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<document id="A9D20E7818E49ADA81B803D5B655B3CD" ID-DOI="10.5281/zenodo.5174820" ID-ISSN="1942-1354" ID-Zenodo-Dep="5174820" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628338230777" checkinUser="felipe" docAuthor="Miller, Robert B. &amp; Stange, Lionel A." docDate="2012" docId="2D0587A25457FFC0FF7FBD00FA0EFBA2" docLanguage="en" docName="InsectaMundi.0250.1-65.pdf" docOrigin="Insecta Mundi 2012 (250)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Speleon Miller and Stange 2012, new genus" docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="D13CFFDA5445FFD3FFBFB822FFAFFFD8" masterDocTitle="The cave mouth antlions of Australia (Neuroptera: Myrmeleontidae)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="17" updateTime="1699045397822" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="13E9B75D2D2277D3083A45122F0EA0EA">The cave mouth antlions of Australia (Neuroptera: Myrmeleontidae)</mods:title>
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<mods:namePart id="078D6AC7303B8E072D795CC1CF4FD945">Miller, Robert B.</mods:namePart>
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<mods:namePart id="8CBC3729F8B6A80B6D629CA6D48FE3D0">Stange, Lionel A.</mods:namePart>
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<mods:title id="289E22A5623755977F9EA209BE73EE81">Insecta Mundi</mods:title>
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<mods:date id="16EC225016675663ED49FFBDC933D3A5">2012</mods:date>
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<mods:number id="9BE514AF87E13EB0A2F264B50C03EBB9">2012-10-26</mods:number>
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<treatment id="2D0587A25457FFC0FF7FBD00FA0EFBA2" ID-DOI="http://doi.org/10.5281/zenodo.5187531" ID-Zenodo-Dep="5187531" LSID="urn:lsid:plazi:treatment:2D0587A25457FFC0FF7FBD00FA0EFBA2" httpUri="http://treatment.plazi.org/id/2D0587A25457FFC0FF7FBD00FA0EFBA2" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
<subSubSection id="EDB6653F5457FFC1FF7FBD00FD6AFAE2" box="[192,709,1314,1338]" pageId="18" pageNumber="17" type="nomenclature">
<paragraph id="A51336B45457FFC1FF7FBD00FD6AFAE2" blockId="18.[192,709,1314,1338]" box="[192,709,1314,1338]" pageId="18" pageNumber="17">
<heading id="FE5B81D85457FFC1FF7FBD00FD6AFAE2" bold="true" box="[192,709,1314,1338]" fontSize="10" level="2" pageId="18" pageNumber="17" reason="3">
<emphasis id="97D8EAA65457FFC1FF7FBD00FD6AFAE2" bold="true" box="[192,709,1314,1338]" pageId="18" pageNumber="17">
<taxonomicName id="62AC4D375457FFC1FF7FBD00FD86FAE2" ID-CoL="7K8K" authority="Miller and Stange" authorityName="Miller and Stange" authorityYear="2012" box="[192,553,1314,1338]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="97D8EAA65457FFC1FF7FBD00FE83FAE2" bold="true" box="[192,300,1314,1338]" italics="true" pageId="18" pageNumber="17">Speleon</emphasis>
Miller and Stange
</taxonomicName>
,
<taxonomicNameLabel id="8CEB57DD5457FFC1FD89BD00FD6AFAE2" box="[566,709,1314,1338]" pageId="18" pageNumber="17" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EDB6653F5457FFC1FF7FBD40FBFBFAA2" box="[192,1108,1378,1402]" pageId="18" pageNumber="17" type="type_taxon">
<paragraph id="A51336B45457FFC1FF7FBD40FBFBFAA2" blockId="18.[192,1108,1378,1402]" box="[192,1108,1378,1402]" pageId="18" pageNumber="17">
<emphasis id="97D8EAA65457FFC1FF7FBD40FEC3FAA2" bold="true" box="[192,364,1378,1402]" pageId="18" pageNumber="17">
<typeStatus id="7A1788165457FFC1FF7FBD40FEABFAA2" box="[192,260,1378,1402]" pageId="18" pageNumber="17">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="62AC4D375457FFC1FEC7BD40FC9FFAA2" ID-CoL="4YHFJ" authority="Miller and Stange" authorityName="Miller and Stange. Diagnoses" authorityYear="2012" box="[376,816,1378,1402]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="yallingup">
<emphasis id="97D8EAA65457FFC1FEC7BD40FDFFFAA2" box="[376,592,1378,1402]" italics="true" pageId="18" pageNumber="17">Speleon yallingup</emphasis>
Miller and Stange
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="EDB6653F5457FFC1FF7FBD80FDD0FA02" pageId="18" pageNumber="17" type="diagnosis">
<paragraph id="A51336B45457FFC1FF7FBD80FDD0FA02" blockId="18.[192,1440,1442,1498]" pageId="18" pageNumber="17">
<emphasis id="97D8EAA65457FFC1FF7FBD80FE1FFA62" bold="true" box="[192,432,1442,1466]" pageId="18" pageNumber="17">Diagnosis. Adult:</emphasis>
antennal rim with several setae that project over eye; Millers organ developed; midfemoral sense hair not developed.
</paragraph>
</subSubSection>
<subSubSection id="EDB6653F5457FFC0FF7FBE20FBFDFDC2" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17" type="description">
<paragraph id="A51336B45457FFC0FF7FBE20FDBDFEE2" blockId="18.[192,1441,1538,1915]" lastBlockId="19.[192,1441,194,539]" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
<emphasis id="97D8EAA65457FFC1FF7FBE20FE65F9C2" bold="true" box="[192,458,1538,1562]" pageId="18" pageNumber="17">Description. Adult:</emphasis>
body
<quantity id="62549B515457FFC1FDACBE20FD3EF9C2" box="[531,657,1538,1562]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="18" pageNumber="17" unit="mm" value="19.5" valueMax="21.0" valueMin="18.0">18-21 mm</quantity>
, forewing
<quantity id="62549B515457FFC1FCAEBE20FC20F9C2" box="[785,911,1538,1562]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="18" pageNumber="17" unit="mm" value="25.5" valueMax="27.0" valueMin="24.0">24-27 mm</quantity>
, hindwing
<quantity id="62549B515457FFC1FBA8BE20FB39F9C2" box="[1047,1174,1538,1562]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="18" pageNumber="17" unit="mm" value="24.5" valueMax="26.0" valueMin="23.0">23-26 mm</quantity>
.
<emphasis id="97D8EAA65457FFC1FB1EBE20FA85F9C2" bold="true" box="[1185,1322,1538,1562]" pageId="18" pageNumber="17">Structure</emphasis>
: antenna sturdy, moderately long; antennal fossae separated by about lateral width of scape; antennal rim with several short setae that project over eye; palps short, distal palpomere weakly swollen, oval palpomaculum about in center; pronotum longer than greatest width; forecoxa about l.5 longer than greatest diameter as viewed laterally; foreleg shorter than midleg which is shorter than hindleg; forefemoral sense hair short, inconspicuous; midfemur without distinguishable sense hair; forefemur about five times longer than middle diameter, shorter than foretibia; tibial spurs absent or very short (no more than one-half basitarsal length); tarsus shorter than one-half length of tibia, basitarsus of hindleg longer than distal tarsomere; pretarsal claws small, about as long as tarsomere IV, not opposable; male pilula axillaris present; forewings slightly longer than hindwing; forewing vein CuP originates slightly distad of basal crossvein; forewing costal area simple, without interconnected crossveins, highest near middle of wing; Banksian lines absent; hindwing posteriorly at highest point higher than greatest presectoral area; Millers organ present (small gland-like structure) on abdominal sternite I; abdomen shorter than wings; male ectoproct without lobes; male paramere plate-like, gonarcus strongly arched; female terminalia without digging setae; posterior gonapophysis short, incurved (
<figureCitation id="3D972A315456FFC0FC19B920FBB3FEC2" box="[934,1052,258,282]" captionStart="Figures 22-25" captionStartId="54.[192,286,1158,1180]" captionText="Figures 22-25. Speleon yallingup Miller and Stange. 22) Dorsal view, adult body and base of wings. 23) Adult wings. 24) Adult legs. 25) Female terminalia." pageId="19" pageNumber="18">Figure 25</figureCitation>
), anterior gonapophysis present, but usually weakly swollen.
</paragraph>
<paragraph id="A51336B45456FFC0FF4FB960FBFDFDC2" blockId="19.[192,1441,194,539]" pageId="19" pageNumber="18">
<emphasis id="97D8EAA65456FFC0FF4FB960FEEDFE82" bold="true" box="[240,322,322,346]" pageId="19" pageNumber="18">Larva</emphasis>
(based on
<taxonomicName id="62AC4D375456FFC0FE7EB961FDF9FE82" authorityName="Miller and Stange. Diagnoses" authorityYear="2012" box="[449,598,322,346]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="yallingup">
<emphasis id="97D8EAA65456FFC0FE7EB961FDF9FE82" box="[449,598,322,346]" italics="true" pageId="19" pageNumber="18">S. yallingup</emphasis>
</taxonomicName>
): mandible longer than head capsule, three teeth increasing in length distally; distance between teeth much longer than distance between mandibular base and basal tooth; mesoscutum without specialized setae medially; mesothoracic spiracle borne on very short tubercle; anterior thoracic scolus about three times longer than middle diameter, swollen distally; posterior mesothoracic scolus a little longer than middle diameter; metathoracic scoli weakly developed, much broader than long; abdominal scoli absent; abdominal spiracles small, flush with integument; sternite VIII without submedian teeth; abdominal segment IX about one-half as long as wide.
</paragraph>
</subSubSection>
<subSubSection id="EDB6653F5456FFC0FF7FBA60FD96FD82" box="[192,569,578,602]" pageId="19" pageNumber="18" type="distribution">
<paragraph id="A51336B45456FFC0FF7FBA60FD96FD82" blockId="19.[192,569,578,602]" box="[192,569,578,602]" pageId="19" pageNumber="18">
<emphasis id="97D8EAA65456FFC0FF7FBA60FED9FD82" bold="true" box="[192,374,578,602]" pageId="19" pageNumber="18">Distribution.</emphasis>
Only
<collectingCountry id="DDBB76245456FFC0FE01BA60FD9AFD82" box="[446,565,578,602]" name="Australia" pageId="19" pageNumber="18">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="EDB6653F5456FFC0FF7FBAA0FEF9FCC2" pageId="19" pageNumber="18" type="biology_ecology">
<paragraph id="A51336B45456FFC0FF7FBAA0FEF9FCC2" blockId="19.[192,1440,642,794]" pageId="19" pageNumber="18">
<emphasis id="97D8EAA65456FFC0FF7FBAA0FE85FD42" bold="true" box="[192,298,642,666]" pageId="19" pageNumber="18">Biology</emphasis>
. The larvae of
<taxonomicName id="62AC4D375456FFC0FE63BAA0FD94FD42" authorityName="Miller and Stange" authorityYear="2012" box="[476,571,642,666]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FE63BAA0FD94FD42" box="[476,571,642,666]" italics="true" pageId="19" pageNumber="18">Speleon</emphasis>
</taxonomicName>
are found in Zone 2C which is sun and rain protected, but warm enough to support species living in horizontal, sometimes lichen filled (
<taxonomicName id="62AC4D375456FFC0FC6BBA81FB22FD62" authorityName="Miller &amp; Stange" authorityYear="2012" box="[980,1165,674,698]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="97D8EAA65456FFC0FC6BBA81FB22FD62" box="[980,1165,674,698]" italics="true" pageId="19" pageNumber="18">S. cavernicolus</emphasis>
</taxonomicName>
), rock cracks, with 90 degree angles, characteristic of areas with shale. The mostly immobile larvae were nearly exposed because they lived in tight cracks usually oriented horizontally. They only had a partial covering of dust on their bodies.
</paragraph>
</subSubSection>
<subSubSection id="EDB6653F5456FFC0FF7FBB60FD3EFBE3" pageId="19" pageNumber="18" type="discussion">
<paragraph id="A51336B45456FFC0FF7FBB60FD3EFBE3" blockId="19.[192,1441,834,1083]" pageId="19" pageNumber="18">
<emphasis id="97D8EAA65456FFC0FF7FBB60FEFAFC82" bold="true" box="[192,341,834,858]" pageId="19" pageNumber="18">Discussion</emphasis>
.
<taxonomicName id="62AC4D375456FFC0FEDCBB60FE6DFC82" authorityName="Miller and Stange" authorityYear="2012" box="[355,450,834,858]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FEDCBB60FE6DFC82" box="[355,450,834,858]" italics="true" pageId="19" pageNumber="18">Speleon</emphasis>
</taxonomicName>
is distinct from the other 18 genera of
<taxonomicName id="62AC4D375456FFC0FC20BB60FBF4FC82" authorityName="Banks" authorityYear="1899" box="[927,1115,834,858]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Dendroleontini">Dendroleontini</taxonomicName>
in
<collectingCountry id="DDBB76245456FFC0FB3CBB60FB55FC82" box="[1155,1274,834,858]" name="Australia" pageId="19" pageNumber="18">Australia</collectingCountry>
. In the key to genera presented by New (1985), this genus would key out to
<taxonomicName id="62AC4D375456FFC0FC22BB40FB55FCA2" authority="Banks. However" authorityName="Banks. However" authorityYear="1913" box="[925,1274,866,890]" class="Insecta" family="Myrmeleontidae" genus="Compsoleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FC22BB40FB9FFCA2" box="[925,1072,866,890]" italics="true" pageId="19" pageNumber="18">Compsoleon</emphasis>
Banks. However
</taxonomicName>
, the pretarsal claws are opposable in
<taxonomicName id="62AC4D375456FFC0FE67BBA0FDC4FC42" authorityName="Banks. However" authorityYear="1913" box="[472,619,898,922]" class="Insecta" family="Myrmeleontidae" genus="Compsoleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FE67BBA0FDC4FC42" box="[472,619,898,922]" italics="true" pageId="19" pageNumber="18">Compsoleon</emphasis>
</taxonomicName>
which is probably closely related to
<taxonomicName id="62AC4D375456FFC0FB9BBBA1FB11FC42" authorityName="Esben-Petersen" authorityYear="1915" box="[1060,1214,899,922]" class="Insecta" family="Myrmeleontidae" genus="Froggattisca" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FB9BBBA1FB11FC42" box="[1060,1214,899,922]" italics="true" pageId="19" pageNumber="18">Froggattisca</emphasis>
</taxonomicName>
. Several species of the speciose genera
<taxonomicName id="62AC4D375456FFC0FE12BB80FDD8FC62" authority="Banks" authorityName="Banks. One" authorityYear="1913" box="[429,631,930,954]" class="Insecta" family="Myrmeleontidae" genus="Glenoleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FE12BB80FD89FC62" box="[429,550,930,954]" italics="true" pageId="19" pageNumber="18">Glenoleon</emphasis>
Banks
</taxonomicName>
and
<taxonomicName id="62AC4D375456FFC0FD10BB81FBC4FC62" authority="Esben-Petersen" authorityName="Esben-Petersen" authorityYear="1917" box="[687,1131,930,954]" class="Insecta" family="Myrmeleontidae" genus="Austrogymnocnemia" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FD10BB81FC08FC62" box="[687,935,931,954]" italics="true" pageId="19" pageNumber="18">Austrogymnocnemia</emphasis>
Esben-Petersen
</taxonomicName>
are known and the larvae have a mesoscutal setal lure which is not found in the larvae of
<taxonomicName id="62AC4D375456FFC0FC03BBE0FBB4FC02" authorityName="Miller and Stange" authorityYear="2012" box="[956,1051,962,986]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FC03BBE0FBB4FC02" box="[956,1051,962,986]" italics="true" pageId="19" pageNumber="18">Speleon</emphasis>
</taxonomicName>
. The presence of Millers organ, a small gland-like structure located on sternite I (function unknown) is an apomorphy of
<taxonomicName id="62AC4D375456FFC0FB4ABBC0FAFBFC22" authorityName="Miller and Stange" authorityYear="2012" box="[1269,1364,994,1018]" class="Insecta" family="Myrmeleontidae" genus="Speleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FB4ABBC0FAFBFC22" box="[1269,1364,994,1018]" italics="true" pageId="19" pageNumber="18">Speleon</emphasis>
</taxonomicName>
found also in
<taxonomicName id="62AC4D375456FFC0FEACBC21FD61FBC2" authority="Esben-Petersen" authorityName="Esben-Petersen" authorityYear="1917" box="[275,718,1026,1050]" class="Insecta" family="Myrmeleontidae" genus="Austrogymnocnemia" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FEACBC21FDA6FBC2" box="[275,521,1027,1050]" italics="true" pageId="19" pageNumber="18">Austrogymnocnemia</emphasis>
Esben-Petersen
</taxonomicName>
,
<taxonomicName id="62AC4D375456FFC0FD67BC20FBA5FBC2" authority="Esben-Petersen" authorityName="Esben-Petersen" authorityYear="1923" box="[728,1034,1026,1050]" class="Insecta" family="Myrmeleontidae" genus="Platyleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FD67BC20FCE7FBC2" box="[728,840,1026,1050]" italics="true" pageId="19" pageNumber="18">Platyleon</emphasis>
Esben-Petersen
</taxonomicName>
and some
<taxonomicName id="62AC4D375456FFC0FB3BBC20FA0EFBC2" authority="Banks. There" authorityName="Banks. There" box="[1156,1441,1026,1050]" class="Insecta" family="Myrmeleontidae" genus="Glenoleon" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="97D8EAA65456FFC0FB3BBC20FB53FBC2" box="[1156,1276,1026,1050]" italics="true" pageId="19" pageNumber="18">Glenoleon</emphasis>
Banks. There
</taxonomicName>
are three species known in this genus.
</paragraph>
</subSubSection>
<subSubSection id="EDB6653F5456FFC0FF7FBC40FA0EFBA2" box="[192,1441,1122,1147]" pageId="19" pageNumber="18" type="etymology">
<paragraph id="A51336B45456FFC0FF7FBC40FA0EFBA2" blockId="19.[192,1441,1122,1147]" box="[192,1441,1122,1147]" pageId="19" pageNumber="18">
<emphasis id="97D8EAA65456FFC0FF7FBC40FEF7FBA2" bold="true" box="[192,344,1122,1146]" pageId="19" pageNumber="18">Etymology.</emphasis>
The genus name is from the Greek
<emphasis id="97D8EAA65456FFC0FCBCBC40FCE3FBA2" box="[771,844,1122,1146]" italics="true" pageId="19" pageNumber="18">spelos</emphasis>
(cave) and
<emphasis id="97D8EAA65456FFC0FC6CBC40FBA9FBA2" box="[979,1030,1122,1146]" italics="true" pageId="19" pageNumber="18">leon</emphasis>
(lion), and is masculine in gender.
</paragraph>
</subSubSection>
</treatment>
</document>