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<document ID-CLB-Dataset="263454" ID-DOI="http://dx.doi.org/10.3897/zookeys.1174.103324" ID-GBIF-Dataset="7797aa47-8d4c-45a9-a2e8-6fc406472d3c" ID-PMC="PMC10848836" ID-Pensoft-Pub="1313-2970-1174-85" ID-Pensoft-UUID="C382EDC7029755EFA1BC59D090835B37" ID-PubMed="38328308" ID-ZooBank="5C6509E838A34F6A878B08C6868E0D2A" ModsDocID="1313-2970-1174-85" checkinTime="1691559543624" checkinUser="pensoft" docAuthor="McKamey, Stuart H." docDate="2023" docId="0292BFB428F25546BEA9B09FF7E0DEB7" docLanguage="en" docName="ZooKeys 1174: 85-95" docOrigin="ZooKeys 1174" docPubDate="2023-08-08" docSource="http://dx.doi.org/10.3897/zookeys.1174.103324" docTitle="Smilidarnis duocornus McKamey 2023, sp. nov." docType="treatment" docUuid="83FF1199-024D-4397-A31F-A942B5B39E81" docUuidSource="ZooBank" docVersion="4" id="C382EDC7029755EFA1BC59D090835B37" lastPageNumber="85" masterDocId="C382EDC7029755EFA1BC59D090835B37" masterDocTitle="Three new species of the Neotropical genus Smilidarnis Andrade (Hemiptera, Membracidae)" masterLastPageNumber="95" masterPageNumber="85" pageNumber="85" updateTime="1732480995221" updateUser="ExternalLinkService">
<mods:mods id="3EE8DB259935121EF6BEB481A864EF69" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AE132BE5D4154CF080C22330C2550BB9">
<mods:title id="664ABCB473BB38F2D45B3A96B8409839">Three new species of the Neotropical genus Smilidarnis Andrade (Hemiptera, Membracidae)</mods:title>
</mods:titleInfo>
<mods:name id="7A181527D5F13AA544D0C43AABF4D778" type="personal">
<mods:role id="8DAA71EF24FF8F205130DD64EF9D4937">
<mods:roleTerm id="8E157C86F27C30508C64C4C87590336E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="896AF68748E28A7B8BCAD4FC7CF531FC">McKamey, Stuart H.</mods:namePart>
<mods:nameIdentifier id="6E2710E059B0303B3A127C9EDD5C53E4" type="ORCID">https://orcid.org/0000-0002-9617-0594</mods:nameIdentifier>
<mods:affiliation id="F8DA424D085E67663049942AD1778E33">Systematic Entomology Laboratory, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum of Natural History, P. O. Box 37012, Washington, D. C. 20013, USA</mods:affiliation>
<mods:nameIdentifier id="5F0BB3B5B4637EA14355A8C5278075E0" type="email">stuart.mckamey@usda.gov</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="56B5C471CF0B58974832B2994C0E4346">text</mods:typeOfResource>
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<mods:title id="4141C80EAE7FF6ED8E003F1181C4683C">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="B6FC4A397E608BA63681713765A2D015">
<mods:date id="B8588DBE61D021F2F09114A62ACDE28D">2023</mods:date>
<mods:detail id="351EFCD9509BDA35BDC1686B8D8401DA" type="pubDate">
<mods:number id="7DA33CBFF41D45DED7100D33882E09CA">2023-08-08</mods:number>
</mods:detail>
<mods:detail id="04708B7F7FF8B8EDCDEAF9279AD623AA" type="volume">
<mods:number id="D9A86939DD78B222B1408877B12B8B7B">1174</mods:number>
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<mods:start id="3311997461CD58B1FBE17AD4AB850C7F">85</mods:start>
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<mods:location id="7901DEE5A4D86CD8EE084065B187C5FC">
<mods:url id="EDEA01388940355996CBA726B8AC61F7">http://dx.doi.org/10.3897/zookeys.1174.103324</mods:url>
</mods:location>
<mods:classification id="14B31F40E0FBB5355DEFAB506545729B">journal article</mods:classification>
<mods:identifier id="D8E14E23192E2DFE55489E9690E6B09F" type="DOI">http://dx.doi.org/10.3897/zookeys.1174.103324</mods:identifier>
<mods:identifier id="BB5BF27041F97DBAC652824581F1491D" type="Pensoft-Pub">1313-2970-1174-85</mods:identifier>
<mods:identifier id="ED2479D4153763ACE8A64C104C9F83B9" type="ZooBank">5C6509E838A34F6A878B08C6868E0D2A</mods:identifier>
<mods:identifier id="B9483A8A249556C6954BE438DC610141" type="Pensoft-UUID">C382EDC7029755EFA1BC59D090835B37</mods:identifier>
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<treatment id="0292BFB428F25546BEA9B09FF7E0DEB7" ID-GBIF-Taxon="212426716" LSID="urn:lsid:zoobank.org:act:83FF1199-024D-4397-A31F-A942B5B39E81" httpUri="http://treatment.plazi.org/id/0292BFB428F25546BEA9B09FF7E0DEB7" lastPageNumber="85" pageId="0" pageNumber="85" scope_family="Membracidae" scope_order="Hemiptera">
<subSubSection id="B85BD9CA61768D31EFBAC0E01CE5DB47" pageId="0" pageNumber="85" type="nomenclature">
<paragraph id="293B89DD9A9D44A8A7D1BE87B72CEF38" pageId="0" pageNumber="85">
<taxonomicName id="4E847E2E92D4E9CAB3C2CEBAB37D3F10" ID-CoL="CMTLF" LSID="https://zoobank.org/83FF1199-024D-4397-A31F-A942B5B39E81" authority="McKamey, 2023" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis duocornus" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="duocornus" status="sp. nov.">Smilidarnis duocornus</taxonomicName>
<taxonomicNameLabel id="F8E38BB9EE7587E7D53866AC6A8BFB49" pageId="0" pageNumber="85">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B1E8BE411960B0A67F337395B35D6D04" pageId="0" pageNumber="85" type="description">
<paragraph id="CB5FED850C3B31FF0AD37553AD44D2AF" pageId="0" pageNumber="85">
<figureCitation id="532A7792FEFD9DB0A3DA69BB24A24A3B" captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Smilidarnis duocornus sp. nov. 2 - 4 habitus in dorsal, anterior, and lateral views, respectively 5 - 7 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85">Figs 2-7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5F6F6AA6F255328F9F51C037C5EA7572" pageId="0" pageNumber="85" type="diagnosis">
<paragraph id="A5AD7DBC12C9DB2B959DDF44B8C10C68" pageId="0" pageNumber="85">Diagnosis.</paragraph>
<paragraph id="4FA397AF1F6A6C12EE7E829B57ABD302" pageId="0" pageNumber="85">Pronotum with pair of suprahumeral spines; pronotum broader across suprahumeral spines than across posterior spines.</paragraph>
</subSubSection>
<subSubSection id="1E42308B477B2B54785EB2DD14EF7E33" pageId="0" pageNumber="85" type="description">
<paragraph id="EE552068F1A0D6EA4978212EF037D111" pageId="0" pageNumber="85">Description of female.</paragraph>
<paragraph id="C307AA14686D69DB8D63169C636AEE46" pageId="0" pageNumber="85">
Dimensions (mm). Length of pronotum 7.5; length including wings in repose 9.5; width across suprahumeral spine apices 4.5; width across posterolateral spine apices 3.0; height in anterior view 4.3. Head (Fig.
<figureCitation id="DF3F4906B850958A2CA1170A9857788A" captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Smilidarnis duocornus sp. nov. 2 - 4 habitus in dorsal, anterior, and lateral views, respectively 5 - 7 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85">3</figureCitation>
). Ocelli circular, below imaginary midline between eyes; distance to eye 1.2
<normalizedToken id="C3AC7B2785CF2E072847573A4805C049" originalValue="×">x</normalizedToken>
distance between ocelli; vertex with depression ventrolaterally adjacent to ocelli, pair of narrow diagonal depressions dorsally, with distinct sutures and frontoclypeus extending ventrally beyond vertex ventrolateral margins. Pronotum (Figs
<figureCitation id="E9F45C9CD01360F0A2F7CDA1920D1CED" captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Smilidarnis duocornus sp. nov. 2 - 4 habitus in dorsal, anterior, and lateral views, respectively 5 - 7 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85">2-4</figureCitation>
) with pair of stout suprahumeral spines; pronotum distinctly broader across suprahumeral spines than across posterior spines; weakly elevated immediately behind suprahumeral spines, this portion roughly trapezoidal in lateral view, abruptly narrowed laterally and expanding again to apical portion that bears pair of widely separated, stout lateral spines directed posteriorly and slender middle spine. Forewing (as in Fig.
<figureCitation id="B4E864B0226E2395688ED703B5F5E9FF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Smilidarnis robustus sp. nov., forewing and hind wing." figureDoi="10.3897/zookeys.1174.103324.figure1" httpUri="https://binary.pensoft.net/fig/890186" pageId="0" pageNumber="85">1</figureCitation>
) R and M not confluent for short distance near apex. Leg chaetotaxy. Femora lacking cucullate setae and spines; metathoracic tibia row I lacking cucullate setae, row II cucullate setae sparse in single line, row III cucullate setae dense, single row basally, double row distally. Abdominal terga without pits, fossae, or digitate processes. Color. Head pale brown except darker in depressions and along sutures of frontoclypeus; pronotum pale brown throughout except dark brown mottling in anterior half, at bases of suprahumeral spines, and posteriorly across lateral spines.
</paragraph>
<caption id="820FB6A4C1694A506455BC4D0EEBB80E" doi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85" start="Figures 2–7" startId="F2">
<paragraph id="006825F19C661E9F093983D6CA116735" pageId="0" pageNumber="85">
<emphasis id="87A326A323708AA79720C3CA952236D2" bold="true" pageId="0" pageNumber="85">Figures 2-7.</emphasis>
<taxonomicName id="6B3A1E25D4E365D734AB5BADACC26FBE" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis duocornus" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="duocornus">
<emphasis id="CA802FC93A976225C285B9CB4F9BDB02" italics="true" pageId="0" pageNumber="85">Smilidarnis duocornus</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="BADF7F814B1FBE66D294D9ABA4E1EBEB" bold="true" pageId="0" pageNumber="85">2-4</emphasis>
habitus in dorsal, anterior, and lateral views, respectively
<emphasis id="EDBF1CC2E33811045DA64E3443A99B2F" bold="true" pageId="0" pageNumber="85">5-7</emphasis>
valvifers 1, 2, and 3, respectively.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="738CA78C9DBA76D889AE02F488096D62" pageId="0" pageNumber="85" type="female terminalia">
<paragraph id="10CE12BEC947537417164C664462A8B3" pageId="0" pageNumber="85">Female terminalia.</paragraph>
<paragraph id="16D9AB8CB9F5B6847A89775BE81792E7" pageId="0" pageNumber="85">
Sternite VII broadly emarginate medially (as in Fig.
<figureCitation id="8B04C7C4FAD8DB44C5044D9C2BAA312F" captionStart="Figures 14–20" captionStartId="F4" captionText="Figures 14 - 20. Smilidarnis robustus sp. nov. 14 - 16 habitus in dorsal, lateral, and anterior views, respectively 17 abdominal segment VII, ventral view 18 - 20 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures14-20" httpUri="https://binary.pensoft.net/fig/890189" pageId="0" pageNumber="85">17</figureCitation>
); valvula I (Fig.
<figureCitation id="654BA3957A7DF541587D5C2BC94CE221" captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Smilidarnis duocornus sp. nov. 2 - 4 habitus in dorsal, anterior, and lateral views, respectively 5 - 7 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85">5</figureCitation>
) long, apex subacute; valvula II (Fig.
<figureCitation id="5698ECAF57A5C01D71F93528C0BBB0A2" captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Smilidarnis duocornus sp. nov. 2 - 4 habitus in dorsal, anterior, and lateral views, respectively 5 - 7 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85">6</figureCitation>
) in lateral view with dorsal margin linear in basal 2/3, then minutely sinuate, apex rounded; valvula III (Fig.
<figureCitation id="A20192E4A0AA8C42210C65AAE68386D2" captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Smilidarnis duocornus sp. nov. 2 - 4 habitus in dorsal, anterior, and lateral views, respectively 5 - 7 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85">7</figureCitation>
) long, apex rounded, bearing macrosetae along entire ventral margin.
</paragraph>
</subSubSection>
<subSubSection id="D7C198AB6D500EBD7C15DDF2A7F9784F" pageId="0" pageNumber="85" type="male">
<paragraph id="ABBCC82737FC0EAABFC618DEFE9B335C" pageId="0" pageNumber="85">Male.</paragraph>
<paragraph id="B4D16391408626F08FAF6576869A48C4" pageId="0" pageNumber="85">Unknown.</paragraph>
</subSubSection>
<subSubSection id="9D50222D102A372CBCEB53FD8DB56286" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="304A0D24EB3ECB11CF2C6DE4B8B8B19D" pageId="0" pageNumber="85">Material examined.</paragraph>
<paragraph id="BA9919F11AC6B3A50BB60E7F8F41D37F" pageId="0" pageNumber="85">
<materialsCitation id="11D41D337722EF8403660EDAF32C5706" collectingDate="1993-10-05" collectingDateMax="1993-10-17" collectingDateMin="1993-10-05" collectorName="S. H. McKamey" country="BRAZIL" location="Rondonia" specimenCount="♀" stateProvince="Rondonia" typeStatus="Holotype">
<emphasis id="69E0B2347D004A08DD0C6865D6A1300F" bold="true" italics="true" pageId="0" pageNumber="85">
<typeStatus id="3D347F5F2BF6F590040DA3FDE70092D1">Holotype</typeStatus>
</emphasis>
<specimenCount id="359D47CD8B88F4555CB3E0CE719436E9">♀</specimenCount>
(INPA) with labels &quot;
<collectingCountry id="047ADB9D9BCCF4E5DF5A4ED71BF08254" name="Brazil">BRAZIL</collectingCountry>
:
<collectingRegion id="72E4936964AF902AC76DD35D8DAA4C32" country="Brazil" name="Rondonia">Rondonia</collectingRegion>
. 62 | km SW Ariquemes, nr | Fzda. Rancho Grande |
<collectingDate id="EBDA3D01E4689534E22FCEAE6DFF1C8A" value="1993-10-05" valueMax="1993-10-17" valueMin="1993-10-05">5-17-X-1993</collectingDate>
JE Eger | MV &amp; Black Lights&quot; and a red &quot;
<typeStatus id="BA6DD636ACE7A262AB8BCB48AE76A234">HOLOTYPE</typeStatus>
|
<taxonomicName id="2B76D0A5CA4E6CD1BD253881868E4536" authorityName="Andrade" authorityYear="1989" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Smilidarnis</taxonomicName>
|
<taxonomicName id="27079B5E18131087C218C1AE57A3FD2B" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis duocornus" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="duocornus">Smilidarnis duocornus</taxonomicName>
|
<collectorName id="FE36A6C4F442BCB838B99015BFABF096">S.H. McKamey</collectorName>
&quot;.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4E975B4D3E5D9E02B958397A4C1FAC38" pageId="0" pageNumber="85" type="etymology">
<paragraph id="0EC8F8FAAC228F8ADD120CF0B1C3848D" pageId="0" pageNumber="85">Etymology.</paragraph>
<paragraph id="5EDF8D075554832640187E8F60980EA3" pageId="0" pageNumber="85">
The species is a Latin adjective derived from
<normalizedToken id="3A9825F5C727D391B41E3EF83111B52A" originalValue="“duo”">&quot;duo&quot;</normalizedToken>
for two, and
<normalizedToken id="8627ACDA60167E13962EB83419607049" originalValue="“cornu”">&quot;cornu&quot;</normalizedToken>
for horn, referring to the pair of stout suprahumeral spines.
</paragraph>
</subSubSection>
<subSubSection id="E623A063029F572D0A2671729F5C416E" pageId="0" pageNumber="85" type="notes">
<paragraph id="6FB72F4DD9D3EF0E7AD944E415DBB631" pageId="0" pageNumber="85">Notes.</paragraph>
<paragraph id="30558430256F74188A7EE5F0BE89AF85" pageId="0" pageNumber="85">
The pronota of
<taxonomicName id="581FFD39FC41FA94CD78C1A074AA7CB0" lsidName="S. duocornus" pageId="0" pageNumber="85" rank="species" species="duocornus">
<emphasis id="FDD3E3173EE3B69AFB9BAA844BF97242" italics="true" pageId="0" pageNumber="85">S. duocornus</emphasis>
</taxonomicName>
and
<taxonomicName id="62896C9046BCEEE13E5692B245F267A8" lsidName="S. robustus" pageId="0" pageNumber="85" rank="species" species="robustus">
<emphasis id="CE16029964B49E7A49E4161E21D41A28" italics="true" pageId="0" pageNumber="85">S. robustus</emphasis>
</taxonomicName>
, which are both only known from a female holotype, closely resemble each other and have similar coloration. In some treehoppers, such as
<taxonomicName id="D37FFA5639A153FAA0691300F3827D48" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Quinquespinosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quinquespinosa septamacula" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="septamacula">
<emphasis id="78FDA8334A1E8B1947B9B0F819F016EA" italics="true" pageId="0" pageNumber="85">Quinquespinosa septamacula</emphasis>
</taxonomicName>
<bibRefCitation id="5BED06E14FED3DF0DD0ECCD4FDD5B05C" DOI="https://doi.org/10.3897/zookeys.1143.94124" author="McKamey, SH" journalOrPublisher="ZooKeys" pageId="0" pageNumber="85" pagination="189 - 203" refId="B13" refString="McKamey, SH, 2023. Three new monobasic genera and three new species of the New World treehopper tribe Acutalini (Hemiptera, Membracidae, Smiliinae) with a key to all genera. ZooKeys 1143: 189 - 203, DOI: https://doi.org/10.3897/zookeys.1143.94124" title="Three new monobasic genera and three new species of the New World treehopper tribe Acutalini (Hemiptera, Membracidae, Smiliinae) with a key to all genera." url="https://doi.org/10.3897/zookeys.1143.94124" volume="1143" year="2023">McKamey (2023)</bibRefCitation>
, color and surprahumeral spine length is variable, so these differences alone might not suffice for species recognition. Nevertheless, these are considered separate species for several reasons:
<taxonomicName id="B86304D05D8816753FC65B3C1C8DA0C3" lsidName="S. sobustus" pageId="0" pageNumber="85" rank="species" species="sobustus">
<emphasis id="66224713DE6CA861C822C4CF04C66FFE" italics="true" pageId="0" pageNumber="85">S. sobustus</emphasis>
</taxonomicName>
is almost 3 mm larger; its posterior portion is more elevated, and the apical lateral spines are directed more laterally. Futhermore, the setation of valvifer III is distinct; bearing macrosetae along ventral margin in S.
<taxonomicName id="7A89833CA1ACA43DEE229F5AEBE4A7EA" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis duocornus" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="duocornus">
<emphasis id="E07A8088597917A1ED421F1366E9AD43" italics="true" pageId="0" pageNumber="85">Smilidarnis duocornus</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="6E65534F5803E9C68A67B733806C1232" captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Smilidarnis duocornus sp. nov. 2 - 4 habitus in dorsal, anterior, and lateral views, respectively 5 - 7 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures2-7" httpUri="https://binary.pensoft.net/fig/890187" pageId="0" pageNumber="85">7</figureCitation>
), in contrast to that of
<taxonomicName id="9931C745BAE71B6B609521B7D8F60DB3" lsidName="S. robustus" pageId="0" pageNumber="85" rank="species" species="robustus">
<emphasis id="FE4487D83207C4D0E517279B571F36A0" italics="true" pageId="0" pageNumber="85">S. robustus</emphasis>
</taxonomicName>
, which bears fine hairlike setae on entire ventral half (Fig.
<figureCitation id="F371D8233C9DC186B1EAEB146DEADDC7" captionStart="Figures 14–20" captionStartId="F4" captionText="Figures 14 - 20. Smilidarnis robustus sp. nov. 14 - 16 habitus in dorsal, lateral, and anterior views, respectively 17 abdominal segment VII, ventral view 18 - 20 valvifers 1, 2, and 3, respectively." figureDoi="10.3897/zookeys.1174.103324.figures14-20" httpUri="https://binary.pensoft.net/fig/890189" pageId="0" pageNumber="85">20</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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