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<document ID-DOI="http://doi.org/10.5281/zenodo.6167536" ID-GBIF-Dataset="1fd9017d-6a56-470b-aa83-68944cd8ac46" ID-GBIF-Taxon="130509477" ID-Zenodo-Dep="6167536" checkinTime="1492608005255" checkinUser="jeremy" docAuthor="Dankittipakul, Pakawin, Tavano, Maria &amp; Singtripop, Tippawan" docDate="2012" docId="03A6879FA860FF87E5C8FC48022E6740" docLanguage="en" docName="DankittipakulTavano_Singtripop2012FINAL.imf" docOrigin="Journal of Natural History 46" docSource="https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae" docStyle="DocumentStyle{}" docTitle="Teutamus leptothecus Dankittipakul, Tavano &amp; Singtripop, 2012, sp. nov." docType="treatment" docVersion="18" lastPageId="36" lastPageNumber="1725" masterDocId="FF9FFFE7A842FFA3E422FFF90169654C" masterDocTitle="Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae)" masterLastPageNumber="1730" masterPageNumber="1689" pageId="34" pageNumber="1723" updateTime="1645288162421" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dankittipakul, Pakawin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tavano, Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Singtripop, Tippawan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>46</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1689</mods:start>
<mods:end>1730</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.110.1080/00222933.2012.681314</mods:identifier>
<mods:identifier type="GBIF-Dataset">1fd9017d-6a56-470b-aa83-68944cd8ac46</mods:identifier>
<mods:identifier type="Zenodo-Dep">574642</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6167536" ID-GBIF-Taxon="130509477" ID-Zenodo-Dep="6167536" LSID="urn:lsid:plazi:treatment:03A6879FA860FF87E5C8FC48022E6740" httpUri="http://treatment.plazi.org/id/03A6879FA860FF87E5C8FC48022E6740" lastPageId="36" lastPageNumber="1725" pageId="34" pageNumber="1723">
<subSubSection box="[490,818,944,968]" pageId="34" pageNumber="1723" type="nomenclature">
<paragraph blockId="34.[490,818,944,1001]" box="[490,818,944,968]" pageId="34" pageNumber="1723">
<heading box="[490,818,944,968]" centered="true" fontSize="10" level="3" pageId="34" pageNumber="1723" reason="4">
<taxonomicName box="[490,726,944,968]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="1723" phylum="Arthropoda" rank="species" species="leptothecus" status="sp. nov.">
<emphasis bold="true" box="[490,726,944,968]" italics="true" pageId="34" pageNumber="1723">Teutamus leptothecus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[733,818,945,968]" pageId="34" pageNumber="1723" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[539,768,976,1001]" pageId="34" pageNumber="1723" type="description">
<paragraph blockId="34.[490,818,944,1001]" box="[539,768,976,1001]" pageId="34" pageNumber="1723">
<heading box="[539,768,976,1001]" centered="true" fontSize="10" level="3" pageId="34" pageNumber="1723" reason="4">
(
<figureCitation box="[546,653,976,1001]" captionStart="Figure 3" captionStartId="5.[105,172,1238,1260]" captionTargetBox="[105,1065,130,1205]" captionTargetId="figure@5.[105,1065,130,1205]" captionTargetPageId="5" captionText="Figure 3. Teutamus serrulatus, female paratype (A, B, G). Teutamus globularis, female paratype (C). Teutamus apiculatus, female paratype (D, F). Teutamus orthogonus, female paratype (E). Teutamus leptothecus, female holotype (H). (A C) Epigastric region, ventral; (D) anterior portion of spermatheca showing glandular pore and mucus plug; (E) secretory glands; (F) mucus plug; (G) secretory ampulla surrounded by epithelial gland cells (arrowed); (H) gland ductules on spermathecal wall. Abbreviations: MP, mucus plug; PS, post-epigastric scutum." httpUri="https://zenodo.org/record/574645/files/figure.png" pageId="34" pageNumber="1723">Figures 3</figureCitation>
H,
<figureCitation box="[688,716,976,1000]" captionStart="Figure 16" captionText="Figure 16. Teutamus leptothecus sp. nov., female holotype (A, B). Teutamus deelemanae sp. nov., female holotype (C, D). (A, C) Epigastric region, ventral; (B, D) internal genitalia, dorsal. Scale bars 0. 2 5 mm." pageId="34" pageNumber="1723">16</figureCitation>
A,B)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="1723" type="diagnosis">
<paragraph blockId="34.[173,1134,1054,1148]" box="[173,281,1054,1077]" pageId="34" pageNumber="1723">
<emphasis box="[173,281,1054,1077]" italics="true" pageId="34" pageNumber="1723">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="34.[173,1134,1054,1148]" pageId="34" pageNumber="1723">
Females of
<taxonomicName box="[301,452,1093,1117]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="1723" phylum="Arthropoda" rank="species" species="leptothecus" status="sp. nov.">
<emphasis box="[301,452,1093,1117]" italics="true" pageId="34" pageNumber="1723">T. leptothecus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[457,541,1094,1117]" pageId="34" pageNumber="1723" rank="species">sp. nov.</taxonomicNameLabel>
can be recognized by the anteriorly narrowed, elliptic spermathecae and the simple curved ducts (
<figureCitation box="[661,772,1124,1148]" captionStart="Figure 16" captionStartId="35.[105,172,1251,1273]" captionTargetBox="[108,1065,133,1210]" captionTargetId="figure@35.[105,1065,129,1218]" captionTargetPageId="35" captionText="Figure 16. Teutamus leptothecus sp. nov., female holotype (A, B). Teutamus deelemanae sp. nov., female holotype (C, D). (A, C) Epigastric region, ventral; (B, D) internal genitalia, dorsal. Scale bars 0. 2 5 mm." httpUri="https://zenodo.org/record/574658/files/figure.png" pageId="34" pageNumber="1723">Figure 16</figureCitation>
A,B).
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="1723" type="materials_examined">
<paragraph blockId="34.[173,1134,1244,1372]" box="[173,326,1244,1268]" pageId="34" pageNumber="1723">
<emphasis box="[173,326,1244,1268]" italics="true" pageId="34" pageNumber="1723">Type material</emphasis>
</paragraph>
<paragraph blockId="34.[173,1134,1244,1372]" pageId="34" pageNumber="1723">
<materialsCitation ID-GBIF-Occurrence="1502458270" collectingDate="2001-06-24" collectingDateMax="2001-06-26" collectingDateMin="2001-06-24" collectionCode="MHNG" collection_SpecimenCode="SIM-01/07" collectorName="P. J. Schwendinger" country="Malaysia" elevation="170" location="Kota Tinggi Waterfalls, at the foot of Mt Muntahak" pageId="34" pageNumber="1723" specimenCount="1" specimenCount-female="1" stateProvince="Johor State" typeStatus="holotype">
<emphasis box="[173,283,1284,1308]" italics="true" pageId="34" pageNumber="1723">
<typeStatus box="[173,278,1284,1308]" pageId="34" pageNumber="1723">Holotype</typeStatus>
.
</emphasis>
<specimenCount box="[290,304,1284,1308]" pageId="34" pageNumber="1723" type="female"></specimenCount>
,
<collectingCountry box="[327,434,1283,1307]" name="Malaysia" pageId="34" pageNumber="1723" value="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion box="[453,593,1283,1308]" pageId="34" pageNumber="1723">Johor State</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:03A6879FA860FF87E5C8FC48022E6740:8ED06052A860FF81E647FAFD00416077" country="Malaysia" name="Kota Tinggi Waterfalls, at the foot of Mt Muntahak" pageId="34" pageNumber="1723" stateProvince="Johor State">Kota Tinggi Waterfalls, at the foot of Mt Muntahak</location>
,
<elevation box="[310,384,1316,1340]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="34" pageNumber="1723" unit="m" value="170.0">
<quantity box="[310,384,1316,1340]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="34" pageNumber="1723" unit="m" value="170.0">170 m</quantity>
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:03A6879FA860FF87E5C8FC48022E6740:8ED06052A860FF81E5AFFADA03F06077" box="[397,665,1315,1339]" country="Malaysia" name="rainforest along stream" pageId="34" pageNumber="1723" stateProvince="Johor State">rainforest along stream</location>
,
<date box="[678,870,1315,1340]" pageId="34" pageNumber="1723" value="2001-06-24" valueMax="2001-06-26" valueMin="2001-06-24">
<collectingDate box="[678,870,1315,1340]" pageId="34" pageNumber="1723" value="2001-06-24" valueMax="2001-06-26" valueMin="2001-06-24">2426 June 2001</collectingDate>
</date>
, leg.
<collectorName box="[932,1134,1315,1339]" pageId="34" pageNumber="1723">P.J. Schwendinger</collectorName>
(
<collectionCode box="[182,416,1348,1372]" pageId="34" pageNumber="1723">
MHNG, SIM-01
<emphasis box="[374,386,1348,1372]" italics="true" pageId="34" pageNumber="1723">/</emphasis>
07
</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="34.[173,789,1437,1462]" box="[173,789,1437,1462]" pageId="34" pageNumber="1723">
<materialsCitation ID-GBIF-Occurrence="1502458274" box="[173,784,1437,1462]" collectingDate="2001-06-24" collectingDateMax="2001-06-26" collectingDateMin="2001-06-24" collectionCode="MHNG" collection_SpecimenCode="SIM-01/07" collectorName="P. J. Schwendinger" country="Malaysia" elevation="170" location="Kota Tinggi Waterfalls, at the foot of Mt Muntahak" pageId="34" pageNumber="1723" specimenCount="1" specimenCount-female="1" stateProvince="Johor State" typeStatus="paratype">
<emphasis box="[173,279,1438,1461]" italics="true" pageId="34" pageNumber="1723">
<typeStatus box="[173,275,1438,1461]" pageId="34" pageNumber="1723">Paratype</typeStatus>
.
</emphasis>
Data as holotype,
<specimenCount box="[492,519,1437,1462]" pageId="34" pageNumber="1723" type="female">1♀</specimenCount>
, (
<collectionCode box="[542,776,1437,1462]" pageId="34" pageNumber="1723">
MHNG, SIM-01
<emphasis box="[733,745,1437,1461]" italics="true" pageId="34" pageNumber="1723">/</emphasis>
07
</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="1723" type="etymology">
<paragraph blockId="34.[173,1134,1556,1652]" box="[173,293,1556,1580]" pageId="34" pageNumber="1723">
<emphasis box="[173,293,1556,1580]" italics="true" pageId="34" pageNumber="1723">Etymology</emphasis>
</paragraph>
<paragraph blockId="34.[173,1134,1556,1652]" pageId="34" pageNumber="1723">
The specific epithet is a nominative form of an adjective (
<emphasis box="[819,883,1596,1620]" italics="true" pageId="34" pageNumber="1723">leptos</emphasis>
= thin;
<emphasis box="[978,1047,1596,1620]" italics="true" pageId="34" pageNumber="1723">thecus</emphasis>
= sac), and refers to the shape of spermathecae.
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/574658/files/figure.png" pageId="35" pageNumber="1724" targetBox="[108,1065,133,1210]" targetPageId="35">
<paragraph blockId="35.[105,1066,1251,1332]" pageId="35" pageNumber="1724">
Figure 16.
<taxonomicName box="[220,432,1251,1273]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="1724" phylum="Arthropoda" rank="species" species="leptothecus" status="sp. nov.">
<emphasis box="[220,432,1251,1273]" italics="true" pageId="35" pageNumber="1724">Teutamus leptothecus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[437,511,1252,1273]" pageId="35" pageNumber="1724" rank="species">sp. nov.</taxonomicNameLabel>
, female holotype (A, B).
<taxonomicName box="[764,980,1251,1273]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="1724" phylum="Arthropoda" rank="species" species="deelemanae" status="sp. nov.">
<emphasis box="[764,980,1251,1273]" italics="true" pageId="35" pageNumber="1724">Teutamus deelemanae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[985,1059,1252,1273]" pageId="35" pageNumber="1724" rank="species">sp. nov.</taxonomicNameLabel>
, female holotype (C, D). (A, C) Epigastric region, ventral; (B, D) internal genitalia, dorsal. Scale bars 0.2 5 mm.
</paragraph>
</caption>
<subSubSection lastPageId="36" lastPageNumber="1725" pageId="35" pageNumber="1724" type="description">
<paragraph blockId="35.[105,1066,1430,1653]" box="[105,232,1430,1453]" pageId="35" pageNumber="1724">
<emphasis box="[105,232,1430,1453]" italics="true" pageId="35" pageNumber="1724">Description</emphasis>
</paragraph>
<paragraph blockId="35.[105,1066,1430,1653]" lastBlockId="36.[173,1134,129,313]" lastPageId="36" lastPageNumber="1725" pageId="35" pageNumber="1724">
<emphasis box="[105,184,1469,1493]" italics="true" pageId="35" pageNumber="1724">Female</emphasis>
(holotype). Total length 5.86; prosoma 2.62 long, 1.62 wide; opisthosoma 3.24 long, 1.62 wide. Leg formula: 1423; I 11.78 (2.58, 1.00, 3.96, 2.62, 1.62); II 8.94 (2.06, 0.98, 3.04, 1.68, 1.18); III 7.84 (1.72, 0.92, 2.22, 1.82, 1.16); IV 11.30 (2.64, 1.00, 3.22, 2.80, 1.64). Spination: Leg I: Femora p1-1-1; tibiae p9 r9; metatarsi p6 r6. Leg II: Tibiae p8 r8; metatarsi p5 r5. Carapace dark chestnut-brown; submarginal lobes blunt. Sternum brown. Legs hairless, bi-coloured: Tibiae, metatarsi and tarsi yellowish brown, other segments brown. Opisthosoma elongate-ovoid. Dorsum pale, posteriorly with a series of faint chevrons.
</paragraph>
<paragraph blockId="36.[173,1134,129,313]" pageId="36" pageNumber="1725">
Genitalia (
<figureCitation box="[344,464,193,217]" captionStart="Figure 16" captionStartId="35.[105,172,1251,1273]" captionTargetBox="[108,1065,133,1210]" captionTargetId="figure@35.[105,1065,129,1218]" captionTargetPageId="35" captionText="Figure 16. Teutamus leptothecus sp. nov., female holotype (A, B). Teutamus deelemanae sp. nov., female holotype (C, D). (A, C) Epigastric region, ventral; (B, D) internal genitalia, dorsal. Scale bars 0. 2 5 mm." httpUri="https://zenodo.org/record/574658/files/figure.png" pageId="36" pageNumber="1725">Figure 16</figureCitation>
A,B): Epigyne with circular copulatory orifices situated anteriorly. Insemination ducts curving posteriorly, connected to apical portion of spermathecae. Spermathecae elliptic, distinctly narrowed anteriorly, surface provided with numerous pore ductules. Fertilization ducts lanceolated.
</paragraph>
<paragraph blockId="36.[173,367,360,384]" box="[173,367,360,384]" pageId="36" pageNumber="1725">
<emphasis box="[173,239,360,384]" italics="true" pageId="36" pageNumber="1725">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection box="[173,839,460,525]" pageId="36" pageNumber="1725" type="distribution">
<paragraph blockId="36.[173,839,460,525]" box="[173,306,460,484]" pageId="36" pageNumber="1725">
<emphasis box="[173,306,460,484]" italics="true" pageId="36" pageNumber="1725">Distribution</emphasis>
</paragraph>
<paragraph blockId="36.[173,839,460,525]" box="[173,839,500,525]" pageId="36" pageNumber="1725">
Known only from the type locality in Malaysia (
<figureCitation box="[715,826,500,524]" captionStart="Figure 18" captionText="Figure 18. Distribution of new Teutamus species treated in the present study. 1, Teutamus rama sp. nov.; 2, Teutamus serrulatus sp. nov.; 3, Teutamus brachiatus sp. nov.; 4, Teutamus apiculatus sp. nov.; 5, Teutamus lioneli sp. nov.; 6, Teutamus calceolatus sp. nov.; 7, Teutamus orthogonus sp. nov.; 8, Teutamus globularis sp. nov.; 9, Teutamus seculatus sp. nov.; 10, Teutamus hirtellus sp. nov.; 11, Teutamus tortuosus sp. nov.; 12, Teutamus rollardae sp. nov.; 13, Teutamus poggii sp. nov.; 14, Teutamus leptothecus sp. nov.; 15, Teutamus deelemanae sp. nov.; 16, Teutamus spiralis sp. nov.; 17, Teutamus sumatranus sp. nov.; , Teutamus politus Thorell. " pageId="36" pageNumber="1725">Figure 18</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>