treatments-xml/data/03/94/AB/0394AB1DFFAC4D303981F9D23F38FE5F.xml

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<mods:title id="56404B52704B5B6FE0074751A7695AC8">Cambarus ectopistes sp. nov., a new stream-dwelling crayfish (Decapoda: Cambaridae) from the French Broad, Pigeon, and Nolichucky River watersheds in the Appalachian Mountain region of North Carolina and Tennessee, USA</mods:title>
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<mods:namePart id="23E49C2E1EDC8E5E43DC2F2261E369F2">Loughman, Zachary J.</mods:namePart>
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<mods:affiliation id="839CFA30DB70E7898316BD17B4050508">West Liberty University, Department of Biological Sciences, C. S. C Box 139, West Liberty, West Virginia, 26074. zloughman @ westliberty. edu; https: // orcid. org / 0000 - 0002 - 1486 - 4999</mods:affiliation>
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<mods:namePart id="7623D9A2B9EFF756B1D313E280831F51">Williams, Bronwyn W.</mods:namePart>
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<mods:affiliation id="4EB26173B7CAB6DEBC665EF26ED13717">North Carolina Museum of Natural Sciences, Research Laboratory, 1671 Gold Star Drive, Raleigh, NC 27669. bronwyn. williams @ naturalsciences. org; https: // orcid. org / 0000 - 0001 - 6403 - 112 X</mods:affiliation>
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<heading id="D0CAAD67FFAC4D393981F9D23E00F999" bold="true" box="[151,803,1559,1586]" fontSize="11" level="1" pageId="6" pageNumber="328" reason="1">
<emphasis id="B949C619FFAC4D393981F9D23E00F999" bold="true" box="[151,803,1559,1586]" pageId="6" pageNumber="328">
<taxonomicName id="4C3D6188FFAC4D393981F9D23F9AF999" authority="Loughman &amp; Williams" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[151,697,1559,1586]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="328" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFAC4D393981F9D23CB3F999" bold="true" box="[151,400,1559,1586]" italics="true" pageId="6" pageNumber="328">Cambarus ectopistes</emphasis>
Loughman &amp; Williams
</taxonomicName>
, 
<taxonomicNameLabel id="A27A7B62FFAC4D393BD3F9DD3E00F999" box="[709,803,1560,1586]" pageId="6" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B821A0BFFAC4D393981F9F93C4EF9FD" blockId="6.[151,803,1559,1623]" box="[151,365,1596,1623]" pageId="6" pageNumber="328">
<figureCitation id="1306068EFFAC4D393981F9F93C13F9FD" box="[151,304,1596,1622]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="6" pageNumber="328">Figures 4A–J</figureCitation>
, 
<figureCitation id="1306068EFFAC4D39382DF9F93C68F9FC" box="[315,331,1596,1623]" captionStart="FIGURE 5" captionStartId="9.[151,250,1097,1121]" captionTargetBox="[151,1435,181,1072]" captionTargetId="figure-28@9.[151,1435,181,1073]" captionTargetPageId="9" captionText="FIGURE 5. Live color of A) adult male and B) adult female C. ectopistes sp. nov. from the Pigeon River, NC. Photos by Michael Perkins and W.T. Russ (North Carolina Wildlife Resources Commission)." figureDoi="http://doi.org/10.5281/zenodo.5792769" httpUri="https://zenodo.org/record/5792769/files/figure.png" pageId="6" pageNumber="328">5</figureCitation>
, 
<figureCitation id="1306068EFFAC4D393841F9F93C44F9FD" box="[343,359,1596,1622]" captionStart="FIGURE 8" captionStartId="14.[151,250,875,899]" captionTargetBox="[155,1433,181,851]" captionTargetId="figure-17@14.[155,1433,181,851]" captionTargetPageId="14" captionText="FIGURE 8. Intraspecific variation in C. ectopistes sp. nov. occurs most notably in the shape of the rostrum.A, Nolichucky and Cane Rivers; B, French Broad and Pigeon Rivers." figureDoi="http://doi.org/10.5281/zenodo.5792777" httpUri="https://zenodo.org/record/5792777/files/figure.png" pageId="6" pageNumber="328">8</figureCitation>
.
</paragraph>
<paragraph id="8B821A0BFFAC4D393981F9453E3AF91D" blockId="6.[151,1436,1664,1811]" pageId="6" pageNumber="328">
<taxonomicName id="4C3D6188FFAC4D393981F9453F03F933" box="[151,544,1664,1688]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="328" phylum="Arthropoda" rank="species" species="robustus" subGenus="Puncticambarus">
<emphasis id="B949C619FFAC4D393981F9453C22F933" box="[151,257,1664,1688]" italics="true" pageId="6" pageNumber="328">Cambarus</emphasis>
(
<emphasis id="B949C619FFAC4D393806F9453C95F933" box="[272,438,1664,1688]" italics="true" pageId="6" pageNumber="328">Puncticambarus</emphasis>
) 
<emphasis id="B949C619FFAC4D3938D0F9453F03F933" box="[454,544,1664,1688]" italics="true" pageId="6" pageNumber="328">robustus</emphasis>
</taxonomicName>
<emphasis id="B949C619FFAC4D393B36F9453F07F933" box="[544,548,1664,1688]" italics="true" pageId="6" pageNumber="328">.</emphasis>
—Cooper 2010: 72, 74, 75 [all in part].—Fullerton, 2002: 13, 16, 21, 27, 31, 40 [all in part].— 
<bibRefCitation id="EFAC67FAFFAC4D393801F95A3F1AF91C" author="Simmons, J. W. &amp; Fraley, S. J." box="[279,569,1695,1719]" pageId="6" pageNumber="328" pagination="79 - 126" refId="ref12705" refString="Simmons, J. W. &amp; Fraley, S. J. (2010) Distribution, Status, and Life-history Observations of Crayfishes in Western North Carolina. Conservation, Biology, and Natural History of Crayfishes from the Southern US. Southeastern Naturalist, 9, 79 - 126. https: // doi. org / 10.1656 / 058.009. s 316" type="journal article" year="2010">Simmons &amp; Fraley 2010: 83</bibRefCitation>
, 111, 112 [all in part].
</paragraph>
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<emphasis id="B949C619FFAC4D393981F97B3C46F97D" box="[151,357,1726,1750]" italics="true" pageId="6" pageNumber="328">
<taxonomicName id="4C3D6188FFAC4D393981F97B3C42F97D" box="[151,353,1726,1750]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="328" phylum="Arthropoda" rank="species" species="robustus">Cambarus robustus</taxonomicName>
.
</emphasis>
—Cooper &amp; Braswell 1995: 116, 126 [all in part].
</paragraph>
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<taxonomicName id="4C3D6188FFAC4D393981F9193C22F95F" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[151,257,1756,1780]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="328" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B949C619FFAC4D393981F9193C22F95F" box="[151,257,1756,1780]" italics="true" pageId="6" pageNumber="328">Cambarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A7B62FFAC4D393811F9183C75F95F" box="[263,342,1757,1780]" pageId="6" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
—Dunn 2010: 9, 27, 33, 53, 60, 61, 64-66.
</paragraph>
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<paragraph id="8B821A0BFFAC4D393981F93E3EE3F8B8" blockId="6.[151,1436,1664,1811]" box="[151,960,1787,1811]" pageId="6" pageNumber="328">
<treatmentCitationGroup id="AB2D3D25FFAC4D393981F93E3EE3F8B8" box="[151,960,1787,1811]" pageId="6" pageNumber="328">
<taxonomicName id="4C3D6188FFAC4D393981F93E3C9DF8B8" ID-CoL="8P427" authority="" box="[151,446,1787,1811]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="328" phylum="Arthropoda" rank="subGenus" subGenus="Puncticambarus">
<emphasis id="B949C619FFAC4D393981F93E3C22F8B8" box="[151,257,1787,1811]" italics="true" pageId="6" pageNumber="328">Cambarus</emphasis>
(
<emphasis id="B949C619FFAC4D393806F93E3C95F8B8" box="[272,438,1787,1811]" italics="true" pageId="6" pageNumber="328">Puncticambarus</emphasis>
)
</taxonomicName>
sp. nov.— 
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<bibRefCitation id="EFAC67FAFFAC4D393B38F93E3E78F8B8" author="Simmons, J. W. &amp; Fraley, S. J." box="[558,859,1787,1811]" pageId="6" pageNumber="328" pagination="79 - 126" refId="ref12705" refString="Simmons, J. W. &amp; Fraley, S. J. (2010) Distribution, Status, and Life-history Observations of Crayfishes in Western North Carolina. Conservation, Biology, and Natural History of Crayfishes from the Southern US. Southeastern Naturalist, 9, 79 - 126. https: // doi. org / 10.1656 / 058.009. s 316" type="journal article" year="2010">Simmons &amp; Fraley 2010: 110</bibRefCitation>
</treatmentCitation>
, 114-115.
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</paragraph>
</subSubSection>
<subSubSection id="C3274980FFAC4D353981F8843FC4FEB6" lastPageId="10" lastPageNumber="332" pageId="6" pageNumber="328" type="diagnosis">
<paragraph id="8B821A0BFFAC4D383981F8843995FD5A" blockId="6.[151,1437,1857,2028]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="329" pageId="6" pageNumber="328">
<emphasis id="B949C619FFAC4D393981F8843C31F8F0" bold="true" box="[151,274,1857,1883]" pageId="6" pageNumber="328">Diagnosis.</emphasis>
Body and eyes pigmented. Base of rostrum broad; anterior portion broad and sub-parallel, moderately excavated, and deflected ventrally, margins thickened to acumen; acumen margins not thickened and strongly converging. Floor of rostrum punctate. Acumen distinctly triangular with prominent dorsally deflected spiniform tubercle at terminus. Areola 2.5–4.9 (
<emphasis id="B949C619FFAC4D393B32F86B3F13F86A" box="[548,560,1966,1985]" italics="true" pageId="6" pageNumber="328">x</emphasis>
= 3.3, 
<emphasis id="B949C619FFAC4D393B92F86B3FB1F86C" box="[644,658,1966,1991]" italics="true" pageId="6" pageNumber="328">n</emphasis>
= 86, s = 0.5) times as long as wide with 7–11 (usually 9) punctations across narrowest point. Juveniles with weak cervical spine; adults with spiniform tubercle along cervical groove. Mandibular, branchiostegal, and orbital regions of carapace with well-developed tubercles. Postorbital ridge not terminating in spine; occasionally terminating in tubercle. Suborbital angle acute, terminating in small tubercle. Total carapace length (TCL) 1.7 – 2.1 (
<emphasis id="B949C619FFAD4D383B02FF253F03FF58" box="[532,544,224,243]" italics="true" pageId="7" pageNumber="329">x</emphasis>
= 1.9, 
<emphasis id="B949C619FFAD4D383B60FF1A3FA7FF53" box="[630,644,223,248]" italics="true" pageId="7" pageNumber="329">n</emphasis>
= 86, s = 0.1) times longer than width. Form I and II males with hook on ischium of third pereopods only; hook gently curved at apex, overarching basioischial joint in form I males, not reaching basioischial joint in form II males; hooks not opposed by tubercle on basis. Mesial surface of chela possessing two rows of tubercles; mesial most row with 5–9 (
<emphasis id="B949C619FFAD4D383A38FE893E19FEF4" box="[814,826,332,351]" italics="true" pageId="7" pageNumber="329">x</emphasis>
= 7.4, n = 86, s = 0.1) tubercles, second (dorsal) row with 4–8 (= 7.4, n = 86, s = 0.9) tubercles. Chela normally lacking sub-palmar tubercles, rarely with one. Dorsomedian ridge of fixed finger of propodus pronounced. Lateral impression at base of fixed finger present. Mesial margin of palm punctate. Dactyl dorsal margin with well-developed tubercles, especially near junction with propodus. Dactyl and fixed finger each with sharp corneous tip. Form I male palm length 82.5–114.5% (
<emphasis id="B949C619FFAD4D383DF4FE1939CDFE44" box="[1250,1262,476,495]" italics="true" pageId="7" pageNumber="329">x</emphasis>
= 95.9, 
<emphasis id="B949C619FFAD4D383C59FE1E387EFE5F" box="[1359,1373,475,500]" italics="true" pageId="7" pageNumber="329">n</emphasis>
= 16, s = 9.3) of palm width, form I male palm length 26.1–30.8% (
<emphasis id="B949C619FFAD4D383A4AFDC53E4BFDB8" box="[860,872,512,531]" italics="true" pageId="7" pageNumber="329">x</emphasis>
= 28.6, 
<emphasis id="B949C619FFAD4D383ADBFE3A3EF8FDB3" box="[973,987,511,536]" italics="true" pageId="7" pageNumber="329">n</emphasis>
= 16, s = 3.7) of total propodus length; female dactyl length 56.8–66.5% (
<emphasis id="B949C619FFAD4D383B0AFDE13F0BFD9C" box="[540,552,548,567]" italics="true" pageId="7" pageNumber="329">x</emphasis>
= 61.0, 
<emphasis id="B949C619FFAD4D383B9DFDE63FBAFD97" box="[651,665,547,572]" italics="true" pageId="7" pageNumber="329">n</emphasis>
= 37, s = 2.5) of total propodus length. Ventral margin of merus with 7–10 (
<emphasis id="B949C619FFAD4D3839F4FD8D3DCDFDF0" box="[226,238,584,603]" italics="true" pageId="7" pageNumber="329">x</emphasis>
= 8.4, 
<emphasis id="B949C619FFAD4D383854FD823C73FDCB" box="[322,336,583,608]" italics="true" pageId="7" pageNumber="329">n</emphasis>
= 86, s = 2.1) spines. First pleopod of form I male with long terminal elements. Central projection not tapering distally; recurved&gt;90º to main shaft of gonopod, possessing well developed subapical notch. Caudal knob absent at lateral base of central projection. Mesial process directed 90° to shaft; nondescript and typical for the genus. Annulus ventralis immovable; distinctly asymmetrical caudally; cephalic portion with median trough leading to strongly sculptured central fossa; exaggerated “S” bend in sinus terminating at caudal edge.
</paragraph>
<paragraph id="8B821A0BFFAD4D3839D1FD3F3F41F94A" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D3839D1FD3F3E34FCBF" bold="true" box="[199,791,762,789]" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D3839D1FD3F3CEDFCBE" bold="true" box="[199,462,762,789]" italics="true" pageId="7" pageNumber="329">Holotypic male, form I</emphasis>
(
<figureCitation id="1306068EFFAD4D3838CBFD3F3F80FCBF" box="[477,675,762,788]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">Figs. 4A–C, G–J</figureCitation>
, 
<tableCitation id="C6BF2FB0FFAD4D383BA6FD3E3E24FCBF" box="[688,775,762,789]" captionStart="TABLE 1" captionStartId="5.[151,239,154,178]" captionTargetPageId="5" captionText="TABLE 1. Morphometric variables compared between C. ectopistes sp. nov. and C. robustus s.s., including mean (x) and standard deviations (s), test statistics (either Wilcoxon rank-sum test, W, or Welch’s t-test, t, with subscript = degrees of freedom), and associated p-values. Statistically significant differences between the two species (P &lt;0.05) are denoted using an asterisk *. Length (L), width (W), depth (D), total carapace length (TCL), carapace width (CW)." httpUri="http://table.plazi.org/id/DF424A83FFAF4D3A3981FF5F39DFFEB5" pageId="7" pageNumber="329" tableUuid="DF424A83FFAF4D3A3981FF5F39DFFEB5">Table 1</tableCitation>
).
</emphasis>
Body compressed dorsoventrally (
<figureCitation id="1306068EFFAD4D383D89FD3F39D6FCBF" box="[1183,1269,762,788]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383D89FD3F39D6FCBF" bold="true" box="[1183,1269,762,788]" pageId="7" pageNumber="329">Fig. 4A</emphasis>
</figureCitation>
); thoracic section of carapace slightly wider than abdomen. Carapace depth less than carapace width at caudodorsal margin of cervical groove. Total carapace length 
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; postorbital carapace length (PCL) 
<quantity id="4CC5B7EEFFAD4D383D28FC863986FCF6" box="[1086,1189,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="4.24" pageId="7" pageNumber="329" unit="mm" value="42.4">42.4 mm</quantity>
. Areola 4.1 times longer than wide, with 10 punctations across narrowest part (
<figureCitation id="1306068EFFAD4D383A0DFCA33E53FC2B" box="[795,880,870,896]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383A0DFCA33E53FC2B" bold="true" box="[795,880,870,896]" pageId="7" pageNumber="329">Fig. 4B</emphasis>
</figureCitation>
); length of areola 35.7% of TCL (42.2% of PCL). Rostrum broad and excavated equally along length; margins thickened and subparallel to acumen; floor of rostrum punctate. Rostrum 1.5 times longer than wide. Acumen distinctly triangular, ending in dorsally deflected corneous tip (
<figureCitation id="1306068EFFAD4D3839D3FC173C1DFC46" box="[197,318,978,1005]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D3839D3FC173C1DFC46" bold="true" box="[197,318,978,1005]" pageId="7" pageNumber="329">Fig. 4A, B</emphasis>
</figureCitation>
). Postorbital ridges long, terminating in small tubercle. Suborbital angle acute, lacking pronounced spine or tubercle (
<figureCitation id="1306068EFFAD4D38387FFC333CE2FBBB" box="[361,449,1014,1040]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D38387FFC333CE2FBBB" bold="true" box="[361,449,1014,1040]" pageId="7" pageNumber="329">Fig. 4A</emphasis>
</figureCitation>
). Cervical spine absent; small spiniform tubercle present. Mandibular, branchiostegal, and orbital regions of carapace with several well-developed tubercles; greatest tubercle density in hepatic region. Abdomen slightly longer than carapace, pleura rounded both cephaloventrally and caudoventrally. Cephalic section of telson with 1 large spine in each caudolateral corner. Proximal podomere of uropod with 8 laterodistal spines on mesial lobe; median spine, originating from ventral surface; mesial ramus of uropod with median ridge ending distally in distomedian spine not overreaching margin of ramus. Distal margin of proximal segment of lateral ramus of right uropod having 10 immovable, small spines. Cephalomedian lobe of epistome subtriangular, forming weak angle at junction with endostyle; cephalolateral margins not thickened; zygoma moderately arched (
<figureCitation id="1306068EFFAD4D383DE3FB37386FFAA7" box="[1269,1356,1266,1292]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383DE3FB37386FFAA7" bold="true" box="[1269,1356,1266,1292]" pageId="7" pageNumber="329">Fig. 4C</emphasis>
</figureCitation>
); main body lacking prominent cephalomedian fovea. Antennal scale broadest anteriorly; lateral margin thickened, terminating in large corneous spine; mesial margin setiferous (
<figureCitation id="1306068EFFAD4D383A02FAFF3E49FAFF" box="[788,874,1338,1364]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383A02FAFF3E49FAFF" bold="true" box="[788,874,1338,1364]" pageId="7" pageNumber="329">Fig. 4D</emphasis>
</figureCitation>
). Right antennal scale 
<quantity id="4CC5B7EEFFAD4D383D7FFAFE399DFAFE" box="[1129,1214,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="7" pageNumber="329" unit="mm" value="7.9">7.9 mm</quantity>
long, 
<quantity id="4CC5B7EEFFAD4D383C15FAFE387AFAFE" box="[1283,1369,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="329" unit="mm" value="4.1">4.1 mm</quantity>
wide. Tip of right antenna reaching first abdominal terga when adpressed. Mesial surface of palm of chela with two well defined tubercle rows; mesial most row possessing 7 tubercles and second row with 5 (
<figureCitation id="1306068EFFAD4D383D4AFA473992FA37" box="[1116,1201,1410,1436]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383D4AFA473992FA37" bold="true" box="[1116,1201,1410,1436]" pageId="7" pageNumber="329">Fig. 4E</emphasis>
</figureCitation>
). Palm length 69.9% of palm width; depth of palm 
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. Ventral surface of palm lacking tubercles. Dorsal longitudinal ridge of dactyl moderately developed, punctate, possessing well defined tubercles (
<emphasis id="B949C619FFAD4D383A86FA0F3ED6FA4E" bold="true" box="[912,1013,1482,1509]" pageId="7" pageNumber="329">
<figureCitation id="1306068EFFAD4D383A86FA0F3EC6FA4F" box="[912,997,1482,1508]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">Fig. 4E</figureCitation>
-
</emphasis>
punctations difficult to see in figure). Dorsomedian ridge of fixed finger of propodus well-developed. Possessing a lateral impression at the junction of fixed finger and palm. Dactyl and fixed finger of propodus both with sharp, corneous tips. All measurements and counts from right chela. Carpus with prominent dorsal furrow (
<figureCitation id="1306068EFFAD4D383A43F9F33E88F9FB" box="[853,939,1590,1616]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383A43F9F33E88F9FB" bold="true" box="[853,939,1590,1616]" pageId="7" pageNumber="329">Fig. 4E</emphasis>
</figureCitation>
), surface lacking extensive punctations; mesial margin with large, procurved spine at about midlength. Distodorsal surface of merus without spines or tubercles. Hook on ischium of third pereopods only; hook gently curved at apex, overarching basioischial joint, not opposed by tubercle on basis. Form I gonopod as described in diagnosis (
<figureCitation id="1306068EFFAD4D383A7FF9673EC2F917" box="[873,993,1698,1724]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383A7FF9673EC2F917" bold="true" box="[873,993,1698,1724]" pageId="7" pageNumber="329">Fig. 4F–G</emphasis>
</figureCitation>
); tip reaching anterior margin of fourth caudomesial boss when abdomen flexed.
</paragraph>
<paragraph id="8B821A0BFFAD4D3839D1F92F3CB5F876" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D3839D1F92F3F46F8AF" bold="true" box="[199,613,1770,1797]" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D3839D1F92F3CA7F8AE" bold="true" box="[199,388,1770,1797]" italics="true" pageId="7" pageNumber="329">Allotypic female</emphasis>
(
<figureCitation id="1306068EFFAD4D383885F92F3CD3F8AF" box="[403,496,1770,1796]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">Fig. 4H</figureCitation>
, 
<tableCitation id="C6BF2FB0FFAD4D3838EBF92E3F76F8AF" box="[509,597,1770,1797]" captionStart="TABLE 1" captionStartId="5.[151,239,154,178]" captionTargetPageId="5" captionText="TABLE 1. Morphometric variables compared between C. ectopistes sp. nov. and C. robustus s.s., including mean (x) and standard deviations (s), test statistics (either Wilcoxon rank-sum test, W, or Welch’s t-test, t, with subscript = degrees of freedom), and associated p-values. Statistically significant differences between the two species (P &lt;0.05) are denoted using an asterisk *. Length (L), width (W), depth (D), total carapace length (TCL), carapace width (CW)." httpUri="http://table.plazi.org/id/DF424A83FFAF4D3A3981FF5F39DFFEB5" pageId="7" pageNumber="329" tableUuid="DF424A83FFAF4D3A3981FF5F39DFFEB5">Table 1</tableCitation>
).
</emphasis>
Differing from 
<typeStatus id="5486A4A9FFAD4D383A0DF92E3E5EF8AE" box="[795,893,1771,1797]" pageId="7" pageNumber="329" type="holotype">holotype</typeStatus>
in following respects: TCL 
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; PCL 
<quantity id="4CC5B7EEFFAD4D383C7DF92E3DE5F882" metricMagnitude="-2" metricUnit="m" metricValue="3.63" pageId="7" pageNumber="329" unit="mm" value="36.3">36.3 mm</quantity>
; areola length 34.6% of TCL (41.6% of PCL); 3.2 times as long as wide; rostrum 1.8 times longer than wide; abdomen length 
<quantity id="4CC5B7EEFFAD4D383847F8F63C94F8E6" box="[337,439,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="4.91" pageId="7" pageNumber="329" unit="mm" value="49.1">49.1 mm</quantity>
; antennal scale 
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long, 
<quantity id="4CC5B7EEFFAD4D383A16F8F63E6CF8E6" box="[768,847,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="329" unit="mm" value="3.6">3.6mm</quantity>
wide. Mesial surface of palm of chela with two rows of 7 tubercles each; palm length (
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) 68.8% of palm width (
<quantity id="4CC5B7EEFFAD4D383A92F8933ECCF8DA" box="[900,1007,1878,1905]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="7" pageNumber="329" unit="mm" value="15.4">15.4 mm</quantity>
); depth of palm 
<quantity id="4CC5B7EEFFAD4D383DBCF8923826F8DA" box="[1194,1285,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="329" unit="mm" value="9.1">9.1 mm</quantity>
; all measurements and counts from right chela. Annulus ventralis as described in diagnosis (
<figureCitation id="1306068EFFAD4D383D2EF8BF39B7F83F" box="[1080,1172,1914,1940]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="7" pageNumber="329">
<emphasis id="B949C619FFAD4D383D2EF8BF39B7F83F" bold="true" box="[1080,1172,1914,1940]" pageId="7" pageNumber="329">Fig. 4H</emphasis>
</figureCitation>
); width of postannular sclerite half total width of annulus ventralis; first pleopods uniramous, reaching central region of annulus ventralis when abdomen flexed.
</paragraph>
<caption id="DF424A83FFA24D373981F8A93CB6F85A" ID-DOI="http://doi.org/10.5281/zenodo.5792767" ID-Zenodo-Dep="5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="8" pageNumber="330" startId="8.[151,250,1900,1924]" targetBox="[218,1369,182,1875]" targetPageId="8">
<paragraph id="8B821A0BFFA24D373981F8A93CB6F85A" blockId="8.[151,1436,1900,2033]" pageId="8" pageNumber="330">
<emphasis id="B949C619FFA24D373981F8A93C2CF82E" bold="true" box="[151,271,1900,1925]" pageId="8" pageNumber="330">FIGURE 4</emphasis>
. 
<taxonomicName id="4C3D6188FFA24D373808F8A83CD6F82F" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[286,501,1901,1925]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="330" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA24D373808F8A83CD6F82F" box="[286,501,1901,1925]" italics="true" pageId="8" pageNumber="330">Cambarus ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA24D3738EBF8A83F73F82E" bold="true" box="[509,592,1901,1925]" pageId="8" pageNumber="330">
<taxonomicNameLabel id="A27A7B62FFA24D3738EBF8A83F73F82E" box="[509,592,1901,1925]" pageId="8" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod. A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195).
</paragraph>
</caption>
<caption id="DF424A83FFA34D363981FB8C3EF9FB2E" ID-DOI="http://doi.org/10.5281/zenodo.5792769" ID-Zenodo-Dep="5792769" httpUri="https://zenodo.org/record/5792769/files/figure.png" pageId="9" pageNumber="331" startId="9.[151,250,1097,1121]" targetBox="[151,1435,181,1072]" targetPageId="9">
<paragraph id="8B821A0BFFA34D363981FB8C3EF9FB2E" blockId="9.[151,1436,1097,1157]" pageId="9" pageNumber="331">
<emphasis id="B949C619FFA34D363981FB8C3C33FBCA" bold="true" box="[151,272,1097,1121]" pageId="9" pageNumber="331">FIGURE 5</emphasis>
. Live color of A) adult male and B) adult female 
<taxonomicName id="4C3D6188FFA34D363A34FB8F3E85FBCA" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[802,934,1098,1121]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="331" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA34D363A34FB8F3E85FBCA" box="[802,934,1098,1121]" italics="true" pageId="9" pageNumber="331">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA34D363AB8FB8C3921FBCA" bold="true" box="[942,1026,1097,1121]" pageId="9" pageNumber="331">
<taxonomicNameLabel id="A27A7B62FFA34D363AB8FB8C3921FBCA" box="[942,1026,1097,1121]" pageId="9" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the Pigeon River, NC. Photos by Michael Perkins and W.T. Russ (North Carolina Wildlife Resources Commission).
</paragraph>
</caption>
<paragraph id="8B821A0BFFA34D3639D1FB773CD3FA37" blockId="9.[151,1437,1202,2029]" pageId="9" pageNumber="331">
<emphasis id="B949C619FFA34D3639D1FB773FCDFB67" bold="true" box="[199,750,1202,1229]" pageId="9" pageNumber="331">
<emphasis id="B949C619FFA34D3639D1FB773CDDFB67" bold="true" box="[199,510,1202,1229]" italics="true" pageId="9" pageNumber="331">Morphotypic male, form II</emphasis>
(
<figureCitation id="1306068EFFA34D363B1BFB773F59FB67" box="[525,634,1202,1228]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="9" pageNumber="331">Fig. 4I–J</figureCitation>
, 
<tableCitation id="C6BF2FB0FFA34D363B91FB773FFDFB67" box="[647,734,1202,1228]" captionStart="TABLE 1" captionStartId="5.[151,239,154,178]" captionTargetPageId="5" captionText="TABLE 1. Morphometric variables compared between C. ectopistes sp. nov. and C. robustus s.s., including mean (x) and standard deviations (s), test statistics (either Wilcoxon rank-sum test, W, or Welch’s t-test, t, with subscript = degrees of freedom), and associated p-values. Statistically significant differences between the two species (P &lt;0.05) are denoted using an asterisk *. Length (L), width (W), depth (D), total carapace length (TCL), carapace width (CW)." httpUri="http://table.plazi.org/id/DF424A83FFAF4D3A3981FF5F39DFFEB5" pageId="9" pageNumber="331" tableUuid="DF424A83FFAF4D3A3981FF5F39DFFEB5">Table 1</tableCitation>
).
</emphasis>
Differing from 
<typeStatus id="5486A4A9FFA34D363AB5FB773926FB67" box="[931,1029,1202,1228]" pageId="9" pageNumber="331" type="holotype">holotype</typeStatus>
in the following respects: TCL 
<quantity id="4CC5B7EEFFA34D363C7DFB773DE0FB44" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="9" pageNumber="331" unit="mm" value="40.6">40.6 mm</quantity>
and PCL 
<quantity id="4CC5B7EEFFA34D363825FB103CB8FB44" box="[307,411,1237,1263]" metricMagnitude="-2" metricUnit="m" metricValue="3.42" pageId="9" pageNumber="331" unit="mm" value="34.2">34.2 mm</quantity>
; areola length 35.2% of TCL (41.8% of PCL), 3.4 times longer than wide; rostrum margins subparallel and thickened; ventrally deflected and moderately excavated, 1.6 times as long as wide; abdomen 
<quantity id="4CC5B7EEFFA34D363C7DFB323DE0FA9E" metricMagnitude="-2" metricUnit="m" metricValue="4.16" pageId="9" pageNumber="331" unit="mm" value="41.6">41.6 mm</quantity>
long; antennal scale 
<quantity id="4CC5B7EEFFA34D3638A6FADF3CDCFA9E" box="[432,511,1306,1333]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="331" unit="mm" value="6.5">6.5mm</quantity>
long, 
<quantity id="4CC5B7EEFFA34D363B50FADF3FBFFA9E" box="[582,668,1306,1333]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="331" unit="mm" value="3.6">3.6 mm</quantity>
wide. Mesial-most row of palm with 7 tubercles; second row with 5 tubercles; palm length (
<quantity id="4CC5B7EEFFA34D36388AFAF83F21FAFC" box="[412,514,1341,1367]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="9" pageNumber="331" unit="mm" value="10.8">10.8 mm</quantity>
) 74.4% of width (
<quantity id="4CC5B7EEFFA34D363BDFFAF83E11FAFC" box="[713,818,1341,1368]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="9" pageNumber="331" unit="mm" value="14.5">14.5 mm</quantity>
); all measurements and counts from right chela. Central projection curved 90° to shaft, with complete apex, rounded (
<figureCitation id="1306068EFFA34D363A5CFA9A3E97FAD2" box="[842,948,1375,1401]" captionStart="FIGURE 4" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[218,1369,182,1875]" captionTargetId="figure-17@8.[217,1370,181,1876]" captionTargetPageId="8" captionText="FIGURE 4. Cambarus ectopistes sp. nov.: (A) lateral cephalothorax; (B) dorsal cephalothorax; (C) epistome; (D) antennal scale; (E) dorsal right chela; (F) mesial form I gonopod; (G) lateral form I gonopod; (H) annulus ventralis; (I) mesial form II gonopod; (J) lateral form II gonopod.A–G from holotype (NCSM 90193); H from allotype (NCSM 90194); and I–J from morphotype (NCSM 90195)." figureDoi="http://doi.org/10.5281/zenodo.5792767" httpUri="https://zenodo.org/record/5792767/files/figure.png" pageId="9" pageNumber="331">
<emphasis id="B949C619FFA34D363A5CFA9A3E97FAD2" bold="true" box="[842,948,1375,1401]" pageId="9" pageNumber="331">Fig. 4I–J</emphasis>
</figureCitation>
); hook on ischium of third pereopod small, not reaching basioischial joint.
</paragraph>
<paragraph id="8B821A0BFFA34D3639D1FA603E9BF98C" blockId="9.[151,1437,1202,2029]" pageId="9" pageNumber="331">
<emphasis id="B949C619FFA34D3639D1FA603DD5FA14" bold="true" box="[199,246,1445,1471]" italics="true" pageId="9" pageNumber="331">Size</emphasis>
Form I male (n = 15) TCL ranges from 
<quantity id="4CC5B7EEFFA34D363BD4FA603E5EFA14" box="[706,893,1445,1472]" metricMagnitude="-2" metricUnit="m" metricValue="4.505" metricValueMax="5.18" metricValueMin="3.8299999999999996" pageId="9" pageNumber="331" unit="mm" value="45.05" valueMax="51.8" valueMin="38.3">38.3 to 51.8 mm</quantity>
(PCL 
<quantity id="4CC5B7EEFFA34D363ADFFA60394DFA14" box="[969,1134,1445,1471]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" metricValueMax="4.22" metricValueMin="2.32" pageId="9" pageNumber="331" unit="mm" value="32.7" valueMax="42.2" valueMin="23.2">23.2–42.2 mm</quantity>
) with a mean TCL of 
<quantity id="4CC5B7EEFFA34D363C7DFA603DE5FA49" metricMagnitude="-2" metricUnit="m" metricValue="4.46" pageId="9" pageNumber="331" unit="mm" value="44.6">44.6 mm</quantity>
. Mean TCL of form II male (n = 26) is 
<quantity id="4CC5B7EEFFA34D363B90FA0D3FCFFA49" box="[646,748,1480,1506]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" pageId="9" pageNumber="331" unit="mm" value="38.4">38.4 mm</quantity>
, ranging in size from 
<quantity id="4CC5B7EEFFA34D363AF2FA0D39BCFA49" box="[996,1183,1480,1506]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.93" metricValueMin="3.07" pageId="9" pageNumber="331" unit="mm" value="40.0" valueMax="49.3" valueMin="30.7">30.7 to 49.3 mm</quantity>
(PCL 
<quantity id="4CC5B7EEFFA34D363DFFFA0D38B3FA49" box="[1257,1424,1480,1506]" metricMagnitude="-2" metricUnit="m" metricValue="3.215" metricValueMax="3.9899999999999998" metricValueMin="2.44" pageId="9" pageNumber="331" unit="mm" value="32.15" valueMax="39.9" valueMin="24.4">24.4–39.9 mm</quantity>
). Female (n = 78) mean TCL is 
<quantity id="4CC5B7EEFFA34D3638F4FA2F3F66F9AE" box="[482,581,1514,1541]" metricMagnitude="-2" metricUnit="m" metricValue="4.33" pageId="9" pageNumber="331" unit="mm" value="43.3">43.3 mm</quantity>
and ranges from 29.0 to 
<quantity id="4CC5B7EEFFA34D363A40FA2F3E9AF9AE" box="[854,953,1514,1541]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="9" pageNumber="331" unit="mm" value="53.6">53.6 mm</quantity>
(PCL 
<quantity id="4CC5B7EEFFA34D363D16FA2F3982F9AE" box="[1024,1185,1514,1541]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" metricValueMax="4.58" metricValueMin="2.38" pageId="9" pageNumber="331" unit="mm" value="34.8" valueMax="45.8" valueMin="23.8">23.8–45.8mm</quantity>
). The largest specimen examined was a form I female with a TCL of 
<quantity id="4CC5B7EEFFA34D363B81F9C83FD8F98C" box="[663,763,1549,1576]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="9" pageNumber="331" unit="mm" value="53.6">53.6 mm</quantity>
(PCL 
<quantity id="4CC5B7EEFFA34D363A52F9C83E8FF98C" box="[836,940,1549,1576]" metricMagnitude="-2" metricUnit="m" metricValue="4.58" pageId="9" pageNumber="331" unit="mm" value="45.8">45.8 mm</quantity>
).
</paragraph>
<paragraph id="8B821A0BFFA34D3539D1F9EA3FC4FEB6" blockId="9.[151,1437,1202,2029]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="332" pageId="9" pageNumber="331">
<emphasis id="B949C619FFA34D3639D1F9EA3C2DF9E1" bold="true" box="[199,270,1583,1610]" italics="true" pageId="9" pageNumber="331">Color.</emphasis>
Carapace ground color of 
<taxonomicName id="4C3D6188FFA34D363B21F9F53FE6F9E2" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[567,709,1584,1610]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="331" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA34D363B21F9F53FE6F9E2" box="[567,709,1584,1610]" italics="true" pageId="9" pageNumber="331">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA34D363BDAF9F53E06F9E1" bold="true" box="[716,805,1584,1610]" pageId="9" pageNumber="331">
<taxonomicNameLabel id="A27A7B62FFA34D363BDAF9F53E06F9E1" box="[716,805,1584,1610]" pageId="9" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1306068EFFA34D363A23F9EA3E54F9E1" box="[821,887,1583,1610]" captionStart="FIGURE 5" captionStartId="9.[151,250,1097,1121]" captionTargetBox="[151,1435,181,1072]" captionTargetId="figure-28@9.[151,1435,181,1073]" captionTargetPageId="9" captionText="FIGURE 5. Live color of A) adult male and B) adult female C. ectopistes sp. nov. from the Pigeon River, NC. Photos by Michael Perkins and W.T. Russ (North Carolina Wildlife Resources Commission)." figureDoi="http://doi.org/10.5281/zenodo.5792769" httpUri="https://zenodo.org/record/5792769/files/figure.png" pageId="9" pageNumber="331">
<emphasis id="B949C619FFA34D363A23F9EA3E54F9E1" bold="true" box="[821,887,1583,1610]" pageId="9" pageNumber="331">Fig. 5</emphasis>
</figureCitation>
) olivaceous brown to brown; posterior margin of carapace same as ground color, with weak saddle. Hepatic and antennal region of carapace with beige punctations. Postorbital ridge maroon or red-brown. Rostrum margins maroon, red-brown, or brown; acumen olivaceous brown. Cephalic section of carapace immediately anterior to and including cervical groove light olive-brown, brown, rarely green; mandibular abductor scars mottled, ranging from light-brown, brown, to dark brown. Lateral margin of antennal scale olive to light brown; body of antennal scale brown to cream. Antennal flagellum and antennules olivaceous, brown, with bluish hue; dorsal surface of lamella tan to brown; ventral surface light green to olivaceous. Dorsal surface of chela generally olivaceous or light brown, with beige punctations. Denticles on opposable surfaces of fingers yellow, white, or tan. Ventral surface of chela cream or tan. Tubercles on mesial margin of palm rust, redbrown, or maroon. Dorsal surface of carpus orange-brown or olivaceous; region adjacent to and including furrow light brown to olive; large mesial spine cream. Merus olive-brown or olive. Podomeres of pereopods olivaceous, blue-brown, or green-brown; joints of pereopod podomeres orange. Dorsal and dorsolateral surface of abdomen heavily mottled and same colors as carapace; tergal margins cream; abdomen lacking dorsal stripe. Uropod same colors as abdomen. Ventral surface of abdomen and carapace cream. Dorsal ridge of form I gonopod central projection amber; body of central projection, gonopod, and mesial process tan. Form II gonopod and all associated processes cream. Cephalic portion of annulus ventralis pink to pink-cream; ridge of fossa pink; caudal region of annulus ventralis ranges from pink to cream colored.
</paragraph>
</subSubSection>
<subSubSection id="C3274980FFA04D3339D1FEE23E2EF986" lastPageId="12" lastPageNumber="334" pageId="10" pageNumber="332" type="materials_examined">
<paragraph id="8B821A0BFFA04D3539D1FEE23870FE06" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="332">
<emphasis id="B949C619FFA04D3539D1FEE23C9BFEEA" bold="true" box="[199,440,294,321]" italics="true" pageId="10" pageNumber="332">Type series localities</emphasis>
<materialsCitation id="3B551056FFA04D3538D7FEE23DECFE22" ID-GBIF-Occurrence="3420008334" collectingDate="2009-10-23" collectorName="Both &amp; C. Dunn" county="Haywood County" location="Hurricane Creek" municipality="The" pageId="10" pageNumber="332" specimenCount="2" stateProvince="North Carolina" typeStatus="holotype">
<collectingMunicipality id="6BE68071FFA04D3538D7FEE23CCEFEEA" box="[449,493,295,321]" pageId="10" pageNumber="332">The</collectingMunicipality>
<typeStatus id="5486A4A9FFA04D3538E0FEE23F7BFEEA" box="[502,600,295,321]" pageId="10" pageNumber="332" type="holotype">holotype</typeStatus>
and 
<typeStatus id="5486A4A9FFA04D353B84FEE23FCFFEEA" box="[658,748,295,321]" pageId="10" pageNumber="332" type="allotype">allotype</typeStatus>
were collected from 
<location id="8EE24CD0FFA04D353AF4FEE239BDFEEA" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353AF4FEE239BDFEEA" box="[994,1182,295,321]" county="Haywood County" municipality="The" name="Hurricane Creek" pageId="10" pageNumber="332" stateProvince="North Carolina">Hurricane Creek</location>
at I-40 west crossing, 
<collectingCounty id="62E36287FFA04D353981FE8E3C42FECE" box="[151,353,331,357]" pageId="10" pageNumber="332">Haywood County</collectingCounty>
, 
<collectingRegion id="49F9D4E9FFA04D353878FE8E3F3FFECE" box="[366,540,331,357]" country="United States of America" name="North Carolina" pageId="10" pageNumber="332">North Carolina</collectingRegion>
. 
<collectorName id="26C87FDDFFA04D353B3FFE8E3F7CFECE" box="[553,607,331,357]" pageId="10" pageNumber="332">Both</collectorName>
specimens were selected from a collection made by 
<collectorName id="26C87FDDFFA04D353DD6FE8E3803FECE" box="[1216,1312,331,357]" pageId="10" pageNumber="332">C. Dunn</collectorName>
on 
<date id="FF833CCBFFA04D353C59FE8E3DECFE22" pageId="10" pageNumber="332" value="2009-10-23">
<collectingDate id="EFC7C523FFA04D353C59FE8E3DECFE22" pageId="10" pageNumber="332" value="2009-10-23">23 Oct 2009</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="3B551056FFA04D3539C1FEAA386CFE06" ID-GBIF-Occurrence="3420008337" collectingDate="2017-06-09" collectorName="B. W. Williams &amp; Z. J. Loughman &amp; E. M. Delekta" county="Unicoi County" location="Tobes Creek" municipality="The" pageId="10" pageNumber="332" specimenCount="1" stateProvince="North Carolina" typeStatus="holotype">
<collectingMethod id="527C621CFFA04D3539C1FEAA3C99FE22" box="[215,442,367,393]" pageId="10" pageNumber="332">using minnow traps</collectingMethod>
. 
<collectingMunicipality id="6BE68071FFA04D3538D0FEAA3CD1FE22" box="[454,498,367,393]" pageId="10" pageNumber="332">The</collectingMunicipality>
morphotype was collected from 
<location id="8EE24CD0FFA04D353A7EFEAA3ED5FE22" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353A7EFEAA3ED5FE22" box="[872,1014,367,393]" county="Unicoi County" municipality="The" name="Tobes Creek" pageId="10" pageNumber="332" stateProvince="North Carolina">Tobes Creek</location>
at the confluence with 
<location id="8EE24CD0FFA04D353C14FEAA38BBFE22" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353C14FEAA38BBFE22" box="[1282,1432,367,393]" county="Unicoi County" municipality="The" name="Pigeon River" pageId="10" pageNumber="332" stateProvince="North Carolina">Pigeon River</location>
, below 
<location id="8EE24CD0FFA04D3539F4FE563CB8FE06" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D3539F4FE563CB8FE06" box="[226,411,403,429]" county="Unicoi County" municipality="The" name="Waterville Road" pageId="10" pageNumber="332" stateProvince="North Carolina">Waterville Road</location>
, 
<collectingCounty id="62E36287FFA04D3538B0FE563F68FE06" box="[422,587,403,429]" pageId="10" pageNumber="332">Unicoi County</collectingCounty>
by 
<collectorName id="26C87FDDFFA04D353B60FE563E3EFE06" box="[630,797,403,429]" pageId="10" pageNumber="332">B.W. Williams</collectorName>
, 
<collectorName id="26C87FDDFFA04D353A3EFE563EF6FE06" box="[808,981,403,429]" pageId="10" pageNumber="332">Z.J. Loughman</collectorName>
, and 
<collectorName id="26C87FDDFFA04D353D06FE563985FE06" box="[1040,1190,403,429]" pageId="10" pageNumber="332">E.M. Delekta</collectorName>
on 
<date id="FF833CCBFFA04D353DC6FE56386CFE06" box="[1232,1359,403,429]" pageId="10" pageNumber="332" value="2017-06-09">
<collectingDate id="EFC7C523FFA04D353DC6FE56386CFE06" box="[1232,1359,403,429]" pageId="10" pageNumber="332" value="2017-06-09">9 Jun 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B821A0BFFA04D3539D1FE72386EFD95" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="332">
<emphasis id="B949C619FFA04D3539D1FE723C8CFE7A" bold="true" box="[199,431,438,465]" italics="true" pageId="10" pageNumber="332">Disposition of types</emphasis>
<materialsCitation id="3B551056FFA04D3538AFFE723869FD95" ID-GBIF-Occurrence="3420008311" collectionCode="NCSM, USNM" county="Museum of Natural Sciences Non-Molluscan Invertebrate Collection" location="Smithsonian National Museum of Natural History Invertebrate Zoology Collection" municipality="Two" pageId="10" pageNumber="332" specimenCode="NCSM 90193, 90194, 90195, USNM 1661660" specimenCount="2" stateProvince="North Carolina" typeStatus="holotype">
The 
<typeStatus id="5486A4A9FFA04D3538F8FE723F70FE7A" box="[494,595,439,465]" pageId="10" pageNumber="332" type="holotype">holotype</typeStatus>
, 
<typeStatus id="5486A4A9FFA04D353B77FE723F9DFE7A" box="[609,702,439,465]" pageId="10" pageNumber="332" type="allotype">allotype</typeStatus>
, morphotype, and 
<specimenCount id="9D3BD182FFA04D353A81FE723939FE7A" box="[919,1050,439,465]" pageId="10" pageNumber="332" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
are deposited in the 
<collectingRegion id="49F9D4E9FFA04D353C07FE723DE2FE5E" country="United States of America" name="North Carolina" pageId="10" pageNumber="332">North Carolina</collectingRegion>
<collectingCounty id="62E36287FFA04D3539DFFE1E3EF1FE5E" box="[201,978,475,501]" pageId="10" pageNumber="332">Museum of Natural Sciences Non-Molluscan Invertebrate Collection</collectingCounty>
(catalog numbers 
<specimenCode id="DB9BB270FFA04D353DB5FE1E3867FE5E" box="[1187,1348,475,501]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 90193</specimenCode>
, 
<specimenCode id="DB9BB270FFA04D353C59FE1E38BBFE5E" box="[1359,1432,475,501]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">90194</specimenCode>
, 
<specimenCode id="DB9BB270FFA04D353981FE3B3DC3FDB2" box="[151,224,510,537]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">90195</specimenCode>
, and 90196; respectively). 
<collectingMunicipality id="6BE68071FFA04D353B1CFE3A3F18FDB2" box="[522,571,511,537]" pageId="10" pageNumber="332">Two</collectingMunicipality>
additional 
<typeStatus id="5486A4A9FFA04D353BAEFE3A3E05FDB2" box="[696,806,511,537]" pageId="10" pageNumber="332" type="paratype">paratypes</typeStatus>
, 1 MI and 1 F, are deposited in the 
<location id="8EE24CD0FFA04D353DBCFE3A3E1DFD96" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353DBCFE3A3E1DFD96" county="Museum of Natural Sciences Non-Molluscan Invertebrate Collection" municipality="Two" name="Smithsonian National Museum of Natural History Invertebrate Zoology Collection" pageId="10" pageNumber="332" stateProvince="North Carolina">Smithsonian National Museum of Natural History Invertebrate Zoology Collection</location>
(
<specimenCode id="DB9BB270FFA04D353A58FDE63929FD96" box="[846,1034,547,573]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="332" type="Museum">USNM 1661660</specimenCode>
, and 1661661, respectively)
</materialsCitation>
.
</paragraph>
<paragraph id="8B821A0BFFA04D3539D1FD82386CFCF6" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="332">
<emphasis id="B949C619FFA04D3539D1FD823E0EFDCB" box="[199,813,582,609]" italics="true" pageId="10" pageNumber="332">
<emphasis id="B949C619FFA04D3539D1FD823F1EFDCA" bold="true" box="[199,573,582,609]" italics="true" pageId="10" pageNumber="332">Range and specimens measured.</emphasis>
<taxonomicName id="4C3D6188FFA04D353B52FD823E0EFDCB" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[580,813,583,609]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="332" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">Cambarus ectopistes</taxonomicName>
</emphasis>
<emphasis id="B949C619FFA04D353A22FD823EAEFDCA" bold="true" box="[820,909,583,609]" pageId="10" pageNumber="332">
<taxonomicNameLabel id="A27A7B62FFA04D353A22FD823EAEFDCA" box="[820,909,583,609]" pageId="10" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occupies a narrow and noncontiguous distribution extending across three major tributaries in the Upper 
<collectingRegion id="49F9D4E9FFA04D353A34FDAE3EB6FD2E" box="[802,917,619,645]" country="United States of America" name="Tennessee" pageId="10" pageNumber="332">Tennessee</collectingRegion>
River basin, namely the Pigeon River, French Broad River, and Nolichucky River (
<figureCitation id="1306068EFFA04D353B25FD4B3FA9FD03" box="[563,650,654,680]" captionStart="FIGURE 6" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[151,1436,185,1909]" captionTargetId="figure-28@11.[151,1436,181,1909]" captionTargetPageId="11" captionText="FIGURE 6. Maps (A. relief, B. HUC 10 boundaries) showing the distribution of C. ectopistes sp. nov. in the Pigeon River (PR), French Broad River (FB), Nolichucky River (NOL), and Cane River (CANE) systems. * Denotes the sampling locality on Paint Creek, Tennessee." figureDoi="http://doi.org/10.5281/zenodo.5792771" httpUri="https://zenodo.org/record/5792771/files/figure.png" pageId="10" pageNumber="332">
<emphasis id="B949C619FFA04D353B25FD4B3FA9FD03" bold="true" box="[563,650,654,680]" pageId="10" pageNumber="332">Fig. 6A</emphasis>
</figureCitation>
). The western, or downstream, limit to the current range of the species approximates the transition between the Blue Ridge Mountain and Ridge and Valley ecoregions. The eastern—upstream—limit is less clearly delimited by ecological or geomorphic boundaries but is tightly constrained by 10-digit Hydrologic Unit Boundaries (Hydrologic Unit Code [HUC] 10 = watersheds) (
<figureCitation id="1306068EFFA04D353D10FD3F3978FCBF" box="[1030,1115,762,788]" captionStart="FIGURE 6" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[151,1436,185,1909]" captionTargetId="figure-28@11.[151,1436,181,1909]" captionTargetPageId="11" captionText="FIGURE 6. Maps (A. relief, B. HUC 10 boundaries) showing the distribution of C. ectopistes sp. nov. in the Pigeon River (PR), French Broad River (FB), Nolichucky River (NOL), and Cane River (CANE) systems. * Denotes the sampling locality on Paint Creek, Tennessee." figureDoi="http://doi.org/10.5281/zenodo.5792771" httpUri="https://zenodo.org/record/5792771/files/figure.png" pageId="10" pageNumber="332">
<emphasis id="B949C619FFA04D353D10FD3F3978FCBF" bold="true" box="[1030,1115,762,788]" pageId="10" pageNumber="332">Fig. 6B</emphasis>
</figureCitation>
). This distribution is characterized via extensive sampling in and adjacent to the Pigeon, French Broad, and Nolichucky River watersheds, and examination of historic specimen lots housed in the 
<collectionCode id="ED2C82CEFFA04D353BC8FC863E0EFCF6" box="[734,813,835,861]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM</collectionCode>
Non-molluscan Invertebrate Collection (
<figureCitation id="1306068EFFA04D353DEAFC87381CFCF7" box="[1276,1343,834,860]" captionStart="FIGURE 1" captionStartId="2.[151,250,1281,1305]" captionTargetBox="[151,1436,181,1257]" captionTargetId="figure-17@2.[151,1436,181,1257]" captionTargetPageId="2" captionText="FIGURE 1. Map showing localities of all specimens examined for this study, including newly sampled material and individuals in the North Carolina Museum of Natural Sciences (NCSM) Non-molluscan Invertebrates Collection originally identified as Cambarus robustus, C. cf. robustus, or C. (Puncticambarus) sp." figureDoi="http://doi.org/10.5281/zenodo.5792761" httpUri="https://zenodo.org/record/5792761/files/figure.png" pageId="10" pageNumber="332">
<emphasis id="B949C619FFA04D353DEAFC87381CFCF7" bold="true" box="[1276,1343,834,860]" pageId="10" pageNumber="332">Fig. 1</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8B821A0BFFA04D3339D1FCA23923FD96" blockId="10.[151,1437,151,2013]" lastBlockId="12.[151,1437,150,2013]" lastPageId="12" lastPageNumber="334" pageId="10" pageNumber="332">
We measured 
<specimenCount id="9D3BD182FFA04D353870FCA23CDDFC2A" box="[358,510,871,897]" pageId="10" pageNumber="332" type="generic">81 specimens</specimenCount>
from 
<specimenCount id="9D3BD182FFA04D353B54FCA23FF3FC2A" box="[578,720,871,897]" pageId="10" pageNumber="332" type="generic">33 specimen</specimenCount>
lots, including the 
<typeStatus id="5486A4A9FFA04D353ABEFCA23EFBFC2A" box="[936,984,871,897]" pageId="10" pageNumber="332">type</typeStatus>
series and additional material housed in the 
<collectingRegion id="49F9D4E9FFA04D3539D7FC4E3C48FC0E" box="[193,363,907,933]" country="United States of America" name="North Carolina" pageId="10" pageNumber="332">North Carolina</collectingRegion>
Museum of Natural Sciences Non-Molluscan Invertebrate Collection, West Liberty University Crayfish Conservation Lab. Only adult specimens are reported below. This material is from the following locations: 
<collectingRegion id="49F9D4E9FFA04D353981FC163CB1FC46" box="[151,402,979,1005]" country="United States of America" name="North Carolina" pageId="10" pageNumber="332">NORTH CAROLINA</collectingRegion>
, 
<emphasis id="B949C619FFA04D353889FC163F40FC46" box="[415,611,979,1005]" italics="true" pageId="10" pageNumber="332">Haywood County</emphasis>
: (1) 
<materialsCitation id="3B551056FFA04D353B8BFC163E10FBBA" ID-GBIF-Occurrence="3420008304" collectingDate="2009-06-22" collectionCode="NCSM" collectorName="C. Dunn" location="Cold Springs Creek" municipality="Cold Springs Creek" pageId="10" pageNumber="332" specimenCode="NCSM 90197" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353B8BFC163E19FC46" box="[669,826,979,1005]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 90197</specimenCode>
<collectingMunicipality id="6BE68071FFA04D353A55FC163900FC46" box="[835,1059,979,1005]" pageId="10" pageNumber="332">Cold Springs Creek</collectingMunicipality>
along 
<location id="8EE24CD0FFA04D353D64FC163871FC46" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353D64FC163871FC46" box="[1138,1362,979,1005]" municipality="Cold Springs Creek" name="Cold Springs Creek" pageId="10" pageNumber="332">Cold Springs Creek</location>
Road, 
<locationDeviation id="9A300329FFA04D353981FC323C5BFBBA" box="[151,376,1015,1041]" location="Fines Creek" pageId="10" pageNumber="332">NW of Fines Creek</locationDeviation>
, 1 F, 1 MII, 
<collectorName id="26C87FDDFFA04D353B1EFC323F48FBBA" box="[520,619,1015,1041]" pageId="10" pageNumber="332">C. Dunn</collectorName>
, 
<date id="FF833CCBFFA04D353B61FC323E26FBBA" box="[631,773,1015,1041]" pageId="10" pageNumber="332" value="2009-06-22">
<collectingDate id="EFC7C523FFA04D353B61FC323E26FBBA" box="[631,773,1015,1041]" pageId="10" pageNumber="332" value="2009-06-22">22 Jun 2009</collectingDate>
</date>
; (2)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353A2DFC323E2AFB9E" ID-GBIF-Occurrence="3420008303" collectingDate="2009-06-08" collectionCode="NCSM" collectorName="C. Dunn" location="Cold Springs Creek" municipality="Cold Springs Creek" pageId="10" pageNumber="332" specimenCode="NCSM 5895" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353A2DFC323EEDFBBA" box="[827,974,1014,1041]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 5895</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353ACCFC32399AFBBA" box="[986,1209,1015,1041]" pageId="10" pageNumber="332">Cold Springs Creek</collectingMunicipality>
along 
<location id="8EE24CD0FFA04D353C1EFC323DF9FB9E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353C1EFC323DF9FB9E" municipality="Cold Springs Creek" name="Cold Springs Creek" pageId="10" pageNumber="332">Cold Springs Creek</location>
Road, 
<locationDeviation id="9A300329FFA04D35383BFBDE3F2EFB9E" box="[301,525,1051,1077]" location="Fines Creek" pageId="10" pageNumber="332">NW of Fines Creek</locationDeviation>
, 
<collectorName id="26C87FDDFFA04D353B0FFBDE3F5FFB9E" box="[537,636,1051,1077]" pageId="10" pageNumber="332">C. Dunn</collectorName>
, 
<date id="FF833CCBFFA04D353B9FFBDE3E2AFB9E" box="[649,777,1051,1077]" pageId="10" pageNumber="332" value="2009-06-08">
<collectingDate id="EFC7C523FFA04D353B9FFBDE3E2AFB9E" box="[649,777,1051,1077]" pageId="10" pageNumber="332" value="2009-06-08">8 Jun 2009</collectingDate>
</date>
</materialsCitation>
, 1 MI, 1 MII, 1 F; (3) 
<materialsCitation id="3B551056FFA04D353D1AFBDE3E8FFBF2" ID-GBIF-Occurrence="3420008301" collectingDate="2009-10-23" collectionCode="NCSM" collectorName="F, C. Dunn" location="Hurricane Creek" pageId="10" pageNumber="332" specimenCode="NCSM 90198" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353D1AFBDE398EFB9E" box="[1036,1197,1051,1077]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 90198</specimenCode>
, 
<location id="8EE24CD0FFA04D353DAFFBDE3857FB9E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353DAFFBDE3857FB9E" box="[1209,1396,1051,1077]" name="Hurricane Creek" pageId="10" pageNumber="332">Hurricane Creek</location>
off I-40, 
<locationDeviation id="9A300329FFA04D3539C2FBFA3C91FBF2" box="[212,434,1087,1113]" location="Fines Creek" pageId="10" pageNumber="332">NW of Fines Creek</locationDeviation>
, 3 MI, 1 MII, 6 
<collectorName id="26C87FDDFFA04D353B71FBFA3FC6FBF2" box="[615,741,1087,1113]" pageId="10" pageNumber="332">F, C. Dunn</collectorName>
, 
<date id="FF833CCBFFA04D353BE6FBFA3E5DFBF2" box="[752,894,1087,1113]" pageId="10" pageNumber="332" value="2009-10-23">
<collectingDate id="EFC7C523FFA04D353BE6FBFA3E5DFBF2" box="[752,894,1087,1113]" pageId="10" pageNumber="332" value="2009-10-23">23 Oct 2009</collectingDate>
</date>
; (4)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353AA4FBFA3FCAFBD6" ID-GBIF-Occurrence="3420008331" collectingDate="2009-01-15" collectionCode="NCSM" collectorName="Waterville, C &amp; Dunn" location="Pigeon River" municipality="Big Creek" pageId="10" pageNumber="332" specimenCode="NCSM 90199" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353AA4FBFA3971FBF2" box="[946,1106,1087,1113]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 90199</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353D4BFBFA39F3FBF2" box="[1117,1232,1087,1113]" pageId="10" pageNumber="332">Big Creek</collectingMunicipality>
above confluence with 
<location id="8EE24CD0FFA04D3539C6FBA63C46FBD6" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D3539C6FBA63C46FBD6" box="[208,357,1123,1149]" municipality="Big Creek" name="Pigeon River" pageId="10" pageNumber="332">Pigeon River</location>
, 
<collectorName id="26C87FDDFFA04D353879FBA63F24FBD6" box="[367,519,1123,1149]" pageId="10" pageNumber="332">Waterville, C</collectorName>
. 
<collectorName id="26C87FDDFFA04D353B04FBA63F70FBD6" box="[530,595,1123,1149]" pageId="10" pageNumber="332">Dunn</collectorName>
, 
<date id="FF833CCBFFA04D353B48FBA73FCAFBD6" box="[606,745,1122,1149]" pageId="10" pageNumber="332" value="2009-01-15">
<collectingDate id="EFC7C523FFA04D353B48FBA73FCAFBD6" box="[606,745,1122,1149]" pageId="10" pageNumber="332" value="2009-01-15">15 Jan 2009</collectingDate>
</date>
</materialsCitation>
, 4 MII; (5) 
<materialsCitation id="3B551056FFA04D353A7DFBA63FCAFB09" ID-GBIF-Occurrence="3420008344" collectingDate="2009-06-18" collectionCode="NCSM" collectorName="F, C. Dunn" county="Big Creek" location="Waterville" municipality="Waterville Road" pageId="10" pageNumber="332" specimenCode="NCSM 5884" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353A7DFBA63EDFFBD6" box="[875,1020,1122,1149]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 5884</specimenCode>
, 
<collectingCounty id="62E36287FFA04D353D11FBA63959FBD6" box="[1031,1146,1123,1149]" pageId="10" pageNumber="332">Big Creek</collectingCounty>
at 
<collectingMunicipality id="6BE68071FFA04D353D8DFBA63877FBD6" box="[1179,1364,1123,1149]" pageId="10" pageNumber="332">Waterville Road</collectingMunicipality>
, south of 
<location id="8EE24CD0FFA04D3539A0FB423C0EFB0A" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D3539A0FB423C0EFB0A" box="[182,301,1159,1185]" county="Big Creek" municipality="Waterville Road" name="Waterville" pageId="10" pageNumber="332">Waterville</location>
, 1 MII, 1 
<collectorName id="26C87FDDFFA04D3538B7FB423F03FB0A" box="[417,544,1159,1185]" pageId="10" pageNumber="332">F, C. Dunn</collectorName>
, 
<date id="FF833CCBFFA04D353B3BFB423F98FB0A" box="[557,699,1159,1185]" pageId="10" pageNumber="332" value="2009-06-18">
<collectingDate id="EFC7C523FFA04D353B3BFB423F98FB0A" box="[557,699,1159,1185]" pageId="10" pageNumber="332" value="2009-06-18">18 Jun 2009</collectingDate>
</date>
; (6)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353BE7FB423E7FFB6E" ID-GBIF-Occurrence="3420008308" collectingDate="2009-10-11" collectionCode="NCSM" collectorName="C. Dunn" location="Walters Lake" municipality="Cataloochee Creek" pageId="10" pageNumber="332" specimenCode="NCSM 5882" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353BE7FB423EA7FB0A" box="[753,900,1158,1185]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 5882</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353A86FB423945FB0A" box="[912,1126,1159,1185]" pageId="10" pageNumber="332">Cataloochee Creek</collectingMunicipality>
at confluence with 
<location id="8EE24CD0FFA04D353C51FB423DF1FB6E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353C51FB423DF1FB6E" municipality="Cataloochee Creek" name="Walters Lake" pageId="10" pageNumber="332">Walters Lake</location>
, 
<locationDeviation id="9A300329FFA04D3539C8FB6E3CF1FB6E" box="[222,466,1195,1221]" location="Cove Creek" pageId="10" pageNumber="332">NNW of Cove Creek</locationDeviation>
, 2 MII, 
<collectorName id="26C87FDDFFA04D353B26FB6E3FB0FB6E" box="[560,659,1195,1221]" pageId="10" pageNumber="332">C. Dunn</collectorName>
, 
<date id="FF833CCBFFA04D353B89FB6E3E0DFB6E" box="[671,814,1195,1221]" pageId="10" pageNumber="332" value="2009-10-11">
<collectingDate id="EFC7C523FFA04D353B89FB6E3E0DFB6E" box="[671,814,1195,1221]" pageId="10" pageNumber="332" value="2009-10-11">11 Oct 2009</collectingDate>
</date>
; (7)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353A73FB6E3823FB42" ID-GBIF-Occurrence="3420008341" collectingDate="1997-07-23" collectionCode="NCSM" collectorName="F, D. Lenat, L &amp; Eaton, B &amp; Tracy" location="Old Cataloochee Turnpike" municipality="Cataloochee Creek" pageId="10" pageNumber="332" specimenCode="NCSM 4335" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353A73FB6E3ED4FB6E" box="[869,1015,1194,1221]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 4335</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353D12FB6E39FAFB6E" box="[1028,1241,1195,1221]" pageId="10" pageNumber="332">Cataloochee Creek</collectingMunicipality>
at SR 1395 (
<location id="8EE24CD0FFA04D353C67FB6E3CB1FB41" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353C67FB6E3CB1FB41" municipality="Cataloochee Creek" name="Old Cataloochee Turnpike" pageId="10" pageNumber="332">Old Cataloochee Turnpike</location>
), 
<locationDeviation id="9A300329FFA04D3538B2FB0A3F5CFB42" box="[420,639,1231,1257]" location="Cove Creek" pageId="10" pageNumber="332">NW of Cove Creek</locationDeviation>
, 2 MII, 2 
<collectorName id="26C87FDDFFA04D353BFAFB0A3EA9FB42" box="[748,906,1231,1257]" pageId="10" pageNumber="332">F, D. Lenat, L</collectorName>
. 
<collectorName id="26C87FDDFFA04D353A83FB0A3ED9FB42" box="[917,1018,1231,1257]" pageId="10" pageNumber="332">Eaton, B</collectorName>
. 
<collectorName id="26C87FDDFFA04D353D15FB0A3967FB42" box="[1027,1092,1231,1257]" pageId="10" pageNumber="332">Tracy</collectorName>
, 
<date id="FF833CCBFFA04D353D46FB0A39F6FB42" box="[1104,1237,1231,1257]" pageId="10" pageNumber="332" value="1997-07-23">
<collectingDate id="EFC7C523FFA04D353D46FB0A39F6FB42" box="[1104,1237,1231,1257]" pageId="10" pageNumber="332" value="1997-07-23">23 Jul 1997</collectingDate>
</date>
; (8)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353C11FB0A3870FAA6" ID-GBIF-Occurrence="3420008343" collectingDate="2001-10-10" collectionCode="NCSM" collectorName="F, A. H. Fullerton, J &amp; Smith" location="Cataloochee Entrance Road" municipality="Cataloochee Creek" pageId="10" pageNumber="332" specimenCode="NCSM 7981" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353C11FB0A38BBFB42" box="[1287,1432,1231,1257]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 7981</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353981FB363C4DFAA6" box="[151,366,1267,1293]" pageId="10" pageNumber="332">Cataloochee Creek</collectingMunicipality>
at 
<location id="8EE24CD0FFA04D353880FB363FF6FAA6" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353880FB363FF6FAA6" box="[406,725,1267,1293]" municipality="Cataloochee Creek" name="Cataloochee Entrance Road" pageId="10" pageNumber="332">Cataloochee Entrance Road</location>
, 1 MII, 3 
<collectorName id="26C87FDDFFA04D353A58FB363913FAA6" box="[846,1072,1267,1293]" pageId="10" pageNumber="332">F, A.H. Fullerton, J</collectorName>
. 
<collectorName id="26C87FDDFFA04D353D28FB3639A7FAA6" box="[1086,1156,1267,1293]" pageId="10" pageNumber="332">Smith</collectorName>
, 
<date id="FF833CCBFFA04D353D87FB363807FAA6" box="[1169,1316,1267,1293]" pageId="10" pageNumber="332" value="2001-10-10">
<collectingDate id="EFC7C523FFA04D353D87FB363807FAA6" box="[1169,1316,1267,1293]" pageId="10" pageNumber="332" value="2001-10-10">10 Oct 2001</collectingDate>
</date>
; (9)
</materialsCitation>
WLU 
<materialsCitation id="3B551056FFA04D353981FAD23FC0FAFE" ID-GBIF-Occurrence="3420008345" collectingDate="2016-11-16" collectionCode="XXXXX" collectorName="F, Z. J. Loughman, B. W &amp; Williams" county="Cataloochee Creek" location="Great Smoky Mountains National Park" municipality="Jarvis Palmer Barn" pageId="10" pageNumber="332" specimenCount="1">
<collectionCode id="ED2C82CEFFA04D353981FAD23DDFFA9A" box="[151,252,1303,1329]" pageId="10" pageNumber="332">XXXXX</collectionCode>
<collectingCounty id="62E36287FFA04D353814FAD23CF6FA9A" box="[258,469,1303,1329]" pageId="10" pageNumber="332">Cataloochee Creek</collectingCounty>
across from 
<collectingMunicipality id="6BE68071FFA04D353B75FAD23E1AFA9A" box="[611,825,1303,1329]" pageId="10" pageNumber="332">Jarvis Palmer Barn</collectingMunicipality>
, 
<location id="8EE24CD0FFA04D353A55FAD239D4FA9A" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353A55FAD239D4FA9A" box="[835,1271,1303,1329]" county="Cataloochee Creek" municipality="Jarvis Palmer Barn" name="Great Smoky Mountains National Park" pageId="10" pageNumber="332">Great Smoky Mountains National Park</location>
, 2 MI, 2 
<collectorName id="26C87FDDFFA04D353C4EFAD23C70FAFE" pageId="10" pageNumber="332">F, Z.J. Loughman, B.W</collectorName>
. 
<collectorName id="26C87FDDFFA04D353848FAFE3CE7FAFE" box="[350,452,1339,1365]" pageId="10" pageNumber="332">Williams</collectorName>
<emphasis id="B949C619FFA04D3538DDFAFE3CDEFAFE" box="[459,509,1339,1365]" italics="true" pageId="10" pageNumber="332">et al</emphasis>
., 
<date id="FF833CCBFFA04D353B04FAFE3F8AFAFE" box="[530,681,1339,1365]" pageId="10" pageNumber="332" value="2016-11-16">
<collectingDate id="EFC7C523FFA04D353B04FAFE3F8AFAFE" box="[530,681,1339,1365]" pageId="10" pageNumber="332" value="2016-11-16">16 Nov 2016</collectingDate>
</date>
; (10)
</materialsCitation>
WLU 
<materialsCitation id="3B551056FFA04D353A27FAFE3DE6FA36" ID-GBIF-Occurrence="3420008321" collectingDate="2016-11-16" collectionCode="XXXXX" collectorName="F, Z. J. Loughman, B. W &amp; Williams" county="Cataloochee Creek" location="Great Smoky Mountains National Park" municipality="Jarvis Palmer Barn" pageId="10" pageNumber="332" specimenCount="1">
<collectionCode id="ED2C82CEFFA04D353A27FAFE3EB5FAFE" box="[817,918,1339,1365]" pageId="10" pageNumber="332">XXXXX</collectionCode>
<collectingCounty id="62E36287FFA04D353A8BFAFE3951FAFE" box="[925,1138,1339,1365]" pageId="10" pageNumber="332">Cataloochee Creek</collectingCounty>
across from 
<collectingMunicipality id="6BE68071FFA04D353C12FAFE3DF3FAD2" pageId="10" pageNumber="332">Jarvis Palmer Barn</collectingMunicipality>
, 
<location id="8EE24CD0FFA04D3539CDFA9A3FB2FAD2" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D3539CDFA9A3FB2FAD2" box="[219,657,1375,1401]" county="Cataloochee Creek" municipality="Jarvis Palmer Barn" name="Great Smoky Mountains National Park" pageId="10" pageNumber="332">Great Smoky Mountains National Park</location>
, 2 MI, 1 MII, 2 
<collectorName id="26C87FDDFFA04D353A54FA9A396BFAD2" box="[834,1096,1375,1401]" pageId="10" pageNumber="332">F, Z.J. Loughman, B.W</collectorName>
. 
<collectorName id="26C87FDDFFA04D353D44FA9A399BFAD2" box="[1106,1208,1375,1401]" pageId="10" pageNumber="332">Williams</collectorName>
<emphasis id="B949C619FFA04D353DA8FA9A39CCFAD2" box="[1214,1263,1375,1401]" italics="true" pageId="10" pageNumber="332">et al</emphasis>
., 
<date id="FF833CCBFFA04D353C15FA9A38BBFAD2" box="[1283,1432,1375,1401]" pageId="10" pageNumber="332" value="2016-11-16">
<collectingDate id="EFC7C523FFA04D353C15FA9A38BBFAD2" box="[1283,1432,1375,1401]" pageId="10" pageNumber="332" value="2016-11-16">16 Nov 2016</collectingDate>
</date>
; (11)
</materialsCitation>
<materialsCitation id="3B551056FFA04D3539DAFA463C0CFA6A" ID-GBIF-Occurrence="3420008320" collectingDate="2016-11-17" collectionCode="NCSM" collectorName="F, B. W. Williams" location="Forest Service Road" municipality="Pigeon River" pageId="10" pageNumber="332" specimenCode="NCSM 90191" specimenCount="1">
<specimenCode id="DB9BB270FFA04D3539DAFA463C4FFA36" box="[204,364,1411,1437]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 90191</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353861FA463F29FA36" box="[375,522,1411,1437]" pageId="10" pageNumber="332">Pigeon River</collectingMunicipality>
at 
<location id="8EE24CD0FFA04D353B3BFA463E32FA36" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353B3BFA463E32FA36" box="[557,785,1411,1437]" municipality="Pigeon River" name="Forest Service Road" pageId="10" pageNumber="332">Forest Service Road</location>
288, 
<locationDeviation id="9A300329FFA04D353A47FA46390CFA36" box="[849,1071,1411,1437]" location="Fines Creek" pageId="10" pageNumber="332">NW of Fines Creek</locationDeviation>
, 1 MI, 2 
<collectorName id="26C87FDDFFA04D353D83FA463875FA36" box="[1173,1366,1411,1437]" pageId="10" pageNumber="332">F, B.W. Williams</collectorName>
<emphasis id="B949C619FFA04D353C4BFA4638ADFA36" box="[1373,1422,1411,1437]" italics="true" pageId="10" pageNumber="332">et al</emphasis>
., 
<date id="FF833CCBFFA04D353981FA623C0CFA6A" box="[151,303,1447,1473]" pageId="10" pageNumber="332" value="2016-11-17">
<collectingDate id="EFC7C523FFA04D353981FA623C0CFA6A" box="[151,303,1447,1473]" pageId="10" pageNumber="332" value="2016-11-17">17 Nov 2016</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B551056FFA04D35382DFA623F15FA6A" ID-GBIF-Occurrence="3420008323" box="[315,566,1447,1473]" county="Madison County" location="Madison County" pageId="10" pageNumber="332" specimenCount="1">
<collectingCounty id="62E36287FFA04D35382DFA623CDFFA6A" box="[315,508,1447,1473]" pageId="10" pageNumber="332">Madison County</collectingCounty>
: (12)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353B28FA623CE0FA4E" ID-GBIF-Occurrence="3420008336" collectingDate="2006-09-18" collectionCode="NCSM" collectorName="F, E. Fleek, W &amp; Crouch" location="Big Laurel Road" municipality="Little Creek" pageId="10" pageNumber="332" specimenCode="NCSM 25667" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353B28FA623FFDFA6A" box="[574,734,1446,1473]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 25667</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353BFCFA623E51FA6A" box="[746,882,1447,1473]" pageId="10" pageNumber="332">Little Creek</collectingMunicipality>
at SR 1318 (
<location id="8EE24CD0FFA04D353D1EFA6239E9FA69" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353D1EFA6239E9FA69" box="[1032,1226,1447,1474]" municipality="Little Creek" name="Big Laurel Road" pageId="10" pageNumber="332">Big Laurel Road</location>
), 1 
<collectorName id="26C87FDDFFA04D353DE2FA6238BAFA6A" box="[1268,1433,1447,1473]" pageId="10" pageNumber="332">F, E. Fleek, W</collectorName>
. 
<collectorName id="26C87FDDFFA04D353981FA0E3DCFFA4E" box="[151,236,1483,1509]" pageId="10" pageNumber="332">Crouch</collectorName>
, 
<date id="FF833CCBFFA04D3539EEFA0E3CAAFA4E" box="[248,393,1483,1509]" pageId="10" pageNumber="332" value="2006-09-18">
<collectingDate id="EFC7C523FFA04D3539EEFA0E3CAAFA4E" box="[248,393,1483,1509]" pageId="10" pageNumber="332" value="2006-09-18">18 Sep 2006</collectingDate>
</date>
; (13)
</materialsCitation>
<materialsCitation id="3B551056FFA04D3538DCFA0E3E61F9A2" ID-GBIF-Occurrence="3420008324" collectingDate="1984-09-29" collectionCode="NCSM" collectorName="F, A. L. Braswell, J. E &amp; Cooper" location="Spring Creek" pageId="10" pageNumber="332" specimenCode="NCSM 2130" specimenCount="1">
<specimenCode id="DB9BB270FFA04D3538DCFA0E3F7FFA4E" box="[458,604,1483,1509]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 2130</specimenCode>
, 
<location id="8EE24CD0FFA04D353B7EFA0E3FDEFA4E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353B7EFA0E3FDEFA4E" box="[616,765,1483,1509]" name="Spring Creek" pageId="10" pageNumber="332">Spring Creek</location>
along NC 209, 
<quantity id="4CC5B7EEFFA04D353AA7FA0E393BFA4E" box="[945,1048,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="4.828032" pageId="10" pageNumber="332" unit="mi" value="0.3">0.3 miles</quantity>
south of junction with SR 1166, 1 MII, 1 F, 1 
<collectorName id="26C87FDDFFA04D35380CFA2A3F2AF9A2" box="[282,521,1519,1545]" pageId="10" pageNumber="332">F, A.L. Braswell, J.E</collectorName>
. 
<collectorName id="26C87FDDFFA04D353B03FA2A3F4AF9A2" box="[533,617,1519,1545]" pageId="10" pageNumber="332">Cooper</collectorName>
, 
<date id="FF833CCBFFA04D353B60FA2A3E24F9A2" box="[630,775,1519,1545]" pageId="10" pageNumber="332" value="1984-09-29">
<collectingDate id="EFC7C523FFA04D353B60FA2A3E24F9A2" box="[630,775,1519,1545]" pageId="10" pageNumber="332" value="1984-09-29">29 Sep 1984</collectingDate>
</date>
; (14)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353A5CFA2A3982F986" ID-GBIF-Occurrence="3420008316" collectingDate="1984-09-29" collectionCode="NCSM" collectorName="F, A. L. Braswell, J. E &amp; Cooper" location="Spring Creek" pageId="10" pageNumber="332" specimenCode="NCSM 2133" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353A5CFA2A3EFEF9A2" box="[842,989,1519,1545]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 2133</specimenCode>
, 
<location id="8EE24CD0FFA04D353AFFFA2A395DF9A2" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353AFFFA2A395DF9A2" box="[1001,1150,1519,1545]" name="Spring Creek" pageId="10" pageNumber="332">Spring Creek</location>
along NC 209, 
<quantity id="4CC5B7EEFFA04D353C23FA2A38BEF9A2" box="[1333,1437,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="4.828032" pageId="10" pageNumber="332" unit="mi" value="0.3">0.3 miles</quantity>
south of junction with SR 1166, 1 MI, 2 
<collectorName id="26C87FDDFFA04D353B67F9D63E40F986" box="[625,867,1555,1581]" pageId="10" pageNumber="332">F, A.L. Braswell, J.E</collectorName>
. 
<collectorName id="26C87FDDFFA04D353A66F9D63EE7F986" box="[880,964,1555,1581]" pageId="10" pageNumber="332">Cooper</collectorName>
, 
<date id="FF833CCBFFA04D353AC7F9D63946F986" box="[977,1125,1555,1581]" pageId="10" pageNumber="332" value="1984-09-29">
<collectingDate id="EFC7C523FFA04D353AC7F9D63946F986" box="[977,1125,1555,1581]" pageId="10" pageNumber="332" value="1984-09-29">29 Sep 1984</collectingDate>
</date>
; (15)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353DBCF9D63F28F9DD" ID-GBIF-Occurrence="3420008329" collectingDate="1984-09-25" collectionCode="NCSM" collectorName="J. E. Cooper &amp; A. L. Braswell" country="United States of America" location="Hot Springs" municipality="Shut-in Creek" pageId="10" pageNumber="332" specimenCode="NCSM 2118" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353DBCF9D6381EF986" box="[1194,1341,1555,1581]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 2118</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353C5CF9D63DF9F9FA" pageId="10" pageNumber="332">Shut-in Creek</collectingMunicipality>
along SR 1183, 1.3 road miles south of junction with 
<collectingCountry id="F32A5A9BFFA04D353A25F9F23E74F9FA" box="[819,855,1591,1617]" name="United States of America" pageId="10" pageNumber="332">US</collectingCountry>
25/70, west of 
<location id="8EE24CD0FFA04D353D15F9F239A8F9FA" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353D15F9F239A8F9FA" box="[1027,1163,1591,1617]" country="United States of America" municipality="Shut-in Creek" name="Hot Springs" pageId="10" pageNumber="332">Hot Springs</location>
, 1 F, 2 MII, 
<collectorName id="26C87FDDFFA04D353C05F9F238BBF9FA" box="[1299,1432,1591,1617]" pageId="10" pageNumber="332">J.E. Cooper</collectorName>
, 
<collectorName id="26C87FDDFFA04D353981F99E3C14F9DE" box="[151,311,1627,1653]" pageId="10" pageNumber="332">A.L. Braswell</collectorName>
, 
<date id="FF833CCBFFA04D353854F99F3CF2F9DE" box="[322,465,1626,1653]" pageId="10" pageNumber="332" value="1984-09-25">
<collectingDate id="EFC7C523FFA04D353854F99F3CF2F9DE" box="[322,465,1626,1653]" pageId="10" pageNumber="332" value="1984-09-25">25 Sep 1984</collectingDate>
</date>
; (16)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353B07F99E397BF932" ID-GBIF-Occurrence="3420008312" collectingDate="1984-09-25" collectionCode="NCSM" collectorName="J. E. Cooper &amp; A. L. Braswell" country="United States of America" location="Hot Springs" municipality="Shut-in Creek" pageId="10" pageNumber="332" specimenCode="NCSM 2119" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353B07F99E3F81F9DE" box="[529,674,1627,1653]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 2119</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353BBAF99E3E6BF9DE" box="[684,840,1627,1653]" pageId="10" pageNumber="332">Shut-in Creek</collectingMunicipality>
along SR 1183, 1.3 road miles south of junction with 
<collectingCountry id="F32A5A9BFFA04D353981F9BA3D98F932" box="[151,187,1663,1689]" name="United States of America" pageId="10" pageNumber="332">US</collectingCountry>
25/70, west of 
<location id="8EE24CD0FFA04D35387BF9BA3CD4F932" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D35387BF9BA3CD4F932" box="[365,503,1663,1689]" country="United States of America" municipality="Shut-in Creek" name="Hot Springs" pageId="10" pageNumber="332">Hot Springs</location>
, 1 MI, 
<collectorName id="26C87FDDFFA04D353B5AF9BA3FF1F932" box="[588,722,1663,1689]" pageId="10" pageNumber="332">J.E. Cooper</collectorName>
, 
<collectorName id="26C87FDDFFA04D353BCBF9BA3E5DF932" box="[733,894,1663,1689]" pageId="10" pageNumber="332">A.L. Braswell</collectorName>
, 
<date id="FF833CCBFFA04D353A9DF9BB393FF932" box="[907,1052,1662,1689]" pageId="10" pageNumber="332" value="1984-09-25">
<collectingDate id="EFC7C523FFA04D353A9DF9BB393FF932" box="[907,1052,1662,1689]" pageId="10" pageNumber="332" value="1984-09-25">25 Sep 1984</collectingDate>
</date>
; (17)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353D76F9BA3C9EF94A" ID-GBIF-Occurrence="3420008328" collectingDate="1984-09-25" collectionCode="NCSM" collectorName="A. L. Braswell &amp; J. E. Cooper" country="United States of America" location="Hot Springs" municipality="Shut-in Creek" pageId="10" pageNumber="332" specimenCode="NCSM 2215" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353D76F9BA39D0F932" box="[1120,1267,1662,1689]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 2215</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353DE9F9BA38BFF932" box="[1279,1436,1663,1689]" pageId="10" pageNumber="332">Shut-in Creek</collectingMunicipality>
along SR 1183, 1.3 road miles south of junction with 
<collectingCountry id="F32A5A9BFFA04D353BF9F9663E30F916" box="[751,787,1699,1725]" name="United States of America" pageId="10" pageNumber="332">US</collectingCountry>
25/70, west of 
<location id="8EE24CD0FFA04D353AD7F9663969F916" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353AD7F9663969F916" box="[961,1098,1699,1725]" country="United States of America" municipality="Shut-in Creek" name="Hot Springs" pageId="10" pageNumber="332">Hot Springs</location>
, 1 F ovig., 
<collectorName id="26C87FDDFFA04D353DD0F9663844F916" box="[1222,1383,1699,1725]" pageId="10" pageNumber="332">A.L. Braswell</collectorName>
, 
<collectorName id="26C87FDDFFA04D353C64F9663DC8F94A" pageId="10" pageNumber="332">J.E. Cooper</collectorName>
, 
<date id="FF833CCBFFA04D3539E0F9033CA0F94A" box="[246,387,1734,1761]" pageId="10" pageNumber="332" value="1984-09-25">
<collectingDate id="EFC7C523FFA04D3539E0F9033CA0F94A" box="[246,387,1734,1761]" pageId="10" pageNumber="332" value="1984-09-25">25 Sep 1984</collectingDate>
</date>
; (18)
</materialsCitation>
<materialsCitation id="3B551056FFA04D3538D5F9023FE2F8AE" ID-GBIF-Occurrence="3420008317" collectingDate="1984-07-23" collectionCode="NCSM" collectorName="F, A. L. Braswell, J. E &amp; Cooper" country="United States of America" location="Lower Shutin Road" municipality="Shut-in Creek" pageId="10" pageNumber="332" specimenCode="NCSM 1805" specimenCount="1">
<specimenCode id="DB9BB270FFA04D3538D5F9023F70F94A" box="[451,595,1734,1761]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 1805</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353B48F9023FDAF94A" box="[606,761,1735,1761]" pageId="10" pageNumber="332">Shut-in Creek</collectingMunicipality>
along 
<location id="8EE24CD0FFA04D353A52F902393EF94A" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353A52F902393EF94A" box="[836,1053,1735,1761]" country="United States of America" municipality="Shut-in Creek" name="Lower Shutin Road" pageId="10" pageNumber="332">Lower Shutin Road</location>
(SR 1303), 
<locationDeviation id="9A300329FFA04D353DB5F90238BBF94A" box="[1187,1432,1735,1761]" location="Hot Springs" pageId="10" pageNumber="332">WNW of Hot Springs</locationDeviation>
, 2 
<collectorName id="26C87FDDFFA04D3539BAF92E3CB4F8AE" box="[172,407,1771,1797]" pageId="10" pageNumber="332">F, A.L. Braswell, J.E</collectorName>
. 
<collectorName id="26C87FDDFFA04D3538B4F92E3CD5F8AE" box="[418,502,1771,1797]" pageId="10" pageNumber="332">Cooper</collectorName>
, 
<date id="FF833CCBFFA04D353B17F92E3FA5F8AE" box="[513,646,1771,1797]" pageId="10" pageNumber="332" value="1984-07-23">
<collectingDate id="EFC7C523FFA04D353B17F92E3FA5F8AE" box="[513,646,1771,1797]" pageId="10" pageNumber="332" value="1984-07-23">23 Jul 1984</collectingDate>
</date>
; (19)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353BDEF92E39DAF882" ID-GBIF-Occurrence="3420008302" collectingDate="1984-09-25" collectionCode="NCSM" collectorName="J. E. Cooper &amp; A. L. Braswell" country="United States of America" location="Hot Springs" municipality="Shut-in Creek" pageId="10" pageNumber="332" specimenCode="NCSM 2203" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353BDEF92E3E7AF8AE" box="[712,857,1771,1797]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 2203</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D353A72F92E3923F8AE" box="[868,1024,1771,1797]" pageId="10" pageNumber="332">Shut-in Creek</collectingMunicipality>
along SR 1183, 1.3 road miles south of junction with 
<collectingCountry id="F32A5A9BFFA04D353846F8CA3C57F882" box="[336,372,1807,1833]" name="United States of America" pageId="10" pageNumber="332">US</collectingCountry>
25/70, west of 
<location id="8EE24CD0FFA04D353B36F8CA3F8BF882" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353B36F8CA3F8BF882" box="[544,680,1807,1833]" country="United States of America" municipality="Shut-in Creek" name="Hot Springs" pageId="10" pageNumber="332">Hot Springs</location>
, 1 MI, 
<collectorName id="26C87FDDFFA04D353BEEF8CA3E5EF882" box="[760,893,1807,1833]" pageId="10" pageNumber="332">J.E. Cooper</collectorName>
, 
<collectorName id="26C87FDDFFA04D353A90F8CA3905F882" box="[902,1062,1807,1833]" pageId="10" pageNumber="332">A.L. Braswell</collectorName>
, 
<date id="FF833CCBFFA04D353D27F8CB399CF882" box="[1073,1215,1806,1833]" pageId="10" pageNumber="332" value="1984-09-25">
<collectingDate id="EFC7C523FFA04D353D27F8CB399CF882" box="[1073,1215,1806,1833]" pageId="10" pageNumber="332" value="1984-09-25">25 Sep 1984</collectingDate>
</date>
; (20)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353C16F8CA3866F8E6" ID-GBIF-Occurrence="3420008318" collectingDate="2001-09-07" collectionCode="NCSM" collectorName="F, A. H. Fullerton, J &amp; Smith" country="United States of America" location="Allenstand" municipality="Little Laurel Creek" pageId="10" pageNumber="332" specimenCode="NCSM 24160" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353C16F8CA38BFF882" box="[1280,1436,1807,1833]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 24160</specimenCode>
<collectingMunicipality id="6BE68071FFA04D353981F8F63C52F8E6" box="[151,369,1843,1869]" pageId="10" pageNumber="332">Little Laurel Creek</collectingMunicipality>
at NC 208, south of 
<location id="8EE24CD0FFA04D353B77F8F63FFEF8E6" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353B77F8F63FFEF8E6" box="[609,733,1843,1869]" country="United States of America" municipality="Little Laurel Creek" name="Allenstand" pageId="10" pageNumber="332">Allenstand</location>
, 1 MI, 1 
<collectorName id="26C87FDDFFA04D353A5EF8F63905F8E6" box="[840,1062,1843,1869]" pageId="10" pageNumber="332">F, A.H. Fullerton, J</collectorName>
. 
<collectorName id="26C87FDDFFA04D353D25F8F6395AF8E6" box="[1075,1145,1843,1869]" pageId="10" pageNumber="332">Smith</collectorName>
, 
<date id="FF833CCBFFA04D353D93F8F6382AF8E6" box="[1157,1289,1843,1869]" pageId="10" pageNumber="332" value="2001-09-07">
<collectingDate id="EFC7C523FFA04D353D93F8F6382AF8E6" box="[1157,1289,1843,1869]" pageId="10" pageNumber="332" value="2001-09-07">7 Sep 2001</collectingDate>
</date>
; (21)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353C5BF8F63C71F83E" ID-GBIF-Occurrence="3420008335" collectingDate="2001-09-06" collectionCode="NCSM" collectorName="A. H. Fullerton &amp; J. Smith" country="United States of America" location="Mead Fork Road" municipality="Meadow Fork Creek" pageId="10" pageNumber="332" specimenCode="NCSM 24165" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353C5BF8F63DC3F8DA" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 24165</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA04D3539FAF8923CF7F8DA" box="[236,468,1879,1905]" pageId="10" pageNumber="332">Meadow Fork Creek</collectingMunicipality>
at 
<location id="8EE24CD0FFA04D3538E1F8923F95F8DA" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D3538E1F8923F95F8DA" box="[503,694,1879,1905]" country="United States of America" municipality="Meadow Fork Creek" name="Mead Fork Road" pageId="10" pageNumber="332">Mead Fork Road</location>
(SR 1175), 
<locationDeviation id="9A300329FFA04D353A29F8923915F8DA" box="[831,1078,1879,1905]" location="Joe" pageId="10" pageNumber="332">
<quantity id="4CC5B7EEFFA04D353A29F8923E84F8DA" box="[831,935,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="2.5749504" pageId="10" pageNumber="332" unit="mi" value="1.6">1.6 miles</quantity>
NNE of Joe
</locationDeviation>
, 1 MI, 
<collectorName id="26C87FDDFFA04D353D90F892380DF8DA" box="[1158,1326,1879,1905]" pageId="10" pageNumber="332">A.H. Fullerton</collectorName>
, 
<collectorName id="26C87FDDFFA04D353C2FF89238BBF8DA" box="[1337,1432,1879,1905]" pageId="10" pageNumber="332">J. Smith</collectorName>
, 
<date id="FF833CCBFFA04D353981F8BE3C3BF83E" box="[151,280,1915,1941]" pageId="10" pageNumber="332" value="2001-09-06">
<collectingDate id="EFC7C523FFA04D353981F8BE3C3BF83E" box="[151,280,1915,1941]" pageId="10" pageNumber="332" value="2001-09-06">6 Sep 2001</collectingDate>
</date>
; (22)
</materialsCitation>
<materialsCitation id="3B551056FFA04D35384FF8BE3822F83E" ID-GBIF-Occurrence="3420008330" box="[345,1281,1914,1941]" collectingDate="2006-09-22" collectionCode="NCSM" collectorName="W. B. Crouch" country="United States of America" location="Big Creek" pageId="10" pageNumber="332" specimenCode="NCSM 25618" specimenCount="1">
<specimenCode id="DB9BB270FFA04D35384FF8BE3CDBF83E" box="[345,504,1914,1941]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 25618</specimenCode>
, 
<location id="8EE24CD0FFA04D353B15F8BE3F56F83E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353B15F8BE3F56F83E" box="[515,629,1915,1941]" country="United States of America" name="Big Creek" pageId="10" pageNumber="332">Big Creek</location>
at SR 1312, 
<locationDeviation id="9A300329FFA04D353A10F8BE3EEAF83E" box="[774,969,1915,1941]" location="Carmen" pageId="10" pageNumber="332">NNW of Carmen</locationDeviation>
, 
<collectorName id="26C87FDDFFA04D353AC5F8BE3944F83E" box="[979,1127,1915,1941]" pageId="10" pageNumber="332">W.B. Crouch</collectorName>
, 
<date id="FF833CCBFFA04D353D64F8BE3822F83E" box="[1138,1281,1915,1941]" pageId="10" pageNumber="332" value="2006-09-22">
<collectingDate id="EFC7C523FFA04D353D64F8BE3822F83E" box="[1138,1281,1915,1941]" pageId="10" pageNumber="332" value="2006-09-22">22 Sep 2006</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B551056FFA04D353C1AF8BE3C6EF812" ID-GBIF-Occurrence="3420008309" country="United States of America" county="Mitchell County" location="Mitchell County" pageId="10" pageNumber="332" specimenCount="1">
1 MI; 
<emphasis id="B949C619FFA04D353C44F8BE3C2DF812" italics="true" pageId="10" pageNumber="332">
<collectingCounty id="62E36287FFA04D353C44F8BE3C2DF812" pageId="10" pageNumber="332">Mitchell County</collectingCounty>
</emphasis>
: (23)
</materialsCitation>
<materialsCitation id="3B551056FFA04D353841F85A3CF9F876" ID-GBIF-Occurrence="3420008310" collectingDate="2007-06-21" collectionCode="NCSM" collectorName="B. Tracy &amp; J. DeBerardinis &amp; A. Rominger &amp; Simonson" country="United States of America" location="Hollow Poplar Creek" pageId="10" pageNumber="332" specimenCode="NCSM 5733" specimenCount="1">
<specimenCode id="DB9BB270FFA04D353841F85A3CC8F812" box="[343,491,1950,1977]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM 5733</specimenCode>
, 
<location id="8EE24CD0FFA04D3538EEF85A3FC8F812" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D3538EEF85A3FC8F812" box="[504,747,1951,1977]" country="United States of America" name="Hollow Poplar Creek" pageId="10" pageNumber="332">Hollow Poplar Creek</location>
at NC 197, 3 MII, 
<collectorName id="26C87FDDFFA04D353AC7F85A3917F812" box="[977,1076,1951,1977]" pageId="10" pageNumber="332">B. Tracy</collectorName>
, 
<collectorName id="26C87FDDFFA04D353D54F85A39D6F812" box="[1090,1269,1951,1977]" pageId="10" pageNumber="332">J. DeBerardinis</collectorName>
, 
<collectorName id="26C87FDDFFA04D353C14F85A38BBF812" box="[1282,1432,1951,1977]" pageId="10" pageNumber="332">A. Rominger</collectorName>
, M. 
<collectorName id="26C87FDDFFA04D3539D5F8063C15F876" box="[195,310,1987,2013]" pageId="10" pageNumber="332">Simonson</collectorName>
, 
<date id="FF833CCBFFA04D353853F8063CF9F876" box="[325,474,1987,2013]" pageId="10" pageNumber="332" value="2007-06-21">
<collectingDate id="EFC7C523FFA04D353853F8063CF9F876" box="[325,474,1987,2013]" pageId="10" pageNumber="332" value="2007-06-21">21 Jun 2007</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B551056FFA04D3538FDF8063FF4F876" ID-GBIF-Occurrence="3420008305" box="[491,727,1987,2013]" country="United States of America" county="Yancey County" location="Yancey County" pageId="10" pageNumber="332" specimenCount="1">
<emphasis id="B949C619FFA04D3538FDF8063FB6F876" box="[491,661,1987,2013]" italics="true" pageId="10" pageNumber="332">
<collectingCounty id="62E36287FFA04D3538FDF8063FB6F876" box="[491,661,1987,2013]" pageId="10" pageNumber="332">Yancey County</collectingCounty>
</emphasis>
: (24)
</materialsCitation>
<materialsCitation id="3B551056FFA04D333BF5F8063F07FF1A" ID-GBIF-Occurrence="3420008306" collectingDate="2008-01-29" collectionCode="NCSM" collectorName="Ramseytown" country="United States of America" lastPageId="12" lastPageNumber="334" location="Will Higgins Road" municipality="Big Creek" pageId="10" pageNumber="332" specimenCount="1">
<collectionCode id="ED2C82CEFFA04D353BF5F8063E11F876" box="[739,818,1987,2013]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="332" type="Museum">NCSM</collectionCode>
uncat., 
<collectingMunicipality id="6BE68071FFA04D353A82F8063928F876" box="[916,1035,1987,2013]" pageId="10" pageNumber="332">Big Creek</collectingMunicipality>
at 
<location id="8EE24CD0FFA04D353D20F8063832F876" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA04D353D20F8063832F876" box="[1078,1297,1987,2013]" country="United States of America" municipality="Big Creek" name="Will Higgins Road" pageId="10" pageNumber="332">Will Higgins Road</location>
(SR 1444), 
<collectorName id="26C87FDDFFA64D333981FF523C0FFF1A" box="[151,300,151,177]" pageId="12" pageNumber="334">Ramseytown</collectorName>
, 1 F, 
<date id="FF833CCBFFA64D333872FF523CC8FF1A" box="[356,491,151,177]" pageId="12" pageNumber="334" value="2008-01-29">
<collectingDate id="EFC7C523FFA64D333872FF523CC8FF1A" box="[356,491,151,177]" pageId="12" pageNumber="334" value="2008-01-29">29 Jan 2008</collectingDate>
</date>
; (25)
</materialsCitation>
<materialsCitation id="3B551056FFA64D333B3CFF523963FF1A" ID-GBIF-Occurrence="3420008313" box="[554,1088,151,177]" collectionCode="NCSM" collectorName="Riverside" country="United States of America" location="Cane River" pageId="12" pageNumber="334" specimenCode="NCSM 23831" specimenCount="1">
<specimenCode id="DB9BB270FFA64D333B3CFF523FEBFF1A" box="[554,712,151,177]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 23831</specimenCode>
, 
<location id="8EE24CD0FFA64D333BC4FF523E6CFF1A" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333BC4FF523E6CFF1A" box="[722,847,151,177]" country="United States of America" name="Cane River" pageId="12" pageNumber="334">Cane River</location>
at 
<collectingCountry id="F32A5A9BFFA64D333A79FF523EB0FF1A" box="[879,915,151,177]" name="United States of America" pageId="12" pageNumber="334">US</collectingCountry>
19E, 
<collectorName id="26C87FDDFFA64D333AC4FF523963FF1A" box="[978,1088,151,177]" pageId="12" pageNumber="334">Riverside</collectorName>
</materialsCitation>
, 
<materialsCitation id="3B551056FFA64D333D5CFF523DE5FF7D" ID-GBIF-Occurrence="3420008327" collectingDate="2002-09-09" collectionCode="T" country="United States of America" location="Savidge" pageId="12" pageNumber="334" specimenCount="1">
<collectionCode id="ED2C82CEFFA64D333D5CFF52397FFF1A" box="[1098,1116,151,177]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="334">T</collectionCode>
.W. 
<location id="8EE24CD0FFA64D333D92FF5239C2FF1A" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333D92FF5239C2FF1A" box="[1156,1249,151,177]" country="United States of America" name="Savidge" pageId="12" pageNumber="334">Savidge</location>
, 1 F, 
<date id="FF833CCBFFA64D333C0FFF5238BBFF1A" box="[1305,1432,151,177]" pageId="12" pageNumber="334" value="2002-09-09">
<collectingDate id="EFC7C523FFA64D333C0FFF5238BBFF1A" box="[1305,1432,151,177]" pageId="12" pageNumber="334" value="2002-09-09">9 Sep 2002</collectingDate>
</date>
; (26)
</materialsCitation>
<materialsCitation id="3B551056FFA64D3339DBFF7E3C28FF52" ID-GBIF-Occurrence="3420008338" collectingDate="2003-06-18" collectionCode="NCSM" collectorName="S. J. Fraley" country="United States of America" location="Old Ferguson Mill Dam" municipality="Cane River" pageId="12" pageNumber="334" specimenCode="NCSM 24438" specimenCount="1">
<specimenCode id="DB9BB270FFA64D3339DBFF7E3C4EFF7E" box="[205,365,187,213]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 24438</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA64D33386EFF7E3CD4FF7E" box="[376,503,187,213]" pageId="12" pageNumber="334">Cane River</collectingMunicipality>
at 
<location id="8EE24CD0FFA64D333B0CFF7E3E0BFF7E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333B0CFF7E3E0BFF7E" box="[538,808,187,213]" country="United States of America" municipality="Cane River" name="Old Ferguson Mill Dam" pageId="12" pageNumber="334">Old Ferguson Mill Dam</location>
upstream of 
<collectingCountry id="F32A5A9BFFA64D333AABFF7E3EC2FF7E" box="[957,993,187,213]" name="United States of America" pageId="12" pageNumber="334">US</collectingCountry>
19E crossing, 1 MI, 
<collectorName id="26C87FDDFFA64D333DD8FF7E3864FF7E" box="[1230,1351,187,213]" pageId="12" pageNumber="334">S.J. Fraley</collectorName>
, 
<date id="FF833CCBFFA64D333C44FF7E3DF1FF52" pageId="12" pageNumber="334" value="2003-06-18">
<collectingDate id="EFC7C523FFA64D333C44FF7E3DF1FF52" pageId="12" pageNumber="334" value="2003-06-18">18 Jun 2003</collectingDate>
</date>
; (27)
</materialsCitation>
<materialsCitation id="3B551056FFA64D333804FF1A3E8DFEB6" ID-GBIF-Occurrence="3420008348" collectingDate="1980-09-07" collectionCode="NCSM" collectorName="E. F. Menhinick &amp; Limnology" country="United States of America" county="Cane River" location="Bloody Fork Road" municipality="Miller Chapel" pageId="12" pageNumber="334" specimenCode="NCSM 3922" specimenCount="1">
<specimenCode id="DB9BB270FFA64D333804FF1A3C81FF52" box="[274,418,223,249]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 3922</specimenCode>
, 
<collectingCounty id="62E36287FFA64D3338BAFF1A3F09FF52" box="[428,554,223,249]" pageId="12" pageNumber="334">Cane River</collectingCounty>
at 
<collectingMunicipality id="6BE68071FFA64D333B5CFF1A3FCAFF52" box="[586,745,223,249]" pageId="12" pageNumber="334">Miller Chapel</collectingMunicipality>
, 
<quantity id="4CC5B7EEFFA64D333BE5FF1A3E7AFF52" box="[755,857,223,249]" metricMagnitude="2" metricUnit="m" metricValue="4.828032" pageId="12" pageNumber="334" unit="mi" value="0.3">0.3 miles</quantity>
north of 
<location id="8EE24CD0FFA64D333AABFF1A39AFFF52" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333AABFF1A39AFFF52" box="[957,1164,223,249]" country="United States of America" county="Cane River" municipality="Miller Chapel" name="Bloody Fork Road" pageId="12" pageNumber="334">Bloody Fork Road</location>
(SR 1387), east of 
<location id="8EE24CD0FFA64D333C77FF1A3DFCFEB6" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333C77FF1A3DFCFEB6" country="United States of America" county="Cane River" municipality="Miller Chapel" name="Lewisburg" pageId="12" pageNumber="334">Lewisburg</location>
, 1 MII, 
<collectorName id="26C87FDDFFA64D33382BFEC63CC8FEB6" box="[317,491,259,285]" pageId="12" pageNumber="334">E.F. Menhinick</collectorName>
and 
<collectorName id="26C87FDDFFA64D333B32FEC63F83FEB6" box="[548,672,259,285]" pageId="12" pageNumber="334">Limnology</collectorName>
class, 
<date id="FF833CCBFFA64D333BF9FEC63E51FEB6" box="[751,882,259,285]" pageId="12" pageNumber="334" value="1980-09-07">
<collectingDate id="EFC7C523FFA64D333BF9FEC63E51FEB6" box="[751,882,259,285]" pageId="12" pageNumber="334" value="1980-09-07">7 Sep 1980</collectingDate>
</date>
; (28)
</materialsCitation>
<materialsCitation id="3B551056FFA64D333AA0FEC63F7AFEEA" ID-GBIF-Occurrence="3420008325" collectionCode="NCSM" collectorName="Cane &amp; F, B. Tracy, D" country="United States of America" location="Prices Creek" municipality="Indian Creek" pageId="12" pageNumber="334" specimenCode="NCSM 4474" specimenCount="1">
<specimenCode id="DB9BB270FFA64D333AA0FEC6396AFEB6" box="[950,1097,259,285]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 4474</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA64D333D43FEC639CBFEB6" box="[1109,1256,259,285]" pageId="12" pageNumber="334">Indian Creek</collectingMunicipality>
at 
<location id="8EE24CD0FFA64D333C1BFEC638BFFEB6" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333C1BFEC638BFFEB6" box="[1293,1436,259,285]" country="United States of America" municipality="Indian Creek" name="Prices Creek" pageId="12" pageNumber="334">Prices Creek</location>
Road (SR 1126), 
<collectorName id="26C87FDDFFA64D33384AFEE23CBBFEEA" box="[348,408,295,321]" pageId="12" pageNumber="334">Cane</collectorName>
, 2 
<collectorName id="26C87FDDFFA64D3338AFFEE23F7AFEEA" box="[441,601,295,321]" pageId="12" pageNumber="334">F, B. Tracy, D</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B551056FFA64D333B4BFEE23EB4FEEA" ID-GBIF-Occurrence="3420008333" box="[605,919,295,321]" collectingDate="1996-02-23" collectionCode="R" country="United States of America" location="Lenat" pageId="12" pageNumber="334" specimenCount="1">
<collectionCode id="ED2C82CEFFA64D333B4BFEE23F50FEEA" box="[605,627,295,321]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="334">R</collectionCode>
. 
<location id="8EE24CD0FFA64D333B69FEE23FE1FEEA" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333B69FEE23FE1FEEA" box="[639,706,295,321]" country="United States of America" name="Lenat" pageId="12" pageNumber="334">Lenat</location>
, 
<date id="FF833CCBFFA64D333BDBFEE23E7EFEEA" box="[717,861,295,321]" pageId="12" pageNumber="334" value="1996-02-23">
<collectingDate id="EFC7C523FFA64D333BDBFEE23E7EFEEA" box="[717,861,295,321]" pageId="12" pageNumber="334" value="1996-02-23">23 Feb 1996</collectingDate>
</date>
; (29)
</materialsCitation>
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<specimenCode id="DB9BB270FFA64D333A89FEE2391CFEEA" box="[927,1087,295,321]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 90187</specimenCode>
, 
<collectingCounty id="62E36287FFA64D333D5CFEE2399EFEEA" box="[1098,1213,295,321]" pageId="12" pageNumber="334">Big Creek</collectingCounty>
along 
<collectingCountry id="F32A5A9BFFA64D333C1CFEE2380DFEEA" box="[1290,1326,295,321]" name="United States of America" pageId="12" pageNumber="334">US</collectingCountry>
<collectingMunicipality id="6BE68071FFA64D333C23FEE238BFFEEA" box="[1333,1436,295,321]" pageId="12" pageNumber="334">Highway</collectingMunicipality>
19W, north of 
<location id="8EE24CD0FFA64D33382AFE8E3C95FECE" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D33382AFE8E3C95FECE" box="[316,438,331,357]" country="United States of America" county="Big Creek" municipality="Highway" name="Lewisburg" pageId="12" pageNumber="334">Lewisburg</location>
, 2 MII, 
<collectorName id="26C87FDDFFA64D333B04FE8E3F99FECE" box="[530,698,331,357]" pageId="12" pageNumber="334">B.W. Williams</collectorName>
, 
<collectorName id="26C87FDDFFA64D333BD0FE8E3E70FECE" box="[710,851,331,357]" pageId="12" pageNumber="334">P.G. Weaver</collectorName>
, D
</materialsCitation>
. 
<materialsCitation id="3B551056FFA64D333A6FFE8E399BFECE" ID-GBIF-Occurrence="3420008307" box="[889,1208,331,357]" collectingDate="2019-03-24" collectionCode="R" country="United States of America" location="Jones" pageId="12" pageNumber="334" specimenCount="1">
<collectionCode id="ED2C82CEFFA64D333A6FFE8E3EACFECE" box="[889,911,331,357]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="334">R</collectionCode>
. 
<location id="8EE24CD0FFA64D333A8DFE8E3EFFFECE" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333A8DFE8E3EFFFECE" box="[923,988,331,357]" country="United States of America" name="Jones" pageId="12" pageNumber="334">Jones</location>
, 
<date id="FF833CCBFFA64D333AFEFE8E395EFECE" box="[1000,1149,331,357]" pageId="12" pageNumber="334" value="2019-03-24">
<collectingDate id="EFC7C523FFA64D333AFEFE8E395EFECE" box="[1000,1149,331,357]" pageId="12" pageNumber="334" value="2019-03-24">24 Mar 2019</collectingDate>
</date>
; (30)
</materialsCitation>
<materialsCitation id="3B551056FFA64D333DD6FE8E382DFE22" ID-GBIF-Occurrence="3420008347" collectionCode="NCSM" collectorName="F, B. W. Williams, P. G &amp; Weaver, D" country="United States of America" county="Cattail Creek" location="United Methodist Church" municipality="Highway" pageId="12" pageNumber="334" specimenCode="NCSM 90188" specimenCount="1">
<specimenCode id="DB9BB270FFA64D333DD6FE8E3843FECE" box="[1216,1376,331,357]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 90188</specimenCode>
, 
<collectingCounty id="62E36287FFA64D333C7AFE8E3C26FE22" pageId="12" pageNumber="334">Cattail Creek</collectingCounty>
at 
<collectingMunicipality id="6BE68071FFA64D33383EFEAA3CACFE22" box="[296,399,367,393]" pageId="12" pageNumber="334">Highway</collectingMunicipality>
197, 
<location id="8EE24CD0FFA64D3338D9FEAA3FD2FE22" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D3338D9FEAA3FD2FE22" box="[463,753,367,393]" country="United States of America" county="Cattail Creek" municipality="Highway" name="United Methodist Church" pageId="12" pageNumber="334">United Methodist Church</location>
, 
<location id="8EE24CD0FFA64D333BEAFEAA3E4CFE22" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333BEAFEAA3E4CFE22" box="[764,879,367,393]" country="United States of America" county="Cattail Creek" municipality="Highway" name="Pensacola" pageId="12" pageNumber="334">Pensacola</location>
, 2 
<collectorName id="26C87FDDFFA64D333A86FEAA39A9FE22" box="[912,1162,367,393]" pageId="12" pageNumber="334">F, B.W. Williams, P.G</collectorName>
. 
<collectorName id="26C87FDDFFA64D333D83FEAA382DFE22" box="[1173,1294,367,393]" pageId="12" pageNumber="334">Weaver, D</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B551056FFA64D333C04FEAA3C2BFE06" ID-GBIF-Occurrence="3420008340" collectingDate="2019-03-25" collectionCode="R" country="United States of America" location="Jones" pageId="12" pageNumber="334" specimenCount="1">
<collectionCode id="ED2C82CEFFA64D333C04FEAA380BFE22" box="[1298,1320,367,393]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="334">R</collectionCode>
. 
<location id="8EE24CD0FFA64D333C22FEAA3856FE22" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333C22FEAA3856FE22" box="[1332,1397,367,393]" country="United States of America" name="Jones" pageId="12" pageNumber="334">Jones</location>
, 
<date id="FF833CCBFFA64D333C96FEAB3C2BFE06" pageId="12" pageNumber="334" value="2019-03-25">
<collectingDate id="EFC7C523FFA64D333C96FEAB3C2BFE06" pageId="12" pageNumber="334" value="2019-03-25">25 Mar 2019</collectingDate>
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; 
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<collectingRegion id="49F9D4E9FFA64D333802FE563C97FE06" box="[276,436,403,429]" country="United States of America" name="Tennessee" pageId="12" pageNumber="334">TENNESSEE</collectingRegion>
, 
<emphasis id="B949C619FFA64D3338A9FE563F7EFE06" box="[447,605,403,429]" italics="true" pageId="12" pageNumber="334">
<collectingCounty id="62E36287FFA64D3338A9FE563F7EFE06" box="[447,605,403,429]" pageId="12" pageNumber="334">Cocke County</collectingCounty>
</emphasis>
: (31)
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<materialsCitation id="3B551056FFA64D333BB4FE563E57FE7A" ID-GBIF-Occurrence="3420008319" collectingDate="2017-06-09" collectionCode="NCSM" collectorName="Browns &amp; B. W. Williams" country="United States of America" county="Tobes Creek" location="Waterville Road" municipality="Pigeon River" pageId="12" pageNumber="334" specimenCode="NCSM 90192" specimenCount="1" stateProvince="Tennessee">
<specimenCode id="DB9BB270FFA64D333BB4FE563E61FE06" box="[674,834,403,429]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 90192</specimenCode>
, 
<collectingCounty id="62E36287FFA64D333A5BFE563EF9FE06" box="[845,986,403,429]" pageId="12" pageNumber="334">Tobes Creek</collectingCounty>
at confluence with 
<collectingMunicipality id="6BE68071FFA64D333DA1FE56386FFE06" box="[1207,1356,403,429]" pageId="12" pageNumber="334">Pigeon River</collectingMunicipality>
, below 
<location id="8EE24CD0FFA64D333981FE723C6CFE7A" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333981FE723C6CFE7A" box="[151,335,439,465]" country="United States of America" county="Tobes Creek" municipality="Pigeon River" name="Waterville Road" pageId="12" pageNumber="334" stateProvince="Tennessee">Waterville Road</location>
, 
<collectorName id="26C87FDDFFA64D33384FFE723C90FE7A" box="[345,435,439,465]" pageId="12" pageNumber="334">Browns</collectorName>
, 1 MII, 
<collectorName id="26C87FDDFFA64D333B1CFE723F8DFE7A" box="[522,686,439,465]" pageId="12" pageNumber="334">B.W. Williams</collectorName>
<emphasis id="B949C619FFA64D333BA2FE723FC7FE7A" box="[692,740,439,465]" italics="true" pageId="12" pageNumber="334">et al</emphasis>
., 
<date id="FF833CCBFFA64D333BEEFE723E57FE7A" box="[760,884,439,465]" pageId="12" pageNumber="334" value="2017-06-09">
<collectingDate id="EFC7C523FFA64D333BEEFE723E57FE7A" box="[760,884,439,465]" pageId="12" pageNumber="334" value="2017-06-09">9 Jun 2017</collectingDate>
</date>
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; 
<materialsCitation id="3B551056FFA64D333A69FE72397DFE7A" ID-GBIF-Occurrence="3420008339" box="[895,1118,439,465]" country="United States of America" county="Unicoi County" location="Unicoi County" pageId="12" pageNumber="334" specimenCount="1" stateProvince="Tennessee">
<emphasis id="B949C619FFA64D333A69FE723901FE7A" box="[895,1058,439,465]" italics="true" pageId="12" pageNumber="334">
<collectingCounty id="62E36287FFA64D333A69FE723901FE7A" box="[895,1058,439,465]" pageId="12" pageNumber="334">Unicoi County</collectingCounty>
</emphasis>
: (32)
</materialsCitation>
<materialsCitation id="3B551056FFA64D333D73FE723EDFFE5E" ID-GBIF-Occurrence="3420008346" collectionCode="NCSM" collectorName="F, B. W. Williams, P. G &amp; Weaver, D" country="United States of America" location="Highway" municipality="North Indian Creek" pageId="12" pageNumber="334" specimenCode="NCSM 90189" specimenCount="1" stateProvince="Tennessee">
<specimenCode id="DB9BB270FFA64D333D73FE723820FE7A" box="[1125,1283,439,465]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 90189</specimenCode>
, 
<collectingMunicipality id="6BE68071FFA64D333C1BFE723DF9FE5E" pageId="12" pageNumber="334">North Indian Creek</collectingMunicipality>
at 
<location id="8EE24CD0FFA64D333816FE1E3C44FE5E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333816FE1E3C44FE5E" box="[256,359,475,501]" country="United States of America" municipality="North Indian Creek" name="Highway" pageId="12" pageNumber="334" stateProvince="Tennessee">Highway</location>
107, 
<locationDeviation id="9A300329FFA64D3338BFFE1E3F70FE5E" box="[425,595,475,501]" location="Unicoi" pageId="12" pageNumber="334">ESE of Unicoi</locationDeviation>
, 1 
<collectorName id="26C87FDDFFA64D333B61FE1E3E56FE5E" box="[631,885,475,501]" pageId="12" pageNumber="334">F, B.W. Williams, P.G</collectorName>
. 
<collectorName id="26C87FDDFFA64D333A97FE1E3EDFFE5E" box="[897,1020,475,501]" pageId="12" pageNumber="334">Weaver, D</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B551056FFA64D333D16FE1E3867FE5E" ID-GBIF-Occurrence="3420008315" box="[1024,1348,475,501]" collectingDate="2019-03-24" collectionCode="R" country="United States of America" location="Jones" pageId="12" pageNumber="334" specimenCount="1" stateProvince="Tennessee">
<collectionCode id="ED2C82CEFFA64D333D16FE1E3935FE5E" box="[1024,1046,475,501]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="334">R</collectionCode>
. 
<location id="8EE24CD0FFA64D333D34FE1E3940FE5E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333D34FE1E3940FE5E" box="[1058,1123,475,501]" country="United States of America" name="Jones" pageId="12" pageNumber="334" stateProvince="Tennessee">Jones</location>
, 
<date id="FF833CCBFFA64D333D66FE1E382BFE5E" box="[1136,1288,475,501]" pageId="12" pageNumber="334" value="2019-03-24">
<collectingDate id="EFC7C523FFA64D333D66FE1E382BFE5E" box="[1136,1288,475,501]" pageId="12" pageNumber="334" value="2019-03-24">24 Mar 2019</collectingDate>
</date>
; (33)
</materialsCitation>
<materialsCitation id="3B551056FFA64D333C5BFE1E3FD4FD96" ID-GBIF-Occurrence="3420008314" collectionCode="NCSM" collectorName="F, B. W. Williams, P. G &amp; Weaver, D" country="United States of America" county="Spivey Creek" location="Spivey Mountain" municipality="Tilson Mountain" pageId="12" pageNumber="334" specimenCode="NCSM 90190" specimenCount="1" stateProvince="Tennessee">
<specimenCode id="DB9BB270FFA64D333C5BFE1E3DC3FDB2" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="12" pageNumber="334" type="Museum">NCSM 90190</specimenCode>
, 
<collectingCounty id="62E36287FFA64D3339FDFE3A3CA5FDB2" box="[235,390,511,537]" pageId="12" pageNumber="334">Spivey Creek</collectingCounty>
, at intersection of 
<collectingMunicipality id="6BE68071FFA64D333B42FE3A3E2CFDB2" box="[596,783,511,537]" pageId="12" pageNumber="334">Tilson Mountain</collectingMunicipality>
Road and 
<location id="8EE24CD0FFA64D333A9EFE3A3969FDB2" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333A9EFE3A3969FDB2" box="[904,1098,511,537]" country="United States of America" county="Spivey Creek" municipality="Tilson Mountain" name="Spivey Mountain" pageId="12" pageNumber="334" stateProvince="Tennessee">Spivey Mountain</location>
Road (
<location id="8EE24CD0FFA64D333D8DFE3A39C3FDB2" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333D8DFE3A39C3FDB2" box="[1179,1248,511,537]" country="United States of America" county="Spivey Creek" municipality="Tilson Mountain" name="Route" pageId="12" pageNumber="334" stateProvince="Tennessee">Route</location>
19W/36), 
<locationDeviation id="9A300329FFA64D333C4EFE3A3C2FFD96" location="Flag Pond" pageId="12" pageNumber="334">NE of Flag Pond</locationDeviation>
, 1 MII, 1 
<collectorName id="26C87FDDFFA64D33386DFDE63F57FD96" box="[379,628,547,573]" pageId="12" pageNumber="334">F, B.W. Williams, P.G</collectorName>
. 
<collectorName id="26C87FDDFFA64D333B68FDE63FD4FD96" box="[638,759,547,573]" pageId="12" pageNumber="334">Weaver, D</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B551056FFA64D333BEDFDE63EDEFD96" ID-GBIF-Occurrence="3420008322" box="[763,1021,547,573]" collectingDate="2019-03-24" collectionCode="R" country="United States of America" location="Jones" pageId="12" pageNumber="334" specimenCount="1" stateProvince="Tennessee">
<collectionCode id="ED2C82CEFFA64D333BEDFDE63E32FD96" box="[763,785,547,573]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="334">R</collectionCode>
. 
<location id="8EE24CD0FFA64D333A0AFDE63E7EFD96" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333A0AFDE63E7EFD96" box="[796,861,547,573]" country="United States of America" name="Jones" pageId="12" pageNumber="334" stateProvince="Tennessee">Jones</location>
, 
<date id="FF833CCBFFA64D333A7EFDE63EDEFD96" box="[872,1021,547,573]" pageId="12" pageNumber="334" value="2019-03-24">
<collectingDate id="EFC7C523FFA64D333A7EFDE63EDEFD96" box="[872,1021,547,573]" pageId="12" pageNumber="334" value="2019-03-24">24 Mar 2019</collectingDate>
</date>
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.
</paragraph>
<caption id="DF424A83FFA14D343981F8483C6CF846" ID-DOI="http://doi.org/10.5281/zenodo.5792771" ID-Zenodo-Dep="5792771" httpUri="https://zenodo.org/record/5792771/files/figure.png" pageId="11" pageNumber="333" startId="11.[151,250,1933,1957]" targetBox="[151,1436,185,1909]" targetPageId="11">
<paragraph id="8B821A0BFFA14D343981F8483C6CF846" blockId="11.[151,1436,1933,2029]" pageId="11" pageNumber="333">
<emphasis id="B949C619FFA14D343981F8483C31F80E" bold="true" box="[151,274,1933,1957]" pageId="11" pageNumber="333">FIGURE 6.</emphasis>
Maps (A. relief, B. HUC 10 boundaries) showing the distribution 
<emphasis id="B949C619FFA14D343ABCF8483967F80E" box="[938,1092,1933,1957]" italics="true" pageId="11" pageNumber="333">
of 
<taxonomicName id="4C3D6188FFA14D343AD5F8483967F80E" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[963,1092,1933,1957]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="333" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">C. ectopistes</taxonomicName>
</emphasis>
<emphasis id="B949C619FFA14D343D5FF84839BAF80E" bold="true" box="[1097,1177,1933,1957]" pageId="11" pageNumber="333">
<taxonomicNameLabel id="A27A7B62FFA14D343D5FF84839BAF80E" box="[1097,1177,1933,1957]" pageId="11" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the Pigeon River (PR), French Broad River (FB), Nolichucky River (NOL), and Cane River (CANE) systems. * Denotes the sampling locality on Paint Creek, Tennessee.
</paragraph>
</caption>
<paragraph id="8B821A0BFFA64D3339D1FD833F2CFC2A" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D3339D1FD833E29FDCB" box="[199,778,582,609]" italics="true" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D3339D1FD833F3FFDCA" bold="true" box="[199,540,582,609]" italics="true" pageId="12" pageNumber="334">Habitat and life history notes.</emphasis>
<taxonomicName id="4C3D6188FFA64D333B37FD823E29FDCB" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[545,778,583,609]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">Cambarus ectopistes</taxonomicName>
</emphasis>
<emphasis id="B949C619FFA64D333A07FD823E49FDCA" bold="true" box="[785,874,583,609]" pageId="12" pageNumber="334">
<taxonomicNameLabel id="A27A7B62FFA64D333A07FD823E49FDCA" box="[785,874,583,609]" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occupies moderate to large perennial streams with substrates composed primarily of large cobbles and boulders (
<figureCitation id="1306068EFFA64D333A49FDAF3EE7FD2E" box="[863,964,618,645]" captionStart="FIGURE 7" captionStartId="13.[151,250,1394,1418]" captionTargetBox="[173,1413,181,1368]" captionTargetId="figure-28@13.[173,1413,181,1370]" captionTargetPageId="13" captionText="FIGURE 7. Examples of stream habitat in which C. ectopistes sp. nov. was encountered." figureDoi="http://doi.org/10.5281/zenodo.5792775" httpUri="https://zenodo.org/record/5792775/files/figure.png" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D333A49FDAF3EE7FD2E" bold="true" box="[863,964,618,645]" pageId="12" pageNumber="334">Figure 7</emphasis>
</figureCitation>
). It uses both stream channel and stream edge habitats and is encountered most often in runs and moderate velocity riffles. This species appears to use pool habitats infrequently in larger streams, whereas in smaller, more moderate-sized streams 
<taxonomicName id="4C3D6188FFA64D333D6CFD76382AFD67" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1146,1289,691,717]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA64D333D6CFD76382AFD67" box="[1146,1289,691,717]" italics="true" pageId="12" pageNumber="334">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA64D333C06FD763849FD66" bold="true" box="[1296,1386,691,717]" pageId="12" pageNumber="334">
<taxonomicNameLabel id="A27A7B62FFA64D333C06FD763849FD66" box="[1296,1386,691,717]" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was encountered primarily in plunge pools associated with waterfalls, and frequently occurred in pool thalwegs amongst leaf packs. Although 
<taxonomicName id="4C3D6188FFA64D333893FD3E3F30FCBF" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[389,531,763,789]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA64D333893FD3E3F30FCBF" box="[389,531,763,789]" italics="true" pageId="12" pageNumber="334">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA64D333B0CFD3E3F57FCBE" bold="true" box="[538,628,763,789]" pageId="12" pageNumber="334">
<taxonomicNameLabel id="A27A7B62FFA64D333B0CFD3E3F57FCBE" box="[538,628,763,789]" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was not usually found in steep, fast-flowing streams—where 
<taxonomicName id="4C3D6188FFA64D333C3EFD3E3C34FC92" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA64D333C3EFD3E38BEFCBE" box="[1320,1437,763,789]" italics="true" pageId="12" pageNumber="334">Cambarus</emphasis>
cf. 
<emphasis id="B949C619FFA64D3339ABFCDA3C34FC92" box="[189,279,799,825]" italics="true" pageId="12" pageNumber="334">bartonii</emphasis>
</taxonomicName>
was predominant—we did encounter several individuals of the species while sampling high gradient second ordered streams in tributaries to the Pigeon River, including Cataloochee and Big Creek drainages in Great Smoky Mountains National Park.
</paragraph>
<paragraph id="8B821A0BFFA64D3339D1FC4E3962FB6E" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="334">
<materialsCitation id="3B551056FFA64D3339D1FC4E391EFB6E" ID-GBIF-Occurrence="3420008332" collectingDate="2017-06-09" collectingDateMax="2017-11-16" collectingDateMin="2017-06-09" collectionCode="TCL" collectorName="Gulf Fork Big Creek, TN" county="Gulf Fork Big Creek" location="Based" municipality="Among" pageId="12" pageNumber="334" specimenCount="116" specimenCount-egg="116">
<location id="8EE24CD0FFA64D3339D1FC4E3C28FC0E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D3339D1FC4E3C28FC0E" box="[199,267,907,933]" county="Gulf Fork Big Creek" municipality="Among" name="Based" pageId="12" pageNumber="334">Based</location>
on specimens examined for this description, both form I and form II males are present during all months of the year. 
<location id="8EE24CD0FFA64D333809FC6A3C68FC62" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333809FC6A3C68FC62" box="[287,331,943,969]" county="Gulf Fork Big Creek" municipality="Among" name="The" pageId="12" pageNumber="334">The</location>
incidence of form I males increased from 
<location id="8EE24CD0FFA64D333A24FC6A3E89FC62" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333A24FC6A3E89FC62" box="[818,938,943,969]" county="Gulf Fork Big Creek" municipality="Among" name="September" pageId="12" pageNumber="334">September</location>
through 
<location id="8EE24CD0FFA64D333D04FC6A3965FC62" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333D04FC6A3965FC62" box="[1042,1094,943,969]" county="Gulf Fork Big Creek" municipality="Among" name="May" pageId="12" pageNumber="334">May</location>
, and form II males were most prevalent from 
<location id="8EE24CD0FFA64D333855FC163C55FC46" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333855FC163C55FC46" box="[323,374,979,1005]" county="Gulf Fork Big Creek" municipality="Among" name="May" pageId="12" pageNumber="334">May</location>
through October. Ovigerous females were encountered in 
<location id="8EE24CD0FFA64D333D17FC163917FC46" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333D17FC163917FC46" box="[1025,1076,979,1005]" county="Gulf Fork Big Creek" municipality="Among" name="May" pageId="12" pageNumber="334">May</location>
and 
<location id="8EE24CD0FFA64D333D7CFC163983FC46" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333D7CFC163983FC46" box="[1130,1184,979,1005]" county="Gulf Fork Big Creek" municipality="Among" name="June" pageId="12" pageNumber="334">June</location>
, females bearing stage I juveniles were collected in late 
<location id="8EE24CD0FFA64D333B03FC323F68FBBA" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333B03FC323F68FBBA" box="[533,587,1015,1041]" county="Gulf Fork Big Creek" municipality="Among" name="June" pageId="12" pageNumber="334">June</location>
, stage II juveniles were present on females collected in 
<location id="8EE24CD0FFA64D333DD9FC3239DCFBBA" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333DD9FC3239DCFBBA" box="[1231,1279,1015,1041]" county="Gulf Fork Big Creek" municipality="Among" name="July" pageId="12" pageNumber="334">July</location>
, and stage IV juveniles were present in 
<location id="8EE24CD0FFA64D3338AAFBDE3F2EFB9E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D3338AAFBDE3F2EFB9E" box="[444,525,1051,1077]" county="Gulf Fork Big Creek" municipality="Among" name="August" pageId="12" pageNumber="334">August</location>
and November. Juveniles collected on females in 
<location id="8EE24CD0FFA64D333D46FBDE39E5FB9E" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333D46FBDE39E5FB9E" box="[1104,1222,1051,1077]" county="Gulf Fork Big Creek" municipality="Among" name="November" pageId="12" pageNumber="334">November</location>
likely would have overwintered with their mothers and would have been recruited into the population the following spring. 
<location id="8EE24CD0FFA64D333C2BFBFA384EFBF2" LSID="urn:lsid:plazi:treatment:0394AB1DFFAC4D303981F9D23F38FE5F:8EE24CD0FFA64D333C2BFBFA384EFBF2" box="[1341,1389,1087,1113]" county="Gulf Fork Big Creek" municipality="Among" name="Few" pageId="12" pageNumber="334">Few</location>
full complements of eggs and juveniles were present. 
<collectingMunicipality id="6BE68071FFA64D333BDEFBA63E3FFBD6" box="[712,796,1123,1149]" pageId="12" pageNumber="334">Among</collectingMunicipality>
those examined were a female collected from 
<collectorName id="26C87FDDFFA64D333C3AFBA63C19FB0A" pageId="12" pageNumber="334">
<collectingCounty id="62E36287FFA64D333C3AFBA63C28FB0A" pageId="12" pageNumber="334">Gulf Fork Big Creek</collectingCounty>
, TN
</collectorName>
on 
<date id="FF833CCBFFA64D333874FB423CC4FB0A" box="[354,487,1159,1185]" pageId="12" pageNumber="334" value="2017-06-09">
<collectingDate id="EFC7C523FFA64D333874FB423CC4FB0A" box="[354,487,1159,1185]" pageId="12" pageNumber="334" value="2017-06-09">9 June 2017</collectingDate>
</date>
with a 
<collectionCode id="ED2C82CEFFA64D333B20FB423F49FB0A" box="[566,618,1159,1185]" pageId="12" pageNumber="334">TCL</collectionCode>
of 
<quantity id="4CC5B7EEFFA64D333B9BFB423FCCFB0A" box="[653,751,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="3.42" pageId="12" pageNumber="334" unit="mm" value="34.2">34.2 mm</quantity>
and bearing 
<specimenCount id="9D3BD182FFA64D333A6BFB423EFCFB0A" box="[893,991,1159,1185]" pageId="12" pageNumber="334" type="egg">116 eggs</specimenCount>
and a female collected on 
<date id="FF833CCBFFA64D333C12FB423DECFB6E" pageId="12" pageNumber="334" value="2017-11-16">
<collectingDate id="EFC7C523FFA64D333C12FB423DECFB6E" pageId="12" pageNumber="334" value="2017-11-16">16 November 2017</collectingDate>
</date>
from the 
<typeStatus id="5486A4A9FFA64D33382BFB6E3C4EFB6E" box="[317,365,1195,1221]" pageId="12" pageNumber="334">type</typeStatus>
locality with a 
<collectionCode id="ED2C82CEFFA64D333B0DFB6E3F6CFB6E" box="[539,591,1195,1221]" pageId="12" pageNumber="334">TCL</collectionCode>
of 45.0 mm harboring 142 stage I juveniles
</materialsCitation>
.
</paragraph>
<paragraph id="8B821A0BFFA64D3339D1FB0A3F60FA4E" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D3339D1FB0A3C91FB42" bold="true" box="[199,434,1231,1257]" italics="true" pageId="12" pageNumber="334">Conservation status.</emphasis>
We suggest that 
<taxonomicName id="4C3D6188FFA64D333B62FB0A3E27FB43" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[628,772,1231,1257]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA64D333B62FB0A3FA4FB42" box="[628,647,1231,1257]" italics="true" pageId="12" pageNumber="334">C</emphasis>
. 
<emphasis id="B949C619FFA64D333B80FB0A3E27FB43" box="[662,772,1231,1256]" italics="true" pageId="12" pageNumber="334">ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA64D333A1AFB0A3E45FB42" bold="true" box="[780,870,1231,1257]" pageId="12" pageNumber="334">
<taxonomicNameLabel id="A27A7B62FFA64D333A1AFB0A3E45FB42" box="[780,870,1231,1257]" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
be assigned the status of Special Concern (SC) in 
<collectingRegion id="49F9D4E9FFA64D333981FB363C61FAA6" box="[151,322,1267,1293]" country="United States of America" name="North Carolina" pageId="12" pageNumber="334">North Carolina</collectingRegion>
, vulnerable (G3) using the NatureServe criteria, and vulnerable (
<collectionCode id="ED2C82CEFFA64D333D00FB363909FAA6" box="[1046,1066,1267,1293]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="334" type="Museum">V</collectionCode>
) using the criteria of the International Union for the Conservation of Nature (
<collectionCode id="ED2C82CEFFA64D333B9EFAD23FEDFA9A" box="[648,718,1303,1329]" pageId="12" pageNumber="334">IUCN</collectionCode>
2001). Although the distribution of the species spans three major tributaries of the French Broad River basin, the area occupied in each is narrow, ranging from two to four HUC 10s. There is no current connectivity among these three populations. If, for example, the population in the French Broad River declines, there will not be recolonization or immigration from either the Pigeon or Nolichucky. Consequently, habitat alteration, habitat degradation, and fluctuations in water quality in any of these tributaries may have drastic and detrimental effects on the species.
</paragraph>
<paragraph id="8B821A0BFFA64D3339D1FA2B3E2EF986" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D3339D1FA2B3FBEF9A3" box="[199,669,1518,1545]" italics="true" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D3339D1FA2B3C8FF9A2" bold="true" box="[199,428,1518,1545]" italics="true" pageId="12" pageNumber="334">Crayfish associates.</emphasis>
<taxonomicName id="4C3D6188FFA64D3338A5FA2A3FBEF9A3" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[435,669,1519,1545]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">Cambarus ectopistes</taxonomicName>
</emphasis>
<emphasis id="B949C619FFA64D333BB3FA2A3FDDF9A2" bold="true" box="[677,766,1519,1545]" pageId="12" pageNumber="334">
<taxonomicNameLabel id="A27A7B62FFA64D333BB3FA2A3FDDF9A2" box="[677,766,1519,1545]" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
co-occurs with 
<taxonomicName id="4C3D6188FFA64D333AA5FA2A398FF9A2" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[947,1196,1519,1545]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA64D333AA5FA2A390BF9A2" box="[947,1064,1519,1545]" italics="true" pageId="12" pageNumber="334">Cambarus</emphasis>
cf. 
<emphasis id="B949C619FFA64D333D44FA2A398FF9A2" box="[1106,1196,1519,1545]" italics="true" pageId="12" pageNumber="334">bartonii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3D6188FFA64D333DADFA2A3C47F986" authority="(Faxon, 1885)" baseAuthorityName="Faxon" baseAuthorityYear="1885" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="B949C619FFA64D333DADFA2A3D9FF987" italics="true" pageId="12" pageNumber="334">Cambarus longirostris</emphasis>
(Faxon, 1885)
</taxonomicName>
, and 
<taxonomicName id="4C3D6188FFA64D3338B6F9D63E2AF986" authority="(Faxon, 1884)" baseAuthorityName="Faxon" baseAuthorityYear="1884" box="[416,777,1555,1581]" class="Malacostraca" family="Cambaridae" genus="Faxonius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="forceps">
<emphasis id="B949C619FFA64D3338B6F9D63F43F986" box="[416,608,1555,1581]" italics="true" pageId="12" pageNumber="334">Faxonius forceps</emphasis>
(Faxon, 1884)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3274980FFA64D3039D1F9F23F81FEEB" lastPageId="15" lastPageNumber="337" pageId="12" pageNumber="334" type="description">
<paragraph id="8B821A0BFFA64D3339D1F9F23954F882" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D3339D1F9F23C1AF9FA" bold="true" box="[199,313,1591,1617]" italics="true" pageId="12" pageNumber="334">Variation.</emphasis>
Several characters, including those of the rostrum and chela, and overall appearance, i.e., gestalt, vary geographically within the range of 
<taxonomicName id="4C3D6188FFA64D333B4EF99E3FC4F9DF" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[600,743,1627,1653]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA64D333B4EF99E3FC4F9DF" box="[600,743,1627,1653]" italics="true" pageId="12" pageNumber="334">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA64D333BF8F99E3E6BF9DE" bold="true" box="[750,840,1627,1653]" pageId="12" pageNumber="334">
<taxonomicNameLabel id="A27A7B62FFA64D333BF8F99E3E6BF9DE" box="[750,840,1627,1653]" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Individuals from the Pigeon and French Broad River systems are generally similar to each other, but are noticeably different from those in the Nolichucky and Cane River systems. Specimens from the Nolichucky and Cane Rivers possess a narrower rostrum with reduced margins compared to specimens from the Pigeon and French Broad rivers, which possess a broad rostrum with more inflated, subparallel margins that terminate in a wide short acumen (
<figureCitation id="1306068EFFA64D333A24F92F3E88F8AE" box="[818,939,1770,1797]" captionStart="FIGURE 8" captionStartId="14.[151,250,875,899]" captionTargetBox="[155,1433,181,851]" captionTargetId="figure-17@14.[155,1433,181,851]" captionTargetPageId="14" captionText="FIGURE 8. Intraspecific variation in C. ectopistes sp. nov. occurs most notably in the shape of the rostrum.A, Nolichucky and Cane Rivers; B, French Broad and Pigeon Rivers." figureDoi="http://doi.org/10.5281/zenodo.5792777" httpUri="https://zenodo.org/record/5792777/files/figure.png" pageId="12" pageNumber="334">
<emphasis id="B949C619FFA64D333A24F92F3E88F8AE" bold="true" box="[818,939,1770,1797]" pageId="12" pageNumber="334">Fig. 8A, B</emphasis>
</figureCitation>
). Despite this variation, all 
<taxonomicName id="4C3D6188FFA64D333DF2F92E3850F8AF" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1252,1395,1771,1797]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA64D333DF2F92E3850F8AF" box="[1252,1395,1771,1797]" italics="true" pageId="12" pageNumber="334">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA64D333C6DF92E3DEBF882" bold="true" pageId="12" pageNumber="334">
<taxonomicNameLabel id="A27A7B62FFA64D333C6DF92E3DEBF882" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
exhibit thickened and pronounced rostral margins, which are absent on the acumen.
</paragraph>
<paragraph id="8B821A0BFFA64D3339D1F8F63F7DF876" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="334">The chela of the French Broad and Pigeon River specimens is robust, with a wide palm, and is subrectangular in shape, whereas that of the Nolichucky and Cane River specimens is comparatively more elongate in appearance, with a narrower palm, and is more subtriangular, and therefore less robust in appearance. Taken as a whole, these differences in the rostrum and chela result in the French Broad and Pigeon River animals seeming more burly than the Nolichucky and Cane River animals.</paragraph>
<caption id="DF424A83FFA74D323981FAB73901FA21" ID-DOI="http://doi.org/10.5281/zenodo.5792775" ID-Zenodo-Dep="5792775" box="[151,1058,1394,1418]" httpUri="https://zenodo.org/record/5792775/files/figure.png" pageId="13" pageNumber="335" startId="13.[151,250,1394,1418]" targetBox="[173,1413,181,1368]" targetPageId="13">
<paragraph id="8B821A0BFFA74D323981FAB73901FA21" blockId="13.[151,1058,1394,1418]" box="[151,1058,1394,1418]" pageId="13" pageNumber="335">
<emphasis id="B949C619FFA74D323981FAB73C2EFA21" bold="true" box="[151,269,1394,1418]" pageId="13" pageNumber="335">FIGURE 7</emphasis>
. Examples of stream habitat in which 
<taxonomicName id="4C3D6188FFA74D323B84FAB73E37FA21" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[658,788,1394,1418]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA74D323B84FAB73E37FA21" box="[658,788,1394,1418]" italics="true" pageId="13" pageNumber="335">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA74D323A0CFAB73E48FA21" bold="true" box="[794,875,1394,1418]" pageId="13" pageNumber="335">
<taxonomicNameLabel id="A27A7B62FFA74D323A0CFAB73E48FA21" box="[794,875,1394,1418]" pageId="13" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was encountered.
</paragraph>
</caption>
<paragraph id="8B821A0BFFA74D3239D1FA7D3CE7F995" blockId="13.[151,1437,1464,2030]" pageId="13" pageNumber="335">
Juvenile and subadult 
<taxonomicName id="4C3D6188FFA74D3238D6FA7C3F6CFA79" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[448,591,1465,1491]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA74D3238D6FA7C3F6CFA79" box="[448,591,1465,1491]" italics="true" pageId="13" pageNumber="335">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA74D323B40FA7D3F8CFA79" bold="true" box="[598,687,1464,1490]" pageId="13" pageNumber="335">
<taxonomicNameLabel id="A27A7B62FFA74D323B40FA7D3F8CFA79" box="[598,687,1464,1490]" pageId="13" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differ from adults in both spination and coloration. The postorbital and cervical tubercles can appear sharply spiniform in juveniles and subadults, but are more rounded, or dulled, in adults. The coloration of juveniles and subadults is dominated by olive greens, greys, and blues, in contrast to the browns observed in adults.
</paragraph>
<paragraph id="8B821A0BFFA74D3239D1F98D3F1EF845" blockId="13.[151,1437,1464,2030]" pageId="13" pageNumber="335">
<emphasis id="B949C619FFA74D3239D1F98D3E00F9C9" box="[199,803,1608,1635]" italics="true" pageId="13" pageNumber="335">
<emphasis id="B949C619FFA74D3239D1F98D3F17F9C9" bold="true" box="[199,564,1608,1635]" italics="true" pageId="13" pageNumber="335">Relationships and comparisons.</emphasis>
<taxonomicName id="4C3D6188FFA74D323B2DF98D3E00F9C9" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[571,803,1608,1634]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">Cambarus ectopistes</taxonomicName>
</emphasis>
<emphasis id="B949C619FFA74D323A3FF98D3EA1F9C9" bold="true" box="[809,898,1608,1634]" pageId="13" pageNumber="335">
<taxonomicNameLabel id="A27A7B62FFA74D323A3FF98D3EA1F9C9" box="[809,898,1608,1634]" pageId="13" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically similar to undescribed members of the 
<taxonomicName id="4C3D6188FFA74D323800F9A83CB4F92D" box="[278,407,1644,1671]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B949C619FFA74D323800F9A83CB4F92D" box="[278,407,1644,1671]" italics="true" pageId="13" pageNumber="335">C. robustus</emphasis>
</taxonomicName>
species complex that occur in the southern Appalachians, 
<taxonomicName id="4C3D6188FFA74D323D3AF9A839EFF92D" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1068,1228,1644,1671]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA74D323D3AF9A83965F92C" box="[1068,1094,1645,1671]" italics="true" pageId="13" pageNumber="335">C.</emphasis>
cf. 
<emphasis id="B949C619FFA74D323D64F9A939EFF92D" box="[1138,1228,1644,1670]" italics="true" pageId="13" pageNumber="335">bartonii</emphasis>
</taxonomicName>
from the southern Appalachians, and 
<taxonomicName id="4C3D6188FFA74D323865F9553FC0F901" authority="Prins, 1968" authorityName="Prins" authorityYear="1968" box="[371,739,1680,1706]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="reburrus">
<emphasis id="B949C619FFA74D323865F9553F70F901" box="[371,595,1680,1706]" italics="true" pageId="13" pageNumber="335">Cambarus reburrus</emphasis>
Prins, 1968
</taxonomicName>
. 
<taxonomicName id="4C3D6188FFA74D323BE7F9553EFDF901" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[753,990,1680,1706]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA74D323BE7F9553EFDF901" box="[753,990,1680,1706]" italics="true" pageId="13" pageNumber="335">Cambarus ectopistes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A7B62FFA74D323AFFF9543961F900" box="[1001,1090,1681,1707]" pageId="13" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
can be differentiated from 
<emphasis id="B949C619FFA74D323C95F9543C07F965" italics="true" pageId="13" pageNumber="335">
<taxonomicName id="4C3D6188FFA74D323C95F9543DD5F965" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" sensu="stricto" species="robustus">C. robustus</taxonomicName>
<taxonomicNameLabel id="A27A7B62FFA74D323817F9703C07F965" box="[257,292,1717,1742]" pageId="13" pageNumber="335" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
—which occurs in northwestern Pennsylvania, New 
<collectingRegion id="49F9D4E9FFA74D323A94F9713E9FF965" box="[898,956,1716,1742]" country="United Kingdom" name="York" pageId="13" pageNumber="335">York</collectingRegion>
, New 
<collectingCountry id="F32A5A9BFFA74D323D1FF971394AF965" box="[1033,1129,1716,1742]" name="United Kingdom" pageId="13" pageNumber="335">England</collectingCountry>
, and 
<collectingRegion id="49F9D4E9FFA74D323DBCF9713821F965" box="[1194,1282,1716,1742]" country="Canada" name="Ontario" pageId="13" pageNumber="335">Ontario</collectingRegion>
, 
<collectingCountry id="F32A5A9BFFA74D323C06F9713847F965" box="[1296,1380,1716,1742]" name="Canada" pageId="13" pageNumber="335">Canada</collectingCountry>
—by characters associated with the chela, rostrum, and overall adult color pattern. 
<emphasis id="B949C619FFA74D323AE6F91D39D7F959" box="[1008,1268,1752,1778]" italics="true" pageId="13" pageNumber="335">
<taxonomicName id="4C3D6188FFA74D323AE6F91D39E9F959" box="[1008,1226,1752,1778]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" sensu="stricto" species="robustus">Cambarus robustus</taxonomicName>
<taxonomicNameLabel id="A27A7B62FFA74D323DC7F91C39D7F959" box="[1233,1268,1753,1778]" pageId="13" pageNumber="335" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
bears 2–4 subpalmar tubercles on the chela, and typically has rostral margins that are entire to the tip of the acumen; the rostral margins are not thickened. In contrast, the chela of 
<taxonomicName id="4C3D6188FFA74D323BF6F8E43E4CF891" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[736,879,1825,1851]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA74D323BF6F8E43E4CF891" box="[736,879,1825,1851]" italics="true" pageId="13" pageNumber="335">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA74D323A6EF8E53EF0F891" bold="true" box="[888,979,1824,1850]" pageId="13" pageNumber="335">
<taxonomicNameLabel id="A27A7B62FFA74D323A6EF8E53EF0F891" box="[888,979,1824,1850]" pageId="13" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
normally lacks subpalmar tubercles and is more streamlined in appearance than 
<taxonomicName id="4C3D6188FFA74D323B43F8803FF6F8F5" box="[597,725,1860,1887]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" sensu="stricto" species="robustus">
<emphasis id="B949C619FFA74D323B43F8803FF6F8F5" box="[597,725,1860,1887]" italics="true" pageId="13" pageNumber="335">C. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A7B62FFA74D323BCAF8803E23F8F4" box="[732,768,1861,1887]" pageId="13" pageNumber="335" sensu="stricto">
<emphasis id="B949C619FFA74D323BCAF8803FDAF8F5" box="[732,761,1861,1886]" italics="true" pageId="13" pageNumber="335">s.s</emphasis>
.
</taxonomicNameLabel>
The rostral margins of 
<taxonomicName id="4C3D6188FFA74D323D1EF88039B4F8F5" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1032,1175,1861,1887]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA74D323D1EF88039B4F8F5" box="[1032,1175,1861,1887]" italics="true" pageId="13" pageNumber="335">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA74D323D88F88139DBF8F5" bold="true" box="[1182,1272,1860,1886]" pageId="13" pageNumber="335">
<taxonomicNameLabel id="A27A7B62FFA74D323D88F88139DBF8F5" box="[1182,1272,1860,1886]" pageId="13" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are thickened, creating a concavity to the rostrum; this thickness terminates prior to the junction with the acumen, and the margins of the acumen are noticeably not inflated. The coloration of adult 
<taxonomicName id="4C3D6188FFA74D323A67F8483ED3F80D" box="[881,1008,1932,1959]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" sensu="stricto" species="robustus">
<emphasis id="B949C619FFA74D323A67F8483ED3F80D" box="[881,1008,1932,1959]" italics="true" pageId="13" pageNumber="335">C. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A7B62FFA74D323AE1F8483938F80C" box="[1015,1051,1933,1959]" pageId="13" pageNumber="335" sensu="stricto">s.s.</taxonomicNameLabel>
is monochromatic on the cephalothorax and abdomen, without mottling. In contrast, adult 
<taxonomicName id="4C3D6188FFA74D323A0CF8743E8AF861" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[794,937,1969,1995]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="335" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA74D323A0CF8743E8AF861" box="[794,937,1969,1995]" italics="true" pageId="13" pageNumber="335">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA74D323AA7F8753928F861" bold="true" box="[945,1035,1968,1994]" pageId="13" pageNumber="335">
<taxonomicNameLabel id="A27A7B62FFA74D323AA7F8753928F861" box="[945,1035,1968,1994]" pageId="13" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is mottled on the dorsal surfaces of both the cephalothorax and abdomen.
</paragraph>
<caption id="DF424A83FFA44D313981FCAE3FB2FC0C" ID-DOI="http://doi.org/10.5281/zenodo.5792777" ID-Zenodo-Dep="5792777" httpUri="https://zenodo.org/record/5792777/files/figure.png" pageId="14" pageNumber="336" startId="14.[151,250,875,899]" targetBox="[155,1433,181,851]" targetPageId="14">
<paragraph id="8B821A0BFFA44D313981FCAE3FB2FC0C" blockId="14.[151,1437,875,935]" pageId="14" pageNumber="336">
<emphasis id="B949C619FFA44D313981FCAE3C31FC28" bold="true" box="[151,274,875,899]" pageId="14" pageNumber="336">FIGURE 8.</emphasis>
Intraspecific variation in 
<taxonomicName id="4C3D6188FFA44D313B01FCA93FBBFC28" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[535,664,876,899]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313B01FCA93FBBFC28" box="[535,664,876,899]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313B88FCAE3FCDFC28" bold="true" box="[670,750,875,899]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313B88FCAE3FCDFC28" box="[670,750,875,899]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs most notably in the shape of the rostrum. A, Nolichucky and Cane Rivers; B, French Broad and Pigeon Rivers.
</paragraph>
</caption>
<paragraph id="8B821A0BFFA44D3139D1FC133EEAFAA4" blockId="14.[151,1437,981,2016]" pageId="14" pageNumber="336">
In both morphology and color pattern 
<taxonomicName id="4C3D6188FFA44D313B61FC133E24FC44" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[631,775,982,1008]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313B61FC133E24FC44" box="[631,775,982,1008]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313A19FC103E4AFC44" bold="true" box="[783,873,981,1007]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313A19FC103E4AFC44" box="[783,873,981,1007]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to undescribed members of the 
<taxonomicName id="4C3D6188FFA44D313C95FC133DD5FBB8" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B949C619FFA44D313C95FC133DD5FBB8" italics="true" pageId="14" pageNumber="336">C. robustus</emphasis>
</taxonomicName>
complex that occur in the greater Toe (upper Nolichucky) and Watauga (Holston) watersheds in 
<collectingRegion id="49F9D4E9FFA44D313C32FC3C3DF1FB9C" country="United States of America" name="North Carolina" pageId="14" pageNumber="336">North Carolina</collectingRegion>
, and 
<collectingRegion id="49F9D4E9FFA44D31381BFBDB3CA0FB93" box="[269,387,1054,1080]" country="United States of America" name="Tennessee" pageId="14" pageNumber="336">Tennessee</collectingRegion>
, 
<collectingRegion id="49F9D4E9FFA44D313899FBD83F18FB9C" box="[399,571,1053,1079]" country="United States of America" name="North Carolina" pageId="14" pageNumber="336">North Carolina</collectingRegion>
, and 
<collectingRegion id="49F9D4E9FFA44D313B60FBD83FF7FB9C" box="[630,724,1053,1079]" country="United States of America" name="Virginia" pageId="14" pageNumber="336">Virginia</collectingRegion>
, respectively. This is unsurprising given the close phylogenetic relationships among these taxa and the similar habitats in which each occurs. Whereas the chela of 
<taxonomicName id="4C3D6188FFA44D313C1DFB8738BFFBF0" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1291,1436,1090,1116]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313C1DFB8738BFFBF0" box="[1291,1436,1090,1116]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313981FBA03DD2FBD4" bold="true" box="[151,241,1125,1151]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313981FBA03DD2FBD4" box="[151,241,1125,1151]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lacks subpalmar tubercles, and individuals exhibit a wide rostrum, 
<taxonomicName id="4C3D6188FFA44D313AFEFBA339A8FBD4" box="[1000,1163,1125,1152]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B949C619FFA44D313AFEFBA33ED8FB2B" box="[1000,1019,1126,1152]" italics="true" pageId="14" pageNumber="336">C</emphasis>
. cf. 
<emphasis id="B949C619FFA44D313D3AFBA039A8FBD4" box="[1068,1163,1125,1151]" italics="true" pageId="14" pageNumber="336">robustus</emphasis>
</taxonomicName>
from the Toe watershed exhibit subpalmar tubercles on the chela, and have a noticeably more acuminate rostrum. Similarly, 
<taxonomicName id="4C3D6188FFA44D313C1AFB4F38BFFB08" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1292,1436,1162,1188]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313C1AFB4F38BFFB08" box="[1292,1436,1162,1188]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313981FB683DD2FB6C" bold="true" box="[151,241,1197,1223]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313981FB683DD2FB6C" box="[151,241,1197,1223]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C3D6188FFA44D313892FB6B3F0BFB6C" box="[388,552,1197,1224]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B949C619FFA44D313892FB6B3CBDFB63" box="[388,414,1198,1224]" italics="true" pageId="14" pageNumber="336">C.</emphasis>
cf. 
<emphasis id="B949C619FFA44D3138DFFB683F0BFB6C" box="[457,552,1197,1223]" italics="true" pageId="14" pageNumber="336">robustus</emphasis>
</taxonomicName>
from the Watauga watershed in both rostrum and chela morphology, as well as coloration. The rostrum of 
<taxonomicName id="4C3D6188FFA44D3138D2FB173F70FB40" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[452,595,1234,1260]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D3138D2FB173F70FB40" box="[452,595,1234,1260]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313B4CFB143F97FB40" bold="true" box="[602,692,1233,1259]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313B4CFB143F97FB40" box="[602,692,1233,1259]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is proportionately longer relative to total carapace length than that of the Watauga 
<taxonomicName id="4C3D6188FFA44D313850FB333CC9FAA4" box="[326,490,1269,1296]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B949C619FFA44D313850FB333C43FABB" box="[326,352,1270,1296]" italics="true" pageId="14" pageNumber="336">C.</emphasis>
cf. 
<emphasis id="B949C619FFA44D31389DFB303CC9FAA4" box="[395,490,1269,1295]" italics="true" pageId="14" pageNumber="336">robustus</emphasis>
</taxonomicName>
, and the chela of the former is less rotund.
</paragraph>
<paragraph id="8B821A0BFFA44D3139D1FADC3F5EFA4C" blockId="14.[151,1437,981,2016]" pageId="14" pageNumber="336">
<taxonomicName id="4C3D6188FFA44D3139D1FADC3C85FA98" authorityName="Prins" authorityYear="1968" box="[199,422,1305,1331]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="reburrus">
<emphasis id="B949C619FFA44D3139D1FADC3C85FA98" box="[199,422,1305,1331]" italics="true" pageId="14" pageNumber="336">Cambarus reburrus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D6188FFA44D3138F4FADF3FA6FA98" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[482,645,1305,1332]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D3138F4FADF3CDFFA9F" box="[482,508,1306,1332]" italics="true" pageId="14" pageNumber="336">C.</emphasis>
cf. 
<emphasis id="B949C619FFA44D313B3DFADC3FA6FA98" box="[555,645,1305,1331]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
both occur in the French Broad, and Pigeon, French Broad and Nolichucky River watersheds, respectively. 
<taxonomicName id="4C3D6188FFA44D313B7DFAF83E69FAFC" authorityName="Prins" authorityYear="1968" box="[619,842,1341,1367]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="reburrus">
<emphasis id="B949C619FFA44D313B7DFAF83E69FAFC" box="[619,842,1341,1367]" italics="true" pageId="14" pageNumber="336">Cambarus reburrus</emphasis>
</taxonomicName>
is readily differentiated from 
<taxonomicName id="4C3D6188FFA44D313DB0FAFB3814FAFC" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1190,1335,1342,1368]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313DB0FAFB3814FAFC" box="[1190,1335,1342,1368]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313C56FAF838BFFAFC" bold="true" box="[1344,1436,1341,1367]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313C56FAF838BFFAFC" box="[1344,1436,1341,1367]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by characters of the rostrum, chela, and carapace. 
<taxonomicName id="4C3D6188FFA44D313BC8FAA43E9EFAD0" authorityName="Prins" authorityYear="1968" box="[734,957,1377,1403]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="reburrus">
<emphasis id="B949C619FFA44D313BC8FAA43E9EFAD0" box="[734,957,1377,1403]" italics="true" pageId="14" pageNumber="336">Cambarus reburrus</emphasis>
</taxonomicName>
possesses a narrow acuminate rostrum, a prominent cervical spine, and has a narrow chela that typically terminates in red or orange coloration. In contrast, 
<taxonomicName id="4C3D6188FFA44D313981FA6F3C05FA68" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[151,294,1450,1476]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313981FA6F3C05FA68" box="[151,294,1450,1476]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D31383BFA6C3CA5FA68" bold="true" box="[301,390,1449,1475]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D31383BFA6C3CA5FA68" box="[301,390,1449,1475]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
typically has a broad non-acuminate rostrum, a robust chela that does not terminate in orange or red coloration, and lacks cervical spines.
</paragraph>
<paragraph id="8B821A0BFFA44D3139D1FA343FD2F874" blockId="14.[151,1437,981,2016]" pageId="14" pageNumber="336">
<taxonomicName id="4C3D6188FFA44D3139D1FA343CE7F9A0" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[199,452,1521,1547]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D3139D1FA343C1FF9A0" box="[199,316,1521,1547]" italics="true" pageId="14" pageNumber="336">Cambarus</emphasis>
cf. 
<emphasis id="B949C619FFA44D31387CFA343CE7F9A0" box="[362,452,1521,1547]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
in western 
<collectingRegion id="49F9D4E9FFA44D313B5DFA343FD6F9A0" box="[587,757,1521,1547]" country="United States of America" name="North Carolina" pageId="14" pageNumber="336">North Carolina</collectingRegion>
and eastern 
<collectingRegion id="49F9D4E9FFA44D313A90FA373EDAF9A7" box="[902,1017,1522,1548]" country="United States of America" name="Tennessee" pageId="14" pageNumber="336">Tennessee</collectingRegion>
has frequently been misidentified as 
<taxonomicName id="4C3D6188FFA44D313981F9D33C19F984" box="[151,314,1557,1584]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B949C619FFA44D313981F9D33D92F99B" box="[151,177,1558,1584]" italics="true" pageId="14" pageNumber="336">C.</emphasis>
cf. 
<emphasis id="B949C619FFA44D3139CDF9D03C19F984" box="[219,314,1557,1583]" italics="true" pageId="14" pageNumber="336">robustus</emphasis>
</taxonomicName>
due to presence of two rows of tubercles on the mesial margin of the palm in both taxa—a character typically used to identify members of the 
<taxonomicName id="4C3D6188FFA44D313B67F9FF3FD2F9F8" box="[625,753,1593,1620]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B949C619FFA44D313B67F9FF3FD2F9F8" box="[625,753,1593,1620]" italics="true" pageId="14" pageNumber="336">C. robustus</emphasis>
</taxonomicName>
complex—and color pattern, which is often heavily mottled with both species. 
<taxonomicName id="4C3D6188FFA44D313878F9983F48F9DC" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[366,619,1629,1655]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D313878F9983CC0F9DC" box="[366,483,1629,1655]" italics="true" pageId="14" pageNumber="336">Cambarus</emphasis>
cf. 
<emphasis id="B949C619FFA44D313B07F9983F48F9DC" box="[529,619,1629,1655]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D6188FFA44D313BB0F99B3E15F9DC" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[678,822,1630,1656]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313BB0F99B3E15F9DC" box="[678,822,1630,1656]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313A56F9983EB8F9DC" bold="true" box="[832,923,1629,1655]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313A56F9983EB8F9DC" box="[832,923,1629,1655]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from each other by the overall shape of the chelae, chela morphology, and rostrum morphology. The chela of 
<taxonomicName id="4C3D6188FFA44D313D59F9443864F930" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1103,1351,1665,1691]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D313D59F94439E7F930" box="[1103,1220,1665,1691]" italics="true" pageId="14" pageNumber="336">Cambarus</emphasis>
cf. 
<emphasis id="B949C619FFA44D313DFBF9443864F930" box="[1261,1351,1665,1691]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
is truncated and subrectangular whereas those of 
<taxonomicName id="4C3D6188FFA44D313B60F9633E27F914" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[630,772,1702,1728]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313B60F9633E27F914" box="[630,772,1702,1728]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313A1DF9603E47F914" bold="true" box="[779,868,1701,1727]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313A1DF9603E47F914" box="[779,868,1701,1727]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is more elongate and range from subrectangular to subtriangular. Tubercles on the mesial margin of the palm in 
<taxonomicName id="4C3D6188FFA44D313A5BF90F3ECEF948" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[845,1005,1737,1764]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D313A5BF90F3E45F94F" box="[845,870,1738,1764]" italics="true" pageId="14" pageNumber="336">C.</emphasis>
cf. 
<emphasis id="B949C619FFA44D313A85F90C3ECEF948" box="[915,1005,1737,1763]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
are usually arranged in a manner that gives the appearance of two disorganized rows, whereas in 
<taxonomicName id="4C3D6188FFA44D313A2BF92B3EEEF8AC" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[829,973,1774,1800]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313A2BF92B3EEEF8AC" box="[829,973,1774,1800]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313AC0F9283912F8AC" bold="true" box="[982,1073,1773,1799]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313AC0F9283912F8AC" box="[982,1073,1773,1799]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
they consistently form an organized double row. 
<taxonomicName id="4C3D6188FFA44D313870F8D43F43F880" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[358,608,1809,1835]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D313870F8D43CF8F880" box="[358,475,1809,1835]" italics="true" pageId="14" pageNumber="336">Cambarus</emphasis>
cf. 
<emphasis id="B949C619FFA44D313B10F8D43F43F880" box="[518,608,1809,1835]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
exhibits two pronounced subpalmar tubercles that are directly adjacent to each other; subpalmar tubercles very rarely are found on the chela of 
<taxonomicName id="4C3D6188FFA44D313ABEF8F3391BF8E4" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[936,1080,1846,1872]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313ABEF8F3391BF8E4" box="[936,1080,1846,1872]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313D56F8F039B9F8E4" bold="true" box="[1088,1178,1845,1871]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313D56F8F039B9F8E4" box="[1088,1178,1845,1871]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The rostral margins of 
<taxonomicName id="4C3D6188FFA44D313981F89C3CB2F8D8" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[151,401,1881,1907]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D313981F89C3C2FF8D8" box="[151,268,1881,1907]" italics="true" pageId="14" pageNumber="336">Cambarus</emphasis>
cf. 
<emphasis id="B949C619FFA44D313821F89C3CB2F8D8" box="[311,401,1881,1907]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
are thickened, as in 
<taxonomicName id="4C3D6188FFA44D313B6FF89F3E2BF8D8" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[633,776,1882,1908]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313B6FF89F3E2BF8D8" box="[633,776,1882,1908]" italics="true" pageId="14" pageNumber="336">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313A06F89C3E4AF8D8" bold="true" box="[784,873,1881,1907]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313A06F89C3E4AF8D8" box="[784,873,1881,1907]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, although are continuous in thickness to the distal point of the acumen; in contrast, the rostral margins of 
<taxonomicName id="4C3D6188FFA44D313BEDF8B83EC7F83C" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[763,996,1917,1943]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313BEDF8B83EC7F83C" box="[763,996,1917,1943]" italics="true" pageId="14" pageNumber="336">Cambarus ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313AFDF8B83967F83C" bold="true" box="[1003,1092,1917,1943]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313AFDF8B83967F83C" box="[1003,1092,1917,1943]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are noticeably thicker than the margins of the acumen. In addition, the rostrum of 
<taxonomicName id="4C3D6188FFA44D313BC4F8673E53F810" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[722,880,1953,1980]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis id="B949C619FFA44D313BC4F8673FC6F817" box="[722,741,1954,1980]" italics="true" pageId="14" pageNumber="336">C</emphasis>
. cf. 
<emphasis id="B949C619FFA44D313A00F8643E53F810" box="[790,880,1953,1979]" italics="true" pageId="14" pageNumber="336">bartonii</emphasis>
</taxonomicName>
is shorter and more truncate in comparison to the prominent rostrum exhibited by 
<taxonomicName id="4C3D6188FFA44D313B14F8033FB3F874" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[514,656,1990,2016]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="336" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA44D313B14F8033F36F84B" box="[514,533,1990,2016]" italics="true" pageId="14" pageNumber="336">C</emphasis>
. 
<emphasis id="B949C619FFA44D313B34F8033FB3F874" box="[546,656,1990,2015]" italics="true" pageId="14" pageNumber="336">ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA44D313B8EF8003FD2F874" bold="true" box="[664,753,1989,2015]" pageId="14" pageNumber="336">
<taxonomicNameLabel id="A27A7B62FFA44D313B8EF8003FD2F874" box="[664,753,1989,2015]" pageId="14" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B821A0BFFA54D3039D1FF533F81FEEB" blockId="15.[151,1437,150,500]" pageId="15" pageNumber="337">
<emphasis id="B949C619FFA54D3039D1FF533F57FF1B" box="[199,628,150,177]" italics="true" pageId="15" pageNumber="337">
<emphasis id="B949C619FFA54D3039D1FF533CA3FF1A" bold="true" box="[199,384,150,177]" italics="true" pageId="15" pageNumber="337">Common name.</emphasis>
<taxonomicName id="4C3D6188FFA54D30389FFF533F57FF1B" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[393,628,150,176]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="337" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">Cambarus ectopistes</taxonomicName>
</emphasis>
<emphasis id="B949C619FFA54D303B6BFF533FFAFF1B" bold="true" box="[637,729,150,176]" pageId="15" pageNumber="337">
<taxonomicNameLabel id="A27A7B62FFA54D303B6BFF533FFAFF1B" box="[637,729,150,176]" pageId="15" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was identified as a putative unique taxon nearly 30 years ago by the late Dr. John E. Cooper (former Curator of Crustaceans at the NCSM) and Roger Thoma, both of whom believed it to be narrowly endemic, restricted to Cataloochee Creek and neighboring tributaries of the Pigeon River. The crayfish has since been colloquially referred to as the “Cataloochee morph.” We therefore suggest Cataloochee Crayfish as the common name for this species.
</paragraph>
</subSubSection>
<subSubSection id="C3274980FFA54D3039D1FE8F3F38FE5F" pageId="15" pageNumber="337" type="etymology">
<paragraph id="8B821A0BFFA54D3039D1FE8F3F38FE5F" blockId="15.[151,1437,150,500]" pageId="15" pageNumber="337">
<emphasis id="B949C619FFA54D3039D1FE8F3C6BFECE" bold="true" box="[199,328,330,357]" italics="true" pageId="15" pageNumber="337">Etymology.</emphasis>
The species epithet—as a noun—is derived from the genus name of the extinct passenger pigeon, 
<emphasis id="B949C619FFA54D303981FEAA3CB5FE23" box="[151,406,367,392]" italics="true" pageId="15" pageNumber="337">Ectopistes migratorius</emphasis>
(Linnaeus, 1766), for which the Pigeon River is named. The Greek word 
<taxonomicName id="4C3D6188FFA54D303DC8FEAA386FFE23" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1246,1356,367,392]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="337" phylum="Arthropoda" rank="species" species="ectopistes">
<emphasis id="B949C619FFA54D303DC8FEAA386FFE23" box="[1246,1356,367,392]" italics="true" pageId="15" pageNumber="337">ectopistes</emphasis>
</taxonomicName>
means “a wanderer” (
<bibRefCitation id="EFAC67FAFFA54D30382FFE573CEEFE06" author="Choate, E. A." box="[313,461,402,429]" pageId="15" pageNumber="337" refId="ref10862" refString="Choate, E. A. (1985) The dictionary of American bird names, revised edition. The Harvard Common Press, Boston, Massachusetts." type="book" year="1985">Choate 1985</bibRefCitation>
). We chose this epithet based on the geographic association, in part, of 
<taxonomicName id="4C3D6188FFA54D303DF3FE563857FE07" authority="Loughman &amp; Williams, 2021" authorityName="Loughman &amp; Williams" authorityYear="2021" box="[1253,1396,403,429]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="337" phylum="Arthropoda" rank="species" species="ectopistes" status="sp. nov.">
<emphasis id="B949C619FFA54D303DF3FE563857FE07" box="[1253,1396,403,429]" italics="true" pageId="15" pageNumber="337">C. ectopistes</emphasis>
</taxonomicName>
<emphasis id="B949C619FFA54D303C6DFE573DEBFE7B" bold="true" pageId="15" pageNumber="337">
<taxonomicNameLabel id="A27A7B62FFA54D303C6DFE573DEBFE7B" pageId="15" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with the Pigeon River, and also to emphasize the need for broader education about the conservation concerns and needs of crayfishes in general.
</paragraph>
</subSubSection>
</treatment>
</document>