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<document id="91D17FAF6D1F9C554A342776B89E2286" ID-DOI="10.1016/j.jcz.2022.12.004" ID-ISSN="1873-2674" ID-Zenodo-Dep="8164206" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1689773440848" checkinUser="felipe" docAuthor="González-Casarrubios, Alberto, Cepeda, Diego, Neuhaus, Birger, García-Cobo, Marta, Pardos, Fernando, Ürkmez, Derya &amp; Sánchez, Nuria" docDate="2023" docId="374D87FDB008FFF5FCCFFB80D7503238" docLanguage="en" docName="ZoolAnz.303.90-111.pdf" docOrigin="Zoologischer Anzeiger 303" docSource="http://dx.doi.org/10.1016/j.jcz.2022.12.004" docStyle="DocumentStyle:32483D6D135DE2FF927BA3F1400E0FFD.1:ZoolAnz.2022-.journal_article" docStyleId="32483D6D135DE2FF927BA3F1400E0FFD" docStyleName="ZoolAnz.2022-.journal_article" docStyleVersion="1" docTitle="Setaphyes dentatus" docType="treatment" docVersion="3" lastPageNumber="98" masterDocId="CB74FF85B00AFFFFFFFDFFAED567374E" masterDocTitle="The genus Setaphyes (Kinorhyncha, Pycnophyidae) in European waters: Redescription of Setaphyes dentatus (Reinhard, 1881) and Setaphyes kielensis (Zelinka, 1928), including notes on morphometrics, sexually dimorphic features and reproduction of the genus" masterLastPageNumber="111" masterPageNumber="90" pageNumber="92" updateTime="1689934032965" updateUser="tatiana">
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<mods:title id="9A1CE3663C435001CAB54AA1C64CCF48">The genus Setaphyes (Kinorhyncha, Pycnophyidae) in European waters: Redescription of Setaphyes dentatus (Reinhard, 1881) and Setaphyes kielensis (Zelinka, 1928), including notes on morphometrics, sexually dimorphic features and reproduction of the genus</mods:title>
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<mods:namePart id="108825999C5A0EC9F5549F8D4B8B365D">Neuhaus, Birger</mods:namePart>
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<mods:namePart id="9947B6F856AB457DD79E1A663ED5E622">García-Cobo, Marta</mods:namePart>
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<mods:namePart id="70AB86FED33DD1F0B98656726ED9871A">Sánchez, Nuria</mods:namePart>
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3.1. Redescription of
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Setaphyes dentatus (
<bibRefCitation id="DB754B1AB008FFFDFB4CFB80D059330F" author="Reinhard, W." box="[1201,1342,1070,1089]" pageId="2" pageNumber="92" pagination="588 - 592" refId="ref17444" refString="Reinhard, W., 1881. Uber Echinoderes und Desmoscolex der Umgebung von Odessa. Zool. Anz. 4 (97), 588 - 592." type="journal article" year="1881">Reinhard, 1881</bibRefCitation>
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<paragraph id="BF5B36EBB008FFFDFCACFBC8D17F3337" blockId="2.[849,1048,1126,1145]" box="[849,1048,1126,1145]" pageId="2" pageNumber="92">
<figureCitation id="27DF2A6EB008FFFDFCACFBC8D6C73337" box="[849,928,1126,1145]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="3.[100,130,1763,1780]" captionStartId-1="4.[100,130,1039,1056]" captionStartId-2="5.[100,130,1756,1773]" captionStartId-3="6.[100,130,1452,1469]" captionStartId-4="7.[100,130,1830,1847]" captionStartId-5="8.[100,130,1556,1573]" captionStartId-6="9.[100,130,1718,1735]" captionTargetBox-0="[264,1323,150,1735]" captionTargetBox-1="[144,726,149,1009]" captionTargetBox-2="[151,1437,149,1726]" captionTargetBox-3="[226,1360,149,1424]" captionTargetBox-4="[233,1355,150,1802]" captionTargetBox-5="[107,766,149,1528]" captionTargetBox-6="[267,1322,243,1689]" captionTargetId-0="figure-8@3.[263,1324,148,1736]" captionTargetId-1="figure-1246@4.[142,727,148,1011]" captionTargetId-2="figure-8@5.[150,1438,148,1728]" captionTargetId-3="figure-351@6.[225,1362,148,1425]" captionTargetId-4="figure-8@7.[232,1356,148,1803]" captionTargetId-5="figure-802@8.[106,767,148,1529]" captionTargetId-6="figure-8@9.[265,1323,242,1690]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." captionText-1="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." captionText-2="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." captionText-3="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." captionText-4="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." captionText-5="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." captionText-6="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi-0="http://doi.org/10.5281/zenodo.8164210" figureDoi-1="http://doi.org/10.5281/zenodo.8164212" figureDoi-2="http://doi.org/10.5281/zenodo.8164214" figureDoi-3="http://doi.org/10.5281/zenodo.8164218" figureDoi-4="http://doi.org/10.5281/zenodo.8164220" figureDoi-5="http://doi.org/10.5281/zenodo.8164222" figureDoi-6="http://doi.org/10.5281/zenodo.8164224" httpUri-0="https://zenodo.org/record/8164210/files/figure.png" httpUri-1="https://zenodo.org/record/8164212/files/figure.png" httpUri-2="https://zenodo.org/record/8164214/files/figure.png" httpUri-3="https://zenodo.org/record/8164218/files/figure.png" httpUri-4="https://zenodo.org/record/8164220/files/figure.png" httpUri-5="https://zenodo.org/record/8164222/files/figure.png" httpUri-6="https://zenodo.org/record/8164224/files/figure.png" pageId="2" pageNumber="92">Figs 2–8</figureCitation>
,
<tableCitation id="F2660350B008FFFDFC51FBC8D17F3337" box="[940,1048,1126,1145]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStartId-0="10.[100,150,150,166]" captionStartId-1="10.[100,150,784,800]" captionStartId-2="16.[100,150,1347,1363]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionTargetPageId-2="16" captionText-0="Table 1 Measurements (μm) and proportions (%) of Setaphyes dentatus, including total values from the different studied populations split by sex (values per specimen and population can be found in Supplementary Table 3. Abbreviations: LTS, lateral terminal spine length; MSW5, maximum sternal width (measured at segment 5); n, number of measured specimens; S, segment length (followed by number of corresponding segment); Sd, standard deviation; SW10, standard sternal width (measured at segment 10); TL, total trunk length." captionText-1="Table 2 Summary of nature and arrangement of cuticular elevations, processes, spines, tubes, setae, sensory spots and nephridiopores in Setaphyes dentatus. Abbreviations: ce, cuticular elevation; cp, cuticular process; LD, laterodorsal; lts, lateral terminal spine; LV, lateroventral; MD, middorsal; ne, nephridiopore; PD, paradorsal; ps, penile spine; SD, subdorsal; se, seta; ss, sensory spot; tu, tube; VL, ventrolateral; VM, ventromedial; * indicates unpaired structures; 6 and ♀ indicates sexually dimorphic characters." captionText-2="Table 3 Measurements (μm) and proportions (%) of Setaphyes kielensis, including total values from the different studied populations split by sex (values per specimen and population can be found in Supplementary Table 3. Abbreviations: LTS, lateral terminal spine length; MSW5, maximum sternal width (measured at segment 5); n, number of measured specimens; S, segment length (followed by number of corresponding segment); Sd, standard deviation; SW10, standard sternal width (measured at segment 10); TL, total trunk length." pageId="2" pageNumber="92">Tables 1–2.</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7FE6560B008FFFDFCCFFB30D0B73179" pageId="2" pageNumber="92" type="diagnosis">
<paragraph id="BF5B36EBB008FFFDFCCFFB30D6AF33FF" blockId="2.[818,1488,1182,1592]" box="[818,968,1182,1201]" pageId="2" pageNumber="92">
<emphasis id="8D90EAF9B008FFFDFCCFFB30D6AF33FF" bold="true" box="[818,968,1182,1201]" italics="true" pageId="2" pageNumber="92">3.1.1. Diagnosis</emphasis>
</paragraph>
<paragraph id="BF5B36EBB008FFFDFCACFB17D0B73179" blockId="2.[818,1488,1182,1592]" pageId="2" pageNumber="92">
<taxonomicName id="78E44D68B008FFFDFCACFB17D6CF3382" authorityName="Sanchez, Yamasaki, Pardos, Sorensen &amp; Martinez" authorityYear="2016" box="[849,936,1209,1228]" class="Allomalorhagida" family="Pycnophyidae" genus="Setaphyes" kingdom="Animalia" pageId="2" pageNumber="92" phylum="Kinorhyncha" rank="genus">
<emphasis id="8D90EAF9B008FFFDFCACFB17D6CF3382" bold="true" box="[849,936,1209,1228]" italics="true" pageId="2" pageNumber="92">Setaphyes</emphasis>
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with anterior margin of the tergal plate on segment 1 finely denticulated, posteriorly followed by a transverse area of minute cuticular, net-like ridges. Middorsal elevations on segments 2–6, and middorsal processes on segments 1 and 7–9. Posterior end of middorsal structures covered by tufts of elongated, thick hairs whose tips usually surpass the posterior segment margin. Unpaired paradorsal setae on segments 2–9, alternating left and right position along the trunk and showing intraspecific variability but without any specific pattern. Laterodorsal setae on segments 3, 5 and 7. Lateroventral setae on segments 2–10. Ventromedial setae on segments 4–5 and
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males and 3–5 and
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females. Patch of conspicuous longitudinal cuticular ridges on segment 10, from laterodorsal to ventrolateral positions. Lateral terminal spines short, slender (males LTS:TL average ratio 21.2 %; females LTS:TL average ratio 15.7 %; males and females average ratio 17.8 %).
</paragraph>
</subSubSection>
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<emphasis id="8D90EAF9B008FFFDFCCFF9F2D17B3121" bold="true" box="[818,1052,1628,1647]" italics="true" pageId="2" pageNumber="92">3.1.2. Examined material</emphasis>
</paragraph>
<paragraph id="BF5B36EBB008FFFBFCACF9D6D41333AC" blockId="2.[818,1488,1628,1982]" lastBlockId="4.[100,770,1145,1251]" lastPageId="4" lastPageNumber="94" pageId="2" pageNumber="92">
<materialsCitation id="0F8C3CB6B008FFFDFCACF9D6D0C131B4" collectingDate="2011-02-08" collectingDateMax="2019-07-13" collectingDateMin="2011-02-08" collectionCode="ZMB" elevation="76" longitude="-16.549" pageId="2" pageNumber="92" specimenCode="ZMB 12408-12418, 12616-12740, ZMB 12745, R2, ZMB 12746, ZMB 12747, ZMB 12748-12750, ZMB 12754, ZMB 12753, ZMB 12751, ZMB 12752" specimenCount="278" specimenCount-adult="138" specimenCount-female="86" specimenCount-male="54">
<collectorName id="1211533DB008FFFDFCACF9D6D61731C5" box="[849,880,1656,1675]" pageId="2" pageNumber="92">For</collectorName>
light microscopy,
<specimenCount id="A9E2FD62B008FFFDFBE1F9D6D11231C5" box="[1052,1141,1656,1675]" pageId="2" pageNumber="92" type="adult">136 adult</specimenCount>
specimens,
<specimenCount id="A9E2FD62B008FFFDFB18F9D6D05F31C5" box="[1253,1336,1656,1675]" pageId="2" pageNumber="92" type="male">53 males</specimenCount>
and
<specimenCount id="A9E2FD62B008FFFDFA98F9D6D0AB31C5" box="[1381,1484,1656,1675]" pageId="2" pageNumber="92" type="female">83 females</specimenCount>
, were studied from 17 populations belonging to eight marine ecoregions (
<specimenCode id="EF429E90B008FFFDFCCAF91ED692318D" box="[823,1013,1712,1731]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="92" type="Museum">ZMB 12408–12418</specimenCode>
,
<specimenCode id="EF429E90B008FFFDFBF9F901D1EF318D" box="[1028,1160,1711,1731]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="92" type="Museum">12616–12740</specimenCode>
; see
<collectorName id="1211533DB008FFFDFB3DF901D0F8318D" box="[1216,1439,1711,1731]" pageId="2" pageNumber="92">Supplementary Table I</collectorName>
).
<collectorName id="1211533DB008FFFDFA44F901D0A8318C" box="[1465,1487,1711,1730]" pageId="2" pageNumber="92">Of</collectorName>
those,
<specimenCount id="A9E2FD62B008FFFDFC93F965D6863190" box="[878,993,1739,1758]" pageId="2" pageNumber="92" type="female">four females</specimenCount>
from the
<collectorName id="1211533DB008FFFDFBC7F965D1923190" box="[1082,1269,1739,1758]" pageId="2" pageNumber="92">Anatolian Peninsula</collectorName>
are new records for the region. 11 additional adult specimens were examined with SEM:
</materialsCitation>
<materialsCitation id="0F8C3CB6B008FFFDFA50F946D1F53000" collectingDate="2011-02-08" collectingDateMax="2019-07-13" collectingDateMin="2011-02-08" collectionCode="ZMB" elevation="76" longitude="-16.549" pageId="2" pageNumber="92" specimenCode="ZMB 12745" specimenCount="1" specimenCount-adult="1">
<specimenCount id="A9E2FD62B008FFFDFA50F946D6053058" pageId="2" pageNumber="92" type="adult">one adult</specimenCount>
specimen from the
<collectorName id="1211533DB008FFFDFBDCF8ADD1B83058" box="[1057,1247,1795,1814]" pageId="2" pageNumber="92">Anatolian Peninsula</collectorName>
(indeterminate sex:
<specimenCode id="EF429E90B008FFFDFA5EF8AAD617307C" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="92" type="Museum">ZMB 12745</specimenCode>
; station YSL09
<specimenCode id="EF429E90B008FFFDFC03F8B1D170307C" box="[1022,1047,1823,1842]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="92">R2</specimenCode>
;
<date id="CB5A102BB008FFFDFBDDF8B1D1F9307C" box="[1056,1182,1823,1842]" pageId="2" pageNumber="92" value="2019-07-13">
<collectingDate id="DB1EE9C3B008FFFDFBDDF8B1D1F9307C" box="[1056,1182,1823,1842]" pageId="2" pageNumber="92" value="2019-07-13">13 July 2019</collectingDate>
</date>
, 41
<geoCoordinate id="DAD0502CB008FFFDFB3DF8B3D06C307C" box="[1216,1291,1821,1842]" degrees="24.738" direction="west" orientation="longitude" pageId="2" pageNumber="92" precision="55" value="-24.738">
<superScript id="48919BA3B008FFFDFB3DF8B3D1AF3067" attach="none" box="[1216,1224,1821,1833]" fontSize="6" pageId="2" pageNumber="92">Ǫ</superScript>
24.738
</geoCoordinate>
<superScript id="48919BA3B008FFFDFAF7F8B2D0683067" attach="right" box="[1290,1295,1820,1833]" fontSize="6" pageId="2" pageNumber="92">′</superScript>
N, 036
<geoCoordinate id="DAD0502CB008FFFDFAB2F8B3D0FD307C" box="[1359,1434,1821,1842]" degrees="39.210" direction="west" orientation="longitude" pageId="2" pageNumber="92" precision="55" value="-39.21">
<superScript id="48919BA3B008FFFDFAB2F8B3D0303067" attach="none" box="[1359,1367,1821,1833]" fontSize="6" pageId="2" pageNumber="92">Ǫ</superScript>
39.210
</geoCoordinate>
<superScript id="48919BA3B008FFFDFA64F8B2D0F93067" attach="right" box="[1433,1438,1820,1833]" fontSize="6" pageId="2" pageNumber="92">′</superScript>
E,
<elevation id="34C9D1D8B008FFFDFA4AF8B1D6E03000" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="92" unit="m" value="76.0">
<quantity id="781C9B0EB008FFFDFA4AF8B1D6233000" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="92" unit="m" value="76.0">76 m</quantity>
depth
</elevation>
; collector:
<collectorName id="1211533DB008FFFDFC05F895D1E03000" box="[1016,1159,1848,1870]" pageId="2" pageNumber="92">Derya Ürkmez</collectorName>
),
</materialsCitation>
<materialsCitation id="0F8C3CB6B008FFFDFB60F895D17830F0" collectingDate="2011-02-08" collectingDateMax="2019-07-13" collectingDateMin="2011-02-08" collectionCode="ZMB" elevation="76" longitude="-16.549" pageId="2" pageNumber="92" specimenCode="ZMB 12746-12750" specimenCount="10" specimenCount-adult="3" specimenCount-female="1" specimenCount-male="1">
10 from the Iberian Peninsula, collected by
<collectorName id="1211533DB008FFFDFC53F8F9D16D3024" box="[942,1034,1879,1898]" pageId="2" pageNumber="92">F. Pardos</collectorName>
, M. Herranz and
<collectorName id="1211533DB008FFFDFB4AF8F9D0473024" box="[1207,1312,1879,1898]" pageId="2" pageNumber="92">N. Sanchez</collectorName>
´(
<specimenCount id="A9E2FD62B008FFFDFACDF8F9D0FC3024" box="[1328,1435,1879,1898]" pageId="2" pageNumber="92" type="female">one female</specimenCount>
<specimenCode id="EF429E90B008FFFDFA5EF8F9D61730C8" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="92" type="Museum">ZMB 12746</specimenCode>
,
<specimenCount id="A9E2FD62B008FFFDFC81F8DDD6B330C8" box="[892,980,1907,1926]" pageId="2" pageNumber="92" type="male">one male</specimenCount>
<specimenCode id="EF429E90B008FFFDFC21F8DDD12930C8" box="[988,1102,1907,1926]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="92" type="Museum">ZMB 12747</specimenCode>
, and
<specimenCount id="A9E2FD62B008FFFDFB79F8DDD19130C8" box="[1156,1270,1907,1926]" pageId="2" pageNumber="92" type="adult">three adults</specimenCount>
<specimenCode id="EF429E90B008FFFDFB03F8DDD0D230C8" box="[1278,1461,1907,1926]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="92" type="Museum">ZMB 12748–12750</specimenCode>
of indeterminate sex from Algeciras,
<date id="CB5A102BB008FFFDFB6EF821D02630EC" box="[1171,1345,1935,1954]" pageId="2" pageNumber="92" value="2011-02-08">
<collectingDate id="DB1EE9C3B008FFFDFB6EF821D02630EC" box="[1171,1345,1935,1954]" pageId="2" pageNumber="92" value="2011-02-08">8 February 2011</collectingDate>
</date>
, 36
<geoCoordinate id="DAD0502CB008FFFDFA97F822D0D230EC" box="[1386,1461,1932,1954]" degrees="10.741" direction="west" orientation="longitude" pageId="2" pageNumber="92" precision="55" value="-10.741">
<superScript id="48919BA3B008FFFDFA97F822D01530D6" attach="none" box="[1386,1394,1932,1944]" fontSize="6" pageId="2" pageNumber="92">Ǫ</superScript>
10.741
</geoCoordinate>
<superScript id="48919BA3B008FFFDFA49F822D0DE30D7" attach="right" box="[1460,1465,1932,1945]" fontSize="6" pageId="2" pageNumber="92">′</superScript>
N, 005
<geoCoordinate id="DAD0502CB008FFFDFCABF806D6C730F0" box="[854,928,1960,1982]" degrees="23.243" direction="west" orientation="longitude" pageId="2" pageNumber="92" precision="55" value="-23.243">
<superScript id="48919BA3B008FFFDFCABF806D63930FA" attach="none" box="[854,862,1960,1972]" fontSize="6" pageId="2" pageNumber="92">Ǫ</superScript>
23.243
</geoCoordinate>
<superScript id="48919BA3B008FFFDFC5DF809D6C230FA" attach="right" box="[928,933,1959,1972]" fontSize="6" pageId="2" pageNumber="92">′</superScript>
W, 8 depth;
</materialsCitation>
<materialsCitation id="0F8C3CB6B008FFFBFBD7F805D66533E5" collectingDate="2011-02-08" collectingDateMax="2019-07-13" collectingDateMin="2011-02-08" collectionCode="ZMB" elevation="76" lastPageId="4" lastPageNumber="94" longitude="-16.549" pageId="2" pageNumber="92" specimenCode="ZMB 12754, 12755" specimenCount="2" specimenCount-female="2">
<specimenCount id="A9E2FD62B008FFFDFBD7F805D1C730F0" box="[1066,1184,1963,1982]" pageId="2" pageNumber="92" type="female">two females</specimenCount>
<specimenCode id="EF429E90B008FFFDFB50F805D04730F3" box="[1197,1312,1962,1982]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="92" type="Museum">ZMB 12754</specimenCode>
and 12755 from Huelva,
<date id="CB5A102BB00EFFFBFF48FBD2D45A33C1" box="[181,317,1148,1167]" pageId="4" pageNumber="94" value="2011-04-12">
<collectingDate id="DB1EE9C3B00EFFFBFF48FBD2D45A33C1" box="[181,317,1148,1167]" pageId="4" pageNumber="94" value="2011-04-12">12 April 2011</collectingDate>
</date>
, 37
<geoCoordinate id="DAD0502CB00EFFFBFE9CFBD7D4CC33C1" box="[353,427,1145,1167]" degrees="08.324" direction="west" orientation="longitude" pageId="4" pageNumber="94" precision="55" value="-8.324">
<superScript id="48919BA3B00EFFFBFE9CFBD7D40E33CB" attach="none" box="[353,361,1145,1157]" fontSize="6" pageId="4" pageNumber="94">Ǫ</superScript>
08.324
</geoCoordinate>
<superScript id="48919BA3B00EFFFBFE56FBD7D4D733C8" attach="right" box="[427,432,1145,1158]" fontSize="6" pageId="4" pageNumber="94">′</superScript>
N, 007
<geoCoordinate id="DAD0502CB00EFFFBFE0FFBD7D75B33C1" box="[498,572,1145,1167]" degrees="20.308" direction="west" orientation="longitude" pageId="4" pageNumber="94" precision="55" value="-20.308">
<superScript id="48919BA3B00EFFFBFE0FFBD7D49D33CB" attach="none" box="[498,506,1145,1157]" fontSize="6" pageId="4" pageNumber="94">Ǫ</superScript>
20.308
</geoCoordinate>
<superScript id="48919BA3B00EFFFBFDC1FBD7D72633C8" attach="right" box="[572,577,1145,1158]" fontSize="6" pageId="4" pageNumber="94">′</superScript>
W,
<elevation id="34C9D1D8B00EFFFBFD9EFBD2D7B333C1" box="[611,724,1148,1167]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="94" unit="m" value="15.0">
<quantity id="781C9B0EB00EFFFBFD9EFBD2D7F233C1" box="[611,661,1148,1167]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="94" unit="m" value="15.0">15 m</quantity>
depth
</elevation>
; one additional female
<specimenCode id="EF429E90B00EFFFBFEEDFB36D4E633E5" box="[272,385,1176,1195]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="4" pageNumber="94" type="Museum">ZMB 12753</specimenCode>
, and two indeterminate sex,
<specimenCode id="EF429E90B00EFFFBFD69FB36D66533E5" box="[660,770,1176,1195]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="4" pageNumber="94" type="Museum">ZMB 12751</specimenCode>
</materialsCitation>
<materialsCitation id="0F8C3CB6B00EFFFBFF99FB1DD41733AC" collectingDate="2011-02-08" collectingDateMax="2019-07-13" collectingDateMin="2011-02-08" collectionCode="ZMB" elevation="76" longitude="-16.549" pageId="4" pageNumber="94" specimenCode="ZMB 12752" specimenCount="1" specimenCount-female="1">
and
<specimenCode id="EF429E90B00EFFFBFF66FB1AD47F3388" box="[155,280,1203,1223]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="4" pageNumber="94" type="Museum">ZMB 12752</specimenCode>
, from Huelva,
<date id="CB5A102BB00EFFFBFE30FB1AD70A3389" box="[461,621,1203,1223]" pageId="4" pageNumber="94" value="2011-04-11">
<collectingDate id="DB1EE9C3B00EFFFBFE30FB1AD70A3389" box="[461,621,1203,1223]" pageId="4" pageNumber="94" value="2011-04-11">11 April 2011</collectingDate>
</date>
, 37
<geoCoordinate id="DAD0502CB00EFFFBFD60FB1FD7803388" box="[669,743,1201,1222]" degrees="10.963" direction="west" orientation="longitude" pageId="4" pageNumber="94" precision="55" value="-10.963">
<superScript id="48919BA3B00EFFFBFD60FB1FD7C233F3" attach="none" box="[669,677,1201,1213]" fontSize="6" pageId="4" pageNumber="94">Ǫ</superScript>
10.963
</geoCoordinate>
<superScript id="48919BA3B00EFFFBFD1AFB1ED78B33F3" attach="right" box="[743,748,1200,1213]" fontSize="6" pageId="4" pageNumber="94">′</superScript>
N, 007
<geoCoordinate id="DAD0502CB00EFFFBFF75FB63D5B433AC" box="[136,211,1229,1250]" degrees="16.549" direction="west" orientation="longitude" pageId="4" pageNumber="94" precision="55" value="-16.549">
<superScript id="48919BA3B00EFFFBFF75FB63D5F73397" attach="none" box="[136,144,1229,1241]" fontSize="6" pageId="4" pageNumber="94">Ǫ</superScript>
16.549
</geoCoordinate>
<superScript id="48919BA3B00EFFFBFF2EFB62D5BF3397" attach="right" box="[211,216,1228,1241]" fontSize="6" pageId="4" pageNumber="94">′</superScript>
W,
<elevation id="34C9D1D8B00EFFFBFF05FB7ED40E33AC" box="[248,361,1231,1251]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="94" unit="m" value="11.0">
<quantity id="781C9B0EB00EFFFBFF05FB7ED44F33AD" box="[248,296,1232,1251]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="94" unit="m" value="11.0">11 m</quantity>
depth
</elevation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="EB9B6663B009FFFCFF99F94DD6D3303A" ID-DOI="http://doi.org/10.5281/zenodo.8164210" ID-Zenodo-Dep="8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="3" pageNumber="93" startId="3.[100,130,1763,1780]" targetBox="[264,1323,150,1735]" targetPageId="3" targetType="figure">
<paragraph id="BF5B36EBB009FFFCFF99F94DD6D3303A" blockId="3.[100,1488,1763,1908]" pageId="3" pageNumber="93">
<emphasis id="8D90EAF9B009FFFCFF99F94DD5F831BB" bold="true" box="[100,159,1763,1781]" pageId="3" pageNumber="93">Fig. 2.</emphasis>
Line art illustration of
<emphasis id="8D90EAF9B009FFFCFE90F94DD76B31BA" bold="true" box="[365,524,1763,1780]" italics="true" pageId="3" pageNumber="93">
<taxonomicName id="78E44D68B009FFFCFE90F94DD76031BA" baseAuthorityName="Reinhard" baseAuthorityYear="1881" box="[365,519,1763,1780]" class="Allomalorhagida" family="Pycnophyidae" genus="Setaphyes" kingdom="Animalia" pageId="3" pageNumber="93" phylum="Kinorhyncha" rank="species" species="dentatus">Setaphyes dentatus</taxonomicName>
.
</emphasis>
A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid.
</paragraph>
</caption>
<caption id="EB9B6663B00EFFFBFF99FBA1D7DC331C" ID-DOI="http://doi.org/10.5281/zenodo.8164212" ID-Zenodo-Dep="8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94" startId="4.[100,130,1039,1056]" targetBox="[144,726,149,1009]" targetPageId="4" targetType="figure">
<paragraph id="BF5B36EBB00EFFFBFF99FBA1D7DC331C" blockId="4.[100,770,1038,1107]" pageId="4" pageNumber="94">
<emphasis id="8D90EAF9B00EFFFBFF99FBA1D5FA336E" bold="true" box="[100,157,1039,1056]" pageId="4" pageNumber="94">Fig. 3.</emphasis>
Diagram of mouth cone and introvert in
<taxonomicName id="78E44D68B00EFFFBFE02FBA0D7F23351" baseAuthorityName="Reinhard" baseAuthorityYear="1881" box="[511,661,1038,1055]" class="Allomalorhagida" family="Pycnophyidae" genus="Setaphyes" kingdom="Animalia" pageId="4" pageNumber="94" phylum="Kinorhyncha" rank="species" species="dentatus">
<emphasis id="8D90EAF9B00EFFFBFE02FBA0D7F23351" bold="true" box="[511,661,1038,1055]" italics="true" pageId="4" pageNumber="94">Setaphyes dentatus</emphasis>
</taxonomicName>
showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector.
</paragraph>
</caption>
<subSubSection id="F7FE6560B00EFFF5FF99FAA9D7503238" lastPageId="10" lastPageNumber="100" pageId="4" pageNumber="94" type="description">
<paragraph id="BF5B36EBB00EFFFBFF99FAA9D46D3254" blockId="4.[100,770,1287,1976]" box="[100,266,1287,1306]" pageId="4" pageNumber="94">
<emphasis id="8D90EAF9B00EFFFBFF99FAA9D46D3254" bold="true" box="[100,266,1287,1306]" italics="true" pageId="4" pageNumber="94">3.1.3. Description</emphasis>
</paragraph>
<paragraph id="BF5B36EBB00EFFFBFF79FA8DD7783220" blockId="4.[100,770,1287,1976]" pageId="4" pageNumber="94">
See
<tableCitation id="F2660350B00EFFFBFF56FA8DD5953278" box="[171,242,1315,1334]" captionStart="Table 1" captionStartId="10.[100,150,150,166]" captionTargetPageId="10" captionText="Table 1 Measurements (μm) and proportions (%) of Setaphyes dentatus, including total values from the different studied populations split by sex (values per specimen and population can be found in Supplementary Table 3. Abbreviations: LTS, lateral terminal spine length; MSW5, maximum sternal width (measured at segment 5); n, number of measured specimens; S, segment length (followed by number of corresponding segment); Sd, standard deviation; SW10, standard sternal width (measured at segment 10); TL, total trunk length." pageId="4" pageNumber="94">Table 1</tableCitation>
and Supplementary Table III for measurements and dimensions and
<tableCitation id="F2660350B00EFFFBFF15FA91D44A321C" box="[232,301,1343,1362]" captionStart="Table 2" captionStartId="10.[100,150,784,800]" captionTargetPageId="10" captionText="Table 2 Summary of nature and arrangement of cuticular elevations, processes, spines, tubes, setae, sensory spots and nephridiopores in Setaphyes dentatus. Abbreviations: ce, cuticular elevation; cp, cuticular process; LD, laterodorsal; lts, lateral terminal spine; LV, lateroventral; MD, middorsal; ne, nephridiopore; PD, paradorsal; ps, penile spine; SD, subdorsal; se, seta; ss, sensory spot; tu, tube; VL, ventrolateral; VM, ventromedial; * indicates unpaired structures; 6 and ♀ indicates sexually dimorphic characters." pageId="4" pageNumber="94">Table 2</tableCitation>
for location of middorsal cuticular specializations, setae, tubes, nephridiopores and sensory spots.
</paragraph>
<paragraph id="BF5B36EBB00EFFFBFF79FAD9D754317F" blockId="4.[100,770,1287,1976]" pageId="4" pageNumber="94">
Ring 00 of mouth cone with nine, equally sized outer oral styles composed of a single, flexible unit, wider at base, with a fringed sheath, tapering progressively toward the distal, pointed tip (
<figureCitation id="27DF2A6EB00EFFFBFDABFA01D7F4328C" box="[598,659,1455,1474]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs. 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFD63FA01D7DA328C" box="[670,701,1455,1474]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4D</figureCitation>
). Outer oral styles located anterior to each introvert sector, except in the middorsal position (sector 6) where a style is missing (
<figureCitation id="27DF2A6EB00EFFFBFDAAFA48D7EC32B4" box="[599,651,1510,1530]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Fig. 3</figureCitation>
). Inner rings of mouth cone not observed, herein details on the morphology and distribution of inner oral styles are not provided.
</paragraph>
<paragraph id="BF5B36EBB00EFFFBFF79F994D007367C" blockId="4.[100,770,1287,1976]" lastBlockId="4.[818,1488,148,1980]" pageId="4" pageNumber="94">
Introvert with six rings of spinoscalids and 10 longitudinal sectors, each limited by the position of two adjacent primary spinoscalids (
<figureCitation id="27DF2A6EB00EFFFBFD21F9F8D51731CB" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFF7CF9DCD5F931CB" box="[129,158,1650,1669]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFF56F9DCD58431CB" box="[171,227,1650,1669]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–D</figureCitation>
). Ring 01 with 10 primary spinoscalids, conspicuously larger than the other ones; primary spinoscalids composed of a basal, rectangular, wide sheath and a distal, elongated, flexible, distally pointed end-piece; basal sheath equipped with a row of elongated fringes, longer in the middle region of the spinoscalid than on the lateral margins (
<figureCitation id="27DF2A6EB00EFFFBFF3CF953D59A305E" box="[193,253,1789,1808]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFEF1F953D44E305F" box="[268,297,1789,1809]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFECAF953D40B305E" box="[311,364,1789,1809]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–C</figureCitation>
). Ring 02 with 10 spinoscalids, arranged medially in each sector; spinoscalids on this and following rings are also composed of a basal, rectangular, wide sheath and a distal, elongated, flexible, distally pointed end-piece; basal sheath appears hairy at its proximal end and terminates into a short fringe (
<figureCitation id="27DF2A6EB00EFFFBFDD7F8C3D70630CE" box="[554,609,1901,1920]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFD91F8C3D7EE30CE" box="[620,649,1901,1920]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFD6CF8C3D7A130CE" box="[657,710,1901,1920]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–B</figureCitation>
). Ring 03 with 20 spinoscalids, arranged as two in each sector (
<figureCitation id="27DF2A6EB00EFFFBFD65F827D7B430D2" box="[664,723,1929,1948]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFD1FF827D79830D2" box="[738,767,1929,1948]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFF99F80BD5FE30F6" box="[100,153,1957,1976]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–B</figureCitation>
). Ring 04 with 10 spinoscalids, arranged medially in each sector (
<figureCitation id="27DF2A6EB00EFFFBFCC7FF3AD61737E9" box="[826,880,148,167]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC87FF3AD6F037E9" box="[890,919,148,167]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC63FF3AD6B437E9" box="[926,979,148,167]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–B</figureCitation>
). Ring 05 with 20 spinoscalids, arranged as two in each sector (
<figureCitation id="27DF2A6EB00EFFFBFC84FF1ED6D4378D" box="[889,947,176,195]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC3DFF1ED6BA378D" box="[960,989,176,195]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC14FF1ED17A378D" box="[1001,1053,176,195]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–B</figureCitation>
). Ring 06 with four, smaller spinoscalids, arranged medially in sectors 1, 3, 6 and 9 (
<figureCitation id="27DF2A6EB00EFFFBFB53FF65D1823791" box="[1198,1253,203,223]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB0DFF62D06A3791" box="[1264,1293,204,223]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFAEBFF65D02D3791" box="[1302,1354,203,223]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–B</figureCitation>
). The location of spinoscalids throughout rings 02–06 follows a strict pattern around the introvert: sectors 1, 3, 6 and 9 bear seven spinoscalids, while the remaining sectors carry six spinoscalids (
<figureCitation id="27DF2A6EB00EFFFBFB48FEB1D18A367C" box="[1205,1261,287,306]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB04FEB1D071367C" box="[1273,1302,287,306]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFAE2FEB1D033367C" box="[1311,1364,287,307]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–B</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB00EFFFBFCACFE95D13E36EC" blockId="4.[818,1488,148,1980]" pageId="4" pageNumber="94">
A ring of 14 hairy trichoscalids present posterior to the spinoscalid rings, arranged as two in the odd-numbered sectors (except sector 1 with a single trichoscalid) and one in the even-numbered sectors of the introvert (
<figureCitation id="27DF2A6EB00EFFFBFC6EFE21D6AC36EC" box="[915,971,399,418]" captionStart="Fig" captionStartId="4.[100,130,1039,1056]" captionTargetBox="[144,726,149,1009]" captionTargetId="figure-1246@4.[142,727,148,1011]" captionTargetPageId="4" captionText="Fig. 3. Diagram of mouth cone and introvert in Setaphyes dentatus showing the distribution and type of scalids by ring and sector. Abbreviations:? position of inner oral styles not revealed; S, sector followed by number of sector." figureDoi="http://doi.org/10.5281/zenodo.8164212" httpUri="https://zenodo.org/record/8164212/files/figure.png" pageId="4" pageNumber="94">Figs 3</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC2AFE21D69336EC" box="[983,1012,399,418]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC03FE21D12836EC" box="[1022,1103,399,418]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5A–B, D</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB00EFFFBFCACFE05D1223507" blockId="4.[818,1488,148,1980]" pageId="4" pageNumber="94">
Neck with four dorsal and two ventral sclerotized placids (
<figureCitation id="27DF2A6EB00EFFFBFA90FE05D0AC36F0" box="[1389,1483,427,446]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2A–B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFCCFFE69D6023694" box="[818,869,455,474]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B–C</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC92FE68D6E93694" box="[879,910,454,474]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5D</figureCitation>
). Dorsal placids rectangular, mesial ones broader (ca. 24 μm wide at the base) than lateral ones (ca. 22 μm wide at the base) (
<figureCitation id="27DF2A6EB00EFFFBFA7BFE4CD0AC36B8" box="[1414,1483,482,502]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFCCFFE50D62A355C" box="[818,845,510,530]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFCAAFE50D611355F" box="[855,886,510,529]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5D</figureCitation>
). Ventral placids (ca. 22–23 μm wide at the base) similar to the dorsal ones but much more elongated, becoming thinner towards the lateral sides (
<figureCitation id="27DF2A6EB00EFFFBFC4DFD98D1743507" box="[944,1043,566,585]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2A, C</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFBE0FD98D15E3507" box="[1053,1081,566,585]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4C</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB00EFFFBFCACFDFCD05431B7" blockId="4.[818,1488,148,1980]" pageId="4" pageNumber="94">
Trunk with eleven segments (
<figureCitation id="27DF2A6EB00EFFFBFB8DFDFCD1CE352B" box="[1136,1193,594,613]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2</figureCitation>
AB; 6AF). Segment 1 with one tergal, two episternal and one trapezoidal, midsternal plate; remaining segments with one tergal and two sternal cuticular plates (
<figureCitation id="27DF2A6EB00EFFFBFA8FFD24D0CB35D3" box="[1394,1452,650,669]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2</figureCitation>
AD; 4B–C, F–G; 6A–F). Sternal plates reaching their maximum width at segment 7, almost constant in width throughout the trunk, progressively tapering at the last trunk segments. Sternal cuticular plates relatively narrow (MSW-5:TL average ratio = 24 %). Middorsal elevations on segments 2–6, rectangular, narrow, distally blunted, not projecting beyond the posterior margin of segments (
<figureCitation id="27DF2A6EB00EFFFBFB38FC9FD06A340B" box="[1221,1293,817,837]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFAEAFC9FD02C340B" box="[1303,1355,817,837]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B, F</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFAA8FC9FD0EB340A" box="[1365,1420,817,836]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="4" pageNumber="94">6A, C</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFA6BFC9CD0AC340B" box="[1430,1483,817,837]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="4" pageNumber="94">7C–D</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFCCFFCE3D629342E" box="[818,846,845,864]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="4" pageNumber="94">8B</figureCitation>
). Middorsal processes on segments 1 and 7–9, similar in morphology but exceeding the posterior margin of segment (
<figureCitation id="27DF2A6EB00EFFFBFB10FCC7D0553432" box="[1261,1330,873,892]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFAC6FCC7D00B3432" box="[1339,1388,873,892]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B, F</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFA89FCC7D0CF3432" box="[1396,1448,873,892]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="4" pageNumber="94">6A, E</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFA4DFCC7D63E34D6" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="4" pageNumber="94">7B, F–G</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC99FC2BD6E434D6" box="[868,899,901,920]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="4" pageNumber="94">8H</figureCitation>
). Paradorsal butterfly to trident-like intracuticular atria associated to middorsal structures (
<figureCitation id="27DF2A6EB00EFFFBFB8CFC0FD1D934FA" box="[1137,1214,929,948]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB32FC0FD06E34FA" box="[1231,1289,929,948]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B, F</figureCitation>
). Lateral margin of middorsal structures surrounded by short, thick cuticular hairs; and posterior ends covered by tufts of elongated, thick hairs whose tips usually surpass the posterior margin of segment (
<figureCitation id="27DF2A6EB00EFFFBFB08FC5AD05D3346" box="[1269,1338,1012,1032]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFABFFC5BD0C13346" box="[1346,1446,1013,1032]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="4" pageNumber="94">7C–D, F–G</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFA53FC5BD0AF3346" box="[1454,1480,1013,1032]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="4" pageNumber="94">8B</figureCitation>
) (those of segment 1 remarkably shorter). Middorsal processes progressively longer towards the posterior segments, reaching their maximum length on segment 9
<figureCitation id="27DF2A6EB00EFFFBFC04FBE6D1273312" box="[1017,1088,1096,1116]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFBB7FBE6D1033312" box="[1098,1124,1096,1116]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4F</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB92FBE6D1ED3312" box="[1135,1162,1096,1116]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="4" pageNumber="94">6E</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB69FBE7D1A03315" box="[1172,1223,1096,1116]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="4" pageNumber="94">7F–G</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB2CFBE6D1973315" box="[1233,1264,1096,1115]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="4" pageNumber="94">8H</figureCitation>
). Cuticular scars (likely glandular cell outlets) as minute, dot-shaped, rounded to oval perforations throughout the cuticle on segments 1–11 (
<figureCitation id="27DF2A6EB00EFFFBFB09FB2ED03133DD" box="[1268,1366,1152,1171]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2A–D</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFAA2FB2ED0A333DD" box="[1375,1476,1152,1171]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B–C, F–G</figureCitation>
), also present at the base of the middorsal processes and elevations; the number and arrangement of these structures vary greatly among the specimens, not showing any clear pattern. Up to two pairs of cuticular ridges in subdorsal (one pair) and laterodorsal (one pair) positions on segments 2–4; an unpaired middorsal cuticular ridge and up to three pairs in subdorsal (one pair) and laterodorsal (two pairs) positions on segments 5–10; one pair of ventrolateral cuticular ridges on segments 2–10, with adjacent, minute glandular cell outlets (
<figureCitation id="27DF2A6EB00EFFFBFAD1FAF1D0F7323C" box="[1324,1424,1375,1395]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Fig. 2A–D</figureCitation>
). Conspicuous reticulate pattern of cuticular wrinkles, as a net-like ornamentation, across the most anterior dorsal and ventrolateral margins, at the overlapping area between following segments, on segments 2–10 (
<figureCitation id="27DF2A6EB00EFFFBFCC7FA61D6F932AC" box="[826,926,1487,1506]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2A–D</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC57FA61D17432AC" box="[938,1043,1487,1506]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B–C, F–G</figureCitation>
). Cuticular hairs acicular, non-bracteate, scattered throughout the trunk on segments 1–10, except in ventromedial position, denser at the tergosternal junctions, not following any particular pattern. Pachycycli and ball-and-socket joints conspicuous on segments 2–8, reduced on most posterior segments (
<figureCitation id="27DF2A6EB00EFFFBFB03F990D039311F" box="[1278,1374,1598,1618]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Fig. 2A–D</figureCitation>
). Apodemes on segments 9–10 (
<figureCitation id="27DF2A6EB00EFFFBFC18F9F4D13A3123" box="[997,1117,1626,1645]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2A–B, D</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB9BF9F4D1FC3123" box="[1126,1179,1626,1645]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4F–G</figureCitation>
). Primary pectinate fringes finely serrated (
<figureCitation id="27DF2A6EB00EFFFBFC72F9D8D69731C7" box="[911,1008,1654,1673]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="4" pageNumber="94">Figs 6A–F</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC00F9D8D1E331C7" box="[1021,1156,1654,1673]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="4" pageNumber="94">8B–C, E, G–H</figureCitation>
); secondary pectinate fringes as a wavy, quite inconspicuous single line in laterodorsal and ventrolateral positions at the anterior most region of the segments (
<figureCitation id="27DF2A6EB00EFFFBFAADF900D0FB318F" box="[1360,1436,1710,1729]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2A</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFA54F900D0A2318F" box="[1449,1477,1710,1729]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="4" pageNumber="94">8E</figureCitation>
). Muscular scars as rounded to oval, hairless areas in laterodorsal and ventromedial positions on segments 1–10 (
<figureCitation id="27DF2A6EB00EFFFBFB3AF948D04F31B7" box="[1223,1320,1766,1785]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Fig. 2A–D</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB00EFFF9FCACF8ACD40C3037" blockId="4.[818,1488,148,1980]" lastBlockId="6.[100,771,1587,1969]" lastPageId="6" lastPageNumber="96" pageId="4" pageNumber="94">
Segment 1 with anterior dorsal margin finely denticulated, posteriorly followed by a transverse area of cuticular wrinkles forming a net-like band, broader towards the lateral than in the middle region (
<figureCitation id="27DF2A6EB00EFFFBFA57F897D62A3027" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFCA4F8FBD6133027" box="[857,884,1877,1897]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="4" pageNumber="94">4B</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC82F8FBD6FB3027" box="[895,924,1877,1897]" captionStart="Fig" captionStartId="6.[100,130,1452,1469]" captionTargetBox="[226,1360,149,1424]" captionTargetId="figure-351@6.[225,1362,148,1425]" captionTargetPageId="6" captionText="Fig. 5. SEM photographs of the head of Setaphyes dentatus (ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids." figureDoi="http://doi.org/10.5281/zenodo.8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="4" pageNumber="94">5D</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFC55F8F8D6BA3026" box="[936,989,1877,1897]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="4" pageNumber="94">7A–B</figureCitation>
). Anterolateral margins of the tergal plate as short, wide, distally rounded extensions (
<figureCitation id="27DF2A6EB00EFFFBFB78F8DFD18F30CA" box="[1157,1256,1905,1925]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2A–C</figureCitation>
;
<figureCitation id="27DF2A6EB00EFFFBFB09F8DFD07730CA" box="[1268,1296,1905,1924]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="4" pageNumber="94">6B</figureCitation>
). Middorsal process with paradorsal, butterfly to trident-like atria of associated paradorsal sensory spots located near the posterior margin of the segment (
<figureCitation id="27DF2A6EB00EFFFBFA7BF807D0AC30F2" box="[1414,1483,1961,1980]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="4" pageNumber="94">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FF99F99DD5E73108" box="[100,128,1587,1606]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="6" pageNumber="96">4B</figureCitation>
). Anterior margin of episternal plates with a pair of ridges forming small, quadrangular depressions; anterior margin of midsternal plate with a single, large rectangular depression (
<figureCitation id="27DF2A6EB00CFFF9FDFAF9C5D70C3130" box="[519,619,1643,1662]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2A, C</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FD88F9C5D7F53130" box="[629,658,1643,1662]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="6" pageNumber="96">4C</figureCitation>
). Episternal plates with usually five, scattered, minute, dot-shaped glandular cell outlets forming a quincunx (
<figureCitation id="27DF2A6EB00CFFF9FE83F90DD48031F8" box="[382,487,1699,1718]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2A, C</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FE09F90DD77631F8" box="[500,529,1699,1718]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="6" pageNumber="96">4C</figureCitation>
). Trapezoidal midsternal plate, wider at the base (ca. 34 μm wide at the most anterior margin, ca. 56 μm wide at the most posterior margin; average ratio = 61 %), with wavy lateral margins at the middle region (
<figureCitation id="27DF2A6EB00CFFF9FE0FF958D7363044" box="[498,593,1782,1802]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2A, C</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FDA4F959D7113047" box="[601,630,1782,1802]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="6" pageNumber="96">4C</figureCitation>
). Sensory spots present in paradorsal, subdorsal, laterodorsal and ventrolateral positions (
<figureCitation id="27DF2A6EB00CFFF9FF62F880D599300F" box="[159,254,1838,1858]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2A–C</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FEFBF880D45D300F" box="[262,314,1838,1858]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="6" pageNumber="96">6A–B</figureCitation>
). Sensory spots on this and following segments as oval areas with an oval area of cuticular micropapillae surrounding a single pore (
<figureCitation id="27DF2A6EB00CFFF9FF27F8C8D4463034" box="[218,289,1894,1914]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="6" pageNumber="96">Figs 7E</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FED6F8C8D4073037" box="[299,352,1894,1914]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="6" pageNumber="96">8D–E</figureCitation>
).
</paragraph>
<caption id="EB9B6663B00FFFFAFF99F972D00230CB" ID-DOI="http://doi.org/10.5281/zenodo.8164214" ID-Zenodo-Dep="8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="5" pageNumber="95" startId="5.[100,130,1756,1773]" targetBox="[151,1437,149,1726]" targetPageId="5" targetType="figure">
<paragraph id="BF5B36EBB00FFFFAFF99F972D00230CB" blockId="5.[100,1488,1755,1926]" pageId="5" pageNumber="95">
<emphasis id="8D90EAF9B00FFFFAFF99F972D5FA31A3" bold="true" box="[100,157,1756,1773]" pageId="5" pageNumber="95">Fig. 4.</emphasis>
Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of
<taxonomicName id="78E44D68B00FFFFAFBF2F975D1C131A2" baseAuthorityName="Reinhard" baseAuthorityYear="1881" box="[1039,1190,1755,1772]" class="Allomalorhagida" family="Pycnophyidae" genus="Setaphyes" kingdom="Animalia" pageId="5" pageNumber="95" phylum="Kinorhyncha" rank="species" species="dentatus">
<emphasis id="8D90EAF9B00FFFFAFBF2F975D1C131A2" bold="true" box="[1039,1190,1755,1772]" italics="true" pageId="5" pageNumber="95">Setaphyes dentatus</emphasis>
</taxonomicName>
from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment.
</paragraph>
</caption>
<caption id="EB9B6663B00CFFF9FF99FA02D1D33144" ID-DOI="http://doi.org/10.5281/zenodo.8164218" ID-Zenodo-Dep="8164218" httpUri="https://zenodo.org/record/8164218/files/figure.png" pageId="6" pageNumber="96" startId="6.[100,130,1452,1469]" targetBox="[226,1360,149,1424]" targetPageId="6" targetType="figure">
<paragraph id="BF5B36EBB00CFFF9FF99FA02D1D33144" blockId="6.[100,1488,1451,1546]" pageId="6" pageNumber="96">
<emphasis id="8D90EAF9B00CFFF9FF99FA02D5FB32F3" bold="true" box="[100,156,1452,1469]" pageId="6" pageNumber="96">Fig. 5.</emphasis>
SEM photographs of the head of
<taxonomicName id="78E44D68B00CFFF9FE48FA02D72C32F3" baseAuthorityName="Reinhard" baseAuthorityYear="1881" box="[437,587,1452,1469]" class="Allomalorhagida" family="Pycnophyidae" genus="Setaphyes" kingdom="Animalia" pageId="6" pageNumber="96" phylum="Kinorhyncha" rank="species" species="dentatus">
<emphasis id="8D90EAF9B00CFFF9FE48FA02D72C32F3" bold="true" box="[437,587,1452,1469]" italics="true" pageId="6" pageNumber="96">Setaphyes dentatus</emphasis>
</taxonomicName>
(ZMB 12751). A: Introvert sector 1 (midventral); B: introvert sector 10 (ventrolateral); C: detail of introvert ring 01 showing the primary spinoscalid morphology; D: detail of neck (dorsal view) and anterior half of segment 1 tergal plate with focus on its cuticular ornamentation. Abbreviations: bs, basal sheath; co, cuticular ornamentation; dpl, dorsal placid; ep, end-piece of spinoscalid; psp, primary spinoscalid; S1, segment 1; sp, spinoscalid (followed by number of corresponding ring); ts, trichoscalid. Lambda symbols (Λ) mark attachment points of scalids.
</paragraph>
</caption>
<paragraph id="BF5B36EBB00CFFF9FF79F82CD0DE3047" blockId="6.[100,771,1587,1969]" lastBlockId="6.[817,1488,1587,1969]" pageId="6" pageNumber="96">
Segment 2 with middorsal elevation with paradorsal, butterfly to trident-like atria of associated paradorsal sensory spots (near the posterior margin) (
<figureCitation id="27DF2A6EB00CFFF9FC77F99DD6B53109" box="[906,978,1587,1607]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FC20F99DD6903109" box="[989,1015,1587,1607]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="6" pageNumber="96">4B</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FBFFF99DD1783108" box="[1026,1055,1587,1606]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="6" pageNumber="96">6A</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FBD7F99AD12A3108" box="[1066,1101,1587,1607]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="6" pageNumber="96">7 C</figureCitation>
), surrounded by tufts of elongated, thick hairs with tips usually surpassing the posterior segment margin (
<figureCitation id="27DF2A6EB00CFFF9FA78F9E1D0AB312C" box="[1413,1484,1615,1634]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="6" pageNumber="96">Figs 6A</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FCCCF9C5D6293130" box="[817,846,1643,1662]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="6" pageNumber="96">7C</figureCitation>
). Unpaired seta in paradorsal position, and paired setae in lateroventral position; females with sexual dimorphism, an additional, pair of setae in ventrolateral position lacking in males. Males with sexually dimorphic tubes in ventromedial position lacking in females(
<figureCitation id="27DF2A6EB00CFFF9FA97F911D0AB319C" box="[1386,1484,1727,1746]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2A, C</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FCCCF975D62831A0" box="[817,847,1755,1774]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="6" pageNumber="96">8A</figureCitation>
). Sensory spots in paradorsal (near the posterior margin), subdorsal, laterodorsal and ventromedial positions (
<figureCitation id="27DF2A6EB00CFFF9FB52F958D0683044" box="[1199,1295,1782,1802]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2A–C</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FAE5F959D02C3044" box="[1304,1355,1783,1802]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="6" pageNumber="96">4B–C</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FAA9F958D0EF3044" box="[1364,1416,1782,1802]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="6" pageNumber="96">6A–B</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FA6CF959D0C93047" box="[1425,1454,1782,1802]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="6" pageNumber="96">7C</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB00CFFF7FCACF8BCD58030C7" blockId="6.[817,1488,1587,1969]" lastBlockId="8.[100,770,1882,1957]" lastPageId="8" lastPageNumber="98" pageId="6" pageNumber="96">
Segment 3 with middorsal elevation as on the preceding segment (
<figureCitation id="27DF2A6EB00CFFF9FCC7F880D6E6300C" box="[826,897,1838,1858]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="6" pageNumber="96">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FC76F880D6C1300C" box="[907,934,1838,1858]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="6" pageNumber="96">4B</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FC4DF880D6AA300F" box="[944,973,1838,1857]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="6" pageNumber="96">6A</figureCitation>
;
<figureCitation id="27DF2A6EB00CFFF9FC2AF881D691300F" box="[983,1014,1838,1858]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="6" pageNumber="96">7D</figureCitation>
). Unpaired seta in paradorsal position, and paired setae in laterodorsal and lateroventral positions. Females with an additional, pair of setae in ventromedial position, more mesial than the ventromedial sensory spots. Sensory spots in paradorsal (near the posterior margin), subdorsal, laterodorsal and ventromedial positions; ventromedial sensory spots present in females only (
<figureCitation id="27DF2A6EB002FFF7FDA3F8F4D7A73023" box="[606,704,1882,1901]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–C</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FD36F8F4D7983023" box="[715,767,1882,1901]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4B–C</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FF99F8D8D5D430C7" box="[100,179,1910,1929]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6A–B, D</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FF43F8D8D5BA30C7" box="[190,221,1910,1929]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="8" pageNumber="98">7D</figureCitation>
).
</paragraph>
<caption id="EB9B6663B00DFFF8FF99F888D12430D3" ID-DOI="http://doi.org/10.5281/zenodo.8164220" ID-Zenodo-Dep="8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="7" pageNumber="97" startId="7.[100,130,1830,1847]" targetBox="[233,1355,150,1802]" targetPageId="7" targetType="figure">
<paragraph id="BF5B36EBB00DFFF8FF99F888D12430D3" blockId="7.[100,1488,1829,1949]" pageId="7" pageNumber="97">
<emphasis id="8D90EAF9B00DFFF8FF99F888D5FA3079" bold="true" box="[100,157,1830,1847]" pageId="7" pageNumber="97">Fig. 6.</emphasis>
SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of
<taxonomicName id="78E44D68B00DFFF8FBDEF888D1DD3079" baseAuthorityName="Reinhard" baseAuthorityYear="1881" box="[1059,1210,1830,1847]" class="Allomalorhagida" family="Pycnophyidae" genus="Setaphyes" kingdom="Animalia" pageId="7" pageNumber="97" phylum="Kinorhyncha" rank="species" species="dentatus">
<emphasis id="8D90EAF9B00DFFF8FBDEF888D1DD3079" bold="true" box="[1059,1210,1830,1847]" italics="true" pageId="7" pageNumber="97">Setaphyes dentatus</emphasis>
</taxonomicName>
from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots.
</paragraph>
</caption>
<caption id="EB9B6663B002FFF7FF99F9BAD4533070" ID-DOI="http://doi.org/10.5281/zenodo.8164222" ID-Zenodo-Dep="8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="8" pageNumber="98" startId="8.[100,130,1556,1573]" targetBox="[107,766,149,1528]" targetPageId="8" targetType="figure">
<paragraph id="BF5B36EBB002FFF7FF99F9BAD4533070" blockId="8.[100,771,1556,1854]" pageId="8" pageNumber="98">
<emphasis id="8D90EAF9B002FFF7FF99F9BAD5C5316B" bold="true" box="[100,162,1556,1573]" pageId="8" pageNumber="98">Fig. 7.</emphasis>
SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of
<taxonomicName id="78E44D68B002FFF7FE34F9E9D7073116" baseAuthorityName="Reinhard" baseAuthorityYear="1881" box="[457,608,1607,1624]" class="Allomalorhagida" family="Pycnophyidae" genus="Setaphyes" kingdom="Animalia" pageId="8" pageNumber="98" phylum="Kinorhyncha" rank="species" species="dentatus">
<emphasis id="8D90EAF9B002FFF7FE34F9E9D7073116" bold="true" box="[457,608,1607,1624]" italics="true" pageId="8" pageNumber="98">Setaphyes dentatus</emphasis>
</taxonomicName>
from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots.
</paragraph>
</caption>
<paragraph id="BF5B36EBB002FFF7FF79F83CD1E43659" blockId="8.[100,770,1882,1957]" lastBlockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 4 with middorsal elevation as on the preceding segments (
<figureCitation id="27DF2A6EB002FFF7FCC7FF3AD6E537E9" box="[826,898,148,167]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC70FF3AD6CD37E9" box="[909,938,148,167]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4A</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC48FF3AD68937E9" box="[949,1006,148,167]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6A, C</figureCitation>
). Unpaired seta in paradorsal position, and paired setae in lateroventral and ventromedial positions. Paired sensory spots in paradorsal (near the posterior margin), subdorsal, laterodorsal and ventromedial positions (
<figureCitation id="27DF2A6EB002FFF7FBE2FF49D1E537B5" box="[1055,1154,231,251]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB6DFF46D1A337B5" box="[1168,1220,232,251]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4B–C</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB2CFF49D04237B5" box="[1233,1317,231,251]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6A, C–D</figureCitation>
), the latter more lateral than the ventromedial setae.
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFEB1D07136B8" blockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 5 with middorsal elevation as on the preceding segments (
<figureCitation id="27DF2A6EB002FFF7FCC7FE95D6183600" box="[826,895,315,334]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC7AFE95D6DE3600" box="[903,953,315,334]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4B, F</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC3CFE95D6BA3600" box="[961,989,315,334]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6C</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC18FE95D1663600" box="[997,1025,315,334]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8B</figureCitation>
). Unpaired seta in paradorsal position, and paired setae in laterodorsal, lateroventral and ventromedial positions; laterodorsal setae longitudinally aligned with those of segment 3. Laterodorsal setae longitudinally aligned with those of segment 3. Paired sensory spots in paradorsal (near the posterior margin), subdorsal, laterodorsal and ventromedial positions (
<figureCitation id="27DF2A6EB002FFF7FB59FE69D0623694" box="[1188,1285,455,474]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FAEDFE69D0233694" box="[1296,1348,455,474]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4B–C</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FAB2FE69D0E33694" box="[1359,1412,455,474]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6C–D</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FA72FE69D0A33694" box="[1423,1476,455,474]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8B–D</figureCitation>
), the latter more lateral than the ventromedial setae.
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFE50D1F635CF" blockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 6 with middorsal elevation as on the preceding segments (
<figureCitation id="27DF2A6EB002FFF7FCC7FDB4D6183560" box="[826,895,538,558]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC7AFDB4D6C63560" box="[903,929,538,558]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4F</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC57FDB4D6A13563" box="[938,966,538,557]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6C</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC33FDB4D68D3563" box="[974,1002,538,557]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8B</figureCitation>
). Unpaired seta in paradorsal position, and paired in lateroventral position. Paired sensory spots in paradorsal (near the posterior margin), subdorsal, laterodorsal and ventromedial positions (
<figureCitation id="27DF2A6EB002FFF7FCC7FDC0D6FD35CF" box="[826,922,622,641]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC59FDC0D69735CF" box="[932,1008,622,641]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4C, F–G</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC07FDC0D14835CF" box="[1018,1071,622,641]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6C–D</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FBC7FDC0D1E135CF" box="[1082,1158,622,641]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8B–C, F</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFD24D0E8359B" blockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 7 with middorsal process extending beyond the posterior margin of segment (
<figureCitation id="27DF2A6EB002FFF7FC0EFD08D15C35F7" box="[1011,1083,678,697]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FBBBFD08D10635F7" box="[1094,1121,678,697]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="8" pageNumber="98">7F</figureCitation>
). Similar to segment
<quantity id="781C9B0EB002FFF7FAD3FD08D03435F7" box="[1326,1363,678,697]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="98" unit="in" value="5.0">5 in</quantity>
the arrangement of setae and sensory spots (
<figureCitation id="27DF2A6EB002FFF7FB97FD6CD1AD359B" box="[1130,1226,706,725]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB29FD6CD060359B" box="[1236,1287,706,725]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4F–G</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FAEFFD6CD023359B" box="[1298,1348,706,725]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6C–E</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FAB3FD6CD0E3359B" box="[1358,1412,706,725]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8C, E</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFD70D0673466" blockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 8 with middorsal process longer than that of the preceding segment (
<figureCitation id="27DF2A6EB002FFF7FC72FD57D6B13443" box="[911,982,761,781]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC1CFD57D69C3443" box="[993,1019,761,781]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4F</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FBFBFD54D1463442" box="[1030,1057,761,781]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="8" pageNumber="98">7F</figureCitation>
). Similar to segment
<quantity id="781C9B0EB002FFF7FB11FD57D0763442" box="[1260,1297,761,780]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="98" unit="in" value="4.0">4 in</quantity>
the arrangement of setae and sensory spots (
<figureCitation id="27DF2A6EB002FFF7FBE6FCBBD11C3467" box="[1051,1147,789,809]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB78FCBBD1DE3466" box="[1157,1209,789,809]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4F–G</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB3EFCBBD1923466" box="[1219,1269,789,809]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6E–F</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFC9FD0D934A2" blockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 9 with middorsal process longer than that of the preceding segment (
<figureCitation id="27DF2A6EB002FFF7FC73FCE3D6B2342F" box="[910,981,845,865]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2B</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC22FCE3D69E342F" box="[991,1017,845,865]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4F</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FBFEFCE3D179342F" box="[1027,1054,845,865]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6E</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FBD5FCE0D13C342E" box="[1064,1115,845,865]" captionStart="Fig" captionStartId="8.[100,130,1556,1573]" captionTargetBox="[107,766,149,1528]" captionTargetId="figure-802@8.[106,767,148,1529]" captionTargetPageId="8" captionText="Fig. 7. SEM photographs of two females (110412 Huelva_sp1: A; 110412 Huelva_sp2: B, D; specimens damaged during study and not deposited) from Huelva and adults of indeterminate sex of Setaphyes dentatus from Huelva (ZMB 12751: E; ZMB 12752: F), and Algeciras (ZMB 12750: C; G) populations. A: detail of the reticular, net-like ornamentation on the dorsal, anterior margin of segment 1; B: middorsal to lateroventral view of segment 1; C: middorsal elevation of segment 2; D: middorsal elevation of segment 3; E: detail of laterodorsal sensory spot of segment 1; F: middorsal view of segments 8–9; G: middorsal process of segment 9. Abbreviations: co, cuticular ornamentation; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164222" httpUri="https://zenodo.org/record/8164222/files/figure.png" pageId="8" pageNumber="98">7F–G</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB9BFCE3D1E2342E" box="[1126,1157,845,864]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8H</figureCitation>
). Unpaired seta in paradorsal position, and paired in lateroventral position; females with an additional pair of setae in ventromedial position lacking in males (
<figureCitation id="27DF2A6EB002FFF7FAD2FC2BD0C134D6" box="[1327,1446,901,920]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–B, D</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FA53FC2BD0AC34D6" box="[1454,1483,901,920]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4G</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FCCFFC0FD62A34FA" box="[818,845,929,948]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6F</figureCitation>
). Sensory spots in paradorsal (near the posterior margin), subdorsal, laterodorsal, and ventrolateral positions (
<figureCitation id="27DF2A6EB002FFF7FB46FC13D051349E" box="[1211,1334,957,976]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Fig. 2A–B, D</figureCitation>
). Nephridiopore as small opening surrounded by short tubes in lateroventral position.
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFC5BD12833E1" blockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 10 without middorsal cuticular specialization. Setae in lateroventral position (
<figureCitation id="27DF2A6EB002FFF7FBF1FBBED117336A" box="[1036,1136,1040,1060]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A, D</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB86FBBED1F2336A" box="[1147,1173,1040,1060]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6F</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB62FBBFD1D9336D" box="[1183,1214,1040,1060]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8H</figureCitation>
). Sensory spots in subdorsal, laterodorsal and ventrolateral positions, the two latter between patches of conspicuous, parallel cuticular ridges, extending throughout the laterodorsal to ventrolateral areas (
<figureCitation id="27DF2A6EB002FFF7FB9AFBCAD1AC3339" box="[1127,1227,1124,1143]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A, D</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FB28FBCAD06C3339" box="[1237,1291,1124,1143]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4E, G</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FAE8FBCAD02A3339" box="[1301,1357,1124,1143]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8G–H</figureCitation>
). Tergal plate with rounded posterior margin; sternal plates distally straight in females, more pointed in males.
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFB16D6B73274" blockId="8.[818,1488,148,1339]" pageId="8" pageNumber="98">
Segment 11 without middorsal cuticular specialization. Males with two pairs of stout, thick penile spines (
<figureCitation id="27DF2A6EB002FFF7FB51FB7AD19133A9" box="[1196,1270,1236,1255]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FAFFFB7AD07933A9" box="[1282,1310,1236,1255]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4E</figureCitation>
). Short lateral terminal spines, longer in males than in females (males LTS:TL average ratio 21.21 %; females LTS:TL average ratio 15.66 %) (
<figureCitation id="27DF2A6EB002FFF7FAB0FAA5D0AC3251" box="[1357,1483,1291,1311]" captionStart="Fig" captionStartId="3.[100,130,1763,1780]" captionTargetBox="[264,1323,150,1735]" captionTargetId="figure-8@3.[263,1324,148,1736]" captionTargetPageId="3" captionText="Fig. 2. Line art illustration of Setaphyes dentatus. A: Male, ventral view; B: Male, dorsal view; C: Female, segments 1–3, ventral view; D: female, segments 9–11, ventral view. Scale bar: 100 μm. Abbreviations: ap, apodeme; co, cuticular ornamentation, reticular, net-like structure; cr, cuticular ridge; cs, cuticular scar; cw, cuticular wrinkles; dpl, dorsal placid; ica, intracuticlular atria; ldse, laterodorsal setae; ldss, laterodorsal sensory spot; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; ms, muscular scar; ne, nephridiopore; pcr, patch of cuticular ridges; pdse, paradorsal setae; pdss, paradorsal sensory spot; ps, penile spine; sdss, subdorsal sensory spot; spf, secondary pectinate fringe; vlse, ventrolateral setae; vlss, ventrolateral sensory spot; vmse, ventromedial setae; vmtu, ventromedial tube; vmss, ventromedial sensory spot; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.8164210" httpUri="https://zenodo.org/record/8164210/files/figure.png" pageId="8" pageNumber="98">Figs 2A–B, D</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FCCFFA89D6063275" box="[818,865,1319,1339]" captionStart="Fig" captionStartId="5.[100,130,1756,1773]" captionTargetBox="[151,1437,149,1726]" captionTargetId="figure-8@5.[150,1438,148,1728]" captionTargetPageId="5" captionText="Fig. 4. Light micrographs of females (ZMB 12617: A, D; ZMB 12409: B–C, F–H) and male (ZMB 12418: E, I) of Setaphyes dentatus from Sylt. A: Sector 1 (midventral) of the introvert; B: Dorsal view on segments 1–5; C: Ventral view on segments 1–6; D: Mouth cone, outer oral styles (ring 01); E: Cuticular ornamentation on segment 10. Note that punctuate appearance represents preservation artefact. F: Dorsal view on segments 5–11; G: Ventral view on segments 6–11. H: Female LTS; I: Male LTS. Lambda symbols (Λ) mark anterior attachment points of spinoscalids on introvert (A). Abbreviations: co, cuticular ornamentation; cs, cuticular scars; dpl, dorsal placid; ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; oos, outer oral style; pcr, patch of cuticular ridges; pdse, paradorsal setae; ps, penile spine; psp, primary spinoscalid; sp, spinoscalid (followed by the number of corresponding ring); ts, trichoscalid; vlse, ventrolateral setae; vmse, ventromedial setae; vpl, ventral placid. Numbers after abbreviations indicate the corresponding segment." figureDoi="http://doi.org/10.5281/zenodo.8164214" httpUri="https://zenodo.org/record/8164214/files/figure.png" pageId="8" pageNumber="98">4H–I</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC96FA89D6FB3275" box="[875,924,1319,1339]" captionStart="Fig" captionStartId="7.[100,130,1830,1847]" captionTargetBox="[233,1355,150,1802]" captionTargetId="figure-8@7.[232,1356,148,1803]" captionTargetPageId="7" captionText="Fig. 6. SEM photographs of female (ZMB 12746: B, D, F) and adult of indeterminate sex (ZMB 12750: A, C, E) of Setaphyes dentatus from the Mediterranean coast of the Iberian Peninsula, Algeciras population. A: dorsal view of segments 1–4; B: ventral view of segments 1–3; C: dorsal view of segments 4–7; D: ventral view of segments 3–7; E: dorsal view of segments 7–10; F: ventral view of segments 8–10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pdse, paradorsal setae; ppf, primary pectinate fringe; vlse, ventrolateral setae; vmse, ventromedial setae. Numbers after abbreviations indicate the number of the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164220" httpUri="https://zenodo.org/record/8164220/files/figure.png" pageId="8" pageNumber="98">6E–F</figureCitation>
;
<figureCitation id="27DF2A6EB002FFF7FC5BFA89D6A23274" box="[934,965,1319,1338]" captionStart="Fig" captionStartId="9.[100,130,1718,1735]" captionTargetBox="[267,1322,243,1689]" captionTargetId="figure-8@9.[265,1323,242,1690]" captionTargetPageId="9" captionText="Fig. 8. SEM photographs of female (ZMB 12746: G), male (ZMB 12747: A, C, F) and adults of indeterminate sex (ZMB 12748: B, D; ZMB 12750: E; ZMB 12745: H) of Setaphyes dentatus from the Atlantic and the Mediterranean coasts of the Iberian Peninsula and from the Black Sea coast of Türkiye, Huelva (G), Algeciras (A–F), Türkiye (H) populations. A: ventral view of segments 1–2; B: left side of the tergal plate, segments 5–6; C: right side of segments 5–7; D: left side of the tergal plate, segment 5, laterodorsal setae and sensory spot; E: laterodorsal side of the tergal plate, segment 7; F: right lateroventral seta of segment 6; G: ventral view of segment 10; H: left side of the tergal plate, segment 10. Abbreviations: ldse, laterodorsal setae; lts, lateral terminal spine; lvse, lateroventral setae; mde, middorsal elevation; mdp, middorsal process; pcr, patch of cuticular ridges; pdse, paradorsal setae; vmtu, ventromedial tube. Numbers after abbreviations indicate the corresponding segment. Dashed circles mark sensory spots." figureDoi="http://doi.org/10.5281/zenodo.8164224" httpUri="https://zenodo.org/record/8164224/files/figure.png" pageId="8" pageNumber="98">8G</figureCitation>
).
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCCFFAF1D150323C" blockId="8.[818,1488,1375,1980]" box="[818,1079,1375,1394]" pageId="8" pageNumber="98">
<emphasis id="8D90EAF9B002FFF7FCCFFAF1D150323C" bold="true" box="[818,1079,1375,1394]" italics="true" pageId="8" pageNumber="98">3.1.4. Intraspecific variation</emphasis>
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACFAD5D1C331EB" blockId="8.[818,1488,1375,1980]" pageId="8" pageNumber="98">
Due to the preservation conditions of the studied material, the pattern of sensory spots throughout the trunk could not be fully confirmed in all the LM specimens. The remaining cuticular characters of taxonomic relevance for pycnophyids (i.e. setae, middorsal cuticular specializations, spines, glandular cell outlets, and ornamentation) could be examined in detail in the Anatolian population (
<specimenCount id="A9E2FD62B002FFF7FADBF9A8D0C63157" box="[1318,1441,1542,1561]" pageId="8" pageNumber="98" type="female">four females</specimenCount>
,
<specimenCount id="A9E2FD62B002FFF7FA50F9A9D605317B" pageId="8" pageNumber="98" type="adult">one adult</specimenCount>
specimen for SEM),
<location id="BA3B6030B002FFF7FBCAF98CD1DF317B" LSID="urn:lsid:plazi:treatment:374D87FDB008FFF5FCCFFB80D7503238:BA3B6030B002FFF7FBCAF98CD1DF317B" box="[1079,1208,1570,1589]" name="North Frisian" pageId="8" pageNumber="98">North Frisian</location>
(
<specimenCount id="A9E2FD62B002FFF7FB34F98CD028317B" box="[1225,1359,1570,1589]" pageId="8" pageNumber="98" type="female">three females</specimenCount>
,
<specimenCount id="A9E2FD62B002FFF7FAA0F98DD0A2317B" box="[1373,1477,1570,1590]" pageId="8" pageNumber="98" type="male">two males</specimenCount>
), Italian (
<specimenCount id="A9E2FD62B002FFF7FC86F990D6B2311F" box="[891,981,1598,1617]" pageId="8" pageNumber="98" type="adult">one adult</specimenCount>
specimen) and Iberian (Pontevedra,
<specimenCount id="A9E2FD62B002FFF7FAC9F991D0F8311F" box="[1332,1439,1598,1618]" pageId="8" pageNumber="98" type="female">one female</specimenCount>
,
<specimenCount id="A9E2FD62B002FFF7FA55F990D60C3123" pageId="8" pageNumber="98" type="male">four males</specimenCount>
; Algeciras,
<specimenCount id="A9E2FD62B002FFF7FC28F9F4D14A3123" box="[981,1069,1626,1645]" pageId="8" pageNumber="98" type="male">one male</specimenCount>
,
<specimenCount id="A9E2FD62B002FFF7FBCAF9F4D1C73123" box="[1079,1184,1626,1645]" pageId="8" pageNumber="98" type="female">one female</specimenCount>
,
<specimenCount id="A9E2FD62B002FFF7FB54F9F4D0773123" box="[1193,1296,1626,1645]" pageId="8" pageNumber="98" type="adult">three adult</specimenCount>
specimens mounted for SEM; Huelva,
<specimenCount id="A9E2FD62B002FFF7FC2BF9D8D14931C7" box="[982,1070,1654,1673]" pageId="8" pageNumber="98" type="adult">one adult</specimenCount>
specimen mounted for SEM; and Tarragona,
<specimenCount id="A9E2FD62B002FFF7FCCFF93CD6CE31EB" box="[818,937,1682,1701]" pageId="8" pageNumber="98" type="female">four females</specimenCount>
,
<specimenCount id="A9E2FD62B002FFF7FC4EF93CD14531EB" box="[947,1058,1682,1701]" pageId="8" pageNumber="98" type="male">three males</specimenCount>
) populations.
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACF900D6D13193" blockId="8.[818,1488,1375,1980]" pageId="8" pageNumber="98">
<emphasis id="8D90EAF9B002FFF7FCACF900D155318F" bold="true" box="[849,1074,1710,1729]" pageId="8" pageNumber="98">Anatolian population:</emphasis>
All specimens lack the ventromedial setae on segment 8.
</paragraph>
<paragraph id="BF5B36EBB002FFF7FCACF948D0D93002" blockId="8.[818,1488,1375,1980]" pageId="8" pageNumber="98">
<emphasis id="8D90EAF9B002FFF7FCACF948D13231B6" bold="true" box="[849,1109,1765,1785]" pageId="8" pageNumber="98">North Frisian population:</emphasis>
ventromedial seta on segment 2 absent on one sternal plate in
<specimenCount id="A9E2FD62B002FFF7FBECF8ACD10E305A" box="[1041,1129,1793,1813]" pageId="8" pageNumber="98" type="male">one male</specimenCount>
(
<specimenCode id="EF429E90B002FFF7FB8BF8ACD18D305B" box="[1142,1258,1794,1813]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="98" type="Museum">ZMB 12418</specimenCode>
); ventromedial setae on segment 6 present on one sternal plate in
<specimenCount id="A9E2FD62B002FFF7FB28F8B0D058307E" box="[1237,1343,1821,1841]" pageId="8" pageNumber="98" type="female">one female</specimenCount>
(
<specimenCode id="EF429E90B002FFF7FAB3F8B0D0A3307E" box="[1358,1476,1821,1841]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="98" type="Museum">ZMB 12413</specimenCode>
); ventromedial setae on segment 9 absent in
<specimenCount id="A9E2FD62B002FFF7FB36F894D0553002" box="[1227,1330,1849,1869]" pageId="8" pageNumber="98" type="female">one female</specimenCount>
(
<specimenCode id="EF429E90B002FFF7FAC3F897D0D43002" box="[1342,1459,1849,1868]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="98" type="Museum">ZMB 12408</specimenCode>
).
</paragraph>
<paragraph id="BF5B36EBB002FFF5FCACF8FBD7503238" blockId="8.[818,1488,1375,1980]" lastBlockId="10.[100,770,1239,1398]" lastPageId="10" lastPageNumber="100" pageId="8" pageNumber="98">
<emphasis id="8D90EAF9B002FFF7FCACF8FBD1723026" bold="true" box="[849,1045,1877,1896]" pageId="8" pageNumber="98">Iberian population:</emphasis>
laterodorsal seta on one side of the tergal plate on segment 2 present in
<specimenCount id="A9E2FD62B002FFF7FBD9F8DCD11B30CA" box="[1060,1148,1905,1925]" pageId="8" pageNumber="98" type="male">one male</specimenCount>
(
<specimenCode id="EF429E90B002FFF7FB77F8DFD19830CA" box="[1162,1279,1905,1924]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="98" type="Museum">ZMB 12715</specimenCode>
); laterodorsal seta on segment 3 absent on one side of the tergal plate in
<specimenCount id="A9E2FD62B002FFF7FAC4F823D0F530EE" box="[1337,1426,1933,1952]" pageId="8" pageNumber="98" type="male">one male</specimenCount>
(
<specimenCode id="EF429E90B002FFF7FA5CF823D60830F2" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="98" type="Museum">ZMB 12640</specimenCode>
) and in one SEM specimen (
<specimenCode id="EF429E90B002FFF7FB8CF807D18430F2" box="[1137,1251,1961,1980]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="98" type="Museum">ZMB 12751</specimenCode>
); laterodorsal seta on one side of the tergal plate on segment 4 present in
<specimenCount id="A9E2FD62B000FFF5FDEBFB76D71C33A4" box="[534,635,1239,1259]" pageId="10" pageNumber="100" type="female">one female</specimenCount>
(
<specimenCode id="EF429E90B000FFF5FD78FB76D79033A4" box="[645,759,1239,1259]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="100" type="Museum">ZMB 12716</specimenCode>
); ventromedial seta on one sternal plate on segment 4 not detected in
<specimenCount id="A9E2FD62B000FFF5FD1DFB5AD5F5326C" pageId="10" pageNumber="100" type="male">one male</specimenCount>
(
<specimenCode id="EF429E90B000FFF5FF5CFAA1D470326C" box="[161,279,1295,1314]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="100" type="Museum">ZMB 12640</specimenCode>
); ventromedial setae on segment 6 present in
<specimenCount id="A9E2FD62B000FFF5FD1DFABED5F53270" pageId="10" pageNumber="100" type="male">one male</specimenCount>
and
<specimenCount id="A9E2FD62B000FFF5FF3EFA82D44C3270" box="[195,299,1323,1343]" pageId="10" pageNumber="100" type="female">one female</specimenCount>
(
<specimenCode id="EF429E90B000FFF5FEC5FA85D4CE3270" box="[312,425,1323,1342]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="100" type="Museum">ZMB 12638</specimenCode>
and
<specimenCode id="EF429E90B000FFF5FE27FA82D72D3270" box="[474,586,1323,1343]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="100" type="Museum">ZMB 12714</specimenCode>
) and on the lateral half of the sternal plate in
<specimenCount id="A9E2FD62B000FFF5FEA6FAE9D4A53214" box="[347,450,1351,1370]" pageId="10" pageNumber="100" type="female">one female</specimenCount>
(
<specimenCode id="EF429E90B000FFF5FE30FAE9D7273214" box="[461,576,1351,1370]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="100" type="Museum">ZMB 12716</specimenCode>
); ventromedial setae on segment 8 absent in
<specimenCount id="A9E2FD62B000FFF5FEB8FACDD4CB3238" box="[325,428,1379,1398]" pageId="10" pageNumber="100" type="female">one female</specimenCount>
(
<specimenCode id="EF429E90B000FFF5FE45FACDD74A3238" box="[440,557,1379,1398]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="100" type="Museum">ZMB 12643</specimenCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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