321 lines
36 KiB
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321 lines
36 KiB
XML
<document ID-CLB-Dataset="28281" ID-DOI="http://dx.doi.org/10.3897/phytokeys.114.28799" ID-GBIF-Dataset="175ade4f-e320-4f41-ab85-7811f4147be4" ID-PMC="PMC6323114" ID-Pensoft-Pub="1314-2003-114-123" ID-Pensoft-UUID="CB70FFE82A46DE6DFFEBFFF8F43A6230" ID-PubMed="30627045" ID-Zenodo-Dep="2533527" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2003-114-123" ModsDocOrigin="PhytoKeys " ModsDocTitle="Revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Peru: classification, phylogeny, and a new species, M.romaschenkoi" checkinTime="1553125316505" checkinUser="pensoft" docAuthor="Peterson, Paul M., Vega, Isidoro Sanchez, Romaschenko, Konstantin, Giraldo-Canas, Diego & Rodriguez, Nancy F. Refulio" docDate="2018" docId="02D4E5D47361A3A950391441C6A017FA" docLanguage="en" docName="PhytoKeys 114: 123-206" docOrigin="PhytoKeys 114" docSource="http://dx.doi.org/10.3897/phytokeys.114.28799" docTitle="Muhlenbergia phalaroides P. M. Peterson, Caldasia 31 (2): 294 - 296, f. 7 A-B. 2009" docType="treatment" docVersion="8" id="CB70FFE82A46DE6DFFEBFFF8F43A6230" lastPageNumber="173" masterDocId="CB70FFE82A46DE6DFFEBFFF8F43A6230" masterDocTitle="Revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Peru: classification, phylogeny, and a new species, M. romaschenkoi" masterLastPageNumber="206" masterPageNumber="123" pageNumber="170" updateTime="1732930826153" updateUser="ExternalLinkService">
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<mods:titleInfo id="DDF7AF8EDFBBCBEBB5D8CE53E932B1FD">
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<mods:title id="CF050C1C37E96094132F0BC6C0895E2E">Revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Peru: classification, phylogeny, and a new species, M. romaschenkoi</mods:title>
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<mods:namePart id="63B0197D2995512469F5BF98BC851F3E">Peterson, Paul M.</mods:namePart>
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<mods:roleTerm id="74E9003CCA458CCD4D9EF18923C4ECC5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="16A847A0446507B9092A5758FCA885B0">Vega, Isidoro Sanchez</mods:namePart>
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</mods:name>
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<mods:name id="8B83FB97CCEF7542D40CB163DACC20B2" type="personal">
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<mods:roleTerm id="8C4035397BE14B58F6CF6AD892906691">Author</mods:roleTerm>
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<mods:namePart id="287C1B1D818C7196A70EF8BCC184E1C7">Romaschenko, Konstantin</mods:namePart>
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<mods:namePart id="5783BC09F4D1E0B3530CCE4FEF365D03">Giraldo-Canas, Diego</mods:namePart>
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<mods:namePart id="AD7075B77E30065551E08154FE94CBD6">Rodriguez, Nancy F. Refulio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="67DA14FD3DE16247AC3727090CBDE05C">text</mods:typeOfResource>
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<mods:title id="D9D33F7850709E9D56C24CA91373404D">PhytoKeys</mods:title>
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<mods:date id="3D925A7121240FF8788C21200904BB9A">2018</mods:date>
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<mods:detail id="1D188042EAE35B9A5AED3F42C6FF368A" type="volume">
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<mods:number id="99ABCE61CCBF4D19999128A8B7D6DDCE">114</mods:number>
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<mods:url id="16C94A9B23337A8D76DB3AE43000210A">http://dx.doi.org/10.3897/phytokeys.114.28799</mods:url>
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</mods:location>
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<mods:classification id="D9ACBDC85341FE04F9D64D06355A58A9">journal article</mods:classification>
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<mods:identifier id="70C734B66AEF0AACDA8AB80142FF5228" type="DOI">http://dx.doi.org/10.3897/phytokeys.114.28799</mods:identifier>
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<mods:identifier id="D2F9EE52A8D98282DFA95DC6BCA1353F" type="Pensoft-Pub">1314-2003-114-123</mods:identifier>
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<mods:identifier id="1C291C46675F2255F432A1D4E4692CD2" type="Zenodo-Dep">2533527</mods:identifier>
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<treatment id="02D4E5D47361A3A950391441C6A017FA" ID-GBIF-Taxon="154472464" LSID="urn:lsid:plazi:treatment:02D4E5D47361A3A950391441C6A017FA" httpUri="http://treatment.plazi.org/id/02D4E5D47361A3A950391441C6A017FA" lastPageId="50" lastPageNumber="173" pageId="47" pageNumber="170" scope_family="Poaceae">
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<subSubSection id="1D3F154EC9A43BD071DA948CF93D8CA8" pageId="47" pageNumber="170" type="nomenclature">
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<paragraph id="4C63842419CCEA5DFCB6500522AAC1BC" pageId="47" pageNumber="170">
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16.
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<taxonomicName id="0006965DB23AF77889D4D62F10079ADB" ID-CoL="44JLD" LSID="9C5ABA17-7384-5F3F-B610-7F25B42979CE" authority="(Kunth) P. M. Peterson, Caldasia 31 (2): 294 - 296, f. 7 A-B. 2009." class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia phalaroides" order="Poales" pageId="47" pageNumber="170" phylum="Tracheophyta" rank="species" species="phalaroides">
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Muhlenbergia phalaroides (Kunth) P.M. Peterson, Caldasia 31(2): 294-296, f. 7
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<normalizedToken id="A070181D1CD22D32C61F4E4A8915FF33" originalValue="A–B">A-B</normalizedToken>
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. 2009.
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</taxonomicName>
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<figureCitation id="07E506E837DB5514E2B86F9725BD7DF7" captionStart="Figure 11" captionStartId="F12" captionText="Figure 11. A-D Muhlenbergia peruviana (P. Beauv.) Steud A habit B ligule C glumes D floret E, F Muhlenbergia phalaroides (Kunth) P. M. Peterson E panicle F spikelet. Drawings A-D from Peterson and Annable (1991) drawn from P. M. Peterson & C. R. Annable 4739 (WS) E, F from Giraldo-Canas and Peterson (2009) drawn from S. Laegaard 71419 (AAU)." figureDoi="10.3897/phytokeys.114.28799.figure11" httpUri="https://binary.pensoft.net/fig/254052" pageId="47" pageNumber="170">Fig. 11E, F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6DDB531D6DCEB2B6F283C7E45BE505CF" lastPageId="49" lastPageNumber="172" pageId="48" pageNumber="171" type="reference_group">
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<paragraph id="45F163F0B5646D0EF51CEF5467CDB282" pageId="48" pageNumber="171">
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<taxonomicName id="E670F56AC7F6B5D40C8AD9445A3090A4" class="Liliopsida" family="Poaceae" genus="Lycurus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycurus phalaroides" order="Poales" pageId="48" pageNumber="171" phylum="Tracheophyta" rank="species" species="phalaroides">
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<pageBreakToken id="C1DCCB4F3C9FB94FB671BB6C7FFC1DFC" pageId="48" pageNumber="171" start="start">Lycurus</pageBreakToken>
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phalaroides
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</taxonomicName>
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Kunth, Nov. Gen. Sp. (quarto ed.) 1: 142. 1815 (1816). Type: Mexico,
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<normalizedToken id="9F8FB63EBBAAAA713846130308EAE410" originalValue="Michoacán">Michoacan</normalizedToken>
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, near Valladolid, Alberca de Palangeo and Patzcuaro, Sep,
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<emphasis id="05E647D2BC6AB91CBD9C8E399C858DBC" italics="true" pageId="48" pageNumber="171">F.W.H.A. Humboldt & A.J.A. Bonpland s.n.</emphasis>
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(holotype: P-00669405 [image!]; isotypes: B-W-1630, BM!, BAA-1530!, US-91988 fragm. ex P-BONPL!, US-610837 fragm. ex LE-TRIN!).
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</paragraph>
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<paragraph id="E8154F396543AE9B7E7EFFED1CCE70F8" lastPageId="49" lastPageNumber="172" pageId="48" pageNumber="171">
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<taxonomicName id="88CF883A1C0DBF5D1F62E4AD622D0F73" authorityName="Vasey ex Beal" authorityYear="1896" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia lycuroides" order="Poales" pageId="48" pageNumber="171" phylum="Tracheophyta" rank="species" species="lycuroides">Muhlenbergia lycuroides</taxonomicName>
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<pageBreakToken id="A0D527773CF7E1B3A478722751761939" pageId="49" pageNumber="172" start="start">Vasey</pageBreakToken>
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ex Beal, Grass. N. Amer. 2: 239. 1896. Type: Mexico, Jalisco, Guadalajara, Jul-Oct 1886,
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<emphasis id="17451568D6519754E8C4D318E47AA70E" italics="true" pageId="49" pageNumber="172">E. Palmer 489</emphasis>
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(holotype: MSC; isotypes: GH-00023916 [image!], LE, MEXU, MO-2972929!, NDG-07247 [image!], NY, P-00644181 [image!], P-00644182 [image!], S14-29628 [image!], US-822925!, US-81642!, YU-000898 [image!]).
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</paragraph>
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<paragraph id="79BA8A414DD3605ED34993FCE436A288" pageId="49" pageNumber="172">
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<taxonomicName id="BB658B00E2744B44212236664EEACAF1" authorityName="Scribn ex Beal" authorityYear="1896" class="Liliopsida" family="Poaceae" genus="Lycurus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Lycurus phleoides var. brevifolius" order="Poales" pageId="49" pageNumber="172" phylum="Tracheophyta" rank="variety" species="phleoides" variety="brevifolius">Lycurus phleoides var. brevifolius</taxonomicName>
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Scribn. ex Beal, Grass. N. Amer. 2: 271. 1896. Type: Mexico, Jalisco, plains of Guadalajara, 23 Oct 1889,
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<emphasis id="F45722CB07965B4FC20E8556E3C38096" italics="true" pageId="49" pageNumber="172">C.G. Pringle 2470</emphasis>
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(lectotype: MSC, designated by C. Reeder, Phytologia 57(4): 288. 1985; isolectotypes: BAA!, GH, MEXU, MO-2972926!, NY!, P-00644183 [image!], P-00644184 [image!], US-996049!, W-18900000580 [image!], W-19160029092 [image!]).
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</paragraph>
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</subSubSection>
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<subSubSection id="E57A97128314A140D0E000FFFFBDA0A8" pageId="49" pageNumber="172" type="description">
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<paragraph id="BBBB16662D4CEC60EBA2E7B39BF9729F" pageId="49" pageNumber="172">Description.</paragraph>
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<paragraph id="A1F32AA98A600AD9350FF7BC00F245DB" pageId="49" pageNumber="172">
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<taxonomicName id="C1028EF33D58260AC40E41326DE2812E" genus="Perennials" lsidName="Perennials" pageId="49" pageNumber="172" rank="genus">
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<emphasis id="772C7E1C86C2B0F71853703E9D2DD4E6" italics="true" pageId="49" pageNumber="172">Perennials</emphasis>
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</taxonomicName>
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, intricately branched near base.
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<emphasis id="EF74B35E8C8C149DEC177B80B5B87EE6" italics="true" pageId="49" pageNumber="172">Culms</emphasis>
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10-30 cm tall, erect, mostly glabrous, usually decumbent and sprawling below, bent at the pubescent to short pilose nodes;
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<emphasis id="ADBB79E270C6909FC2203D3B32C76DBC" italics="true" pageId="49" pageNumber="172">internodes</emphasis>
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0.4-10 (-15) cm long, pubescent to short pilose.
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<emphasis id="7F85E9157F6EFD91C83DC26786E8741C" italics="true" pageId="49" pageNumber="172">Leaf sheaths</emphasis>
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much shorter than the internodes above, hyaline near the margins, pilose near summit;
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<emphasis id="42E69B07FF8456D63F2582E75E0BEA13" italics="true" pageId="49" pageNumber="172">ligules</emphasis>
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0.4-1 mm long, membranous, apex truncate to deltoid, often erose and lacerate;
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<emphasis id="E1E7886E05F96D9EDDA0E903ADD73050" italics="true" pageId="49" pageNumber="172">blades</emphasis>
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0.5-6.5 cm long, 0.5-1.2 mm wide, shorter near the base of culms, flat, folded or loosely involute, lanate above and glabrous or with scattered, short appressed hairs below, margins whitish-thickened, apex navicular, occasionally with a short seta, seta usually less than 2 mm long.
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<emphasis id="A8CB61278C25CE5A396BA5AFBA765B8C" italics="true" pageId="49" pageNumber="172">Panicles</emphasis>
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1.5-6.5 cm long, 3-8 mm wide, spiciform and spike-like, densely flowered, often interrupted below with only a few spikelets, terminal or axillary;
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<emphasis id="0D93DEDAE770761C217EFF9400DD6598" italics="true" pageId="49" pageNumber="172">rachis</emphasis>
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lanate to hispid, the short hairs antrose or appressed;
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<emphasis id="319F58590DBF5121BC96F319593805D7" italics="true" pageId="49" pageNumber="172">branches</emphasis>
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1.5-7 mm long, very short, the spiklets usually in pairs, rarely 1 or 3 per terminal branch, when in pairs the lower short-pedicelled spikelet perfect, staminate or sterile and the upper longer-pedicelled spikelet usually perfect;
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<emphasis id="FC4C558E3280B6E5F924C6DB88750237" italics="true" pageId="49" pageNumber="172">pedicels</emphasis>
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0.3-1.4 mm long;
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<emphasis id="CBEE4517305B7B699E0EED68A41420E3" italics="true" pageId="49" pageNumber="172">disarticulation</emphasis>
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usually at the base of the pedicel, each spikelet falling as a unit leaving a small cup-like tip.
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<emphasis id="609ABDEDE3E4B8A015D5AA161FBFE24F" italics="true" pageId="49" pageNumber="172">Spikelets</emphasis>
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3-4 mm long, stramineous with plumbeous mottles, sometimes additionally with purplish mottles;
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<emphasis id="D284E8A713DC261DE008F44B1846D24B" italics="true" pageId="49" pageNumber="172">glumes</emphasis>
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1-2.1 mm long, shorter than the lemma, subequal, 1-3-veined;
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<emphasis id="1BF2C53EC3D0B07E6F29819E74068E45" italics="true" pageId="49" pageNumber="172">lower glumes</emphasis>
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commonly 2 or 3-veined, usually 2-awned, ocassionally 1 or 3 awned, the awns 1-3 mm long, equal or subequal, scabrous, recurved;
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<emphasis id="FD0B0363D62E052D1B66B49ABCE395F0" italics="true" pageId="49" pageNumber="172">upper glumes</emphasis>
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commonly 1-veined, usually 1-awned, the awns 1-2.5 mm long;
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<emphasis id="C5FD0CC0B31EF26FBCBF6B0E2C429ADC" italics="true" pageId="49" pageNumber="172">lemmas</emphasis>
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3-4 mm long, narrowly lanceolate, 3-veined, margins hirsute to lanate and occasionally the lower
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<normalizedToken id="8CA90D665109EF76E44C0A960F7F3D7B" originalValue="½">1/2</normalizedToken>
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sparsely hairy, the hairs 0.1-0.3 mm long, apex usually awned, occasionally unawned or mucronate, the awns 1-3 mm long;
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<emphasis id="E242B1D041B4CEDE71884C303DC44AD2" italics="true" pageId="49" pageNumber="172">paleas</emphasis>
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2.8-3.8 mm long, hairy between the veins, the veins occasionally extending as mucros; anthers 1.3-2 mm long, yellowish.
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<emphasis id="D603A309E14496C8105D440741A67FFA" italics="true" pageId="49" pageNumber="172">Caryopses</emphasis>
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1.7-2 mm long, fusiform, brownish.
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</paragraph>
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</subSubSection>
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<subSubSection id="BEF9A321FBAD324EC234A5137342408F" pageId="49" pageNumber="172" type="distribution">
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<paragraph id="830BB5C5EA01CED88A77258088E47CE3" pageId="49" pageNumber="172">Distribution.</paragraph>
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<paragraph id="CAEA1A1E75878B82DE3C82BDD297F8C3" pageId="49" pageNumber="172">
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<taxonomicName id="CED1B660FC4E79CBF4E562DEFD6EA46A" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia phalaroides" order="Poales" pageId="49" pageNumber="172" phylum="Tracheophyta" rank="species" species="phalaroides">
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<emphasis id="92C6C29DE9FCF0209AF1F5B7F64352F1" italics="true" pageId="49" pageNumber="172">Muhlenbergia phalaroides</emphasis>
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</taxonomicName>
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ranges from Mexico to South America where it is found in Argentina, Bolivia, Columbia, Ecuador and Peru (
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<bibRefCitation id="48B93DFB6B3CAF8616099B52333792FB" author="Reeder, CG" journalOrPublisher="Phytologia" pageId="59" pageNumber="182" pagination="283 - 291" refId="B49" refString="Reeder, CG, 1985. The genus Lycurus (Gramineae) in North America. Phytologia 57: 283 - 291" title="The genus Lycurus (Gramineae) in North America." volume="57" year="1985">Reeder 1985</bibRefCitation>
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;
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<normalizedToken id="C0D54C35FC39E84FCB3B483B2C0D3731" originalValue="Sánchez">Sanchez</normalizedToken>
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and
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<normalizedToken id="CA8ACA6B133CEC3338753C31A9E95D9E" originalValue="Rúgolo">Rugolo</normalizedToken>
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de Agrasar 1986;
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<bibRefCitation id="5B99FF0306D5D72D11F9B7D0E7866228" author="Davidse, G" journalOrPublisher="Memoirs of the Torrey Botanical Club" pageId="58" pageNumber="181" refId="B4" refString="Davidse, G, Pohl, RW, 1994. 262. Poaceae. In: Davidse G, Sousa SM, Chater AO (Eds) Flora Mesoamericana, Volumen 6, Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Instituto de Biologia, Mexico, D.F., 184-402." title="262. Poaceae. In: Davidse G, Sousa SM, Chater AO (Eds) Flora Mesoamericana, Volumen 6, Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Instituto de Biologia, Mexico, D. F., 184 - 402." year="1994">Davidse and Pohl 1994</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="AF1616B0E1F90B1FDE662DD1514994B2" lastPageId="50" lastPageNumber="173" pageId="49" pageNumber="172" type="ecology">
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<paragraph id="E10B31F38E707BE5FDD59A9CD772D4CE" pageId="49" pageNumber="172">Ecology.</paragraph>
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<paragraph id="CA1E80B76EF6C54A5E04469792522D38" lastPageId="50" lastPageNumber="173" pageId="49" pageNumber="172">
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This species occurs in open grasslands and savannahs on steep rocky slopes flats and along disturbed irrigation canals in deep clayish-loam to sandy soils associated with
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<taxonomicName id="B7714788E933B30DA8E7284A7F71BAD0" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Baccharis" order="Asterales" pageId="49" pageNumber="172" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="ECC792CCA012B5ABED8984855AE9EA70" italics="true" pageId="49" pageNumber="172">Baccharis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F1B5B1448A07A5B224BB59BB6BAADEA8" class="Dicotyledoneae" family="Berberidaceae" genus="Berberis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Berberis" order="Ranunculales" pageId="49" pageNumber="172" phylum="Angiospermae" rank="genus">
|
||
<emphasis id="8321AC42CAB84EDE20E7B520BE08A127" italics="true" pageId="49" pageNumber="172">Berberis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="06FA9CD43C63C483B7D0F31C0EF8213C" class="Polypodiopsida" family="Pteridaceae" genus="Cheilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cheilanthes" order="Polypodiales" pageId="49" pageNumber="172" phylum="Polypodiopsida" rank="genus">
|
||
<emphasis id="8BF96FBA529D0FFD478119B80B212007" italics="true" pageId="49" pageNumber="172">Cheilanthes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="357403789AC023D901B75694CEA08D2A" class="Magnoliopsida" family="Rhamnaceae" genus="Condalia" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Condalia" order="Rosales" pageId="49" pageNumber="172" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="509C4F3F2DB735B2148C67324E060D55" italics="true" pageId="49" pageNumber="172">Condalia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5A40B6724A1BA4C63428F3E8730ED70E" class="Dicotyledoneae" family="Sapindaceae" genus="Dodonaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Dodonaea viscosa" order="Sapindales" pageId="49" pageNumber="172" phylum="Angiospermae" rank="species" species="viscosa">
|
||
<emphasis id="EAF7596C4B9255680F6C70ADDE75BF8C" italics="true" pageId="49" pageNumber="172">Dodonaea viscosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6271A111BDE552F566E1B09330DD53A7" class="Equisetopsida" family="Poaceae" genus="Eragrostis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Eragrostis" order="Poales" pageId="49" pageNumber="172" phylum="Magnoliophyta" rank="genus">
|
||
<emphasis id="0BF01ACC3675E4C72C052E4E9EAFC542" italics="true" pageId="49" pageNumber="172">Eragrostis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="19C5D2BDED9093DD10142E94F27DD68A" italics="true" pageId="50" pageNumber="173">
|
||
<taxonomicName id="4CE09704056025598A5524AC3E157FE2" authorityName="Ruiz & Pav" authorityYear="1798" class="Liliopsida" family="Poaceae" genus="Jarava" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Jarava ichu" order="Poales" pageId="50" pageNumber="173" phylum="Magnoliophyta" rank="species" species="ichu">
|
||
<pageBreakToken id="FACA05FCECF5A5A432F23F3291E676E8" pageId="50" pageNumber="173" start="start">Jarava</pageBreakToken>
|
||
ichu
|
||
</taxonomicName>
|
||
<taxonomicName id="7BA5A36211F5A18490DAB3A8E0F7D77D" class="Dicotyledoneae" family="Cactaceae" genus="Opuntia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Opuntia" order="Cactales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="genus">Opuntia</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="C4638347DCAE9D754FC757BBF475605A" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia cenchroides" order="Poales" pageId="50" pageNumber="173" phylum="Tracheophyta" rank="species" species="cenchroides">
|
||
<emphasis id="664A1BE695937C24FBAA218464DEB340" italics="true" pageId="50" pageNumber="173">Muhlenbergia cenchroides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0E685FEF12B69A4F136BA17D0C56CCD0" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. rigida" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="rigida">
|
||
<emphasis id="1B36BC86D455420C0873DEDC2A42D920" italics="true" pageId="50" pageNumber="173">M. rigida</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8BD4CC7355B52BD82A8F5524A5BBCF05" class="Liliopsida" family="Poaceae" genus="Nassella" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Nassella" order="Poales" pageId="50" pageNumber="173" phylum="Magnoliophyta" rank="genus">
|
||
<emphasis id="87E55046AAE48ABE3A36A3D3369E38F2" italics="true" pageId="50" pageNumber="173">Nassella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="184FBCE1927E47C48BD3764344F5D444" class="Dicotyledoneae" family="Plantaginaceae" genus="Plantago" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Plantago" order="Plantaginales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="genus">
|
||
<emphasis id="A403D64609E5944C7B55E1644FFB64B8" italics="true" pageId="50" pageNumber="173">Plantago</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="32CF0ED095A7110C4DBCC473D434F98E" authorityName="Molina" authorityYear="1782" class="Liliopsida" family="Bromeliaceae" genus="Puya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Puya" order="Poales" pageId="50" pageNumber="173" phylum="Liliopsida" rank="genus">
|
||
<emphasis id="3E8BD901318858ADA16EDD6C8D8D5C18" italics="true" pageId="50" pageNumber="173">Puya</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="377D806553FA784DB238A8C2708438FE" class="Liliopsida" family="Poaceae" genus="Sporobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sporobolus indicus" order="Poales" pageId="50" pageNumber="173" phylum="Tracheophyta" rank="species" species="indicus">
|
||
<emphasis id="D936E133BE166F20B456D9B9B0832EEC" italics="true" pageId="50" pageNumber="173">Sporobolus indicus</emphasis>
|
||
</taxonomicName>
|
||
(L.) R. Br.; 2800-3500 m.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70C7F5B017CA3465FC1427A21FBB4DE9" pageId="50" pageNumber="173" type="comments">
|
||
<paragraph id="282105342B680C8BFE3C28AFC753AAEF" pageId="50" pageNumber="173">Comments.</paragraph>
|
||
<paragraph id="F71C275321D54A34BA42508AB86978CA" pageId="50" pageNumber="173">
|
||
<taxonomicName id="D384D799F6274AC7E00EF99AC05E8A3F" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia phalaroides" order="Poales" pageId="50" pageNumber="173" phylum="Tracheophyta" rank="species" species="phalaroides">
|
||
<emphasis id="A9C5B7CA1BF6DB212355F9D641423665" italics="true" pageId="50" pageNumber="173">Muhlenbergia phalaroides</emphasis>
|
||
</taxonomicName>
|
||
is morphologically similar to
|
||
<taxonomicName id="439CA83B02C067C1E4B5EB5E24D01E60" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. phleoides" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="phleoides">
|
||
<emphasis id="4B979FECAF349DFA5BCF65C1AFE94821" italics="true" pageId="50" pageNumber="173">M. phleoides</emphasis>
|
||
</taxonomicName>
|
||
(Kunth) Columbus, known in south-western U.S.A. and Mexico and
|
||
<taxonomicName id="D356B5494E552F663E4D2FB981D8135C" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. alopecuroides" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="alopecuroides">
|
||
<emphasis id="11E8D14BE98FAA59A53614370BA9E0A0" italics="true" pageId="50" pageNumber="173">M. alopecuroides</emphasis>
|
||
</taxonomicName>
|
||
found in south-western U.S.A., Mexico and disjunct in Argentina and Bolivia (
|
||
<bibRefCitation id="FCCD15EF0FD40CFCDB02395F185E0722" author="Davidse, G" journalOrPublisher="Memoirs of the Torrey Botanical Club" pageId="58" pageNumber="181" refId="B4" refString="Davidse, G, Pohl, RW, 1994. 262. Poaceae. In: Davidse G, Sousa SM, Chater AO (Eds) Flora Mesoamericana, Volumen 6, Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Instituto de Biologia, Mexico, D.F., 184-402." title="262. Poaceae. In: Davidse G, Sousa SM, Chater AO (Eds) Flora Mesoamericana, Volumen 6, Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Instituto de Biologia, Mexico, D. F., 184 - 402." year="1994">Davidse and Pohl 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="078BA2A0BCA38A872D925F710DD5C7D5" author="Renvoize, SA" journalOrPublisher="Phytologia" pageId="59" pageNumber="182" refId="B52" refString="Renvoize, SA, 1998. Gramineas de Bolivia. The Royal Botanic Gardens, Kew, 1-644." title="Gramineas de Bolivia. The Royal Botanic Gardens, Kew, 1 - 644." year="1998">Renvoize 1998</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="AB1A0D0402E234780AED63C5D14163BF" author="Reeder, CG" journalOrPublisher="Phytologia" pageId="59" pageNumber="182" refId="B50" refString="Reeder, CG, 2003. 17.34 Lycurus Kunth. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 2 Flora of North America North of Mexico, Vol 25. Oxford University Press, New York, 200-203." title="17.34 Lycurus Kunth. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 2 Flora of North America North of Mexico, Vol 25. Oxford University Press, New York, 200 - 203." year="2003">Reeder 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="6E14DC8E575E1F6F244BEF15338D1446" author="Peterson, PM" journalOrPublisher="Caldasia" pageId="58" pageNumber="181" refId="B29" refString="Peterson, PM, Giraldo-Canas, D, 2012. Muhlenbergia Schreb. In: Zuloaga FO, Rugolo ZE, Anton AMR (Eds) Flora Argentina, Flora Vascular de la Republica Argentina, Vol. 3, Tomo 1, Monocotyledoneae, Poaceae: Aristidoideae a Pharoideae. IBODA-IMBIV, CONICET, Buenos Aires, Argentina, 140-154." title="Muhlenbergia Schreb. In: Zuloaga FO, Rugolo ZE, Anton AMR (Eds) Flora Argentina, Flora Vascular de la Republica Argentina, Vol. 3, Tomo 1, Monocotyledoneae, Poaceae: Aristidoideae a Pharoideae. IBODA-IMBIV, CONICET, Buenos Aires, Argentina, 140 - 154." year="2012">
|
||
Peterson and
|
||
<normalizedToken id="7962631937C543BC16BD0150F972B312" originalValue="Giraldo-Cañas">Giraldo-Canas</normalizedToken>
|
||
2012
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="83F3BCD6C7DCAE3DCB7176D4781DAF91" authorityName="P. M. Peterson & Columbus" authorityYear="2008" baseAuthorityName="Griseb." class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia alopecuroides" order="Poales" pageId="50" pageNumber="173" phylum="Tracheophyta" rank="species" species="alopecuroides">
|
||
<emphasis id="76410AC29754D8DD71E94C12914FD596" italics="true" pageId="50" pageNumber="173">Muhlenbergia alopecuroides</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="BA1D5D8101318A7F34C2FA8F8140ADE4" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. phalaroides" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="phalaroides">
|
||
<emphasis id="CF6CAAA78BFDA7D250CCF392FA95E697" italics="true" pageId="50" pageNumber="173">M. phalaroides</emphasis>
|
||
</taxonomicName>
|
||
in having leaf blades with terminal seta (3-)4-7(-12) mm long and ligules(2-)3-12 mm long whereas
|
||
<taxonomicName id="6055CFB7297D093086A07F8357EB3FB2" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. phleoides" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="phleoides">
|
||
<emphasis id="4E5E8C1756E973C1AF0F39C6A2998A82" italics="true" pageId="50" pageNumber="173">M. phleoides</emphasis>
|
||
</taxonomicName>
|
||
differs in having auriculate ligules 1-2 mm long (
|
||
<bibRefCitation id="55C6C96803FA3A8F78FA638E0F1F46C0" author="Reeder, CG" journalOrPublisher="Phytologia" pageId="59" pageNumber="182" pagination="283 - 291" refId="B49" refString="Reeder, CG, 1985. The genus Lycurus (Gramineae) in North America. Phytologia 57: 283 - 291" title="The genus Lycurus (Gramineae) in North America." volume="57" year="1985">Reeder 1985</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7B0AEA472F8568F33D9EDEDD2325D92D" author="Reeder, CG" journalOrPublisher="Phytologia" pageId="59" pageNumber="182" refId="B50" refString="Reeder, CG, 2003. 17.34 Lycurus Kunth. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 2 Flora of North America North of Mexico, Vol 25. Oxford University Press, New York, 200-203." title="17.34 Lycurus Kunth. In: Barkworth ME, Capels KM, Long S, Piep MB (Eds) Magnoliophyta: Commelinidae (in part): Poaceae, part 2 Flora of North America North of Mexico, Vol 25. Oxford University Press, New York, 200 - 203." year="2003">2003</bibRefCitation>
|
||
). These morphological differences are perhaps better recognised at the subspecific level but there have been no population studies comparing these three species, other than
|
||
<bibRefCitation id="4902A52A5DF0418BAA807904ADB0ED6E" author="Peterson, PM" journalOrPublisher="Madrono" pageId="58" pageNumber="181" pagination="334 - 346" refId="B31" refString="Peterson, PM, Morrone, O, 1997. Allelic variation in the amphitropical disjunct Lycurus setosus (Poaceae: Muhlenbergiinae). Madrono 44: 334 - 346" title="Allelic variation in the amphitropical disjunct Lycurus setosus (Poaceae: Muhlenbergiinae)." volume="44" year="1997">Peterson and Morrone (1997)</bibRefCitation>
|
||
who investigated populations of only the amphitropical,
|
||
<taxonomicName id="17AC6B04B71A82605E85211AB7A71BA5" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. alocpecuroides" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="alocpecuroides">
|
||
<emphasis id="DE70A62A184F13FFAFF1E493A27CDB1E" italics="true" pageId="50" pageNumber="173">M. alocpecuroides</emphasis>
|
||
</taxonomicName>
|
||
[as
|
||
<taxonomicName id="193642F40753B22429DACD081E039B59" authorityName="C. Reeder" authorityYear="1985" baseAuthorityName="Nutt." class="Liliopsida" family="Gramineae" genus="Lycurus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycurus setosus" order="Poales" pageId="50" pageNumber="173" phylum="Tracheophyta" rank="species" species="setosus">
|
||
<emphasis id="BD02E216599663B2CE892A822F8A89CC" italics="true" pageId="50" pageNumber="173">Lycurus setosus</emphasis>
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="FD24EBED5ADD176456D635EFFBC71748" pageId="50" pageNumber="173">
|
||
<taxonomicName id="979B0D29E65CF5AC64369CE88CD01242" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia phalaroides" order="Poales" pageId="50" pageNumber="173" phylum="Tracheophyta" rank="species" species="phalaroides">
|
||
<emphasis id="A1DDDFD9AA09D42C0FA97ACC876FADA4" italics="true" pageId="50" pageNumber="173">Muhlenbergia phalaroides</emphasis>
|
||
</taxonomicName>
|
||
lies within
|
||
<taxonomicName id="465C2CDABA853A4AD6CC2280D494894C" authority="subg. Pseudosporobolus" class="Magnoliopsida" family="Poaceae" genus="M." higherTaxonomySource="GBIF" infraspecific-rank="subg." kingdom="Plantae" lsidName="M." order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="genus">M. subg. Pseudosporobolus</taxonomicName>
|
||
, aligning with
|
||
<taxonomicName id="8B2EBA1CA43708917D9894465075EEC2" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. alopecuroides" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="alopecuroides">
|
||
<emphasis id="86DB7DC1D78F3CD3D992DF2F644B3E8C" italics="true" pageId="50" pageNumber="173">M. alopecuroides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7D3E7183979EAF00E5B2B50DAD73F181" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. phleoides" order="Poales" pageId="50" pageNumber="173" phylum="Angiospermae" rank="species" species="phleoides">
|
||
<emphasis id="F3A2C1CDF8584B630ED3E6A0BD18E1C2" italics="true" pageId="50" pageNumber="173">M. phleoides</emphasis>
|
||
</taxonomicName>
|
||
(Peterson and Romaschenko, in prep.). Many members of this subgenus have narrow
|
||
<emphasis id="535550F81F5CD29889585A107BE9774E" italics="true" pageId="50" pageNumber="173">panicles</emphasis>
|
||
, plumbeous spikelets with unawned, mucronate or short-awned lemmas (
|
||
<bibRefCitation id="D097E0B30AA670460623FE49128EA94C" DOI="https://doi.org/10.3732/ajb.0900359" author="Peterson, PM" journalOrPublisher="American Journal of Botany" pageId="59" pageNumber="182" pagination="1532 - 1554" refId="B38" refString="Peterson, PM, Romaschenko, K, Johnson, G, 2010b. A phylogeny and classification of the Muhlenbergiinae (Poaceae: Chloridoideae: Cynodonteae) based on plastid and nuclear DNA sequences. American Journal of Botany 97 (9): 1532 - 1554, DOI: https://doi.org/10.3732/ajb.0900359" title="A phylogeny and classification of the Muhlenbergiinae (Poaceae: Chloridoideae: Cynodonteae) based on plastid and nuclear DNA sequences." url="https://doi.org/10.3732/ajb.0900359" volume="97" year="2010 b">Peterson et al. 2010b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAE84DAEEAC7C1F9A84837AC19ADD688" pageId="50" pageNumber="173" type="materials_examined">
|
||
<paragraph id="6D8DB56A334965C7786438F58397881C" pageId="50" pageNumber="173">Specimens examined.</paragraph>
|
||
<paragraph id="D9489BF70BBF4FC96D1F5E86472E10EF" pageId="50" pageNumber="173">
|
||
Peru.
|
||
<emphasis id="CD0D13EEB259E81F388E2EF58902B8E7" bold="true" pageId="50" pageNumber="173">Ancash</emphasis>
|
||
: 15 Mar 1947, J. Infantes Vera 1170 (LIL).
|
||
<emphasis id="C9089BBE7A57408F112E41DEDBC276EA" bold="true" pageId="50" pageNumber="173">Ayacucho</emphasis>
|
||
: Huanca Sancos, 27 km NW of Putajasa and 3 km S of Sacsamarca,
|
||
<geoCoordinate id="B95F50E8951F78F93B4DA25E4C5DAFB0" degrees="13" direction="south" minutes="57" orientation="latitude" precision="1" seconds="51.1" value="-13.964194">13°57'51.1"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E6CB89F4AA1C9B1CD682A7C2FD972A5F" degrees="74" direction="west" minutes="18" orientation="longitude" precision="1" seconds="41.5" value="-74.31153">74°18'41.5"W</geoCoordinate>
|
||
, 3650 m, 25 Feb 2002, P.M. Peterson 16277, A.A. Cano, M.I. LaTorre, A. Ramirez & D. Susanibar (US, USM).
|
||
<emphasis id="DAD9009367469B06194F1F9FDCB3ECE1" bold="true" pageId="50" pageNumber="173">Cusco</emphasis>
|
||
: Canchis, Hacienda Occobamba, 3450 m, 4 Apr 1950, C. Vargas C. 9410 (CUZ, US); Prov. Quispicanchis, Pikillacta, 3250 m, 28 Apr 1946, C. Vargas C. 6033 (CUZ, US).
|
||
<emphasis id="03355B8D742C743B925DFC86A433F2BB" bold="true" pageId="50" pageNumber="173">Huancavelica</emphasis>
|
||
: near Huancayo, 3270 m, 1 Apr 1952, O. Tovar 344 (US, USM); Sacxhahuajta, a 6 km de Conaica, 3600 m, 3 Apr 1952, O. Tovar 939 (US, USM).
|
||
<emphasis id="279D5B10A088A0B10EDAEF31D251D757" bold="true" pageId="50" pageNumber="173">
|
||
<normalizedToken id="9BAE06CBE749F256DD976D0D6289BAA9" originalValue="Junín">Junin</normalizedToken>
|
||
</emphasis>
|
||
: Huancayo, 3317 m, Mar 1943, J. Soukup 1900 (COL, LIL, US); Prov. Huancayo, Agua de las Virgenes, 3270 m, 1 Apr 1951, O. Tovar 344 (MOL, USM); Tarma, between La Oroya and La Merced, 3000 m, 24 Oct 1923, A.S. Hitchcock 22169 (US).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |