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<document id="A176FC9225925C5632F62524ADD0A144" ID-DOI="10.11646/zootaxa.3736.5.1" ID-GBIF-Dataset="495ee224-dd4f-4c96-aa06-094b23e135a1" ID-ISSN="1175-5326" ID-Zenodo-Dep="248431" ID-ZooBank="2A05A2A2-CF49-4585-A75D-7086B9DDD7B3" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460698653243" checkinUser="plazi" docAuthor="Sinclair, Bradley J., Brooks, Scott E. &amp; Cumming, Jeffrey M." docDate="2013" docId="03A68780FFF1FFE356A0FC966D52FE99" docLanguage="en" docName="zt03736p456.pdf" docOrigin="Zootaxa 3736 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empis" docType="treatment" docVersion="11" lastPageNumber="404" masterDocId="FF9FFFF8FFF3FFE05637FFDF691EFFFC" masterDocTitle="Revision of the Empis subgenus Enoplempis Bigot, east of the Rocky Mountains (Diptera: Empididae)" masterLastPageNumber="456" masterPageNumber="401" pageNumber="403" updateTime="1698354041895" updateUser="plazi">
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<mods:title id="CB302C36816B6DB6559E3C43AB1662F0">Revision of the Empis subgenus Enoplempis Bigot, east of the Rocky Mountains (Diptera: Empididae)</mods:title>
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<mods:namePart id="FCACDE3E52D8E2A645EBA178D4710AED">Sinclair, Bradley J.</mods:namePart>
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<mods:namePart id="DF02C18F0899B5A94CA92468ADBE8AFE">Brooks, Scott E.</mods:namePart>
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<mods:namePart id="E03E8820493F7D325901FB005764F111">Cumming, Jeffrey M.</mods:namePart>
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<mods:title id="67DF5A9EF823392768CD7E2F72CD30A3">Zootaxa</mods:title>
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<mods:date id="A8A67B0BB2DF0DBC20E8A18E58B10F55">2013</mods:date>
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<treatment id="03A68780FFF1FFE356A0FC966D52FE99" ID-DOI="http://doi.org/10.5281/zenodo.6157554" ID-GBIF-Taxon="119582592" ID-Zenodo-Dep="6157554" LSID="urn:lsid:plazi:treatment:03A68780FFF1FFE356A0FC966D52FE99" httpUri="http://treatment.plazi.org/id/03A68780FFF1FFE356A0FC966D52FE99" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<subSubSection id="C315651DFFF1FFE256A0FC966DE4FC5A" pageId="2" pageNumber="403" type="nomenclature">
<paragraph id="8BB03696FFF1FFE256A0FC9668C9FC9F" blockId="2.[151,471,841,867]" box="[151,471,841,867]" pageId="2" pageNumber="403">
<heading id="D0F881FAFFF1FFE256A0FC9668C9FC9F" bold="true" box="[151,471,841,867]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B97BEA84FFF1FFE256A0FC9668C9FC9F" bold="true" box="[151,471,841,867]" pageId="2" pageNumber="403">
<taxonomicName id="4C0F4D15FFF1FFE256A0FC9669FDFC9F" ID-CoL="9CJJ3" box="[151,227,841,867]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE256A0FC9669FDFC9F" bold="true" box="[151,227,841,867]" italics="true" pageId="2" pageNumber="403">Empis</emphasis>
</taxonomicName>
(
<taxonomicName id="4C0F4D15FFF1FFE256C3FC9668CEFC9F" authority="Bigot" authorityName="Bigot" box="[244,464,841,867]" class="Insecta" family="Empididae" genus="Enoplempis" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE256C3FC96689DFC9F" bold="true" box="[244,387,841,867]" italics="true" pageId="2" pageNumber="403">Enoplempis</emphasis>
Bigot
</taxonomicName>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB03696FFF1FFE256A0FC4F6DE4FC5A" blockId="2.[151,1274,912,934]" box="[151,1274,912,934]" pageId="2" pageNumber="403">
<taxonomicName id="4C0F4D15FFF1FFE256A0FC4F68A0FC5A" authority="Bigot, 1880a: 63" authorityName="Bigot" authorityPageNumber="63" authorityYear="1880" box="[151,446,912,934]" class="Insecta" family="Empididae" genus="Enoplempis" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE256A0FC4F6811FC5A" box="[151,271,912,934]" italics="true" pageId="2" pageNumber="403">Enoplempis</emphasis>
Bigot, 1880a: 63
</taxonomicName>
[1880b: xlvii]. Type-species,
<taxonomicName id="4C0F4D15FFF1FFE254DAFC4E6A66FC5A" authority="Bigot" authorityName="Bigot" box="[749,888,912,934]" class="Insecta" family="Empididae" genus="Enoplempis" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B97BEA84FFF1FFE254DAFC4E6A24FC5A" box="[749,826,913,934]" italics="true" pageId="2" pageNumber="403">E. mira</emphasis>
Bigot
</taxonomicName>
(monotypy) (Evenhuis &amp; Pont 2004).
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFF1FFE256A0FC0C6BAAFA86" pageId="2" pageNumber="403" type="diagnosis">
<paragraph id="8BB03696FFF1FFE256A0FC0C6BAAFA86" blockId="2.[151,1436,979,1690]" pageId="2" pageNumber="403">
<emphasis id="B97BEA84FFF1FFE256A0FC0C6814FC10" bold="true" box="[151,266,979,1004]" pageId="2" pageNumber="403">Diagnosis</emphasis>
. Typical features of the subgenus are as follows: male head dichoptic to nearly holoptic; face and prosternum bare; labellum setose, short and fleshy, not prolonged; acrostichals and dorsocentrals usually short; apical scutellar setae often widely separated; wing (Figs 2A, B) with M1 and M2 complete to wing margin, anal vein (CuA+CuP) complete (reaching wing margin) or incomplete, axillary excision acute to obtuse, axillary lobe well or little developed; male hindleg frequently modified with tibia and femur variously armed with tubercles, lobes, series of prominent setae, or bristle pencils (e.g.,
<figureCitation id="13342A13FFF1FFE254CFFB566A59FB5E" box="[760,839,1161,1186]" captionStart="FIGURE 6" captionStartId="12.[151,250,1898,1920]" captionTargetBox="[151,1436,208,1872]" captionTargetId="figure@12.[151,1436,193,1876]" captionTargetPageId="12" captionText="FIGURE 6. Male legs of E. (Enoplempis). A, E. arthritica, hindleg, anterior view; B, E. arthritica, apex of hind femur and base of hind tibia, anterior view; C, E. arthritica, apex of hind femur and base of hind tibia, posterior view; D, E. ctenocnema, hindleg, anterior view; E, E. enodis, hindleg, anterior view." httpUri="https://zenodo.org/record/248436/files/figure.png" pageId="2" pageNumber="403">Figs 6</figureCitation>
,
<figureCitation id="13342A13FFF1FFE2556DFB556A74FB5E" box="[858,874,1162,1186]" captionStart="FIGURE 9" captionStartId="18.[151,250,1907,1929]" captionTargetBox="[220,1361,193,1872]" captionTargetId="figure@18.[220,1361,193,1873]" captionTargetPageId="18" captionText="FIGURE 9. Male legs of E. (Enoplempis). A, E. gladiator, apex of hind femur and base of hind tibia, anterior view; B, E. gladiator, apex of hind femur and base of hind tibia, posterior view; C, E. gulosa, hindleg, anterior view; D, E. gulosa, hind trochanter, anterior view; E, E. loripedis, hindleg, anterior view; F, E. loripedis, apex of hind femur and base of hind tibia, anterior view; G, E. loripedis, apex of hind femur and base of hind tibia, posterior view." httpUri="https://zenodo.org/record/248439/files/figure.png" pageId="2" pageNumber="403">9</figureCitation>
,
<figureCitation id="13342A13FFF1FFE2554AFB556A85FB5E" box="[893,923,1162,1186]" captionStart="FIGURE 12" captionStartId="25.[151,250,1821,1843]" captionTargetBox="[151,1436,337,1799]" captionTargetId="figure@25.[151,1436,337,1799]" captionTargetPageId="25" captionText="FIGURE 12. Male legs of E. (Enoplempis). A, E. montywoodi, apex of hind femur and base of hind tibia, anterior view; B, E. montywoodi, apex of hind femur and base of hind tibia, posterior view; C, E. nodipoplitea, apex of hind femur and base of hind tibia, anterior view; D, E. nuda, hind trochanters; E, E. nuda, hindleg, anterior view." httpUri="https://zenodo.org/record/248442/files/figure.png" pageId="2" pageNumber="403">12</figureCitation>
,
<figureCitation id="13342A13FFF1FFE25598FB566AD3FB5E" box="[943,973,1161,1186]" captionStart="FIGURE 16" captionStartId="33.[151,250,1425,1447]" captionTargetBox="[151,1436,359,1409]" captionTargetId="figure@33.[151,1436,359,1410]" captionTargetPageId="33" captionText="FIGURE 16. Male legs of E. (Enoplempis). A, E. pectinata, hindleg, anterior view; B, E. penicillata, hindleg, anterior view; C, E. prodigiosa, hindleg, anterior view; D, E. prodigiosa, apex of hind femur and base of hind tibia, anterior view; E, E. prodigiosa, apex of hind femur and base of hind tibia, posterior view." httpUri="https://zenodo.org/record/248446/files/figure.png" pageId="2" pageNumber="403">16</figureCitation>
,
<figureCitation id="13342A13FFF1FFE255D7FB556AE1FB5E" box="[992,1023,1162,1186]" captionStart="FIGURE 23" captionStartId="49.[151,250,1934,1956]" captionTargetBox="[166,1436,1185,1912]" captionTargetId="figure@49.[151,1436,1170,1912]" captionTargetPageId="49" captionText="FIGURE 23. Male legs of E. (Enoplempis) volsella. A, hindleg, anterior view; B, apex of hind femur and base of hind tibia, anterior view; C, apex of hind femur and base of hind tibia, posterior view." httpUri="https://zenodo.org/record/248453/files/figure.png" pageId="2" pageNumber="403">23</figureCitation>
); right and left hindlegs normally symmetrical; tarsomeres 2–5 of all legs (in both sexes) usually with 2 ventral rows of short spine-like setae (Figs 2C, D, E); tarsomere 4 usually with apicolateral pair of spine-like setae; male pregenital segments generally unmodified, except some western species; ejaculatory apodeme in anterior view, often inverted Y-shaped, rarely Tshaped or flattened; female without most empidine secondary sexual features, such as expanded and darkened wings, pinnate leg scales, and silvery abdominal pruinesence; extensive eversible darkened abdominal pleural membrane observed in females of some species.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFF1FFE356F0FA5A6D52FE99" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403" type="discussion">
<paragraph id="8BB03696FFF1FFE256F0FA5A6B9FF966" blockId="2.[151,1436,979,1690]" pageId="2" pageNumber="403">
<emphasis id="B97BEA84FFF1FFE256F0FA5A682AFA62" bold="true" box="[199,308,1413,1438]" pageId="2" pageNumber="403">Remarks</emphasis>
. The subgenus
<taxonomicName id="4C0F4D15FFF1FFE257D2FA5A6B77FA62" box="[485,617,1413,1438]" class="Insecta" family="Empididae" genus="Enoplempis" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE257D2FA5A6B77FA62" box="[485,617,1413,1438]" italics="true" pageId="2" pageNumber="403">Enoplempis</emphasis>
</taxonomicName>
is most readily identified from other
<taxonomicName id="4C0F4D15FFF1FFE25238FA596D4BFA61" box="[1039,1109,1414,1437]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE25238FA596D4BFA61" box="[1039,1109,1414,1437]" italics="true" pageId="2" pageNumber="403">Empis</emphasis>
</taxonomicName>
subgenera by paired rows of ventral spine-like setae on the tarsi, which appears to be correlated with characteristic post-mortum tarsal curling seen in most museum specimens (Figs 2D, E). These spine-like setae are reduced in some species (see species group description) and are not unique to
<emphasis id="B97BEA84FFF1FFE25456FA2C6B64F9F6" box="[609,634,1523,1546]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<taxonomicName id="4C0F4D15FFF1FFE254BDFA2E6A10F9F6" box="[650,782,1521,1546]" class="Insecta" family="Empididae" genus="Enoplempis" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE254BDFA2E6A10F9F6" box="[650,782,1521,1546]" italics="true" pageId="2" pageNumber="403">Enoplempis</emphasis>
</taxonomicName>
). Rows of ventral tarsal spines also occur in some species of other subgenera of
<taxonomicName id="4C0F4D15FFF1FFE257AFF9C968C0F9D1" box="[408,478,1558,1581]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE257AFF9C968C0F9D1" box="[408,478,1558,1581]" italics="true" pageId="2" pageNumber="403">Empis</emphasis>
</taxonomicName>
, such as various
<emphasis id="B97BEA84FFF1FFE2549CF9C86BDDF9D2" box="[683,707,1559,1582]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<taxonomicName id="4C0F4D15FFF1FFE254E1F9CA6A70F9D2" box="[726,878,1557,1582]" class="Insecta" family="Empididae" genus="Coptophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE254E1F9CA6A70F9D2" box="[726,878,1557,1582]" italics="true" pageId="2" pageNumber="403">Coptophlebia</emphasis>
</taxonomicName>
)
<emphasis id="B97BEA84FFF1FFE255B5F9C86A93F9D2" box="[898,909,1559,1582]" italics="true" pageId="2" pageNumber="403">s</emphasis>
.
<emphasis id="B97BEA84FFF1FFE255A9F9CA6AA5F9D2" box="[926,955,1557,1582]" italics="true" pageId="2" pageNumber="403">lat</emphasis>
. (from North
<collectingCountry id="F3187606FFF1FFE25256F9CA6DDCF9D2" box="[1121,1218,1557,1582]" name="United States of America" pageId="2" pageNumber="403">America</collectingCountry>
and
<collectingCountry id="F3187606FFF1FFE252C9F9C96C21F9D2" box="[1278,1343,1558,1582]" name="Japan" pageId="2" pageNumber="403">Japan</collectingCountry>
) and
<taxonomicName id="4C0F4D15FFF1FFE253B3F9C8687CF9AE" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="livida" subGenus="Kritempis">
<emphasis id="B97BEA84FFF1FFE253B3F9C86C82F9D2" box="[1412,1436,1559,1582]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<emphasis id="B97BEA84FFF1FFE25697F9E46810F9AE" box="[160,270,1595,1618]" italics="true" pageId="2" pageNumber="403">Kritempis</emphasis>
)
<emphasis id="B97BEA84FFF1FFE25715F9E6687CF9AE" box="[290,354,1593,1618]" italics="true" pageId="2" pageNumber="403">livida</emphasis>
</taxonomicName>
L. (not as pronounced in females).
<taxonomicName id="4C0F4D15FFF1FFE25531F9E46A91F9AE" authority="(En.)" baseAuthorityName="En." box="[774,911,1594,1618]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF1FFE25531F9E46A52F9AE" box="[774,844,1595,1618]" italics="true" pageId="2" pageNumber="403">Empis</emphasis>
(
<emphasis id="B97BEA84FFF1FFE25557F9E46A98F9AE" box="[864,902,1595,1618]" italics="true" pageId="2" pageNumber="403">En.</emphasis>
)
</taxonomicName>
can be tentatively divided into the following three broad species groups, although final decisions on the monophyly and relationships of these groups must wait until the western species are fully analyzed.
</paragraph>
<paragraph id="8BB03696FFF1FFE256A0F91568D6F8B2" blockId="2.[151,1436,1737,2014]" pageId="2" pageNumber="403">
<bibRef id="52AFA260FFF1FFE256A0F91568D6F8B2" pageId="2" pageNumber="403" type="book chapter">
(
<pagination id="B2C6C862FFF1FFE25696F91569B3F91E" box="[161,173,1738,1762]" pageId="2" pageNumber="403" type="pagination">1</pagination>
)
<title id="F90BDDA0FFF1FFE256F5F9166AE8F8D6" jopScore="0.3024191" pageId="2" pageNumber="403">
The
<emphasis id="B97BEA84FFF1FFE256CFF914680FF91E" box="[248,273,1739,1762]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<emphasis id="B97BEA84FFF1FFE25713F91668B6F91E" box="[292,424,1737,1762]" italics="true" pageId="2" pageNumber="403">Enoplempis</emphasis>
)
<emphasis id="B97BEA84FFF1FFE2578AF91468ECF91E" box="[445,498,1739,1762]" italics="true" pageId="2" pageNumber="403">mira</emphasis>
-group includes three species with male hindlegs highly modified and bilaterally asymmetrical; hind tibia geniculate at base in both sexes; acrostichals biserial; dorsocentrals uniserial; apical scutellar setae closely positioned; ventral tarsal spine-like setae absent
</title>
.
<journalOrPublisher id="51CE3C5AFFF1FFE25233F8CE68DAF8B2" jopScore="0.82193863" pageId="2" pageNumber="403">This group is recorded from Idaho, Oregon and California</journalOrPublisher>
.
</bibRef>
</paragraph>
<paragraph id="8BB03696FFF1FFE256A0F8856D6CF846" blockId="2.[151,1436,1737,2014]" pageId="2" pageNumber="403">
<bibRef id="52AFA260FFF1FFE256A0F8856D6CF846" pageId="2" pageNumber="403" type="book chapter">
(
<pagination id="B2C6C862FFF1FFE256A8F88569B0F88E" box="[159,174,1882,1906]" pageId="2" pageNumber="403" type="pagination">2</pagination>
)
<title id="F90BDDA0FFF1FFE256F5F8866B26F846" jopScore="0.3273254" pageId="2" pageNumber="403">
The
<emphasis id="B97BEA84FFF1FFE256CFF884680FF88E" box="[248,273,1883,1906]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<emphasis id="B97BEA84FFF1FFE25713F88668B6F88E" box="[292,424,1881,1906]" italics="true" pageId="2" pageNumber="403">Enoplempis</emphasis>
)
<emphasis id="B97BEA84FFF1FFE2578BF8846B07F88E" box="[444,537,1883,1906]" italics="true" pageId="2" pageNumber="403">geneatis</emphasis>
-group includes at least two species and is characterized by unmodified male hindlegs; long setae on scape, palpi and back of head; acrostichals and dorsocentrals multiserial; apical scutellar setae closely positioned
</title>
.
<journalOrPublisher id="51CE3C5AFFF1FFE25474F87E6D73F846" box="[579,1133,1953,1978]" jopScore="0.4814511" pageId="2" pageNumber="403">This group is known from Oregon and California</journalOrPublisher>
.
</bibRef>
</paragraph>
<paragraph id="8BB03696FFF1FFE356A0F819686BFF04" blockId="2.[151,1436,1737,2014]" lastBlockId="3.[197,1436,151,248]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<bibRef id="52AFA260FFF1FFE356A0F819686BFF04" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403" type="journal article">
(
<part id="FFAB9C0BFFF1FFE25697F81969B0F822" box="[160,174,1990,2014]" pageId="2" pageNumber="403" type="issue">3</part>
) The
<emphasis id="B97BEA84FFF1FFE256C0F8186811F822" box="[247,271,1991,2014]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<emphasis id="B97BEA84FFF1FFE25715F81A68B8F822" box="[290,422,1989,2014]" italics="true" pageId="2" pageNumber="403">Enoplempis</emphasis>
)
<emphasis id="B97BEA84FFF1FFE2578DF81A68E5F822" box="[442,507,1989,2014]" italics="true" pageId="2" pageNumber="403">bigoti</emphasis>
-group comprises over
<pagination id="B2C6C862FFF1FFE25530F8196A3DF822" box="[775,803,1990,2014]" pageId="2" pageNumber="403" type="pagination">40</pagination>
<title id="F90BDDA0FFF1FFE3551AF81A6D3DFF29" jopScore="0.26533204" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403" type="_" typeScore="20">species and is usually characterized by modified male hindlegs (tibia, femur and/or trochanter and coxa); apical scutellar setae widely separated; acrostichals and dorsocentrals uniserial and reduced; tarsal spine-like setae very pronounced</title>
.
<journalOrPublisher id="51CE3C5AFFF0FFE35218FF63686FFF04" jopScore="0.3887188" pageId="3" pageNumber="404">
This group is widespread across North
<collectingCountry id="F3187606FFF0FFE3573AFF00686FFF04" box="[269,369,223,248]" name="United States of America" pageId="3" pageNumber="404">America</collectingCountry>
</journalOrPublisher>
.
</bibRef>
</paragraph>
<paragraph id="8BB03696FFF0FFE356F0FEF86D52FE99" blockId="3.[151,1436,295,357]" pageId="3" pageNumber="404">
The eastern species of
<emphasis id="B97BEA84FFF0FFE357E2FEF668F0FEBC" box="[469,494,297,320]" italics="true" pageId="3" pageNumber="404">E.</emphasis>
(
<taxonomicName id="4C0F4D15FFF0FFE35435FEF86B98FEBC" box="[514,646,295,320]" class="Insecta" family="Empididae" genus="Enoplempis" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEA84FFF0FFE35435FEF86B98FEBC" box="[514,646,295,320]" italics="true" pageId="3" pageNumber="404">Enoplempis</emphasis>
</taxonomicName>
) all belong to the large
<taxonomicName id="4C0F4D15FFF0FFE3559FFEF66DB0FEBC" box="[936,1198,295,320]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="bigoti" subGenus="Enoplempis">
<emphasis id="B97BEA84FFF0FFE3559FFEF66ADFFEBC" box="[936,961,297,320]" italics="true" pageId="3" pageNumber="404">E.</emphasis>
(
<emphasis id="B97BEA84FFF0FFE355E2FEF86D47FEBC" box="[981,1113,295,320]" italics="true" pageId="3" pageNumber="404">Enoplempis</emphasis>
)
<emphasis id="B97BEA84FFF0FFE3525AFEF86DB0FEBC" box="[1133,1198,295,320]" italics="true" pageId="3" pageNumber="404">bigoti</emphasis>
</taxonomicName>
-group. They can be further classified into at least four subgroups as indicated below under “Discussion”.
</paragraph>
</subSubSection>
</treatment>
</document>
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