,629]" class="Insecta" family="Psychidae" genus="Chalipecten" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis box="[151,273,604,628]" italics="true" pageId="4" pageNumber="377">Chalipecten</emphasis>
<bibRefCitation author="Strand, E." box="[280,464,604,629]" pageId="4" pageNumber="377" refId="ref15244" refString="Strand, E. (1912) Psychidae. In: Seitz, A. (Ed.), Die Grossschmetterlinge der Erde. Vol. 2. Bombyces &amp; Sphinges, Stuttgart, pp. 353 - 370." type="book" year="1912">Strand, 1912: 355</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<emphasis box="[474,831,604,628]" italics="true" pageId="4" pageNumber="377">Die Gross-Schmetterlinge der Erde</emphasis>
2: 355.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="377" type="materials_examined">
<paragraph blockId="4.[151,1437,543,874]" box="[151,899,635,660]" pageId="4" pageNumber="377">
<typeStatus box="[151,201,636,660]" pageId="4" pageNumber="377">Type</typeStatus>
species:
<taxonomicName authority="Heylaerts, 1889" authorityName="Heylaerts" authorityYear="1889" box="[294,652,635,659]" class="Insecta" family="Eurybrachidae" genus="Chalia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="species" species="staudingeri">
<emphasis box="[294,485,635,659]" italics="true" pageId="4" pageNumber="377">Chalia staudingeri</emphasis>
<bibRefCitation author="Heylaerts, F." box="[491,652,635,659]" pageId="4" pageNumber="377" pagination="27 - 28" refId="ref14528" refString="Heylaerts, F. (1889) Monsieur le Dr. Staudinger vient de m'rnvoyer a l'etude quelces Psychides capturees par lui-meme an Algerie. Bulletin &amp; Annales de la Societe Entomologique de Belgique, 33, 27 - 28." type="journal article" year="1889">Heylaerts, 1889</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
<paragraph blockId="4.[151,1437,543,874]" pageId="4" pageNumber="377">
Syn. ‡
<taxonomicName authority="Gerasimov, 1937: 14" authorityName="Gerasimov" authorityPageNumber="14" authorityYear="1937" box="[225,584,666,690]" class="Insecta" family="Empididae" genus="Amictoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis box="[225,338,666,690]" italics="true" pageId="4" pageNumber="377">Amictoides</emphasis>
<bibRefCitation author="Gerasimov, A. M." box="[352,584,666,690]" pageId="4" pageNumber="377" pagination="7 - 17" refId="ref14346" refString="Gerasimov, A. M. (1937) Beitrag zur Systematik der Psychiden auf Grund der Erforschung der Raupen (Lepid.). Zoologischer Anzeiger, 120 (1 / 2), 7 - 17." type="journal article" year="1937">Gerasimov, 1937: 14</bibRefCitation>
</taxonomicName>
;
<emphasis box="[603,1157,666,690]" italics="true" pageId="4" pageNumber="377">
Annales de la Société entomologique de
<collectingCountry box="[1067,1157,666,690]" name="Belgium" pageId="4" pageNumber="377">Belgique</collectingCountry>
</emphasis>
25: 66. Nomenclaturally unavailable.
</paragraph>
<paragraph blockId="4.[151,1437,543,874]" box="[151,755,727,752]" pageId="4" pageNumber="377">
<typeStatus box="[151,201,728,752]" pageId="4" pageNumber="377">Type</typeStatus>
species:
<taxonomicName authority="Boyer de Fonscolombe, 1835" authorityName="Boyer de Fonscolombe" authorityYear="1835" box="[294,755,727,752]" class="Insecta" family="Psychidae" genus="Psyche" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="species" species="febretta">
<emphasis box="[294,450,727,751]" italics="true" pageId="4" pageNumber="377">Psyche febretta</emphasis>
<bibRefCitation author="Boyer de Fonscolombe" box="[457,755,727,752]" pageId="4" pageNumber="377" pagination="107 - 110" refId="ref14055" refString="Boyer de Fonscolombe (1835) Description de la Psyche febretta, nouvelle espece de Bombycites. Annales de la Societe entomologique de France, 4, 107 - 110, pl. 1." type="journal article" year="1835">Boyer de Fonscolombe, 1835</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="4.[151,1437,543,874]" box="[151,1183,758,782]" pageId="4" pageNumber="377">
Syn.
<taxonomicName authorityName="Gerasimov" authorityYear="1937" box="[203,316,758,782]" class="Insecta" family="Empididae" genus="Amictoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis box="[203,316,758,782]" italics="true" pageId="4" pageNumber="377">Amictoides</emphasis>
</taxonomicName>
<bibRefCitation author="Bourgogne, J." box="[322,552,758,782]" pageId="4" pageNumber="377" pagination="98 - 103" refId="ref14003" refString="Bourgogne, J. (1949) Remarques sur le genre Amicta (sensu lato) et determination de la position systematique d' Amicta ecksteini. Bulletin de la Societe entomologique de France, 54, 98 - 103. https: // doi. org / 10.3406 / bsef. 1949.18384" type="journal article" year="1949">
<collectingRegion box="[322,437,758,782]" country="France" name="Bourgogne" pageId="4" pageNumber="377">Bourgogne</collectingRegion>
, 1949: 100
</bibRefCitation>
;
<emphasis box="[562,1044,758,782]" italics="true" pageId="4" pageNumber="377">
Bulletin de la Société entomologique de
<collectingCountry box="[971,1044,759,782]" name="France" pageId="4" pageNumber="377">France</collectingCountry>
</emphasis>
. 54 (7): 100.
</paragraph>
<paragraph blockId="4.[151,1437,543,874]" box="[151,1002,788,813]" pageId="4" pageNumber="377">
<typeStatus box="[151,201,789,813]" pageId="4" pageNumber="377">Type</typeStatus>
species:
<taxonomicName authority="Boyer de Fonscolombe, 1835" authorityName="Boyer de Fonscolombe" authorityYear="1835" box="[294,755,788,813]" class="Insecta" family="Psychidae" genus="Psyche" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="species" species="febretta">
<emphasis box="[294,450,788,812]" italics="true" pageId="4" pageNumber="377">Psyche febretta</emphasis>
<bibRefCitation author="Boyer de Fonscolombe" box="[457,755,788,813]" pageId="4" pageNumber="377" pagination="107 - 110" refId="ref14055" refString="Boyer de Fonscolombe (1835) Description de la Psyche febretta, nouvelle espece de Bombycites. Annales de la Societe entomologique de France, 4, 107 - 110, pl. 1." type="journal article" year="1835">Boyer de Fonscolombe, 1835</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
<paragraph blockId="4.[151,1437,543,874]" pageId="4" pageNumber="377">
Syn.
<taxonomicName authority="Kocak, 1980: 145" authorityName="Kocak" authorityPageNumber="145" authorityYear="1980" box="[201,534,819,843]" class="Insecta" family="Psychidae" genus="Paramictoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis box="[201,349,819,843]" italics="true" pageId="4" pageNumber="377">Paramictoides</emphasis>
<bibRefCitation author="Kocak, A. O." box="[354,534,819,843]" pageId="4" pageNumber="377" pagination="139 - 146" refId="ref14648" refString="Kocak, A. O. (1980) On the nomenclature of some genus- and species- group names of Lepidoptera. Nota lepidopterologica, 2 (4), 139 - 146." type="journal article" year="1980">Koçak, 1980: 145</bibRefCitation>
</taxonomicName>
;
<emphasis box="[542,782,819,843]" italics="true" pageId="4" pageNumber="377">Nota Lepidopterologica</emphasis>
2 (4): 145. A replacement name for a junior homonym
<taxonomicName authority="Gerasimov, 1937" authorityName="Gerasimov" authorityYear="1937" class="Insecta" family="Empididae" genus="Amictoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis box="[1324,1437,819,843]" italics="true" pageId="4" pageNumber="377">Amictoides</emphasis>
<bibRefCitation author="Gerasimov, A. M." box="[199,375,850,874]" pageId="4" pageNumber="377" pagination="7 - 17" refId="ref14346" refString="Gerasimov, A. M. (1937) Beitrag zur Systematik der Psychiden auf Grund der Erforschung der Raupen (Lepid.). Zoologischer Anzeiger, 120 (1 / 2), 7 - 17." type="journal article" year="1937">Gerasimov, 1937</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="377" type="description">
<paragraph blockId="4.[151,1437,920,1883]" pageId="4" pageNumber="377">
<emphasis bold="true" box="[151,325,920,946]" pageId="4" pageNumber="377">Redescription.</emphasis>
Male.
<emphasis box="[409,470,920,946]" italics="true" pageId="4" pageNumber="377">Head</emphasis>
. Eyes roundish, antennae scapus thickened, pedicellus disc-shaped, antennal pecten densely scaled dorsally, ventrally ciliated. The antennae are bipectinate without scales, long ventrally ciliated, decreasing evenly in length towards the apex. Head densely covered with long, hair-like scales.
</paragraph>
<paragraph blockId="4.[151,1437,920,1883]" pageId="4" pageNumber="377">
<emphasis box="[199,280,1028,1054]" italics="true" pageId="4" pageNumber="377">Thorax</emphasis>
. Legs long and hairy. Foreleg with a long and narrow tibial epiphysis. Tibia of the mid- and hindlegs with tiny spurs distally. Forewings elongated, mostly densely covered with short hair-like scales; venation with 10 veins from dc, r3+r4 and m2+m3 mostly stalked. Dc with a simple median vein (rarely divided). On hindwing m1+m2 are mostly short-stalked, median vein not forked (rarely divided), sc and rr fused and mostly connected to the dc with a bar. Cross connections between the veins are absent.
</paragraph>
<paragraph blockId="4.[151,1437,920,1883]" box="[199,1125,1208,1235]" pageId="4" pageNumber="377">
<emphasis box="[199,305,1208,1234]" italics="true" pageId="4" pageNumber="377">Abdomen</emphasis>
. Often protruding the hindwings. Mostly densely covered with long hair.
</paragraph>
<paragraph blockId="4.[151,1437,920,1883]" pageId="4" pageNumber="377">
<emphasis box="[199,364,1244,1270]" italics="true" pageId="4" pageNumber="377">Male genitalia</emphasis>
. The shape of the male genitalia and the abdominal sclerites is very characteristic for the genus with very little difference between the species. The small differences are difficult to evaluate and probably can only be considered by accurate statistical analysis of a larger series of specimens. The long tegumen is evenly rounded dorsally with an even, inwardly folded edge. The vinculum is long and narrow, at the broadest with small, protruding lobes at the sides, the size of which varies with the species. The vinculum continues evenly into a saccus, which is usually thickened again caudally. The valvae are short and broad and do not protrude beyond the tegumen. The clasper is curved inwards and bears about 6 pointed conical spines distally. The pulvillae at the base of the valvae are weakly set off and bear many very fine-pointed sensilla. The phallus is a uniformly thick tube with barely indicated curvature. The distal end is two-lobed with a ventral inversion. The vesica is up to 1/3 of the penis length filled with many very fine granules.
</paragraph>
<paragraph blockId="4.[151,1437,920,1883]" pageId="4" pageNumber="377">Female. Apterous, maggot-shaped and wingless. The antennae are not developed or indicated as tiny bumps. Instead of the eyes mostly small pigment spots are present. The legs are reduced to two segments and bear a pair of claws distally. The females remain in their pupal shell in the case and in this manner, mating takes place. After the mating procedure, the females lay their eggs directly in the case. The females have no diagnostic features for species identification.</paragraph>
<paragraph blockId="4.[151,1437,920,1883]" pageId="4" pageNumber="377">
Early stages. Mostly unknown. The hitherto known larval cases are generally built of silk and they are covered longitudinally with fragments of twigs and leaves in a very similar way. The caterpillars are polyphagous and feed on different herbs like
<taxonomicName box="[406,516,1857,1883]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="377" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[406,463,1857,1882]" italics="true" pageId="4" pageNumber="377">Vicia</emphasis>
spp.
</taxonomicName>
(de Freina, in litt.,
<figureCitation box="[732,798,1856,1882]" captionStart="FIGURE 2" captionStartId="5.[151,250,1962,1986]" captionTargetBox="[168,1419,1094,1939]" captionTargetId="figure-56@5.[168,1419,1094,1939]" captionTargetPageId="5" captionText="FIGURE 2 Caterpillar of Oiketicoides armena (Heylaerts, 1885), feeding at Viccia sp., Turkey, Tokat-Sivas, 1700 m, Çamlýbel- Pass, 110.vii.1978 (Photo: J. de Freina)." figureDoi="http://doi.org/10.5281/zenodo.7634929" httpUri="https://zenodo.org/record/7634929/files/figure.png" pageId="4" pageNumber="377">Fig. 2</figureCitation>
) and grass.
</paragraph>
</subSubSection>
</treatment>
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