157 lines
27 KiB
XML
157 lines
27 KiB
XML
<document id="CDC728CA5CEDE10651CC8CA9595AB5EF" ID-CLB-Dataset="272041" ID-DOI="10.1093/zoolinnean/zlad013" ID-GBIF-Dataset="effa6181-8326-4c7d-8c91-809f7fabb862" ID-ISSN="0024-4082" ID-Zenodo-Dep="8432389" ID-ZooBank="C3E2DFF9-A0A6-41EA-A149-0F73A2BEE5E6" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1696939870042" checkinUser="plazi" docAuthor="Silva, Felipe De Vasconcelos, Pola, Marta & Cervera, Juan Lucas" docDate="2023" docId="03A6878CFF846503FC81FDA4EC08F9BF" docLanguage="en" docName="zlad013.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad013" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Tritonidoxa Bergh 1907" docType="treatment" docVersion="4" lastPageNumber="465" masterDocId="FF9FFFF4FF906517FC52FF84EE67FF8B" masterDocTitle="A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)" masterLastPageNumber="476" masterPageNumber="445" pageNumber="465" updateTime="1697044790341" updateUser="ExternalLinkService">
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<mods:title id="D3B8324BF946D301A2EC7D95D3782F88">A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)</mods:title>
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<mods:namePart id="749E9D50D5025E6140B3F4B5FF0289EB">Silva, Felipe De Vasconcelos</mods:namePart>
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<mods:affiliation id="F68548BF9FD3A1C13D7B8480DC2E0251">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:nameIdentifier id="EBD0E849F1EE0161085FF2780ABDDB89" type="email">felipe.devasconcelos@uca.es</mods:nameIdentifier>
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<mods:namePart id="8E080FABB3E956EDEF67A289F85D45F5">Pola, Marta</mods:namePart>
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<mods:affiliation id="2647001C706EC37D035E31C4B2145351">Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid; Campus de Excelencia Internacional UAM + CSIC, C / Darwin, 2, 28049 Madrid, Spain</mods:affiliation>
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<mods:namePart id="93759443634F603623CA0689DF211041">Cervera, Juan Lucas</mods:namePart>
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<mods:affiliation id="F28C4AFB14ABCF130BF6052B1C42E032">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain & Instituto Universitario de Investigación Marina (INMAR), Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Universidad de Cádiz, Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:date id="2E77D586D8387019C018D96AD7EA5255">2023</mods:date>
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<treatment id="03A6878CFF846503FC81FDA4EC08F9BF" ID-DOI="http://doi.org/10.5281/zenodo.8426089" ID-Zenodo-Dep="8426089" LSID="urn:lsid:plazi:treatment:03A6878CFF846503FC81FDA4EC08F9BF" httpUri="http://treatment.plazi.org/id/03A6878CFF846503FC81FDA4EC08F9BF" lastPageNumber="465" pageId="20" pageNumber="465">
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<subSubSection id="C3156511FF846503FC81FDA4ECD0FDB1" box="[211,695,544,571]" pageId="20" pageNumber="465" type="nomenclature">
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<paragraph id="8BB0369AFF846503FC81FDA4ECD0FDB1" blockId="20.[211,695,544,571]" box="[211,695,544,571]" pageId="20" pageNumber="465">
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<heading id="D0F881F6FF846503FC81FDA4ECD0FDB1" bold="true" box="[211,695,544,571]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="465" reason="2">
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<emphasis id="B97BEA88FF846503FC81FDA4ECD0FDB1" bold="true" box="[211,695,544,571]" pageId="20" pageNumber="465">
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<taxonomicName id="4C0F4D19FF846503FC81FDA4EFABFDB1" ID-CoL="9G3YC" authority="Bergh 1907" authorityName="Bergh" authorityYear="1907" box="[211,460,544,571]" class="Gastropoda" family="Tritoniidae" genus="Tritonidoxa" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="465" phylum="Mollusca" rank="genus">
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<emphasis id="B97BEA88FF846503FC81FDA4EF2CFDB0" bold="true" box="[211,331,544,571]" italics="true" pageId="20" pageNumber="465">Tritonidoxa</emphasis>
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<bibRefCitation id="EF9E4B6BFF846503FD1DFDA4EFABFDB1" author="Bergh R" box="[335,460,544,570]" pageId="20" pageNumber="465" pagination="1 - 144" refId="ref24498" refString="Bergh R. The Opisthobranchiata of South Africa. Transactions of the South Aflican Philosophical Society 1907; 17: 1 - 144." type="journal article" year="1907">Bergh 1907</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="13342A1FFF846503FD89FDA4EC4BFDB1" box="[475,556,544,570]" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="20" pageNumber="465">Figs 3D</figureCitation>
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,
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<figureCitation id="13342A1FFF846503FE67FDA4EC30FDB1" box="[565,599,544,570]" captionStart="Figure 4" captionStartId="13.[113,178,1528,1552]" captionTargetBox="[309,1263,147,1498]" captionTargetId="figure-134@13.[306,1266,144,1501]" captionTargetPageId="13" captionText="Figure 4. Scanning electron micrographs of radulae of analysed key species of Tritoniinae and Marioniinae subfam. nov.: A, tricuspid rachidian teeth and differentiated first lateral of Tritonia hombergii (ZMBN 83471); B, tricuspid rachidian teeth and differentiated first lateral with single accessory denticle (circled) of Candiella plebeia (MNCN15.05/93393); C, unicuspid rachidian teeth and undifferentiated first lateral of Tritoniopsis elegans (MNHNC/MB28-004506); D, tricuspid rachidian teeth and differentiated first lateral of Tritonidoxa capensis (MNCN15.05/93399); E, unicuspid rachidian teeth and undifferentiated first lateral of Tochuina gigantea (NHMD-115967); F, tricuspid rachidian teeth and differentiated first lateral with several accessory denticles (circled) of Tritonicula hamnerorum (MNCN15.05/93378); G, tricuspid rachidian teeth and differentiated first lateral of Marionia blainvillea (NHMD115968); Scale bars: A, D, E = 60 μm; B, C = 22 μm; F = 20 μm; G = 110 μm." figureDoi="http://doi.org/10.5281/zenodo.8432401" httpUri="https://zenodo.org/record/8432401/files/figure.png" pageId="20" pageNumber="465">4D</figureCitation>
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,
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<figureCitation id="13342A1FFF846503FE32FDA4ECE6FDB1" box="[608,641,544,570]" captionStart="Figure 5" captionStartId="14.[129,194,1262,1286]" captionTargetBox="[164,1437,147,1230]" captionTargetId="figure-262@14.[161,1441,144,1234]" captionTargetPageId="14" captionText="Figure 5. Digestive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view: A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (MNCN15.05/93393); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis(MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations:a, anus; aodg, anterior ODG complex; e, esophagus; i, intestine; odg, ovotestis-digestive gland complex; s, stomach; podg, posterior ODG complex." figureDoi="http://doi.org/10.5281/zenodo.8432405" httpUri="https://zenodo.org/record/8432405/files/figure.png" pageId="20" pageNumber="465">5D</figureCitation>
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,
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<figureCitation id="13342A1FFF846503FED9FDA4ECCBFDB1" box="[651,684,544,570]" captionStart="Figure 6" captionStartId="15.[113,178,1771,1795]" captionTargetBox="[317,1256,144,1743]" captionTargetId="figure-8@15.[316,1256,144,1743]" captionTargetPageId="15" captionText="Figure 6. Reproductive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view:A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (ZMBN 83471); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis (MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations: ag, albumen gland; am, ampulla; bc, bursa copulatrix; fo, female opening; fgm, female gland mass; hd, hermaphrodite duct; o, oviduct; p, penis; pp, prostatic portion; ps, penial sheath; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.8432407" httpUri="https://zenodo.org/record/8432407/files/figure.png" pageId="20" pageNumber="465">6D</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156511FF846503FCD3FDE5EC08F9BF" pageId="20" pageNumber="465" type="materials_examined">
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<paragraph id="8BB0369AFF846503FCD3FDE5EC4AFD13" blockId="20.[129,777,609,664]" pageId="20" pageNumber="465">
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<emphasis id="B97BEA88FF846503FCD3FDE5EE9AFDF3" box="[129,253,609,632]" italics="true" pageId="20" pageNumber="465">
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<typeStatus id="54B48838FF846503FCD3FDE5EED6FDF3" box="[129,177,609,632]" pageId="20" pageNumber="465">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C0F4D19FF846503FD56FDE5EC29FDF2" authority="Bergh 1907" authorityName="Bergh" authorityYear="1907" box="[260,590,609,633]" class="Gastropoda" family="Tritoniidae" genus="Tritonidoxa" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="465" phylum="Mollusca" rank="species" species="capensis">
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<emphasis id="B97BEA88FF846503FD56FDE5EFAAFDF3" box="[260,461,609,633]" italics="true" pageId="20" pageNumber="465">Tritonidoxa capensis</emphasis>
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<bibRefCitation id="EF9E4B6BFF846503FD86FDE5EC29FDF2" author="Bergh R" box="[468,590,609,633]" pageId="20" pageNumber="465" pagination="1 - 144" refId="ref24498" refString="Bergh R. The Opisthobranchiata of South Africa. Transactions of the South Aflican Philosophical Society 1907; 17: 1 - 144." type="journal article" year="1907">Bergh 1907</bibRefCitation>
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</taxonomicName>
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, by original designation. To date, this is a monotypic genus.
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</paragraph>
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<paragraph id="8BB0369AFF846503FCD3FD3EECD0FD7A" blockId="20.[129,777,697,753]" pageId="20" pageNumber="465">
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<emphasis id="B97BEA88FF846503FCD3FD3EEE80FD5A" box="[129,231,698,721]" italics="true" pageId="20" pageNumber="465">Diagnosis:</emphasis>
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Penis long, spiralled with pointed end. Secondary gills abscent; notal margin distinct, broad and wrinkled.
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</paragraph>
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<paragraph id="8BB0369AFF846503FCD3FC96EF93FB4A" blockId="20.[129,778,786,1217]" pageId="20" pageNumber="465">
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<emphasis id="B97BEA88FF846503FCD3FC96EE98FCA1" box="[129,255,786,810]" italics="true" pageId="20" pageNumber="465">Morphology:</emphasis>
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Body slightly broad, length up to
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<quantity id="4CF79B7FFF846503FE3AFC96ECD2FCA2" box="[616,693,786,810]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="20" pageNumber="465" unit="mm" value="32.0">32 mm</quantity>
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. Notum smooth, with transverse white patches from head to tail. Oral veil narrow and entire. Eight simple velar processes. Secondary gills absent. Notal margin distinct, broad and wrinkled (
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<figureCitation id="13342A1FFF846503FEFDFCF4EC9EFC03" box="[687,761,880,904]" captionStart="Figure 3" captionStartId="12.[129,194,1533,1557]" captionTargetBox="[321,1281,144,1505]" captionTargetId="figure-116@12.[321,1281,144,1505]" captionTargetPageId="12" captionText="Figure 3. Representative species of genera of Tritoniidae (with exception of Paratritonia). A, Tritonia hombergii (MNCN15.05/93379) (Portaferry, Ireland); B, Candiella plebeia (MNCN15.05/93393) (Portaferry, Ireland); C, Tritoniopsis elegans (MB28-0045061) (Zavora, Mozambique); D, Tritonidoxa capensis (MNCN15.05/93399) (Scoưburg, South Africa); E, Tritoniella belli (Antarctica); F, Marianina rosea (French Polynesia); G, Myrella challengeriana comb.nov.(Bahía Camarones, Argentina); H, Tochuina gigantea (Agnew Pass, USA); I, Tritonicula khaleesi (MZUSP113165) (Fortaleza, Brazil); J, Marionia cucullata (Golfo Nuevo, Argentina); (K) Marioniopsis levis ‘1’ (MNHNC/ MB28-004895) (Nacala, Mozambique). Photo A and B, courtesy of B. Picton; photo C and K courtesy of Y.Tibiriça; photo D courtesy of V. Fraser; photo E courtesy of M. Caballer; photo F courtesy of G. Rouse; photo G and J, courtesy of N. Baưini; photo H courtesy of S. Thiebaud; photo I by F. Silva." figureDoi="http://doi.org/10.5281/zenodo.8432399" httpUri="https://zenodo.org/record/8432399/files/figure.png" pageId="20" pageNumber="465">Fig. 3D</figureCitation>
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). Ratio jaw/body length: 0.20–0.35. Rachidian tooth tricuspid and smooth with a rectangular base (
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<figureCitation id="13342A1FFF846503FDABFC2BEC23FC4C" box="[505,580,943,967]" captionStart="Figure 4" captionStartId="13.[113,178,1528,1552]" captionTargetBox="[309,1263,147,1498]" captionTargetId="figure-134@13.[306,1266,144,1501]" captionTargetPageId="13" captionText="Figure 4. Scanning electron micrographs of radulae of analysed key species of Tritoniinae and Marioniinae subfam. nov.: A, tricuspid rachidian teeth and differentiated first lateral of Tritonia hombergii (ZMBN 83471); B, tricuspid rachidian teeth and differentiated first lateral with single accessory denticle (circled) of Candiella plebeia (MNCN15.05/93393); C, unicuspid rachidian teeth and undifferentiated first lateral of Tritoniopsis elegans (MNHNC/MB28-004506); D, tricuspid rachidian teeth and differentiated first lateral of Tritonidoxa capensis (MNCN15.05/93399); E, unicuspid rachidian teeth and undifferentiated first lateral of Tochuina gigantea (NHMD-115967); F, tricuspid rachidian teeth and differentiated first lateral with several accessory denticles (circled) of Tritonicula hamnerorum (MNCN15.05/93378); G, tricuspid rachidian teeth and differentiated first lateral of Marionia blainvillea (NHMD115968); Scale bars: A, D, E = 60 μm; B, C = 22 μm; F = 20 μm; G = 110 μm." figureDoi="http://doi.org/10.5281/zenodo.8432401" httpUri="https://zenodo.org/record/8432401/files/figure.png" pageId="20" pageNumber="465">Fig. 4D</figureCitation>
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). First lateral tooth differentiated, broad, smooth and sharp. Masticatory borders of jaws denticulate, with medium-sized (up to 40 µm) denticles with conical bases and sharp cusps. Cuticular folds absent.
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<collectionCode id="ED1EAE5FFF846503FE9CFB89ED6EFBAE" box="[718,777,1037,1061]" pageId="20" pageNumber="465">ODG</collectionCode>
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complex holohepatic (
|
||
<figureCitation id="13342A1FFF846503FD3AFBA8EFD4FBCF" box="[360,435,1068,1092]" captionStart="Figure 5" captionStartId="14.[129,194,1262,1286]" captionTargetBox="[164,1437,147,1230]" captionTargetId="figure-262@14.[161,1441,144,1234]" captionTargetPageId="14" captionText="Figure 5. Digestive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view: A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (MNCN15.05/93393); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis(MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations:a, anus; aodg, anterior ODG complex; e, esophagus; i, intestine; odg, ovotestis-digestive gland complex; s, stomach; podg, posterior ODG complex." figureDoi="http://doi.org/10.5281/zenodo.8432405" httpUri="https://zenodo.org/record/8432405/files/figure.png" pageId="20" pageNumber="465">Fig. 5D</figureCitation>
|
||
). Gonopore at one-third of body length, furthest from rhinophores. Anus and nephroproct aħer middle of body length. Bursa copulatrix reniform. Penis flagelliform long (up to
|
||
<quantity id="4CF79B7FFF846503FD63FB0EEF19FB29" box="[305,382,1162,1186]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="465" unit="mm" value="30.0">30 mm</quantity>
|
||
), spiralled with pointed end (
|
||
<figureCitation id="13342A1FFF846503FEFFFB0EEC9FFB29" box="[685,760,1162,1186]" captionStart="Figure 6" captionStartId="15.[113,178,1771,1795]" captionTargetBox="[317,1256,144,1743]" captionTargetId="figure-8@15.[316,1256,144,1743]" captionTargetPageId="15" captionText="Figure 6. Reproductive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view:A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (ZMBN 83471); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis (MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations: ag, albumen gland; am, ampulla; bc, bursa copulatrix; fo, female opening; fgm, female gland mass; hd, hermaphrodite duct; o, oviduct; p, penis; pp, prostatic portion; ps, penial sheath; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.8432407" httpUri="https://zenodo.org/record/8432407/files/figure.png" pageId="20" pageNumber="465">Fig. 6D</figureCitation>
|
||
). Ampulla slightly pyriform (
|
||
<figureCitation id="13342A1FFF846503FDCAFB2DEF84FB4A" box="[408,483,1193,1217]" captionStart="Figure 6" captionStartId="15.[113,178,1771,1795]" captionTargetBox="[317,1256,144,1743]" captionTargetId="figure-8@15.[316,1256,144,1743]" captionTargetPageId="15" captionText="Figure 6. Reproductive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view:A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (ZMBN 83471); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis (MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations: ag, albumen gland; am, ampulla; bc, bursa copulatrix; fo, female opening; fgm, female gland mass; hd, hermaphrodite duct; o, oviduct; p, penis; pp, prostatic portion; ps, penial sheath; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.8432407" httpUri="https://zenodo.org/record/8432407/files/figure.png" pageId="20" pageNumber="465">Fig. 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF846503FCD3FB67EC08F9BF" blockId="20.[129,778,1251,1588]" pageId="20" pageNumber="465">
|
||
<emphasis id="B97BEA88FF846503FCD3FB67EEB9FB70" box="[129,222,1251,1275]" italics="true" pageId="20" pageNumber="465">Remarks:</emphasis>
|
||
For the first time since its description, the monotypic species
|
||
<taxonomicName id="4C0F4D19FF846503FC9DFA86EFF0FA91" authorityName="Bergh" authorityYear="1907" box="[207,407,1282,1306]" class="Gastropoda" family="Tritoniidae" genus="Tritonidoxa" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="465" phylum="Mollusca" rank="species" species="capensis">
|
||
<emphasis id="B97BEA88FF846503FC9DFA86EFF0FA91" box="[207,407,1282,1306]" italics="true" pageId="20" pageNumber="465">Tritonidoxa capensis</emphasis>
|
||
</taxonomicName>
|
||
is reassessed, and its taxonomic validity confirmed.
|
||
<taxonomicName id="4C0F4D19FF846503FD78FAA5EFF9FAB2" authorityName="Bergh" authorityYear="1907" box="[298,414,1313,1337]" class="Gastropoda" family="Tritoniidae" genus="Tritonidoxa" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="465" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF846503FD78FAA5EFF9FAB2" box="[298,414,1313,1337]" italics="true" pageId="20" pageNumber="465">Tritonidoxa</emphasis>
|
||
</taxonomicName>
|
||
was recovered as the sister-taxon of
|
||
<taxonomicName id="4C0F4D19FF846503FCD3FAC4EE8AFAD3" authorityName="Korshunova and Martynov" authorityYear="2020" box="[129,237,1344,1368]" class="Gastropoda" family="Tritoniidae" genus="Tritonicula" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="465" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF846503FCD3FAC4EE8AFAD3" box="[129,237,1344,1368]" italics="true" pageId="20" pageNumber="465">Tritonicula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED1EAE5FFF846503FCAEFAC5EF7AFAD2" box="[252,285,1345,1369]" pageId="20" pageNumber="465">PP</collectionCode>
|
||
= 1; BS = 98) (
|
||
<figureCitation id="13342A1FFF846503FD98FAC4EC63FAD2" box="[458,516,1344,1369]" captionStart="Figure 2" captionStartId="10.[129,194,1719,1743]" captionTargetBox="[325,1278,148,1689]" captionTargetId="figure-6@10.[321,1281,144,1692]" captionTargetPageId="10" captionText="Figure 2. Bayesian phylogenetic tree based on the concatenated molecular data (COI + 16S + H3) of Tritoniidae (sensu Odhner 1963). Outgroup taxa are depicted in grey.Tritoniidae genera are independently coloured, and branches with several replicates for each species hypothesis were collapsed for clarity.Names in bold are species which sequences were generated for this study with the number of sequenced specimens showed in brackets. PP are shown right/top and BS leħ/boưom.Unsupported branches are not labelled. Green circles indicate maximum support by both analyses. Solid vertical bars indicate specimens grouped as species entities based on COI dataset. Empty vertical bars indicate the specimens grouped as species based on the 16S dataset." figureDoi="http://doi.org/10.5281/zenodo.8432395" httpUri="https://zenodo.org/record/8432395/files/figure.png" pageId="20" pageNumber="465">Fig. 2</figureCitation>
|
||
), but the two genera are morphologically different:
|
||
<taxonomicName id="4C0F4D19FF846503FDC9FAE4EC38FAF3" box="[411,607,1376,1400]" class="Gastropoda" family="Tritoniidae" genus="Tritonicula" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="465" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B97BEA88FF846503FDC9FAE4EC60FAF3" box="[411,519,1376,1400]" italics="true" pageId="20" pageNumber="465">Tritonicula</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
are smaller and have a distinct first lateral tooth and reproductive morphology. The morphological synapomorphies of
|
||
<taxonomicName id="4C0F4D19FF846503FE47FA1AECEEFA3D" authorityName="Bergh" authorityYear="1907" box="[533,649,1438,1462]" class="Gastropoda" family="Tritoniidae" genus="Tritonidoxa" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="465" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF846503FE47FA1AECEEFA3D" box="[533,649,1438,1462]" italics="true" pageId="20" pageNumber="465">Tritonidoxa</emphasis>
|
||
</taxonomicName>
|
||
are the spiralled, long penis (up to
|
||
<quantity id="4CF79B7FFF846503FD38FA3AEFD4FA5D" box="[362,435,1470,1494]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="465" unit="mm" value="30.0">30 mm</quantity>
|
||
) with a pointed end (
|
||
<figureCitation id="13342A1FFF846503FED8FA3AECB3FA5D" box="[650,724,1470,1494]" captionStart="Figure 6" captionStartId="15.[113,178,1771,1795]" captionTargetBox="[317,1256,144,1743]" captionTargetId="figure-8@15.[316,1256,144,1743]" captionTargetPageId="15" captionText="Figure 6. Reproductive system arrangements of analysed key species of Tritoniinae and Marioniinae subfam. nov., dorsal view:A, Tritonia hombergii (ZMBN 83471); B, Candiella plebeia (ZMBN 83471); C, Tritoniopsis flydis (GR7-DMC00040); D, Tritonidoxa capensis (MNCN15.05/93399); E, Tochuina gigantea (NHMD-115967); F, Tritonicula hamnerorum (MNCN15.05/93378); G, Marionia blainvillea (NHMD115968). Abbreviations: ag, albumen gland; am, ampulla; bc, bursa copulatrix; fo, female opening; fgm, female gland mass; hd, hermaphrodite duct; o, oviduct; p, penis; pp, prostatic portion; ps, penial sheath; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.8432407" httpUri="https://zenodo.org/record/8432407/files/figure.png" pageId="20" pageNumber="465">Fig. 6D</figureCitation>
|
||
) and absence of secondary gills, with the notal margin distinct, broad and wrinkled. To date, there is no further knowledge about the distribution and ecology of this aberrant species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |