370 lines
40 KiB
XML
370 lines
40 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1135.93843" ID-Pensoft-Pub="1313-2970-1135-61" ID-Pensoft-UUID="FC0EE843319C574DB0BA3E1125784BD4" ID-ZooBank="7E562C71BD10446E9E102AA52A9888FB" ModsDocID="1313-2970-1135-61" checkinTime="1670952533187" checkinUser="pensoft" docAuthor="Qin, Yuyao, Germann, Christoph & Liang, Hongbin" docDate="2022" docId="B5DB751649F25FBFB9B9635FA0BEBE81" docLanguage="en" docName="ZooKeys 1135: 61-91" docOrigin="ZooKeys 1135" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zookeys.1135.93843" docTitle="Chlaenius (Sphodromimus) davidi Qin, Germann & Liang, 2022, nom. nov." docType="treatment" docVersion="1" id="FC0EE843319C574DB0BA3E1125784BD4" lastPageNumber="61" masterDocId="FC0EE843319C574DB0BA3E1125784BD4" masterDocTitle="Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species" masterLastPageNumber="91" masterPageNumber="61" pageNumber="61" updateTime="1670952533187" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Qin, Yuyao</mods:namePart>
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<mods:affiliation>College of Life Sciences, Hebei University, Baoding 071002, China & Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Germann, Christoph</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8317-3799</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, CH- 4051 Basel, Switzerland</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2022</mods:date>
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||
<mods:detail type="pubDate">
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<mods:number>2022-12-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1135</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>61</mods:start>
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<mods:end>91</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1135.93843</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1135.93843</mods:identifier>
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||
<mods:identifier type="Pensoft-Pub">1313-2970-1135-61</mods:identifier>
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<mods:identifier type="ZooBank">7E562C71BD10446E9E102AA52A9888FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FC0EE843319C574DB0BA3E1125784BD4</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81" httpUri="http://treatment.plazi.org/id/B5DB751649F25FBFB9B9635FA0BEBE81" lastPageNumber="61" pageId="0" pageNumber="61">
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||
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
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||
<paragraph pageId="0" pageNumber="61">
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<taxonomicName LSID="B5DB7516-49F2-5FBF-B9B9-635FA0BEBE81" authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) davidi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="davidi" status=" nom. nov." subGenus="Sphodromimus">Chlaenius (Sphodromimus) davidi</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="61">nom. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="61" type="description">
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<paragraph pageId="0" pageNumber="61">
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<figureCitation captionStart="Figures 9, 10" captionStartId="F6" captionText="Figures 9, 10. 9 A, B Chlaenius (Sphodromimus) davidi nom. nov. (male, Guangdong, Xinyi) 10 A, B C. (Sphodromimus) enleensis (male, Vietnam, Tam Dao). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures9-10" httpUri="https://binary.pensoft.net/fig/780980" pageId="0" pageNumber="61">Figs 9A, B</figureCitation>
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<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">, 15</figureCitation>
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<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">, 23</figureCitation>
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<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">, 31A-E</figureCitation>
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<figureCitation captionStart="Figures 37–39" captionStartId="F13" captionText="Figures 37 - 39. Internal reproductive system of females 37 Chlaenius (Sphodromimus) yinggelingensis sp. nov., paratype 38 C. (Sphodromimus) flavofemoratus Laporte, 1834 39 C. (Sphodromimus) davidi nom. nov. A bursa copulatrix B dorsal view C ventral view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures37-39" httpUri="https://binary.pensoft.net/fig/780987" pageId="0" pageNumber="61">, 39A-C</figureCitation>
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<figureCitation captionStart="Figure 40" captionStartId="F14" captionText="Figure 40. Distribution of Chlaenius (Sphodromimus) species in China and adjacent areas (a possible distribution of C. enleensis, En-le, Yunnan, represented by a question mark)." figureDoi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61">, 40</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="61" type="reference_group">
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<paragraph pageId="0" pageNumber="61">
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<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius (Sphodromimus) wrasei" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="wrasei" status=" nom. nov." subGenus="Sphodromimus">Vachinius (Sphodromimus) wrasei</taxonomicName>
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Kirschenhofer, 2003: 37 (type locality: China, Guangdong);
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
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: 342 (catalogue);
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<bibRefCitation author="Brunk, I" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="61" pagination="51 - 55" refId="B6" refString="Brunk, I, Kirschenhofer, E, 2016. Vachinius (Vachinius) arakanensis sp.n. (Coleoptera: Carabidae: Chlaeniini) from Myanmar. Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 68: 51 - 55" title="Vachinius (Vachinius) arakanensis sp. n. (Coleoptera: Carabidae: Chlaeniini) from Myanmar." volume="68" year="2016">Brunk and Kirschenhofer 2016</bibRefCitation>
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: 54 (record);
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<bibRefCitation DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
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: 1 (genus
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<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status=" nom. nov.">Chlaenius</taxonomicName>
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, subgenus
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<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus)" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" status=" nom. nov." subGenus="Sphodromimus">Chlaenius Sphodromimus</taxonomicName>
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) [nec
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<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="C. (Lithochlaenius) wrasei" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="wrasei" status=" nom. nov." subGenus="Lithochlaenius">C. (Lithochlaenius) wrasei</taxonomicName>
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(Kirschenhofer, 1997)]
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="61" type="type locality">
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<paragraph pageId="0" pageNumber="61">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="61">Guangdong, Xinyi, Datianding.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="61" type="materials_examined">
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<paragraph pageId="0" pageNumber="61">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="61">
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Total
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="China" location="Guangdong" specimenCount="10" stateProvince="Guangdong">
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<specimenCount type="generic">10 specimens</specimenCount>
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.
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<collectingCountry name="China">China</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="61">
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<collectingRegion country="China" name="Guangdong">Guangdong</collectingRegion>
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</emphasis>
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||
</materialsCitation>
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||
:
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<materialsCitation collectingDate="1997-08" collectorName="Li" country="China" county="Guangdong" determinerName="Kirschenhofer" elevation="1500" location="Datianding" municipality="Xinyi" specimenCount="1" typeStatus="Holotype">
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<typeStatus>Holotype</typeStatus>
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female (DWC, photo),
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<collectingCountry name="China">China</collectingCountry>
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,
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<collectingCounty>Guangdong</collectingCounty>
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||
,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
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||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</elevation>
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</quantity>
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||
,
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<collectingMunicipality>Xinyi</collectingMunicipality>
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||
:
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<location LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81:4087E3CD2E6591EC235FD70DAA99A447" country="China" county="Guangdong" municipality="Xinyi" name="Datianding">Datianding</location>
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(22.16/111.15) -
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<collectingDate value="1997-08">VIII-1997</collectingDate>
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- leg.
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<collectorName>Li</collectorName>
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/
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<typeStatus>Holotypus</typeStatus>
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,
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<taxonomicName baseAuthorityName="Kirschenhofer" baseAuthorityYear="1997" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) wrasei" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="wrasei" subGenus="Sphodromimus">Chlaenius (Sphodromimus) wrasei</taxonomicName>
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sp. nov. det.
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<determinerName>Kirschenhofer</determinerName>
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, 2001[red label]/COLL WRASE, BERLIN
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</materialsCitation>
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;
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<materialsCitation collectingDate="2017-05-30" collectorName="Y. Z. Liu, S. P. Yu" country="China" county="Guangdong" elevation="1508" latitude="22.291317" location="Yunkai Shan" longLatPrecision="1" longitude="111.209885" municipality="Xinyi" specimenCount="5" specimenCount-female="4" specimenCount-male="1" typeStatus="holotype">
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<specimenCount type="male">1 ♂</specimenCount>
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and
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<specimenCount type="female">4 ♀♀</specimenCount>
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||
(IZAS),
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<collectingCounty>Guangdong</collectingCounty>
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||
,
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<collectingMunicipality>Xinyi</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81:112247DDFB292C7837FC471C9408CF48" country="China" county="Guangdong" latitude="22.291317" longLatPrecision="1" longitude="111.209885" municipality="Xinyi" name="Yunkai Shan">Yunkai Shan</location>
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||
,
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||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.50821" unit="m" value="1508.21">
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||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.50821" unit="m" value="1508.21">1508.21m</elevation>
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||
</quantity>
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||
/
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||
<geoCoordinate degrees="22.291317" direction="north" orientation="latitude" precision="1" value="22.291317">22.291317°N</geoCoordinate>
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||
,
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||
<geoCoordinate degrees="111.209888" direction="east" orientation="longitude" precision="1" value="111.209885">111.209888°E</geoCoordinate>
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||
,
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||
<collectingDate value="2017-05-30">2017.V.30</collectingDate>
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||
,
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||
<collectorName>Y.Z. Liu</collectorName>
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||
,
|
||
<collectorName>S.P. Yu</collectorName>
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||
leg.,
|
||
<location LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81:1B2DF38BE754A982D547C20ACD7B5683" country="China" county="Guangdong" latitude="22.291317" longLatPrecision="1" longitude="111.209885" municipality="Xinyi" name="Inst. of Zoology">Inst. of Zoology</location>
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||
</materialsCitation>
|
||
;
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||
<materialsCitation collectingDate="2017-05-31" collectorName="Y. Z. Liu, S. P. Yu" country="China" county="Guangdong" elevation="1251" latitude="22.292692" location="Yunkai Shan" longLatPrecision="1" longitude="111.203835" municipality="Xinyi" specimenCount="2" specimenCount-female="2" stateProvince="Guangxi" typeStatus="holotype">
|
||
<specimenCount type="female">2 ♀♀</specimenCount>
|
||
(IZAS),
|
||
<collectingCounty>Guangdong</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Xinyi</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81:49F5BEF872768198B9AB4599DCFCA747" country="China" county="Guangdong" latitude="22.292692" longLatPrecision="1" longitude="111.203835" municipality="Xinyi" name="Yunkai Shan" stateProvince="Guangxi">Yunkai Shan</location>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.25055" unit="m" value="1250.55">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.25055" unit="m" value="1250.55">1250.55 m</elevation>
|
||
</quantity>
|
||
/
|
||
<geoCoordinate degrees="22.292692" direction="north" orientation="latitude" precision="1" value="22.292692">22.292692°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="111.203833" direction="east" orientation="longitude" precision="1" value="111.203835">111.203833°E</geoCoordinate>
|
||
,
|
||
<collectingDate value="2017-05-31">2017.V.31</collectingDate>
|
||
,
|
||
<collectorName>Y.Z. Liu</collectorName>
|
||
,
|
||
<collectorName>S.P. Yu</collectorName>
|
||
leg.,
|
||
<location LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81:7B218E2ECA9A627C8DA2393A1AADD42E" country="China" county="Guangdong" latitude="22.292692" longLatPrecision="1" longitude="111.203835" municipality="Xinyi" name="Inst. of Zoology" stateProvince="Guangxi">Inst. of Zoology</location>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="61">
|
||
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation collectingDate="2011-05-27" collectorName="Xinlei Huang Coll." country="China" county="Guangxi Prov" elevation="1230" location="Tianping Protect Station" municipality="Daming Shan" specimenCount="2" specimenCount-male="2" stateProvince="Guangxi" typeStatus="holotype">
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||
<specimenCount type="male">2 ♂♂</specimenCount>
|
||
(IZAS),
|
||
<collectingCounty>Guangxi Prov</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Daming Shan</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81:257CD2E4B251858B31C753EDDE139563" country="China" county="Guangxi Prov" municipality="Daming Shan" name="Tianping Protect Station" stateProvince="Guangxi">Tianping Protect Station</location>
|
||
, N 23.49811, E 108.43715 /
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.23" unit="m" value="1230.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.23" unit="m" value="1230.0">1230 m</elevation>
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||
</quantity>
|
||
,
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||
<collectingDate value="2011-05-27">2011.V.27</collectingDate>
|
||
N,
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||
<collectorName>Xinlei Huang Coll.</collectorName>
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||
,
|
||
<location LSID="urn:lsid:plazi:treatment:B5DB751649F25FBFB9B9635FA0BEBE81:FCB1978D7E994302A2321A987503D21D" country="China" county="Guangxi Prov" municipality="Daming Shan" name="Inst. of Zoology" stateProvince="Guangxi">Inst. of Zoology</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1135.93843.figures9-10" httpUri="https://binary.pensoft.net/fig/780980" pageId="0" pageNumber="61" start="Figures 9, 10" startId="F6">
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" pageId="0" pageNumber="61">Figures 9, 10.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="61">9A, B</emphasis>
|
||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) davidi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="davidi" subGenus="Sphodromimus">Chlaenius (Sphodromimus) davidi</taxonomicName>
|
||
nom. nov. (male, Guangdong, Xinyi)
|
||
<emphasis bold="true" pageId="0" pageNumber="61">10A, B</emphasis>
|
||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis" subGenus="Sphodromimus">C. (Sphodromimus) enleensis</taxonomicName>
|
||
(male,
|
||
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
|
||
, Tam Dao). Scale bars: 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
Pronotum fully metallic purple, or greenish purple. PW/PL = 1.09-1.11; PAW/PBW = 0.83-0.93 (Fig.
|
||
<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">23</figureCitation>
|
||
); pronotum cordate with posterior angles nearly right angled, rounded at tips; disc sparsely punctate and pubescent, with dense, shallow, transverse rugosities, without glabrous area in the middle. Elytral intervals convex basally, flat apically, densely punctate and pubescent. Hind wings reduced. Metepisterna short, MW/ML = 1.11-1.15 (Fig.
|
||
<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">15</figureCitation>
|
||
). Legs totally black.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="61" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="61">Comparisons.</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
This species is most similar to
|
||
<taxonomicName authorityName="Mandl" authorityYear="1992" baseAuthorityName="Qin & Germann & Liang" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="enleensis" subGenus="Sphodromimus">Chlaenius (Sphodromimus) enleensis</taxonomicName>
|
||
Mandl, 1992 (Figs
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A, B Chlaenius (Sphodromimus) flavofemoratus Laporte, 1834 (male, Yunnan, Menglun) C C. (Sphodromimus) flavofemoratus Laporte, 1834 (female, Yunnan, Menglun) 8 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, Indo Chine) Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures7-8" httpUri="https://binary.pensoft.net/fig/780979" pageId="0" pageNumber="61">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 9, 10" captionStartId="F6" captionText="Figures 9, 10. 9 A, B Chlaenius (Sphodromimus) davidi nom. nov. (male, Guangdong, Xinyi) 10 A, B C. (Sphodromimus) enleensis (male, Vietnam, Tam Dao). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures9-10" httpUri="https://binary.pensoft.net/fig/780980" pageId="0" pageNumber="61">10A, B</figureCitation>
|
||
), sharing the large size, general shape of apical lamella, metepisterna short (Figs
|
||
<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">15</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">16</figureCitation>
|
||
), and reduced hind wings, but can be distinguished from the latter by: (1) pronotum purple or greenish purple (Fig.
|
||
<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">23</figureCitation>
|
||
) (green to coppery, but not purple in
|
||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">24</figureCitation>
|
||
); (2) femora black (middle of mesofemora and metafemora brown in
|
||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||
</taxonomicName>
|
||
); (3) elytral intervals convex basally, flat apically (interval convex throughout in
|
||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||
</taxonomicName>
|
||
); (4) apical lamella wider at base (narrower in
|
||
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">C. enleensis</emphasis>
|
||
</taxonomicName>
|
||
, Figs
|
||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">32A, C, D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">33A, C, D</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="61" type="description">
|
||
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">BL = 21.3-24.4 mm, BW = 8.0-8.7 mm. PL = 4.5-5.0 mm, PW = 5.0-5.5 mm, MW = 2.0-2.1 mm, ML = 1.7-1.9 mm. Head, elytra, venter, and legs dark and black; pronotum fully purple or greenish purple; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown.</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" pageId="0" pageNumber="61">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Vertex finely punctate, pubescent, without a distinct glabrous area; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than antennomere 4.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Pronotum</emphasis>
|
||
cordiform, PW/PL = 1.09-1.11 (Fig.
|
||
<figureCitation captionStart="Figures 17–24" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">23</figureCitation>
|
||
), widest at apical third; anterior margin slightly concave, PAW/PBW = 0.83-0.93; lateral margins distinctly narrowed from middle to base, distinctly sinuate before posterior angles; anterior angles rounded, slightly projected forward; posterior angles nearly right angled, rounded at tips; disc gently convex, sparsely punctate and pubescent, with dense shallow transverse rugosities, without glabrous area in the middle; median line distinct, fine, not reaching anterior margin and base; basal foveae deeply arcuate, punctate and pubescent.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Elytra</emphasis>
|
||
elongate, EL/BW = 1.61-1.75, gently convex near anterior third, widest near posterior third, rounded at apex in males, subtruncate in females; striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals convex at base, flat from middle to apex, densely punctate and pubescent; sutural angles obtuse and right; hind wings reduced.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Venter</emphasis>
|
||
densely punctate, pubescent, metepisterna (Fig.
|
||
<figureCitation captionStart="Figures 11–16" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">15</figureCitation>
|
||
) short, MW/ML = 1.11-1.15; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males, two pairs in females; all abdominal sternites with distinct impressions laterally.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Legs</emphasis>
|
||
long and slender; tarsi nearly smooth dorsally.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" pageId="0" pageNumber="61">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Median lobe (Fig.
|
||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">31B-E</figureCitation>
|
||
) long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella triangular (Fig.
|
||
<figureCitation captionStart="Figures 31–33" captionStartId="F11" captionText="Figures 31 - 33. Aedeagus of Sphodromimus spp. 31 Chlaenius (Sphodromimus) davidi nom. nov. (Guangdong, Xinyi) 32 C. (Sphodromimus) enleensis Mandl, 1992 (Vietnam, Tam Dao) 33 C. (Sphodromimus) enleensis Mandl, 1992 (holotype, " Indo Chine ") A apical lamella B left lateral view C dorsal view D ventral view E right lateral view F endophallus. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-F)." figureDoi="10.3897/zookeys.1135.93843.figures31-33" httpUri="https://binary.pensoft.net/fig/780985" pageId="0" pageNumber="61">31A</figureCitation>
|
||
), distinctly bent to right side, longer than basal width, each side of middle apical lamella with a denticulation or absent on right side; in left lateral view, apical portion slightly bent dorsally at apex, basal orifice ~ 90° relative to preapical shaft; left paramere larger than right paramere, both helically curved; endophallus with flagellum thick and straight; basal part of flagellum with irregular bursa; apical part of flagellum with drop-shaped sclerite.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<emphasis bold="true" pageId="0" pageNumber="61">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">Female genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Bursa copulatrix (Fig.
|
||
<figureCitation captionStart="Figures 37–39" captionStartId="F13" captionText="Figures 37 - 39. Internal reproductive system of females 37 Chlaenius (Sphodromimus) yinggelingensis sp. nov., paratype 38 C. (Sphodromimus) flavofemoratus Laporte, 1834 39 C. (Sphodromimus) davidi nom. nov. A bursa copulatrix B dorsal view C ventral view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures37-39" httpUri="https://binary.pensoft.net/fig/780987" pageId="0" pageNumber="61">39A-C</figureCitation>
|
||
) round, base with a distinct long digitiform protrusion; villous canal long, tortuously contorted, adhered to common oviduct; spermatheca and spermathecal gland absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="61" type="distribution">
|
||
<paragraph pageId="0" pageNumber="61">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 40" captionStartId="F14" captionText="Figure 40. Distribution of Chlaenius (Sphodromimus) species in China and adjacent areas (a possible distribution of C. enleensis, En-le, Yunnan, represented by a question mark)." figureDoi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61">40</figureCitation>
|
||
) China (Guangdong; Guangxi).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="61" type="remarks">
|
||
<paragraph pageId="0" pageNumber="61">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
<taxonomicName baseAuthorityName="Kirschenhofer" baseAuthorityYear="1997" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) wrasei" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="wrasei" subGenus="Sphodromimus">Chlaenius (Sphodromimus) wrasei</taxonomicName>
|
||
was originally described as a member of the genus
|
||
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">Vachinius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer (2019)</bibRefCitation>
|
||
moved it to the genus
|
||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="61">Chlaenius</emphasis>
|
||
</taxonomicName>
|
||
. However, in a result of this treatment, it became a junior homonym of
|
||
<taxonomicName baseAuthorityName="Kirschenhofer" baseAuthorityYear="1997" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Lithochlaenius) wrasei" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="wrasei" subGenus="Lithochlaenius">Chlaenius (Lithochlaenius) wrasei</taxonomicName>
|
||
(Kirschenhofer, 1997). Herein, we propose a new replacement name for the former -
|
||
<taxonomicName authorityName="Qin & Germann & Liang" authorityYear="2022" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) davidi" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="davidi" subGenus="Sphodromimus">Chlaenius (Sphodromimus) davidi</taxonomicName>
|
||
nom. nov., based on the first name of the well-known specialist on the ground beetles, David W. Wrase.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="61">
|
||
This species was described based on a single female from Datianding, Xinyi, Guangdong, China. Recently, one male and six females were collected from the type locality, and those specimens fit well with the original description and illustration of
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<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. davidi" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="davidi">
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||
<emphasis italics="true" pageId="0" pageNumber="61">C. davidi</emphasis>
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||
</taxonomicName>
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||
nom. nov. Two more males were collected in Guangxi, which are identical to the holotype.
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||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |