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<document id="7F5E1ECDF4686FC83FE3A42B80C47409" ID-CLB-Dataset="272041" ID-DOI="10.1093/zoolinnean/zlad013" ID-GBIF-Dataset="effa6181-8326-4c7d-8c91-809f7fabb862" ID-ISSN="0024-4082" ID-Zenodo-Dep="8432389" ID-ZooBank="C3E2DFF9-A0A6-41EA-A149-0F73A2BEE5E6" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1696939870042" checkinUser="plazi" docAuthor="Silva, Felipe De Vasconcelos, Pola, Marta &amp; Cervera, Juan Lucas" docDate="2023" docId="03A6878CFF8A650DFD41FF60ED98FD37" docLanguage="en" docName="zlad013.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad013" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Marioniopsis Odhner 1934" docType="treatment" docVersion="2" lastPageNumber="471" masterDocId="FF9FFFF4FF906517FC52FF84EE67FF8B" masterDocTitle="A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)" masterLastPageNumber="476" masterPageNumber="445" pageNumber="471" updateTime="1697044790341" updateUser="ExternalLinkService">
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<mods:title id="DC9E29C7F5F38DD93E96653834DC83BA">A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)</mods:title>
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<mods:namePart id="74458CBC1C5AF1C53F3123C7736B42A6">Silva, Felipe De Vasconcelos</mods:namePart>
<mods:affiliation id="571B78C8EE3476FDA52460484088BD3C">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:namePart id="92A8F0FA2856ACA54663569561D8C056">Pola, Marta</mods:namePart>
<mods:affiliation id="34DE9D8AA4E9E366542941D0E8D9CE6E">Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid; Campus de Excelencia Internacional UAM + CSIC, C / Darwin, 2, 28049 Madrid, Spain</mods:affiliation>
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<mods:namePart id="678547775D56B34CC2C2D10C8F3F1815">Cervera, Juan Lucas</mods:namePart>
<mods:affiliation id="8BB55B2CEEA3CCFBB8B450913CC5A127">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain &amp; Instituto Universitario de Investigación Marina (INMAR), Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Universidad de Cádiz, Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:title id="9AEAC91E9431DC3ABC2322AAF5B2F16F">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="2F645EE186EC2F898826370EB606FC19">2023</mods:date>
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<mods:title id="9F4D53098E699239DF8661878D5AEDE7">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="20602CCA2556BF25021D805B0240393C">2023-07-11</mods:number>
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<mods:number id="2A0D48F2872C65AB9732F8B871D876A5">199</mods:number>
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<treatment id="03A6878CFF8A650DFD41FF60ED98FD37" ID-DOI="http://doi.org/10.5281/zenodo.8432431" ID-Zenodo-Dep="8432431" LSID="urn:lsid:plazi:treatment:03A6878CFF8A650DFD41FF60ED98FD37" httpUri="http://treatment.plazi.org/id/03A6878CFF8A650DFD41FF60ED98FD37" lastPageNumber="471" pageId="26" pageNumber="471">
<subSubSection id="C3156511FF8A650DFD41FF60EF92FEA9" pageId="26" pageNumber="471" type="nomenclature">
<paragraph id="8BB0369AFF8A650DFD41FF60EF92FEA9" blockId="26.[275,631,228,291]" pageId="26" pageNumber="471">
<emphasis id="B97BEA88FF8A650DFD41FF60EF92FEA9" bold="true" pageId="26" pageNumber="471">
Genus
<taxonomicName id="4C0F4D19FF8A650DFD0FFF60EC10FF75" ID-CoL="BFFS9" authority="Odhner 1934" authorityName="Odhner" authorityYear="1934" box="[349,631,228,254]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFD0FFF60EF86FF75" bold="true" box="[349,481,228,254]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
<bibRefCitation id="EF9E4B6BFF8A650DFDB4FF60EC10FF75" author="Odhner NH" box="[486,631,228,254]" pageId="26" pageNumber="471" pagination="229 - 310" refId="ref26550" refString="Odhner NH. The Nudibranchiata of British Antarctic Expedition. British Antartic (' Terra Nova') Expedition, 1910. Natural History Reports. Zoology 1934; 7: 229 - 310." type="journal article" year="1934">Odhner 1934</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13342A1FFF8A650DFDCDFE8DEF8CFEA8" box="[415,491,265,291]" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="26" pageNumber="471">Fig. 3K</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF8A650DFCD3FEBFEFDDFEFA" pageId="26" pageNumber="471" type="type_taxon">
<paragraph id="8BB0369AFF8A650DFCD3FEBFEFDDFEFA" blockId="26.[129,777,314,369]" pageId="26" pageNumber="471">
<emphasis id="B97BEA88FF8A650DFCD3FEBFEE9DFED9" box="[129,250,315,338]" italics="true" pageId="26" pageNumber="471">
<typeStatus id="54B48838FF8A650DFCD3FEBFEED6FED9" box="[129,177,315,338]" pageId="26" pageNumber="471">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C0F4D19FF8A650DFCACFEBFEEA6FEFA" authority="(Ruppell and Leuckart 1828)" baseAuthorityName="Ruppell and Leuckart" baseAuthorityYear="1828" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="cyanobranchiata">
<emphasis id="B97BEA88FF8A650DFCACFEBFEC46FED9" box="[254,545,314,338]" italics="true" pageId="26" pageNumber="471">Marioniopsis cyanobranchiata</emphasis>
(Rüppell and Leuckart 1828)
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF8A650DFCD3FE17EE8CFD82" pageId="26" pageNumber="471" type="diagnosis">
<paragraph id="8BB0369AFF8A650DFCD3FE17EE8CFD82" blockId="26.[129,778,403,521]" pageId="26" pageNumber="471">
<emphasis id="B97BEA88FF8A650DFCD3FE17EE80FE21" box="[129,231,403,426]" italics="true" pageId="26" pageNumber="471">Diagnosis:</emphasis>
Rachidian tooth tricuspid, denticulate. Masticatory border denticulate, denticles blunt or rod-shaped. Digestive system arrangement cladohepatic or holohepatic, not U- or G-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF8A650DFCD3FDAEEFBCFB44" pageId="26" pageNumber="471" type="description">
<paragraph id="8BB0369AFF8A650DFCD3FDAEEF4CFC11" blockId="26.[129,778,554,922]" pageId="26" pageNumber="471">
<emphasis id="B97BEA88FF8A650DFCD3FDAEEE98FDC9" box="[129,255,554,578]" italics="true" pageId="26" pageNumber="471">Morphology:</emphasis>
Body slender or broad, length up to
<quantity id="4CF79B7FFF8A650DFEDAFDAEECB5FDCA" box="[648,722,554,578]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="26" pageNumber="471" unit="mm" value="88.0">88 mm</quantity>
(
<figureCitation id="13342A1FFF8A650DFEB1FDAEEEC7FDEA" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="26" pageNumber="471">Fig. 3K</figureCitation>
). Oral veil bilobed. Three to seven simple or branched velar processes, with the outermost ones oħen longer. Fully arborescent gills with at least seven pairs. Rachidian tooth tricuspid and denticulate (with accessory denticles or folds). Masticatory border of the jaws denticulate, with large-sized (up to 300 µm) denticles with conical bases and blunt cusps or rod-shaped denticles.
<collectionCode id="ED1EAE5FFF8A650DFCEEFC81EE90FC96" box="[188,247,773,797]" pageId="26" pageNumber="471">ODG</collectionCode>
complex clado- or holohepatic arrangement, with the stomach located in the central or right region of the posterior
<collectionCode id="ED1EAE5FFF8A650DFCD3FCC0EEDBFCD7" box="[129,188,836,860]" pageId="26" pageNumber="471">ODG</collectionCode>
complex (cladohepatic) or posterior
<collectionCode id="ED1EAE5FFF8A650DFE6CFCC0EC1EFCD7" box="[574,633,836,860]" pageId="26" pageNumber="471">ODG</collectionCode>
complex portion/lobe (holohepatic arrangement). Penis flagelliform, cylindrical or conical.
</paragraph>
<paragraph id="8BB0369AFF8A650DFCD3FC38EFBCFB44" blockId="26.[128,778,956,1231]" pageId="26" pageNumber="471">
<emphasis id="B97BEA88FF8A650DFCD3FC38EF29FC5F" box="[129,334,956,980]" italics="true" pageId="26" pageNumber="471">Species composition:</emphasis>
<taxonomicName id="4C0F4D19FF8A650DFD0FFC39ED61FC5F" authority="Eliot 1904" authorityName="Eliot" authorityYear="1904" box="[349,774,956,980]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="albotuberculata">
<emphasis id="B97BEA88FF8A650DFD0FFC39ECE6FC5F" box="[349,641,956,980]" italics="true" pageId="26" pageNumber="471">Marioniopsis albotuberculata</emphasis>
<bibRefCitation id="EF9E4B6BFF8A650DFEC2FC38ED61FC5F" author="Eliot C" box="[656,774,956,980]" pageId="26" pageNumber="471" pagination="268 - 298" refId="ref24969" refString="Eliot C. On some nudibranchs from east Africa and Zanzibar, Part VI. Proceedings of the Zoological Society of London 1904; 74: 268 - 298." type="journal article" year="1904">Eliot 1904</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFCD3FC58EFEDFC78" authorityName="Bergh" authorityYear="1890" box="[129,394,987,1011]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="arborescens">
<emphasis id="B97BEA88FF8A650DFCD3FC58EFEDFC78" box="[129,394,987,1011]" italics="true" pageId="26" pageNumber="471">Marioniopsis arborescens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFDFEFC5FED6DFC78" box="[428,778,987,1011]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="bathycarolinensis">
<emphasis id="B97BEA88FF8A650DFDFEFC5FED6DFC78" bold="true" box="[428,778,987,1011]" italics="true" pageId="26" pageNumber="471">Marioniopsis bathycarolinensis</emphasis>
</taxonomicName>
(
<collectionCode id="ED1EAE5FFF8A650DFCDAFC7EEEFCFB99" box="[136,155,1018,1042]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="471" type="Museum">V</collectionCode>
.G.
<bibRefCitation id="EF9E4B6BFF8A650DFCE6FC7EEFBBFB99" author="Smith VG &amp; Gosliner TM" box="[180,476,1018,1042]" pageId="26" pageNumber="471" pagination="66" refId="ref27653" refString="Smith VG, Gosliner TM. A new species of Marionia (Gastropoda: Nudibranchia) from the Caroline Islands. Proceedings of the California Academy of Sciences 2005; 56: 66." type="journal article" year="2005">Smith and Gosliner 2005</bibRefCitation>
)
<emphasis id="B97BEA88FF8A650DFDA6FC7FEC12FB98" bold="true" box="[500,629,1019,1043]" pageId="26" pageNumber="471">comb. nov.</emphasis>
,
<taxonomicName id="4C0F4D19FF8A650DFED8FC7FEF45FBB9" baseAuthorityName="Ruppell and Leuckart" baseAuthorityYear="1828" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="cyanobranchiata">
<emphasis id="B97BEA88FF8A650DFED8FC7FEF45FBB9" italics="true" pageId="26" pageNumber="471">Marioniopsis cyanobranchiata</emphasis>
</taxonomicName>
,
<emphasis id="B97BEA88FF8A650DFD19FB9EEEC8FBDA" bold="true" pageId="26" pageNumber="471">
<taxonomicName id="4C0F4D19FF8A650DFD19FB9EECC5FBBA" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Silva &amp; Pola &amp; Cervera" baseAuthorityYear="2023" box="[331,674,1049,1074]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="elongoreticulata" status="comb. nov.">
<emphasis id="B97BEA88FF8A650DFD19FB9EECC5FBBA" bold="true" box="[331,674,1049,1074]" italics="true" pageId="26" pageNumber="471">Marioniopsis elongoreticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857F3FF8A650DFE94FB9EEEC8FBDA" pageId="26" pageNumber="471" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B97BEA88FF8A650DFC90FBBDEFE4FBFA" bold="true" pageId="26" pageNumber="471">
<taxonomicName id="4C0F4D19FF8A650DFC90FBBDEF87FBDA" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Silva &amp; Pola &amp; Cervera" baseAuthorityYear="2023" box="[194,480,1081,1105]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="elongoviridis" status="comb. nov.">
<emphasis id="B97BEA88FF8A650DFC90FBBDEF87FBDA" bold="true" box="[194,480,1081,1105]" italics="true" pageId="26" pageNumber="471">Marioniopsis elongoviridis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857F3FF8A650DFDBFFBBDEC0AFBDA" box="[493,621,1081,1105]" pageId="26" pageNumber="471" rank="species">comb. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C0F4D19FF8A650DFE2DFBBDEE9DFBFB" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Pease" baseAuthorityYear="1860" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="hawaiiensis" status="comb. nov.">
<emphasis id="B97BEA88FF8A650DFE2DFBBDEE9DFBFB" bold="true" italics="true" pageId="26" pageNumber="471">Marioniopsis hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857F3FF8A650DFD54FBDDEFE4FBFA" box="[262,387,1113,1137]" pageId="26" pageNumber="471" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0F4D19FF8A650DFDC6FBDDEC2EFBFB" baseAuthorityName="Odhner" baseAuthorityYear="1936" box="[404,585,1112,1136]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="levis">
<emphasis id="B97BEA88FF8A650DFDC6FBDDEC2EFBFB" box="[404,585,1112,1136]" italics="true" pageId="26" pageNumber="471">Marioniopsis levis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFE09FBDDEF5EFB1B" authority="Eliot 1904" authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="pellucida">
<emphasis id="B97BEA88FF8A650DFE09FBDDEEDCFB1B" italics="true" pageId="26" pageNumber="471">Marioniopsis pellucida</emphasis>
<bibRefCitation id="EF9E4B6BFF8A650DFC94FBFCEF5EFB1B" author="Eliot C" box="[198,313,1144,1168]" pageId="26" pageNumber="471" pagination="268 - 298" refId="ref24969" refString="Eliot C. On some nudibranchs from east Africa and Zanzibar, Part VI. Proceedings of the Zoological Society of London 1904; 74: 268 - 298." type="journal article" year="1904">Eliot 1904</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFD15FBFCECDCFB1B" authority="(Eliot 1904)" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Eliot" baseAuthorityYear="1904" box="[327,699,1144,1168]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="ramosa" status="comb. nov.">
<emphasis id="B97BEA88FF8A650DFD15FBFCEC4DFB04" bold="true" box="[327,554,1144,1168]" italics="true" pageId="26" pageNumber="471">Marioniopsis ramosa</emphasis>
(
<bibRefCitation id="EF9E4B6BFF8A650DFE12FBFCECD7FB1B" author="Eliot C" box="[576,688,1144,1168]" pageId="26" pageNumber="471" pagination="268 - 298" refId="ref24969" refString="Eliot C. On some nudibranchs from east Africa and Zanzibar, Part VI. Proceedings of the Zoological Society of London 1904; 74: 268 - 298." type="journal article" year="1904">Eliot 1904</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97BEA88FF8A650DFE94FBFCEEC8FB24" bold="true" pageId="26" pageNumber="471">
<taxonomicNameLabel id="A24857F3FF8A650DFE94FBFCEEC8FB24" pageId="26" pageNumber="471" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0F4D19FF8A650DFCEDFB13EC36FB24" authority="(Ev. Marcus 1983)" authorityName="Silva &amp; Pola &amp; Cervera" authorityYear="2023" baseAuthorityName="Ev. Marcus" baseAuthorityYear="1983" box="[191,593,1175,1199]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="tedi" status="comb. nov.">
<emphasis id="B97BEA88FF8A650DFCEDFB13EF1BFB24" bold="true" box="[191,380,1175,1199]" italics="true" pageId="26" pageNumber="471">Marioniopsis tedi</emphasis>
(Ev.
<bibRefCitation id="EF9E4B6BFF8A650DFDE9FB1CEC21FB24" author="Marcus Ev" box="[443,582,1175,1199]" pageId="26" pageNumber="471" pagination="177 - 214" refId="ref25994" refString="Marcus Ev. The Western Atlantic Tritoniidae. Boletim de Zoologia, Universidade de Sao Paulo 1983; 6: 177 - 214." type="journal article" year="1983">Marcus 1983</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97BEA88FF8A650DFE0EFB13ECBFFB24" bold="true" box="[604,728,1175,1199]" pageId="26" pageNumber="471">
<taxonomicNameLabel id="A24857F3FF8A650DFE0EFB13ECBFFB24" box="[604,728,1175,1199]" pageId="26" pageNumber="471" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0F4D19FF8A650DFCD3FB33EFBCFB44" authority="Eliot 1904" authorityName="Eliot" authorityYear="1904" box="[129,475,1206,1231]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="viridescens">
<emphasis id="B97BEA88FF8A650DFCD3FB33EF0BFB45" box="[129,364,1206,1230]" italics="true" pageId="26" pageNumber="471">Marioniopsis viridescens</emphasis>
<bibRefCitation id="EF9E4B6BFF8A650DFD22FB32EFBCFB44" author="Eliot C" box="[368,475,1206,1231]" pageId="26" pageNumber="471" pagination="268 - 298" refId="ref24969" refString="Eliot C. On some nudibranchs from east Africa and Zanzibar, Part VI. Proceedings of the Zoological Society of London 1904; 74: 268 - 298." type="journal article" year="1904">Eliot 1904</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3156511FF8A650DFCD3FB74ED98FD37" pageId="26" pageNumber="471" type="discussion">
<paragraph id="8BB0369AFF8A650DFCD3FB74ECD9FA68" blockId="26.[129,778,1264,1977]" pageId="26" pageNumber="471">
<emphasis id="B97BEA88FF8A650DFCD3FB74EEBBFA83" box="[129,220,1264,1288]" italics="true" pageId="26" pageNumber="471">Remarks:</emphasis>
The genus
<taxonomicName id="4C0F4D19FF8A650DFD1AFB75EFA4FA83" authorityName="Odhner" authorityYear="1934" box="[328,451,1265,1288]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFD1AFB75EFA4FA83" box="[328,451,1265,1288]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
, originally created to assign plate-bearing tritoniids with holohepatic digestive systems and a single row of denticulate masticatory border of the jaw (
<bibRefCitation id="EF9E4B6BFF8A650DFE22FAAAEC9EFACD" author="Odhner NH" box="[624,761,1326,1350]" pageId="26" pageNumber="471" refId="ref26584" refString="Odhner NH. Nudibranchia dendronotacea: a revision of the system. Brussels: Musee Royal d'Histoire Naturelle de Belgique, 1936." type="book" year="1936">Odhner 1936</bibRefCitation>
), was synonymized with
<taxonomicName id="4C0F4D19FF8A650DFD3FFACBEFA0FAED" authorityName="Vayssiere" authorityYear="1877" box="[365,455,1359,1382]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFD3FFACBEFA0FAED" box="[365,455,1359,1382]" italics="true" pageId="26" pageNumber="471">Marionia</emphasis>
</taxonomicName>
due to taxonomic uncertainties regarding the validity of its morphological characters (
<bibRefCitation id="EF9E4B6BFF8A650DFEF3FAE9EF74FA2F" author="Smith VG &amp; Gosliner TM" pageId="26" pageNumber="471" pagination="66" refId="ref27653" refString="Smith VG, Gosliner TM. A new species of Marionia (Gastropoda: Nudibranchia) from the Caroline Islands. Proceedings of the California Academy of Sciences 2005; 56: 66." type="journal article" year="2005">Smith and Gosliner 2005</bibRefCitation>
). Based on our phylogenetic analyses, the genus
<taxonomicName id="4C0F4D19FF8A650DFCD3FA28EE9AFA48" authorityName="Odhner" authorityYear="1934" box="[129,253,1452,1475]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFCD3FA28EE9AFA48" box="[129,253,1452,1475]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
is reinstated here to assign the plate-bearing species recovered clustered as the sister clade of
<taxonomicName id="4C0F4D19FF8A650DFE5CFA48EC0FFA68" authorityName="Vayssiere" authorityYear="1877" box="[526,616,1484,1507]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFE5CFA48EC0FFA68" box="[526,616,1484,1507]" italics="true" pageId="26" pageNumber="471">Marionia</emphasis>
</taxonomicName>
(
<figureCitation id="13342A1FFF8A650DFE2AFA4FECC9FA68" box="[632,686,1483,1507]" captionStart="Figure 2" captionStartId="10.[129,194,1719,1743]" captionTargetBox="[325,1278,148,1689]" captionTargetId="figure-6@10.[321,1281,144,1692]" captionTargetPageId="10" captionText="Figure 2. Bayesian phylogenetic tree based on the concatenated molecular data (COI + 16S + H3) of Tritoniidae (sensu Odhner 1963). Outgroup taxa are depicted in grey.Tritoniidae genera are independently coloured, and branches with several replicates for each species hypothesis were collapsed for clarity.Names in bold are species which sequences were generated for this study with the number of sequenced specimens showed in brackets. PP are shown right/top and BS leħ/boưom.Unsupported branches are not labelled. Green circles indicate maximum support by both analyses. Solid vertical bars indicate specimens grouped as species entities based on COI dataset. Empty vertical bars indicate the specimens grouped as species based on the 16S dataset." figureDoi="http://doi.org/10.5281/zenodo.8432395" httpUri="https://zenodo.org/record/8432395/files/figure.png" pageId="26" pageNumber="471">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BB0369AFF8A650DFCCEFA6EECC6F8D0" blockId="26.[129,778,1264,1977]" pageId="26" pageNumber="471">
The synapomorphies of
<taxonomicName id="4C0F4D19FF8A650DFDFCFA6FEC4AF989" authorityName="Odhner" authorityYear="1934" box="[430,557,1515,1538]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFDFCFA6FEC4AF989" box="[430,557,1515,1538]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
within
<taxonomicName id="4C0F4D19FF8A650DFEDEFA6EED6DF989" box="[652,778,1514,1538]" pageId="26" pageNumber="467" rank="subFamily" subFamily="Marioniinae">Marioniinae</taxonomicName>
are the denticulate rachidian tooth and blunt and rounded or rod-shaped denticles in the masticatory border. In addition,
<taxonomicName id="4C0F4D19FF8A650DFCD3F9CDEF67F9EB" authorityName="Odhner" authorityYear="1934" box="[129,256,1609,1632]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFCD3F9CDEF67F9EB" box="[129,256,1609,1632]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
has both holohepatic and cladohepatic arrangements, which lack a U- or G-shape, unlike
<taxonomicName id="4C0F4D19FF8A650DFE0CF9ECECDDF9F4" authorityName="Vayssiere" authorityYear="1877" box="[606,698,1640,1663]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFE0CF9ECECDDF9F4" box="[606,698,1640,1663]" italics="true" pageId="26" pageNumber="471">Marionia</emphasis>
</taxonomicName>
, which has only a cladohepatic arrangement. However, the distinction between cladohepatic and holohepatic arrangements can be tricky. For example, the fusion of the right and leħ lobes of the
<collectionCode id="ED1EAE5FFF8A650DFCD3F961EEDBF976" box="[129,188,1765,1789]" pageId="26" pageNumber="471">ODG</collectionCode>
complex in
<taxonomicName id="4C0F4D19FF8A650DFD6AF962EC4DF976" authorityName="Bergh" authorityYear="1890" box="[312,554,1765,1789]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="arborescens">
<emphasis id="B97BEA88FF8A650DFD6AF962EC4DF976" box="[312,554,1765,1789]" italics="true" pageId="26" pageNumber="471">Marioniopsis arborescens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D19FF8A650DFE0EF962ED6EF976" baseAuthorityName="Odhner" baseAuthorityYear="1936" box="[604,777,1765,1789]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="levis">
<emphasis id="B97BEA88FF8A650DFE0EF962ED6EF976" box="[604,777,1765,1789]" italics="true" pageId="26" pageNumber="471">Marioniopsis levis</emphasis>
</taxonomicName>
was initially overlooked in their original descriptions (Rüppell and Leuckart 1828,
<bibRefCitation id="EF9E4B6BFF8A650DFD0BF8A0EFAFF8B7" author="Eliot C" box="[345,456,1828,1852]" pageId="26" pageNumber="471" pagination="268 - 298" refId="ref24969" refString="Eliot C. On some nudibranchs from east Africa and Zanzibar, Part VI. Proceedings of the Zoological Society of London 1904; 74: 268 - 298." type="journal article" year="1904">Eliot 1904</bibRefCitation>
) and only noticed later when
<taxonomicName id="4C0F4D19FF8A650DFCD3F8C0EF67F8D0" authorityName="Odhner" authorityYear="1934" box="[129,256,1860,1883]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFCD3F8C0EF67F8D0" box="[129,256,1860,1883]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
was originally proposed (
<bibRefCitation id="EF9E4B6BFF8A650DFE54F8C7ECF7F8D0" author="Odhner NH" box="[518,656,1859,1883]" pageId="26" pageNumber="471" refId="ref26584" refString="Odhner NH. Nudibranchia dendronotacea: a revision of the system. Brussels: Musee Royal d'Histoire Naturelle de Belgique, 1936." type="book" year="1936">Odhner 1936</bibRefCitation>
).
</paragraph>
<paragraph id="8BB0369AFF8A650DFCCEF8E6ED98FD37" blockId="26.[129,778,1264,1977]" lastBlockId="26.[825,1476,144,700]" pageId="26" pageNumber="471">
Species currently assigned to
<taxonomicName id="4C0F4D19FF8A650DFD92F8E7EC7AF8F1" authorityName="Vayssiere" authorityYear="1877" box="[448,541,1891,1914]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFD92F8E7EC7AF8F1" box="[448,541,1891,1914]" italics="true" pageId="26" pageNumber="471">Marionia</emphasis>
</taxonomicName>
for which no molecular data are available, namely
<taxonomicName id="4C0F4D19FF8A650DFDCEF806ECFBF811" authorityName="Eliot" authorityYear="1904" box="[412,668,1922,1946]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="albotuberculata">
<emphasis id="B97BEA88FF8A650DFDCEF806ECFBF811" box="[412,668,1922,1946]" italics="true" pageId="26" pageNumber="471">Marionia albotuberculata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFEFEF806EF4EF832" authorityName="V. G. Smith &amp; Gosliner" authorityYear="2005" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="bathycarolinensis">
<emphasis id="B97BEA88FF8A650DFEFEF806EF4EF832" italics="true" pageId="26" pageNumber="471">Marionia bathycarolinensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFD64F826EC58F832" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[310,575,1953,1977]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="cyanobranchiata">
<emphasis id="B97BEA88FF8A650DFD64F826EC58F832" box="[310,575,1953,1977]" italics="true" pageId="26" pageNumber="471">Marionia cyanobranchiata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFE1EF826ED63F832" authorityName="Eliot" authorityYear="1904" box="[588,772,1953,1977]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="pelucida">
<emphasis id="B97BEA88FF8A650DFE1EF826ED63F832" box="[588,772,1953,1977]" italics="true" pageId="26" pageNumber="471">Marionia pelucida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFF6BFF14EDA4FF23" authorityName="Ev. Marcus" authorityYear="1983" box="[825,963,144,168]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="tedi">
<emphasis id="B97BEA88FF8A650DFF6BFF14EDA4FF23" box="[825,963,144,168]" italics="true" pageId="26" pageNumber="471">Marionia tedi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D19FF8A650DFF83FF14EA19FF2C" authorityName="Eliot" authorityYear="1904" box="[977,1150,144,167]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="ramosa">
<emphasis id="B97BEA88FF8A650DFF83FF14EA19FF2C" box="[977,1150,144,167]" italics="true" pageId="26" pageNumber="471">Marionia ramosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D19FF8A650DF8E7FF14EBE4FF23" authorityName="Eliot" authorityYear="1904" box="[1205,1411,144,168]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="viridescens">
<emphasis id="B97BEA88FF8A650DF8E7FF14EBE4FF23" box="[1205,1411,144,168]" italics="true" pageId="26" pageNumber="471">Marionia viridescens</emphasis>
</taxonomicName>
, were transferred to
<taxonomicName id="4C0F4D19FF8A650DFF81FF34EA34FF4C" authorityName="Odhner" authorityYear="1934" box="[979,1107,176,199]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFF81FF34EA34FF4C" box="[979,1107,176,199]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
in this study based on their denticulate rachidian tooth, denticulate masticatory border of the jaw, presence of true stomach plates and cladohepatic (in
<taxonomicName id="4C0F4D19FF8A650DFF6BFE8AEDA4FEAE" authorityName="Ev. Marcus" authorityYear="1983" box="[825,963,269,293]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="tedi">
<emphasis id="B97BEA88FF8A650DFF6BFE8AEDA4FEAE" box="[825,963,269,293]" italics="true" pageId="26" pageNumber="471">Marionia tedi</emphasis>
</taxonomicName>
) or holohepatic
<collectionCode id="ED1EAE5FFF8A650DF82AFE89EAD3FEAE" box="[1144,1204,269,293]" pageId="26" pageNumber="471">ODG</collectionCode>
complex (
<bibRefCitation id="EF9E4B6BFF8A650DF975FE89EBF2FEAE" author="Eliot C" box="[1319,1429,269,293]" pageId="26" pageNumber="471" pagination="268 - 298" refId="ref24969" refString="Eliot C. On some nudibranchs from east Africa and Zanzibar, Part VI. Proceedings of the Zoological Society of London 1904; 74: 268 - 298." type="journal article" year="1904">Eliot 1904</bibRefCitation>
, Ev.
<bibRefCitation id="EF9E4B6BFF8A650DFF6BFEA9EDA0FECF" author="Marcus Ev" box="[825,967,300,324]" pageId="26" pageNumber="471" pagination="177 - 214" refId="ref25994" refString="Marcus Ev. The Western Atlantic Tritoniidae. Boletim de Zoologia, Universidade de Sao Paulo 1983; 6: 177 - 214." type="journal article" year="1983">Marcus 1983</bibRefCitation>
,
<bibRefCitation id="EF9E4B6BFF8A650DFF8BFEA8EA96FECF" author="Smith VG &amp; Gosliner TM" box="[985,1265,300,324]" pageId="26" pageNumber="471" pagination="66" refId="ref27653" refString="Smith VG, Gosliner TM. A new species of Marionia (Gastropoda: Nudibranchia) from the Caroline Islands. Proceedings of the California Academy of Sciences 2005; 56: 66." type="journal article" year="2005">Smith and Gosliner 2005</bibRefCitation>
). Not surprisingly, most of these species have been included in
<taxonomicName id="4C0F4D19FF8A650DF952FEC8EBE7FEE8" authorityName="Odhner" authorityYear="1934" box="[1280,1408,332,355]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DF952FEC8EBE7FEE8" box="[1280,1408,332,355]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
in the past due to their holohepatic
<collectionCode id="ED1EAE5FFF8A650DF83AFEEFEAC3FE08" box="[1128,1188,363,387]" pageId="26" pageNumber="471">ODG</collectionCode>
complex and, therefore, are not considered
<emphasis id="B97BEA88FF8A650DFF8CFE0EEAE4FE29" box="[990,1155,394,418]" italics="true" pageId="26" pageNumber="471">combinatio nova</emphasis>
in this study. We have not included species previously assigned to
<taxonomicName id="4C0F4D19FF8A650DF893FE2EEB26FE4A" authorityName="Odhner" authorityYear="1934" box="[1217,1345,426,449]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DF893FE2EEB26FE4A" box="[1217,1345,426,449]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
that did not fit our proposed diagnosis and lack of molecular data, such as the divergent
<taxonomicName id="4C0F4D19FF8A650DFF98FE6DEB5BFD8B" authority="(Willan 1988)" baseAuthorityName="Willan" baseAuthorityYear="1988" box="[970,1340,488,512]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="species" species="platyctenea">
<emphasis id="B97BEA88FF8A650DFF98FE6DEAFAFD8B" box="[970,1181,488,512]" italics="true" pageId="26" pageNumber="471">Marionia platyctenea</emphasis>
(
<bibRefCitation id="EF9E4B6BFF8A650DF8FDFE6CEB48FD8B" author="Willan RC" box="[1199,1327,488,512]" pageId="26" pageNumber="471" pagination="39 - 63" refId="ref27976" refString="Willan RC. The taxonomy of two host-specific, cryptic dendronotoid nudibranch species (Mollusca: Gastropoda) from Australia including a new species description. Zoological Journal of the Linnean Society 1988; 94: 39 - 63." type="journal article" year="1988">Willan 1988</bibRefCitation>
)
</taxonomicName>
. In addition,
<taxonomicName id="4C0F4D19FF8A650DFF6BFD8CEDDEFD94" authorityName="Odhner" authorityYear="1934" box="[825,953,520,543]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B97BEA88FF8A650DFF6BFD8CEDDEFD94" box="[825,953,520,543]" italics="true" pageId="26" pageNumber="471">Marioniopsis</emphasis>
</taxonomicName>
has a high minimum
<emphasis id="B97BEA88FF8A650DF8C9FD8CEAA0FD94" box="[1179,1223,520,543]" italics="true" pageId="26" pageNumber="471">COI</emphasis>
interspecific
<emphasis id="B97BEA88FF8A650DF900FD8CEB38FD94" box="[1362,1375,520,543]" italics="true" pageId="26" pageNumber="471">p</emphasis>
-distance, peaking at 22.5% (Table 3), suggesting the presence of other generic lineages within the genus. Once the various unnamed species are formally described, their internal and external morphology may reveal new characters for further delimitation at the generic level.
</paragraph>
</subSubSection>
</treatment>
</document>
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