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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1186.113701" ID-Pensoft-Pub="1313-2970-1186-105" ID-Pensoft-UUID="B376A41AC8175F4982A4C5B04AA95CC5" ID-ZooBank="A9064FB1EEE64FA3BABD5517DBD77F81" ModsDocID="1313-2970-1186-105" checkinTime="1702462906702" checkinUser="pensoft" docAuthor="Constant, Jerome, Pham, Thai-Hong, Le, Cuong Viet Canh, Vu, Trung Thanh &amp; Nguyen, Hoai Thu Thi" docDate="2023" docId="F762F99D941B58FDAB41B1771CF29320" docLanguage="en" docName="ZooKeys 1186: 105-122" docOrigin="ZooKeys 1186" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zookeys.1186.113701" docTitle="Connelicita phongdienensis Constant &amp; Pham 2023, sp. nov." docType="treatment" docUuid="556D7EFE-52A9-4D76-869B-7C59FDDC754C" docUuidSource="ZooBank" docVersion="1" id="B376A41AC8175F4982A4C5B04AA95CC5" lastPageNumber="105" masterDocId="B376A41AC8175F4982A4C5B04AA95CC5" masterDocTitle="The tropiduchid planthopper genus Connelicita Wang &amp; Bourgoin, 2015: two new species from Central Vietnam and new records (Hemiptera, Fulgoromorpha, Tropiduchidae)" masterLastPageNumber="122" masterPageNumber="105" pageNumber="105" updateTime="1702462906702" updateUser="pensoft">
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<mods:title id="CDCFF41028D290DF6B842CEAA8586E4B">The tropiduchid planthopper genus Connelicita Wang &amp; Bourgoin, 2015: two new species from Central Vietnam and new records (Hemiptera, Fulgoromorpha, Tropiduchidae)</mods:title>
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<mods:namePart id="A4FCEB883733524FB79153704B2D30F2">Constant, Jerome</mods:namePart>
<mods:nameIdentifier id="2D0A501BD7BB624E6B2093F80BE77883" type="ORCID">https://orcid.org/0000-0003-0254-0863</mods:nameIdentifier>
<mods:affiliation id="33B65FC96CAC825C6E359B617D6909F0">Royal Belgian Institute of Natural Sciences, O. D. Taxonomy &amp; Phylogeny - Entomology, Vautier street 29, B- 1000 Brussels, Belgium</mods:affiliation>
<mods:nameIdentifier id="05B65D33039758B58989A6EF7A9F4E1D" type="email">jconstant@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="239248EE7DC44CC9A6767F0CC2F0D98D">Pham, Thai-Hong</mods:namePart>
<mods:nameIdentifier id="53DB854053D1403687E20391FB379264" type="ORCID">https://orcid.org/0000-0002-4763-3679</mods:nameIdentifier>
<mods:affiliation id="66701F9A710EEE8640F94B6A802F0259">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="F5BC4DB3C25EDD0CDDF36AE556C6DE47" type="email">phamthai1976@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="327007A428C27EBC173D079FEDDAD531">Le, Cuong Viet Canh</mods:namePart>
<mods:nameIdentifier id="B7A1A3D49CDFBFE7AA1C95BF7C62F0BD" type="ORCID">https://orcid.org/0000-0002-1430-6305</mods:nameIdentifier>
<mods:affiliation id="26FBC4C98AE8A477D86281898369068D">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam</mods:affiliation>
<mods:nameIdentifier id="DF1BB432EB81BE1FA3A78000AFA79E75" type="email">lcvcuong@misr.vast.vn</mods:nameIdentifier>
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<mods:namePart id="ECA984CA88D200D85683212456D419F3">Vu, Trung Thanh</mods:namePart>
<mods:nameIdentifier id="A55952DEEE55074FEE1511464BF33DFC" type="ORCID">https://orcid.org/0009-0002-8987-1672</mods:nameIdentifier>
<mods:affiliation id="69C36D7A2161042A3B0005597186EF5D">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="FB8B4BBCEE71419AA6A2B51D53796FE8">Nguyen, Hoai Thu Thi</mods:namePart>
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<mods:affiliation id="6783EE1C92A94CDA5F6A373AA6D920CA">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<treatment id="F762F99D941B58FDAB41B1771CF29320" LSID="urn:lsid:zoobank.org:act:556D7EFE-52A9-4D76-869B-7C59FDDC754C" httpUri="http://treatment.plazi.org/id/F762F99D941B58FDAB41B1771CF29320" lastPageNumber="105" pageId="0" pageNumber="105">
<subSubSection id="04661890D892719844DED13CF4B3D802" pageId="0" pageNumber="105" type="nomenclature">
<paragraph id="4881B40637E672BFFEBDCEB2588D6A93" pageId="0" pageNumber="105">
<taxonomicName id="398133CBF9F5CC92A088628A92A07B0B" LSID="https://zoobank.org/556D7EFE-52A9-4D76-869B-7C59FDDC754C" authority="Constant &amp; Pham" authorityName="Constant &amp; Pham" authorityYear="2023" class="Insecta" family="Tropiduchidae" genus="Connelicita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Connelicita phongdienensis" order="Hemiptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="phongdienensis" status="sp. nov.">Connelicita phongdienensis Constant &amp; Pham</taxonomicName>
<taxonomicNameLabel id="C11A69836523FC4B1C6DFEEE5238639E" pageId="0" pageNumber="105">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="B6E3475A3A625F924E17FCDCE6C30DD8" pageId="0" pageNumber="105" type="description">
<paragraph id="6FB2A4378CE0C04581CCE34FC5D59E0C" pageId="0" pageNumber="105">
<figureCitation id="76EC082F64BD02A058146067B28AF692" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of the species of Connelicita of Vietnam." figureDoi="10.3897/zookeys.1186.113701.figure5" httpUri="https://binary.pensoft.net/fig/951194" pageId="0" pageNumber="105">Figs 5</figureCitation>
<figureCitation id="1AC5D1084E6C86C82D4CB2BAA5717F21" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Connelicita bachmaensis sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D habitus, anterolateral view." figureDoi="10.3897/zookeys.1186.113701.figure7" httpUri="https://binary.pensoft.net/fig/951196" pageId="0" pageNumber="105">, 7</figureCitation>
<figureCitation id="47CCA3EE3B0A26C227D1259424892757" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Connelicita phongdienensis sp. nov., holotype ♂ A-C detail of head and thorax A dorsal view B lateral view C perpendicular view of frons D right tegmen E right hind wing F right metatibia and metatarsus, ventral view G left metatarsus, ventral view." figureDoi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105">, 8</figureCitation>
<figureCitation id="D4E53C0A06FE9341F4092A5E8976541B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">, 9</figureCitation>
<figureCitation id="CB8265D035279BA5806395C70F4C63DC" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitat of Connelicita phongdienensis sp. nov., Vietnam, Phong Dien, CCRR, 23 May 2023." figureDoi="10.3897/zookeys.1186.113701.figure10" httpUri="https://binary.pensoft.net/fig/951199" pageId="0" pageNumber="105">, 10</figureCitation>
</paragraph>
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<subSubSection id="45CAF915111559D9771BC0BFAFFBD81A" pageId="0" pageNumber="105" type="type material">
<paragraph id="4D00C3274F929AD83CB7840F0C8841A3" pageId="0" pageNumber="105">Type materials.</paragraph>
<paragraph id="647F9F27625847634B4BA006B9A90B23" pageId="0" pageNumber="105">
<emphasis id="DEE8B346A71EA2D807A71DE13186342D" bold="true" italics="true" pageId="0" pageNumber="105">Holotype</emphasis>
♂, Vietnam -
<emphasis id="D1DD047651DB304470CA0957CB570899" bold="true" pageId="0" pageNumber="105">
Thừa
<normalizedToken id="AAFFD120590C4523D3F21C8638D3EBF6" originalValue="Thiên-Hu">Thien-Hu</normalizedToken>
ế Province
</emphasis>
• Phong Dien Nature Reserve, CCRR;
<geoCoordinate id="0BA8B4ECE0B5E73488E9393A03568D9E" degrees="16" direction="north" minutes="30" orientation="latitude" precision="15" seconds="27" value="16.5075">16°30'27&quot;N</geoCoordinate>
,
<geoCoordinate id="222FC435BCB86A7F8BBEABA10E836A33" degrees="107" direction="east" minutes="16" orientation="longitude" precision="15" seconds="05" value="107.26806">107°16'05&quot;E</geoCoordinate>
; 23 May 2023; alt. 350-400 m; J. Constant &amp; L. Semeraro leg.; I.G.: 34.640; VNMN.
</paragraph>
<paragraph id="A9AA293984F872F9195DBBE4C333B75D" pageId="0" pageNumber="105">
<emphasis id="EC7C39BD3482929EE4E5D3CA3104B2AE" bold="true" italics="true" pageId="0" pageNumber="105">Paratypes</emphasis>
, Vietnam •-
<emphasis id="5FA86777ADD81D325712D1F4CD702B24" bold="true" pageId="0" pageNumber="105">
Thừa
<normalizedToken id="6573C651F4975C1137D0DF3F950F9DFA" originalValue="Thiên-Hu">Thien-Hu</normalizedToken>
ế Province
</emphasis>
• 1 ♂; same collection data as for holotype; RBINS • 1 ♀; Phong Dien Nature Reserve, CCRR;
<geoCoordinate id="13BE55F63A2E534B04B682359492D1C5" degrees="16" direction="north" minutes="30" orientation="latitude" precision="15" seconds="27" value="16.5075">16°30'27&quot;N</geoCoordinate>
,
<geoCoordinate id="FC0F6F5445D6EF3DDB729209CB2777E1" degrees="107" direction="east" minutes="16" orientation="longitude" precision="15" seconds="05" value="107.26806">107°16'05&quot;E</geoCoordinate>
; 23 May 2023; alt. 350-400 m; Trung T. Vu leg.; VNMN • 1 ♀; same collection data as for preceding; RBINS.
</paragraph>
</subSubSection>
<subSubSection id="24B082294A6C5584A82885BCD40E5137" pageId="0" pageNumber="105" type="diagnosis">
<paragraph id="0337673330C5A64FC9AF8A84773864B6" pageId="0" pageNumber="105">Diagnosis.</paragraph>
<paragraph id="9DCBD0177B4FEA6559D41FDDB0431E75" pageId="0" pageNumber="105">
The species can be separated from the other species of
<taxonomicName id="42212B7B32E8EA52C574B2E25766D6DC" authorityName="Wang &amp; Bourgoin" authorityYear="2015" class="Insecta" family="Tropiduchidae" genus="Connelicita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Connelicita" order="Hemiptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="5AB6BA972DF1CE4E48F2CF361F49ACC8" italics="true" pageId="0" pageNumber="105">Connelicita</emphasis>
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by the following characters of the male terminalia: anal tube in lateral view abruptly narrowing in distal half (Fig.
<figureCitation id="8EE4E81DAB7ACFD955D21194F9A7A040" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9A</figureCitation>
), posterior margin of pygofer rounded in lateral view (Fig.
<figureCitation id="30F17031373E47BBD313F4883F3EA98F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9A</figureCitation>
), capitulum of gonostylus with a lateral spine in ventral portion, and a dorsal curved digitiform process (Fig.
<figureCitation id="ABB88BA80309A85AD569686FEB9EF651" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9A, D</figureCitation>
), dorsal process of periandrium tooth-shaped, strong, and projecting dorsocephalad (
<emphasis id="29306CE5A6F14DF358AB1959BAD39347" italics="true" pageId="0" pageNumber="105">dpp</emphasis>
- Fig.
<figureCitation id="3F7391D0F1C7B6341E405B185D976C42" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9E</figureCitation>
) and absence of a strong digitiform process directed dorsally at apex of aedeagus (Fig.
<figureCitation id="771B7F4A129F6DCAF02232AFACD225CC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9E-G</figureCitation>
).
</paragraph>
<paragraph id="3232B5301FE32F50B85DAED8B58B6D1B" pageId="0" pageNumber="105">
The closest species are
<taxonomicName id="59BC75EFDFD9F91033C5CB1FA0D3EBA8" lsidName="C. backyensis" pageId="0" pageNumber="105" rank="species" species="backyensis">
<emphasis id="14D04F15939B830A45C2F2DCC5A261E2" italics="true" pageId="0" pageNumber="105">C. backyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="DD618C3A37AF1EDA1BB08ACCA3A54911" lsidName="C. haiphongensis" pageId="0" pageNumber="105" rank="species" species="haiphongensis">
<emphasis id="05D9D8ADA008C835C2F72639C9BC6547" italics="true" pageId="0" pageNumber="105">C. haiphongensis</emphasis>
</taxonomicName>
, from which
<taxonomicName id="ABFB550E9AD329B6DC3A1E5D99B2AD90" lsidName="C. phongdienensis" pageId="0" pageNumber="105" rank="species" species="phongdienensis">
<emphasis id="7D03F87EF84BABE6420B7CF7A72DB725" italics="true" pageId="0" pageNumber="105">C. phongdienensis</emphasis>
</taxonomicName>
Constant &amp; Pham, sp. nov. can be separated by the dorsal digitiform process of the capitulum of the gonostylus, which is absent in both other species (compare with
<bibRefCitation id="4B398A4A316BB6581C197FEB19B84D4C" DOI="https://doi.org/10.11646/zootaxa.4018.4.5" author="Wang, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="105" pagination="563 - 572" refId="B7" refString="Wang, M, Stroinski, A, Bourgoin, T, Zhang, Y, 2015. A new Asian genus of the tribe Elicini (Hemiptera: Fulgoromorpha: Tropiduchidae) with two new species from Vietnam. Zootaxa 4018 (4): 563 - 572, DOI: https://doi.org/10.11646/zootaxa.4018.4.5" title="A new Asian genus of the tribe Elicini (Hemiptera: Fulgoromorpha: Tropiduchidae) with two new species from Vietnam." url="https://doi.org/10.11646/zootaxa.4018.4.5" volume="4018" year="2015">Wang et al. 2015</bibRefCitation>
: figs 12, 31).
</paragraph>
</subSubSection>
<subSubSection id="6AB643F11680015D1DB7A6DEEEBE26AF" pageId="0" pageNumber="105" type="description">
<paragraph id="8ABBA92AEF84394438EB7CA29ACDA4B4" pageId="0" pageNumber="105">Description.</paragraph>
<paragraph id="DDF9D1D7E1083D336B36E2D2178FDAB9" pageId="0" pageNumber="105">
<emphasis id="B6F41A0E1F43BA2E728EDDA3C3637269" bold="true" italics="true" pageId="0" pageNumber="105">Measurements and ratios</emphasis>
: LT: ♂ (
<emphasis id="F403BD76643210BE2B560F7E21B1DCF0" italics="true" pageId="0" pageNumber="105">n</emphasis>
= 2): 10.15-10.77 mm. LTg/BTg = 2.26; LW/BW = 1.89; LV/BV = 0.85; LF/BF = 1.04.
</paragraph>
<paragraph id="F592C3A011F5E669E2B335BBBF2E90D5" pageId="0" pageNumber="105">
<emphasis id="5693B85DE7771E181524F1DC43FBC7CD" bold="true" italics="true" pageId="0" pageNumber="105">Head</emphasis>
(Fig.
<figureCitation id="8B91A8CB5ACC01C1A9D9BA86BB015219" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Connelicita phongdienensis sp. nov., holotype ♂ A-C detail of head and thorax A dorsal view B lateral view C perpendicular view of frons D right tegmen E right hind wing F right metatibia and metatarsus, ventral view G left metatarsus, ventral view." figureDoi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105">8A-C</figureCitation>
): narrower than thorax and elongate, with ~2/3 of vertex length surpassing eyes and genae largely visible from above. Vertex brown, with median yellowish line, weakly concave, ~1.0
<normalizedToken id="A601ABE159EFAA96E012CA9088FAADB1" originalValue="×">x</normalizedToken>
as long in mid-line as broad basally, concave on each side, with lateral margins subparallel, anterior margin roundly produced anteriorly in middle portion and posterior margin excavate. Frons yellowish brown, with irregular brown markings; convex in lateral view; smooth, with median carina reaching dorsal margin but not frontoclypeal suture. Genae yellowish, with brown marking between eye and anterior margin, brown line between antennal insertion and posterior margin, and a dark brown spot near red ocellus. Clypeus pale yellowish brown, with median carina and one curved line on each side, brown, triangular, with median carina in distal portion and with fronto-clypeal suture rounded. Labium yellowish, elongate, and narrow, reaching metacoxae, with apical segment elongate. Eyes globular (not emarginate) protruding laterally. Antennae yellowish brown, with longitudinal black line along underside of pedicel; scape ring-shaped and pedicel cylindrical, longer than broad.
</paragraph>
<paragraph id="D8D94F24B92B5A2B77AE816540C50CD5" pageId="0" pageNumber="105">
<emphasis id="1FB95FACD1569F72ECDDEDF4DDA7502B" bold="true" italics="true" pageId="0" pageNumber="105">Thorax</emphasis>
(Fig.
<figureCitation id="03F8F8E1D38176CCE2C4D66102CFD170" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Connelicita phongdienensis sp. nov., holotype ♂ A-C detail of head and thorax A dorsal view B lateral view C perpendicular view of frons D right tegmen E right hind wing F right metatibia and metatarsus, ventral view G left metatarsus, ventral view." figureDoi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105">8A, B</figureCitation>
): Pronotum brown, with median carina, posterolateral angles and 3 or 4 small yellowish tubercles on sides of disc, yellowish; paranotal fields yellowish, with 3 black spots, one being behind eye. Mesonotum brown, with median and peridiscal carinae, anterior portion of lateral angles, scutellum, and area before latter yellowish; lateral fields darker and 2 blackish points at base of scutellum. Tegulae yellowish brown.
</paragraph>
<paragraph id="8448E8ACEA4FA98CDCC38E7B3EF7821F" pageId="0" pageNumber="105">
<emphasis id="2BA26AA752C70DEB6BFA225B2668035E" bold="true" italics="true" pageId="0" pageNumber="105">Tegmina</emphasis>
(Figs
<figureCitation id="6177E52D626C6FE36F8993E32EB10298" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Connelicita bachmaensis sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D habitus, anterolateral view." figureDoi="10.3897/zookeys.1186.113701.figure7" httpUri="https://binary.pensoft.net/fig/951196" pageId="0" pageNumber="105">7A-D</figureCitation>
,
<figureCitation id="92FC1D9BD256C512BE11B6765EF126FC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Connelicita phongdienensis sp. nov., holotype ♂ A-C detail of head and thorax A dorsal view B lateral view C perpendicular view of frons D right tegmen E right hind wing F right metatibia and metatarsus, ventral view G left metatarsus, ventral view." figureDoi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105">8D</figureCitation>
): translucent, with brown spot near middle of vein CuP and vein CuA2, extending into clavus and irregular, greyish markings in cells in distal half of tegmen; costal and postclaval margins slightly diverging towards posterior; distal margin widely rounded.
</paragraph>
<caption id="090E1802794C4D170042B878C365E2F3" doi="10.3897/zookeys.1186.113701.figure7" httpUri="https://binary.pensoft.net/fig/951196" pageId="0" pageNumber="105" start="Figure 7" startId="F7">
<paragraph id="B0829B81DA32801F82F7E90546AC6B27" pageId="0" pageNumber="105">
<emphasis id="FDCBCEB547FD23A77C2F66FE02CCB8E1" bold="true" pageId="0" pageNumber="105">Figure 7.</emphasis>
<taxonomicName id="AFDB3449D56D81429A86B0DD44D2C8F1" authorityName="Constant &amp; Pham" authorityYear="2023" class="Insecta" family="Tropiduchidae" genus="Connelicita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Connelicita bachmaensis" order="Hemiptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="bachmaensis">
<emphasis id="23D67C84B32C60149DF9FAD214BBAC15" italics="true" pageId="0" pageNumber="105">Connelicita bachmaensis</emphasis>
</taxonomicName>
sp. nov., holotype ♂
<emphasis id="5390EE37FC642D2096D19D415C809CF6" bold="true" pageId="0" pageNumber="105">A</emphasis>
habitus, dorsal view
<emphasis id="D9D9A03775A1B7BA1288AA35DDB54495" bold="true" pageId="0" pageNumber="105">B</emphasis>
habitus, ventral view
<emphasis id="75A086057668FF7EBF46D1F35E87A627" bold="true" pageId="0" pageNumber="105">C</emphasis>
habitus, lateral view
<emphasis id="4F9EA352F4097CC5F21115FFD641C2E6" bold="true" pageId="0" pageNumber="105">D</emphasis>
habitus, anterolateral view.
</paragraph>
</caption>
<caption id="25AB2E05B03984C8C606943A6A39A417" doi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105" start="Figure 8" startId="F8">
<paragraph id="CBDDFFD700DBF35981A3B8D8E0D62990" pageId="0" pageNumber="105">
<emphasis id="648FD851744A488AFFE1F037474A19C9" bold="true" pageId="0" pageNumber="105">Figure 8.</emphasis>
<taxonomicName id="E7799DEAC966FCDD01E45C430E09D8BE" authorityName="Constant &amp; Pham" authorityYear="2023" class="Insecta" family="Tropiduchidae" genus="Connelicita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Connelicita phongdienensis" order="Hemiptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="phongdienensis">
<emphasis id="ECD10CAC4AD068C6B99E39DCFA0470DF" italics="true" pageId="0" pageNumber="105">Connelicita phongdienensis</emphasis>
</taxonomicName>
sp. nov., holotype ♂
<emphasis id="D2FA266E34B0EE076FA1002293E5F9F2" bold="true" pageId="0" pageNumber="105">A-C</emphasis>
detail of head and thorax
<emphasis id="EB4FC06A847C5E25F630A4FB9267A3C0" bold="true" pageId="0" pageNumber="105">A</emphasis>
dorsal view
<emphasis id="E7BA307B4FF49AA2EE40B8EE8A0F4B3F" bold="true" pageId="0" pageNumber="105">B</emphasis>
lateral view
<emphasis id="B7399B7B333FA14190AF878F8943DA64" bold="true" pageId="0" pageNumber="105">C</emphasis>
perpendicular view of frons
<emphasis id="402A31C1550110595C4033829365C999" bold="true" pageId="0" pageNumber="105">D</emphasis>
right tegmen
<emphasis id="D48987DC12DE9665CE055B72CB88FAC7" bold="true" pageId="0" pageNumber="105">E</emphasis>
right hind wing
<emphasis id="F0912EECD36586D56AA5433DA32429A7" bold="true" pageId="0" pageNumber="105">F</emphasis>
right metatibia and metatarsus, ventral view
<emphasis id="14111EE3F642CBAF5488A6D46A3BC119" bold="true" pageId="0" pageNumber="105">G</emphasis>
left metatarsus, ventral view.
</paragraph>
</caption>
<caption id="3BB14B5C314408F444789049A098326C" doi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105" start="Figure 9" startId="F9">
<paragraph id="7813BD4C4EEDFDF87014D87805CD5B82" pageId="0" pageNumber="105">
<emphasis id="64274E6FDAA65EAF5CCADF5CE17447F6" bold="true" pageId="0" pageNumber="105">Figure 9.</emphasis>
<taxonomicName id="FADFDC04C8DFBFB3CEA2BF39A63D8BD2" authorityName="Constant &amp; Pham" authorityYear="2023" class="Insecta" family="Tropiduchidae" genus="Connelicita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Connelicita phongdienensis" order="Hemiptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="phongdienensis">
<emphasis id="B593162DED926E66EA5D0CC191D522D3" italics="true" pageId="0" pageNumber="105">Connelicita phongdienensis</emphasis>
</taxonomicName>
sp. nov., holotype ♂, terminalia
<emphasis id="741546EC3C662C3148BAB2281456C8D3" bold="true" pageId="0" pageNumber="105">A-C</emphasis>
pygofer, gonostyli and anal tube
<emphasis id="31082CFF6CE4A20C1DD9F6A83AB03066" bold="true" pageId="0" pageNumber="105">A</emphasis>
lateral view
<emphasis id="D4561F63A1BC663596F6D6AC83D97C2E" bold="true" pageId="0" pageNumber="105">B</emphasis>
caudal view
<emphasis id="57CD8F01FCCF6AC95447BA075876DF26" bold="true" pageId="0" pageNumber="105">C</emphasis>
dorsal view
<emphasis id="1EE956DCC0F198CF86746D67B63345F0" bold="true" pageId="0" pageNumber="105">D-E</emphasis>
capitulum of left gonostylus
<emphasis id="0C383448661DD7581A8AA0483E0A7B17" bold="true" pageId="0" pageNumber="105">D</emphasis>
left laterodorsal view
<emphasis id="61A19BC5C600AEA079BC071FB2E24537" bold="true" pageId="0" pageNumber="105">E</emphasis>
right laterodorsal view
<emphasis id="86A4D0998291987E4F9E16D53AB4048A" bold="true" pageId="0" pageNumber="105">F-I</emphasis>
aedeagus, phallobase and connective
<emphasis id="58FA1571939019D5A6E70319F3D4C394" bold="true" pageId="0" pageNumber="105">F</emphasis>
left lateral view
<emphasis id="1D1E84ADDB24D2E3F1E1A52D5721A4F9" bold="true" pageId="0" pageNumber="105">G</emphasis>
anterodorsal view
<emphasis id="7DA69F0FBD00CE8A410B35286F2016B9" bold="true" pageId="0" pageNumber="105">H</emphasis>
posteroventral view dorsal view
<emphasis id="497EE7F5F6AB7B05FFA74BDA2355395B" bold="true" pageId="0" pageNumber="105">I</emphasis>
left lateroventral view.
</paragraph>
</caption>
<paragraph id="CA66F66962EFE0920B2F77C8B66877BF" pageId="0" pageNumber="105">
Venation (Fig.
<figureCitation id="9E4A3A93E578DD12DFDFCBED1CC862A2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Connelicita phongdienensis sp. nov., holotype ♂ A-C detail of head and thorax A dorsal view B lateral view C perpendicular view of frons D right tegmen E right hind wing F right metatibia and metatarsus, ventral view G left metatarsus, ventral view." figureDoi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105">8D</figureCitation>
): costal area well developed, with numerous cross-veins delimitating elongate cells, pterostigma absent; veins ScP+R, MP, and CuA separated at base, first fork of vein ScP+R near base; first fork of vein MP after first fork of vein CuA; oblique, transverse cross-vein between MP and CuA1 in basal portion of latter; clavus long, reaching near 4/5 of tegmina length; veins Pcu and A1 fused at basal half of clavus.
</paragraph>
<paragraph id="F7455769773165A4E1E3333900FB8112" pageId="0" pageNumber="105">
<emphasis id="077B58A52EE711AC7FCDFFF821BFED57" bold="true" italics="true" pageId="0" pageNumber="105">Hind wings</emphasis>
(Fig.
<figureCitation id="F40647C985E8608692D7AF94A6D6A166" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Connelicita phongdienensis sp. nov., holotype ♂ A-C detail of head and thorax A dorsal view B lateral view C perpendicular view of frons D right tegmen E right hind wing F right metatibia and metatarsus, ventral view G left metatarsus, ventral view." figureDoi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105">8E</figureCitation>
): translucent, with veins blackish; CuP-Pcu-A1 lobe infuscate in distal portion, with apical angle largely blackish; A2 lobe weakly infuscate and roundly protruding along postclaval margin.
</paragraph>
<paragraph id="445743059BFC9D06463CE01F492C2717" pageId="0" pageNumber="105">Venation: main veins present; ScP+R, MP, and CuA running more or less parallel, slightly diverging towards posterior; Pcu curved around distal third of wing towards CuP but not reaching the latter; A2 complete, reaching margin.</paragraph>
<paragraph id="454577ECE972D1EFA1688662A3BFCAFC" pageId="0" pageNumber="105">
<emphasis id="8A595A59D640894B22C001168696FDA7" bold="true" italics="true" pageId="0" pageNumber="105">Legs</emphasis>
(Figs
<figureCitation id="F7C8D529FE658F98CABDCB3596070A99" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Connelicita bachmaensis sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D habitus, anterolateral view." figureDoi="10.3897/zookeys.1186.113701.figure7" httpUri="https://binary.pensoft.net/fig/951196" pageId="0" pageNumber="105">7A, B</figureCitation>
,
<figureCitation id="0468E20F6CC6FA13A6B41709EA19524C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Connelicita phongdienensis sp. nov., holotype ♂ A-C detail of head and thorax A dorsal view B lateral view C perpendicular view of frons D right tegmen E right hind wing F right metatibia and metatarsus, ventral view G left metatarsus, ventral view." figureDoi="10.3897/zookeys.1186.113701.figure8" httpUri="https://binary.pensoft.net/fig/951197" pageId="0" pageNumber="105">8F-G</figureCitation>
): pale yellowish brown, with dark brown marking along dorsal portion of metafemora and basiventral portion of metatibiae. Metatibiotarsal formula: (2) 9 / 2 / 2.
</paragraph>
<paragraph id="A77FF7651AA3EF6C6E05CBB5576ADE66" pageId="0" pageNumber="105">
<emphasis id="B117D859B65506AAF46430C1EDE1E1E4" bold="true" italics="true" pageId="0" pageNumber="105">Abdomen</emphasis>
(Fig.
<figureCitation id="D7A900AA6496DCD002D228118A10FC8F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Connelicita bachmaensis sp. nov., holotype ♂ A habitus, dorsal view B habitus, ventral view C habitus, lateral view D habitus, anterolateral view." figureDoi="10.3897/zookeys.1186.113701.figure7" httpUri="https://binary.pensoft.net/fig/951196" pageId="0" pageNumber="105">7B</figureCitation>
): pale yellowish brown, with narrow brown line interrupted in middle, along posterior margin of sternites.
</paragraph>
<paragraph id="4F71371E5FFD4654928D4BB258379EB6" pageId="0" pageNumber="105">
<emphasis id="965182BF3A327DF08077272CAF057F5F" bold="true" italics="true" pageId="0" pageNumber="105">Terminalia</emphasis>
♂ (Fig.
<figureCitation id="2C6522017BB6A5EF9EBC68437BEF6915" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9</figureCitation>
): pygofer (
<emphasis id="CB4CA443E20EC927F79371AED0C5B8A4" italics="true" pageId="0" pageNumber="105">Py</emphasis>
- Fig.
<figureCitation id="7E9521320FBDC0ECB66FA12BF9F44EFC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9A, B</figureCitation>
) ~1.9
<normalizedToken id="CA1D742E96C339F2F963610DEDC932EA" originalValue="×">x</normalizedToken>
as high as wide in caudal view, in lateral view with posterior margin deeply inclined posteriorly at upper 1/3 then broadly rounded, and sinuate at lower 2/3. Gonostyli (
<emphasis id="E5D13668217D4E7BCC4887547EF28B9A" italics="true" pageId="0" pageNumber="105">G</emphasis>
- Fig.
<figureCitation id="48E8A152330617EB2340DA7E369412F8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9A-C</figureCitation>
) rather elongate in lateral view, with capitulum at about 2/3 of gonostylus length, and with large laminate process projecting medially behind capitulum; posterior lobe rounded; capitulum (
<emphasis id="8BFD524DBCE3670E9272AD65E35B53E1" italics="true" pageId="0" pageNumber="105">ca</emphasis>
- Fig.
<figureCitation id="1B9AD4DAA583B55F43559B45F2396BB0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9A-D</figureCitation>
) laterally flattened, with posterior margin rounded in lateral view and bearing a complex lateral process, with upper digitiform process curved cephalodorsad and sinuate ventral spine directed posteroventrad. Aedeagus (Fig.
<figureCitation id="F30452C16BDBDC3AAE227A617EE450A3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9E-I</figureCitation>
) strongly upcurved at mid-length, with pair of slender lateral pointed processes (
<emphasis id="7FC7A027755AC298C7FB922857FC0FDE" italics="true" pageId="0" pageNumber="105">lpa</emphasis>
) not reaching apex of aedeagus, and with apical point directed anterodorsad; 3 small terminal processes, middle one shortly projecting posterad; dorsal process of periandrium (
<emphasis id="AED6AEFB54016C16DAD70660FC885AC3" italics="true" pageId="0" pageNumber="105">dpp</emphasis>
) tooth-shaped, strong and projecting dorsocephalad, placed slightly after mid-length of aedeagus; paired ventral processes of periandrium (
<emphasis id="E0EB8038CCDC5F15C45C251E73A295C1" italics="true" pageId="0" pageNumber="105">vpp</emphasis>
) not reaching
<normalizedToken id="5F7B56F567DDDC3E363530566B4D862E" originalValue="½">1/2</normalizedToken>
length of aedeagus, elongate, sinuate in distal portion, and with pointed apex directed lateroposterad; connective (
<emphasis id="DCE029DC823E7216062989626E4C18C0" italics="true" pageId="0" pageNumber="105">co</emphasis>
) moderately curved. Anal tube (
<emphasis id="183AD0A7686932BE275490A35AC857BA" italics="true" pageId="0" pageNumber="105">An</emphasis>
- Fig.
<figureCitation id="D72F84E796904317640175A8EFEB3844" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Connelicita phongdienensis sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral view B caudal view C dorsal view D-E capitulum of left gonostylus D left laterodorsal view E right laterodorsal view F-I aedeagus, phallobase and connective F left lateral view G anterodorsal view H posteroventral view dorsal view I left lateroventral view." figureDoi="10.3897/zookeys.1186.113701.figure9" httpUri="https://binary.pensoft.net/fig/951198" pageId="0" pageNumber="105">9A-C</figureCitation>
) ~1.26
<normalizedToken id="371700327C81CB165C6ECDA9DCF2345D" originalValue="×">x</normalizedToken>
as long in median line, as wide, in dorsal view; subcylindrical in proximal half, then with dorsal margin excavate and sinuate in lateral view; in dorsal view, lateral margins subparallel in proximal half, then regularly converging in distal half; posterior margin deeply concave in dorsal view, with basal ventral bulge; epiproct rather large, located at
<normalizedToken id="B10DBE3E9CEC8295BC5CAAB5E5393A02" originalValue="½">1/2</normalizedToken>
length of anal tube.
</paragraph>
<paragraph id="B6C4C9F099553BFD8D5A7F703E48EF02" pageId="0" pageNumber="105">
<emphasis id="2CE33009273EDDCD4969170F842E5E4F" bold="true" pageId="0" pageNumber="105">Female.</emphasis>
Similar to male.
</paragraph>
</subSubSection>
<subSubSection id="6B78D0EBFAB29AC0977543BBD8CC1590" pageId="0" pageNumber="105" type="etymology">
<paragraph id="DCF8AF19A084D0929AF500734200BF09" pageId="0" pageNumber="105">Etymology.</paragraph>
<paragraph id="21EDF0A13D1A5956344A1AF8EF9B0018" pageId="0" pageNumber="105">
The species epithet
<emphasis id="1059AE2AA5B4759BDBB4D00D262E8E39" italics="true" pageId="0" pageNumber="105">phongdienensis</emphasis>
refers to Phong Dien District, the locality where the new species was discovered, at the Centre for Conservation of Vietnam Natural Resources and Rescue of Animals and plants.
</paragraph>
</subSubSection>
<subSubSection id="B70F974DAE66743E8843251EDA968136" pageId="0" pageNumber="105" type="biology_ecology">
<paragraph id="D059FD9538C7E69429BF13DD1EE6B95F" pageId="0" pageNumber="105">Biology.</paragraph>
<paragraph id="6091EFB5625EA1C961BDF86E486B86B9" pageId="0" pageNumber="105">
The specimens were found sitting on leaves on the lower vegetation in a subtropical evergreen forest (Fig.
<figureCitation id="766EEECC2D654898B55A571925DB9C72" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitat of Connelicita phongdienensis sp. nov., Vietnam, Phong Dien, CCRR, 23 May 2023." figureDoi="10.3897/zookeys.1186.113701.figure10" httpUri="https://binary.pensoft.net/fig/951199" pageId="0" pageNumber="105">10</figureCitation>
) in the Northern Vietnam lowland rain forests ecoregion at rather low altitude (150-600 m).
</paragraph>
<caption id="D24CCAF1AC6351977FA6A1977E2EA242" doi="10.3897/zookeys.1186.113701.figure10" httpUri="https://binary.pensoft.net/fig/951199" pageId="0" pageNumber="105" start="Figure 10" startId="F10">
<paragraph id="243E6FBB21579AB3665AFAC6B3F9C214" pageId="0" pageNumber="105">
<emphasis id="FC1A753381CAE48E6659E96F3FF08CF2" bold="true" pageId="0" pageNumber="105">Figure 10.</emphasis>
Habitat of
<taxonomicName id="84520A07D1DC5F1D830A3872ED985AEB" authorityName="Constant &amp; Pham" authorityYear="2023" class="Insecta" family="Tropiduchidae" genus="Connelicita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Connelicita phongdienensis" order="Hemiptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="phongdienensis">
<emphasis id="434C5C32CCAC281A6DE8F2436AFA017C" italics="true" pageId="0" pageNumber="105">Connelicita phongdienensis</emphasis>
</taxonomicName>
sp. nov., Vietnam, Phong Dien, CCRR, 23 May 2023.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="18340F6E87220A1A2BDFB9158ACBA0E8" pageId="0" pageNumber="105" type="distribution">
<paragraph id="41FA1940A0BF1499814A0215BB6D9E90" pageId="0" pageNumber="105">Distribution.</paragraph>
<paragraph id="C06F59E0A66DB9054C787CCBAD0D1400" pageId="0" pageNumber="105">
Vietnam, Thua Tinh-Hue Province, Phong Dien District, CCRR (Fig.
<figureCitation id="85D8C2858B6A468363E40DC90550488E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of the species of Connelicita of Vietnam." figureDoi="10.3897/zookeys.1186.113701.figure5" httpUri="https://binary.pensoft.net/fig/951194" pageId="0" pageNumber="105">5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>