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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.4.1408" ID-GBIF-Dataset="f739a3a3-6210-4df3-9e04-a945b6cf7dba" ID-PMC="PMC3174441" ID-Pensoft-Pub="1314-2003-4-149" ID-Pensoft-UUID="CB046E48FFD36843CC56FFC2FFC9FFA0" ID-PubMed="22171187" ID-Zenodo-Dep="576070" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1314-2003-4-149" ModsDocOrigin="PhytoKeys 4" ModsDocTitle="A new species of Meryta (Araliaceae) from the Marquesas Archipelago, French Polynesia" checkinTime="1451252108721" checkinUser="pensoft" docAuthor="Tronchet, Frederic &amp; Lowry II, Porter P." docDate="2011" docId="882013059F97D55CB6C8F1134ED20CA2" docLanguage="en" docName="PhytoKeys 4: 149-156" docOrigin="PhytoKeys 4" docSource="http://dx.doi.org/10.3897/phytokeys.4.1408" docTitle="Meryta pastoralis F. Tronchet &amp; Lowry 2011, sp. nov." docType="treatment" docVersion="5" id="CB046E48FFD36843CC56FFC2FFC9FFA0" lastPageNumber="153" masterDocId="CB046E48FFD36843CC56FFC2FFC9FFA0" masterDocTitle="A new species of Meryta (Araliaceae) from the Marquesas Archipelago, French Polynesia" masterLastPageNumber="156" masterPageNumber="149" pageNumber="150" updateTime="1668141225291" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new species of Meryta (Araliaceae) from the Marquesas Archipelago, French Polynesia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tronchet, Frederic</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lowry II, Porter P.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>149</mods:start>
<mods:end>156</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.4.1408</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.4.1408</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-4-149</mods:identifier>
<mods:identifier type="Pensoft-UUID">CB046E48FFD36843CC56FFC2FFC9FFA0</mods:identifier>
<mods:identifier type="Zenodo-Dep">576070</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152025431" LSID="urn:lsid:plazi:treatment:882013059F97D55CB6C8F1134ED20CA2" httpUri="http://treatment.plazi.org/id/882013059F97D55CB6C8F1134ED20CA2" lastPageId="4" lastPageNumber="153" pageId="1" pageNumber="150">
<subSubSection pageId="1" pageNumber="150" type="nomenclature">
<paragraph pageId="1" pageNumber="150">
<taxonomicName LSID="88201305-9F97-D55C-B6C8-F1134ED20CA2" authority="F. Tronchet &amp; Lowry" authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Meryta pastoralis" order="Apiales" pageId="1" pageNumber="150" phylum="Tracheophyta" rank="species" species="pastoralis" status="sp. nov.">Meryta pastoralis F. Tronchet &amp; Lowry</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="150">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Meryta pastoralis F. Tronchet &amp; Lowry. A branch with leaves and young infructescence B female flowers in bud subtended by bracts C female flowers at receptivity D young fruits. Line drawing by Alain Jouy from herbarium specimens; voucher Perlman 19767 (A, D), Schaefer 5922 (B), Perlman 10206 (C)." pageId="1" pageNumber="150">Figs 1</figureCitation>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Meryta pastoralis F. Tronchet &amp; Lowry. A branch with leaves and young infructescence B female flowers in bud subtended by bracts C female flowers at receptivity D young fruits. Line drawing by Alain Jouy from herbarium specimens; voucher Perlman 19767 (A, D), Schaefer 5922 (B), Perlman 10206 (C)." pageId="1" pageNumber="150">-2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="150" type="latin">
<paragraph pageId="1" pageNumber="150">Latin.</paragraph>
<paragraph pageId="1" pageNumber="150">
<emphasis italics="true" pageId="1" pageNumber="150">Haec species quoad inflorescentiae structuram ac ovaria omnino libera Merytae choristanthae Harms (ex Insulis Australibus) simillima, sed ab ea foliis anguste obovatis usque spathulatis (vs. late ellipticis usque subobovatis), carpellis 5 ad 11 (vs. 5 vel 6 tantum) atque fructu maturitate 7-9</emphasis>
<normalizedToken originalValue="×">x</normalizedToken>
<emphasis italics="true" pageId="1" pageNumber="150">7-10 (vs. 8-12</emphasis>
<normalizedToken originalValue="×">x</normalizedToken>
<emphasis italics="true" pageId="1" pageNumber="150">10-15) mm distinguitur; etiam quoad foliorum aspectum M. raiateensi J.W. Moore et aliquantum M. lanceolatae J.R. Forst. &amp; G. Forst. similis, sed ab eis (atque adeo ab omnibus congeneris Societatis Insularum ac Tuamotu) ovariis omnino liberis distinguitur.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="150" type="type">
<paragraph pageId="1" pageNumber="150">Type.</paragraph>
<paragraph pageId="1" pageNumber="150">
<emphasis bold="true" pageId="1" pageNumber="150">French Polynesia.</emphasis>
Marquesas Islands, Hiva-Oa, Hanamenu valley off Hanamenu trail, in
<emphasis italics="true" pageId="1" pageNumber="150">Metrosideros-Weinmannia-Dicranopteris linearis</emphasis>
wet forest,
<geoCoordinate degrees="09" direction="south" minutes="47" orientation="latitude" precision="15" seconds="50" value="-9.797223">09°47'50&quot;S</geoCoordinate>
,
<geoCoordinate degrees="139" direction="west" minutes="05" orientation="longitude" precision="15" seconds="35" value="-139.09305">139°05'35&quot;W</geoCoordinate>
, 908 m, 2 August 2005 (fr), S. Perlman 19767 (holotype: P [P00398408]!; isotypes, PAP!, PTBG!, US!).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="152" pageId="1" pageNumber="150" type="description">
<paragraph pageId="1" pageNumber="150">Description.</paragraph>
<paragraph lastPageId="3" lastPageNumber="152" pageId="1" pageNumber="150">
Monocaulous to sparsely branched tree, dioecious, Chamberlain architecture (tending toward Leeuwenberg architecture (
<bibRefCitation author="Halle," journalOrPublisher="Montpellier" pageId="4" pageNumber="153" refId="B12" refString="Halle, , 2004. Architecture des plantes. JPC Edition. Montpellier, 162 pp." title="Architecture des plantes. JPC Edition" year="2004">
<normalizedToken originalValue="Hallé">Halle</normalizedToken>
, 2004
</bibRefCitation>
) when very old, fide P.A.
<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
, 1975), 3
<normalizedToken originalValue="6(">-6(-</normalizedToken>
10) m tall, without milky sap. Leaves simple, alternate, grouped at branch ends; petiole 1.5
<normalizedToken originalValue="2.5(">-2.5(-</normalizedToken>
4) cm long, 2-3 mm in diam., base enlarged, slightly clasping, lenticels rarely present abaxially, without dark green transverse striations adaxially when fresh; ligule present adaxially at the base of the petiole, persistent, free portion triangular to widely triangular, 5
<normalizedToken originalValue="11(">-11(-</normalizedToken>
17) mm long, margins entire, membranous, apex obtuse to acute; blade green, shiny adaxially when fresh, slightly lighter green abaxially, narrowly obovate to spatulate, 20.4-40
<normalizedToken originalValue="×">x</normalizedToken>
5.2-9.8 cm (l/w ratio 2.76-5.52), base symmetric to asymmetric, offset 2
<normalizedToken originalValue="5(">-5(-</normalizedToken>
10) mm, mostly positively centered in adaxial view, attenuate, margin entire, undulate distally, minutely revolute, apex acute to obtuse, the extreme tip obtuse to mucronate, chartaceous to coriaceous, glabrous on both surfaces; venation brochidodromous, light green-yellow when fresh, with prominent arches lacking intramarginal veins; midvein strong and massive, straight, without evident abaxial thickenings; secondary veins (12
<normalizedToken originalValue=")17">-)17-</normalizedToken>
22 pairs, diverging from the midvein by (53
<normalizedToken originalValue=")57">-)57-</normalizedToken>
69° (in the widest part of the blade), the distal ones less divergent, weakly recurved near the midvein then strongly recurved toward the margin, joining with the next arch at an obtuse angle without forming a clear intramarginal vein, intersecondary veins present; tertiary veins evident, not reaching the margin, straight to anastomosing at various angles, sometimes forming convex arcs in the intercostals zone; higher order veins visible in dry material, forming a fine, dense reticulum; veinlets forming quadri- to multi-angular areoles 0.5-1.5 mm in size. Juvenile foliage similar to adult, blade with an obtuse apex. Male material unknown (only old inflorescences seen). Female inflorescence terminal, erect, a raceme of spikes, with 1 degree of branching, axes and peduncle light green, bracts brown-red, primary axis 15-20 cm long, 5-8 mm in
<pageBreakToken pageId="2" pageNumber="151" start="start">diam</pageBreakToken>
. at the base when fresh, subtended by persistent or sometimes caducous triangular cataphylls 13-24 mm long, without lenticels, margins entire, mostly to almost entirely membranous, apex obtuse, bearing a dorsal,
<normalizedToken originalValue="±">+/-</normalizedToken>
foliaceous apicule; secondary
<pageBreakToken pageId="3" pageNumber="152" start="start">axes</pageBreakToken>
8
<normalizedToken originalValue="13(">-13(-</normalizedToken>
17), 6-12 cm long (shorter distally), straight or sometimes slightly curved upward in the distal portion, diverging 35-60° from the primary axis, each subtended by a caducous, triangular to narrowly triangular bract 7-20 mm long, without lenticels, margin denticulate, mostly membranous, apex acute to acuminate, sometimes apiculate; spikelets with 7
<normalizedToken originalValue="15(">-15(-</normalizedToken>
27) flowers, the proximal one inserted 3-15 mm from the base of the secondary axis. Female flowers white when fresh, sessile; bractlets caducous or sometimes persistent, broadly triangular, 3-5 mm long, margin weakly dentate, membranous, brown-red, partially covered by the flower and later the fruit, apex acute to obtuse; petals 5-11, caducous in fruit, ovate, 1-2.5 mm long, weakly cucullate, recurved when flowers receptive, with a pronounced adaxial groove, apex acute; androeciuum present, isostemonous, filaments and anthers developed but pollen sacs empty, stamens with filaments 0.5-1 mm long, anthers 0.5 mm long; ovary inferior, (5-)7 or 8(-11)-carpellate, nectar disc epigynous, 2 mm diam., styles weakly differentiated, to 1 mm long, stigmas 1.5-3 mm long at receptivity, often strongly recurved to twisted. Fruit yellow-green when young, light purple at maturity, remains of petals and stamens brown, ovaries entirely free and distinct from one another; drupes globose to sub-globose, 7-9
<normalizedToken originalValue="×">x</normalizedToken>
7-10 mm, smooth and fleshy when fresh, ribbed when dry, the ribs corresponding to the 5-11 pyrenes.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="152" type="distribution">
<paragraph pageId="3" pageNumber="152">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="152">
Known only from the type locality in the Mt. Temetiu/Feani area at ca. 900 m elevation along the Hanamenu trail on the island of Hiva Oa (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Meryta pastoralis F. Tronchet &amp; Lowry on Hiva Oa Island in the Marquesas archipelago (blue lines indicate primary water courses)." pageId="3" pageNumber="152">Fig. 3</figureCitation>
), the largest and highest in the Marquesas archipelago.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="152" type="ecology">
<paragraph pageId="3" pageNumber="152">Ecology.</paragraph>
<paragraph pageId="3" pageNumber="152">
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="3" pageNumber="152" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="3" pageNumber="152">Meryta pastoralis</emphasis>
</taxonomicName>
occurs on slopes and along streams in low stature, humid montane
<emphasis italics="true" pageId="3" pageNumber="152">Metrosideros-Weinmannia</emphasis>
forest at and above 900 m elevation.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="152" type="etymology">
<paragraph pageId="3" pageNumber="152">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="152">
This species is named in honor of P.A.
<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
, who was the first to collect it in 1975 while conducting botanical inventory work in the Marquesas, and who has contributed much to our knowledge of the flora of the archipelago. The name
<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
means shepherd in Alsatian, which when translated into Latin provides the basis for our choice of the epithet
<emphasis italics="true" pageId="3" pageNumber="152">pastoralis</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="152" type="conservation status">
<paragraph pageId="3" pageNumber="152">Conservation status.</paragraph>
<paragraph pageId="3" pageNumber="152">
<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="3" pageNumber="152" phylum="Tracheophyta" rank="species" species="pastorali">
<emphasis italics="true" pageId="3" pageNumber="152">Meryta pastorali</emphasis>
</taxonomicName>
s is known from a single population along a wind-swept ridge below the highest point on Hiva Oa. With an Area of Occupancy of &lt;10 km2 and an Extent of Occurrence that is probably no larger, it meets the area requirements for Critically Endangered status under criteria B1 and B2 of the IUCN Red List Criteria (
<bibRefCitation author="IUCN," journalOrPublisher="IUCN, Gland, Switzerland" pageId="4" pageNumber="153" publicationUrl="http://www.iucnredlist.org/technical-documents/categories-and-criteria/2001-categories-criteria" refId="B3" refString="IUCN, , 2001. IUCN Red List Categories and Criteria Version 3.1. Prepared by the IUCN Species Survival Commission. IUCN, Gland, Switzerland, 33 pp., http://www.iucnredlist.org/technical-documents/categories-and-criteria/2001-categories-criteria" title="IUCN Red List Categories and Criteria Version 3.1. Prepared by the IUCN Species Survival Commission." url="http://www.iucnredlist.org/technical-documents/categories-and-criteria/2001-categories-criteria" year="2001">IUCN 2001</bibRefCitation>
). While the vegetation in the immediate area is still largely intact, increasing impacts from feral pigs, human disturbance and fire during the dry season (D. Lorence, pers. comm.) represent growing threats to
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="3" pageNumber="152" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="3" pageNumber="152">Meryta pastoralis</emphasis>
</taxonomicName>
, prompting us to assign it a provisional threat status of Critically Endangered (CR B1ab(i,ii,iii,v) + 2ab(i,ii,iii,v)).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="153" pageId="3" pageNumber="152" type="other specimens examined">
<paragraph pageId="3" pageNumber="152">Other specimens examined.</paragraph>
<paragraph lastPageId="4" lastPageNumber="153" pageId="3" pageNumber="152">
<emphasis bold="true" pageId="3" pageNumber="152">French Polynesia.</emphasis>
Marquesas Islands, Hiva Oa, trail toward Hanamenu; in gulch off trail, low forest, 3 August 1988 (fr). S. Perlman 10206 (AD!, BISH! [2 sheets], F!, K!, MO!, NY!, P [P00372510]!, PAP!, PTBG!, US!); Teakatu, valley on north side of Hanamenu trail heading down to Hanamenu past summit crest, between Teakatau and Tepuna, along stream bottom in
<emphasis italics="true" pageId="3" pageNumber="152">Metrosideros-Weinmannia-Crossostylis</emphasis>
montane wet forest, [
<geoCoordinate degrees="09" direction="south" minutes="47" orientation="latitude" precision="15" seconds="29" value="-9.791389">09°47'29&quot;S</geoCoordinate>
,
<geoCoordinate degrees="139" direction="west" minutes="05" orientation="longitude" precision="15" seconds="41" value="-139.09471">139°05'41&quot;W</geoCoordinate>
], 933 m, 26
<pageBreakToken pageId="4" pageNumber="153" start="start">August</pageBreakToken>
1995 (st), S. Perlman &amp; J.-Y. Meyer 14894 (BISH!, MO!, PAP!, PTBG!, US!); Feani area, Tepuna, in gulch to north of Hanamenu trail, down west side of summit crest to 1075 meters elev. and drop into gulch, in
<emphasis italics="true" pageId="4" pageNumber="153">Metrosideros-Weinmannia</emphasis>
forest,
<geoCoordinate degrees="09" direction="south" minutes="47" orientation="latitude" precision="15" seconds="49" value="-9.796945">09°47'49&quot;S</geoCoordinate>
,
<geoCoordinate degrees="139" direction="west" minutes="05" orientation="longitude" precision="15" seconds="12" value="-139.08665">139°05'12&quot;W</geoCoordinate>
, 900 m, 29 January 2003 (old ♂fl), S. Perlman 18335 (BISH, P [P00398011]!, PAP, PTBG, US); Feani area, Tepuna, off Hanamenu trail on west side of summit crest, at 1075 meters elev. into gulch north side of trail, scattered plants in
<emphasis italics="true" pageId="4" pageNumber="153">Metrosideros-Weinmannia</emphasis>
forest,
<geoCoordinate degrees="09" direction="south" minutes="47" orientation="latitude" precision="15" seconds="49" value="-9.796945">09°47'49&quot;S</geoCoordinate>
,
<geoCoordinate degrees="139" direction="west" minutes="05" orientation="longitude" precision="15" seconds="12" value="-139.08665">139°05'12&quot;W</geoCoordinate>
, 29 January 2003 (♀fl), S. Perlman 18336 (PAP, PTBG, US); Montagnes NW du Temetiu, entre la haute
<normalizedToken originalValue="vallée">vallee</normalizedToken>
de Hanamenu et la
<normalizedToken originalValue="crête">crete</normalizedToken>
du Temetiu Feani, haute
<normalizedToken originalValue="vallée">vallee</normalizedToken>
,
<normalizedToken originalValue="forêt">foret</normalizedToken>
humide, [
<geoCoordinate degrees="09" direction="south" minutes="48" orientation="latitude" precision="15" seconds="00" value="-9.8">09°48'00&quot;S</geoCoordinate>
,
<geoCoordinate degrees="139" direction="west" minutes="05" orientation="longitude" precision="15" seconds="30" value="-139.09166">139°05'30&quot;W</geoCoordinate>
], 925 m, 23 October 1975 (♀fl), P.A.
<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
5922 (BISH!, K!, MO! [2 sheets], NTBG!, P [P00372508 &amp; P00372509]!, US!).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="153" type="discussion">
<paragraph pageId="4" pageNumber="153">Discussion.</paragraph>
<paragraph pageId="4" pageNumber="153">
Material of our new species most closely resembles specimens of
<taxonomicName authorityName="Harms" authorityYear="1938" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="choristantha">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta choristantha</emphasis>
</taxonomicName>
Harms from the Austral Islands, with which it shares a similarly structured inflorescence and totally free ovaries, but differs in leaf shape (narrowly obovate to spatulate in
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta pastoralis</emphasis>
</taxonomicName>
vs. widely elliptic to slightly obovate in
<taxonomicName authorityName="Harms" authorityYear="1938" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="choristantha">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta choristantha</emphasis>
</taxonomicName>
), number of carpels (5-11 vs. 5 or 6), and fruit size at maturity (7-9
<normalizedToken originalValue="×">x</normalizedToken>
7-10 vs. 8-12
<normalizedToken originalValue="×">x</normalizedToken>
10-15 mm). The leaves of
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta pastoralis</emphasis>
</taxonomicName>
are similar in appearance to those of
<taxonomicName authorityName="J. W. Moore" authorityYear="1963" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="raiateensis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta raiateensis</emphasis>
</taxonomicName>
J.W. Moore, and to a lesser degree
<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="lanceolata">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta lanceolata</emphasis>
</taxonomicName>
J.R. Forst. &amp; G. Forst., both from the Society Islands, but our new taxon differs from these entities, and indeed all other species in the Society Islands and on Tuamotu, by its totally free ovaries.
</paragraph>
<paragraph pageId="4" pageNumber="153">
Results from ongoing molecular phylogenetic work (
<bibRefCitation author="Tronchet, F" journalOrPublisher="Systematic Botany" pageId="4" pageNumber="153" pagination="657 - 670" publicationUrl="http://www.ingentaconnect.com/content/aspt/sb/2005/00000030/00000003/art00016" refId="B10" refString="Tronchet, F, Plunkett, GM, Jeremie, J, Lowry II, PP, 2005a. Monophyly and major clades of Meryta (Araliaceae). Systematic Botany 30: 657 - 670, http://www.ingentaconnect.com/content/aspt/sb/2005/00000030/00000003/art00016" title="Monophyly and major clades of Meryta (Araliaceae)." url="http://www.ingentaconnect.com/content/aspt/sb/2005/00000030/00000003/art00016" volume="30" year="2005 a">Tronchet et al. 2005a</bibRefCitation>
,
<bibRefCitation author="Tronchet, F" journalOrPublisher="Vienna, Austria" pageId="4" pageNumber="153" refId="B11" refString="Tronchet, F, Plunkett, GM, Jeremie, J, Lowry II, PP, 2005b. Phylogeny and biogeography of the Pacific genus Meryta (Araliaceae) based on ITS and ETS sequence data. In: Poster presented at the 17th International Botanical Congress. Vienna, Austria" title="Phylogeny and biogeography of the Pacific genus Meryta (Araliaceae) based on ITS and ETS sequence data." volumeTitle="Poster presented at the 17 th International Botanical Congress." year="2005 b">2005b</bibRefCitation>
, unpubl. data) suggest that
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta pastoralis</emphasis>
</taxonomicName>
is part of a small clade now understood to comprise four species, also including
<taxonomicName authorityName="Seem" authorityYear="1862" baseAuthorityName="Hook. f." class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="sinclairii">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta sinclairii</emphasis>
</taxonomicName>
from New Zealand,
<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="tenuifolia">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta tenuifolia</emphasis>
</taxonomicName>
A.C. Sm. from Fiji, and
<taxonomicName authorityName="Harms" authorityYear="1938" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="choristantha">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta choristantha</emphasis>
</taxonomicName>
, which together correspond to
<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta</emphasis>
</taxonomicName>
sect.
<taxonomicName genus="Choristomeryta" lsidName="" pageId="4" pageNumber="153" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="153">Choristomeryta</emphasis>
</taxonomicName>
Harms (
<bibRefCitation author="Harms, H" journalOrPublisher="Notizblatt des botanischen Gartens und Muzeums zu Berlin-Dahlem" pageId="4" pageNumber="153" pagination="315 - 321" refId="B2" refString="Harms, H, 1938. Zur Kenntnis von Meryta sonchifolia Linden et Andre und einigen anderen Arten des Gattung. Notizblatt des botanischen Gartens und Muzeums zu Berlin-Dahlem 14: 315 - 321" title="Zur Kenntnis von Meryta sonchifolia Linden et Andre und einigen anderen Arten des Gattung." volume="14" year="1938">Harms 1938</bibRefCitation>
). Material here assigned to
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta pastoralis</emphasis>
</taxonomicName>
was referred to informally as 'M.
<normalizedToken originalValue="schaeferi">schaeferi'</normalizedToken>
in
<bibRefCitation author="Tronchet, F" journalOrPublisher="Vienna, Austria" pageId="4" pageNumber="153" refId="B11" refString="Tronchet, F, Plunkett, GM, Jeremie, J, Lowry II, PP, 2005b. Phylogeny and biogeography of the Pacific genus Meryta (Araliaceae) based on ITS and ETS sequence data. In: Poster presented at the 17th International Botanical Congress. Vienna, Austria" title="Phylogeny and biogeography of the Pacific genus Meryta (Araliaceae) based on ITS and ETS sequence data." volumeTitle="Poster presented at the 17 th International Botanical Congress." year="2005 b">Tronchet et al. (2005b)</bibRefCitation>
.
</paragraph>
<caption pageId="4" pageNumber="153" start="Figure 1" startId="F1">
<paragraph pageId="4" pageNumber="153">
<emphasis bold="true" pageId="4" pageNumber="153">Figure 1.</emphasis>
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta pastoralis</emphasis>
</taxonomicName>
F. Tronchet &amp; Lowry.
<emphasis bold="true" pageId="4" pageNumber="153">A</emphasis>
branch with leaves and young infructescence
<emphasis bold="true" pageId="4" pageNumber="153">B</emphasis>
female flowers in bud subtended by bracts
<emphasis bold="true" pageId="4" pageNumber="153">C</emphasis>
female flowers at receptivity
<emphasis bold="true" pageId="4" pageNumber="153">D</emphasis>
young fruits. Line drawing by Alain Jouy from herbarium specimens; voucher Perlman 19767 (
<emphasis bold="true" pageId="4" pageNumber="153">A, D</emphasis>
),
<normalizedToken originalValue="Schäfer">Schaefer</normalizedToken>
5922 (
<emphasis bold="true" pageId="4" pageNumber="153">B</emphasis>
), Perlman 10206 (
<emphasis bold="true" pageId="4" pageNumber="153">C</emphasis>
).
</paragraph>
</caption>
<caption pageId="4" pageNumber="153" start="Figure 2" startId="F2">
<paragraph pageId="4" pageNumber="153">
<emphasis bold="true" pageId="4" pageNumber="153">Figure 2.</emphasis>
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta pastoralis</emphasis>
</taxonomicName>
F. Tronchet &amp; Lowry. Photo of female plant in fruit (Photo by Steve Perlman of Perlman 19767).
</paragraph>
</caption>
<caption pageId="4" pageNumber="153" start="Figure 3" startId="F3">
<paragraph pageId="4" pageNumber="153">
<emphasis bold="true" pageId="4" pageNumber="153">Figure 3.</emphasis>
Distribution map of
<taxonomicName authorityName="F. Tronchet &amp; Lowry" authorityYear="2011" class="Magnoliopsida" family="Araliaceae" genus="Meryta" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Apiales" pageId="4" pageNumber="153" phylum="Tracheophyta" rank="species" species="pastoralis">
<emphasis italics="true" pageId="4" pageNumber="153">Meryta pastoralis</emphasis>
</taxonomicName>
F. Tronchet &amp; Lowry on Hiva Oa Island in the Marquesas archipelago (blue lines indicate primary water courses).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>