1074 lines
134 KiB
XML
1074 lines
134 KiB
XML
<document id="9E7D3FB54D13897F2FB57581F813B670" ID-CLB-Dataset="30928" ID-DOI="10.11646/zootaxa.4375.3.2" ID-GBIF-Dataset="4dab2987-0956-4343-b019-543f793aeaa1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1158870" ID-ZooBank="3363F72A-6A48-4685-98C9-8553F9A79F20" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1516866532209" checkinUser="plazi" docAuthor="Phimmachak, Somphouthone, Sivongxay, Niane, Seateun, Sengvilay, Yodthong, Siriporn, Rujirawan, Attapol, Neang, Thy, Aowphol, Anchalee & Stuart, Bryan L." docDate="2018" docId="6146056EFF98FF82FF3AFAE1FE77A78C" docLanguage="en" docName="zootaxa.4375.3.2.pdf.imf" docOrigin="Zootaxa 4375 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limnonectes coffeatus Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="336" masterDocId="9D7F7D16FF9CFF89FFADFFB1FFCEA460" masterDocTitle="A new Limnonectes (Anura: Dicroglossidae) from southern Laos" masterLastPageNumber="340" masterPageNumber="325" pageNumber="329" updateTime="1698517071176" updateUser="ExternalLinkService">
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<mods:mods id="8CAAD36064A73E9E620637132EAFDD25" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="329E8BF036C2C5EF094CB1A047297FB9">
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<mods:title id="9A042B322B3F714DDC40A4F783209419">A new Limnonectes (Anura: Dicroglossidae) from southern Laos</mods:title>
|
||
</mods:titleInfo>
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<mods:name id="CA618DE4AAD73BDBDC9BB1C89063C39F" type="personal">
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<mods:role id="6AFD1F8692C948973C18D83AD34DC421">
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<mods:roleTerm id="0002A4422E36D3F7A10A373E5B3859FD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2FBD673454672120B6326AD2FE30B1FE">Phimmachak, Somphouthone</mods:namePart>
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</mods:name>
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<mods:name id="9A02C1B160F50D3E044EE5EF6683B2AE" type="personal">
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<mods:roleTerm id="2C72F5E61669D17B3FE4E50B4408FB57">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="918112A2415134D0AA75468044C0B288">Sivongxay, Niane</mods:namePart>
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</mods:name>
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<mods:name id="5861983C894D9567E1DD4797D90AB7E6" type="personal">
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<mods:role id="E7C37FCCA60F4C0FE5D22C26E0154AA5">
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<mods:roleTerm id="FCB778A0207CFFCEAA69FDA064A7CE67">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="974B0D5074711D44DE2128F16974594E">Seateun, Sengvilay</mods:namePart>
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||
</mods:name>
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<mods:name id="9D8E77F21134E0B8130ABF17291A5F0E" type="personal">
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<mods:role id="8C29FE3C9CD7BB79AACA9D7DF7A9DABF">
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<mods:roleTerm id="86F328BF6C0181905485A14F14C20190">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F26AA9E14DFB8E402FF6F57C6ADF8A7">Yodthong, Siriporn</mods:namePart>
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</mods:name>
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<mods:name id="70285960612A9758007B481E080540BA" type="personal">
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<mods:role id="271226BAA886B97C2D17D64FEACC8A43">
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<mods:roleTerm id="A44EA882C9B75FDB83C6E3A795046D4D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6A145D1D8E570D0AB91B66ED6429103C">Rujirawan, Attapol</mods:namePart>
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</mods:name>
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<mods:name id="F58B85104215B2E5267C6BBD5624211E" type="personal">
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<mods:role id="B3E21C9CABB439C53B88EB134F90A54B">
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<mods:roleTerm id="FFA03A7B633E107E5EACC9F8386C0B7D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="55F8DB2FB62DA0727D47D9B531999C9D">Neang, Thy</mods:namePart>
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</mods:name>
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<mods:name id="7EF5EA71E780F5278ACFE36256B4DFDD" type="personal">
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<mods:roleTerm id="C11994EDC7D6FD2468ECB2D242AA7FBE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="54823CAAF54DF6B2A4A92439963D97CC">Aowphol, Anchalee</mods:namePart>
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</mods:name>
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<mods:name id="73B38FB59ECF9FBEDF71E4E1D8ACB8DF" type="personal">
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<mods:roleTerm id="56EC0E32DE168B91ECBFA68BFB7530B5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F169179F72EACA6E5445452D57A2AD9">Stuart, Bryan L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="98499AB4A37CECDAD269976FD49D5620">text</mods:typeOfResource>
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<mods:title id="27BB71405DD4B5E4EB82D0E3C87B106A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="329F72CE9850C8E60A8D31ABFF863282">
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<mods:date id="EBCCD354D135A5B6BD0B41DB4F1F9AE8">2018</mods:date>
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<mods:detail id="957C50485C05C9525D4962C292E6418E" type="pubDate">
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<mods:number id="C6FD03FAFAC23B307CBD0AA36CC5A854">2018-01-25</mods:number>
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</mods:detail>
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<mods:detail id="8B287BCD259D4D4151504098D2B1CDA4" type="volume">
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<mods:number id="7C8E20A6FF61D842BF1735F36D45D655">4375</mods:number>
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</mods:detail>
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<mods:detail id="1CAB53A94B27884F68971F4DA9570FDD" type="issue">
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<mods:number id="26DF03A10CFA99EBA6CDB5D7C7D43C29">3</mods:number>
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</mods:detail>
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<mods:start id="8F140EB915A32702C7FA13FC45C8B272">325</mods:start>
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<mods:end id="916BB2B9214D9FC7783CCECF77A12161">340</mods:end>
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</mods:extent>
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<mods:classification id="0AC136CB03F404867C8A0D1C9FB3F4CD">journal article</mods:classification>
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<mods:identifier id="375C2E057EF7BC3C134469B48E8B441B" type="CLB-Dataset">30928</mods:identifier>
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<mods:identifier id="4474513CD55CD86B5ED32895CDC59631" type="DOI">10.11646/zootaxa.4375.3.2</mods:identifier>
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<mods:identifier id="78F5A8FBF2AD7FCB49535D0DBAF2FACC" type="GBIF-Dataset">4dab2987-0956-4343-b019-543f793aeaa1</mods:identifier>
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<mods:identifier id="944AE44C479A43F5DF5F9D2188970106" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C9237CD84D62988DBC7D0B09EBED76A8" type="Zenodo-Dep">1158870</mods:identifier>
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<mods:identifier id="CD61FB53063C3E8D2AF713B2B731FC10" type="ZooBank">3363F72A-6A48-4685-98C9-8553F9A79F20</mods:identifier>
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</mods:mods>
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<treatment id="6146056EFF98FF82FF3AFAE1FE77A78C" ID-DOI="http://doi.org/10.5281/zenodo.6488965" ID-GBIF-Taxon="140417782" ID-Zenodo-Dep="6488965" LSID="urn:lsid:plazi:treatment:6146056EFF98FF82FF3AFAE1FE77A78C" httpUri="http://treatment.plazi.org/id/6146056EFF98FF82FF3AFAE1FE77A78C" lastPageId="11" lastPageNumber="336" pageId="4" pageNumber="329">
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||
<subSubSection id="A1F5E7F3FF98FF8DFF3AFAE1FDC2A10B" box="[151,524,1360,1387]" pageId="4" pageNumber="329" type="nomenclature">
|
||
<paragraph id="E950B478FF98FF8DFF3AFAE1FDC2A10B" blockId="4.[151,524,1360,1387]" box="[151,524,1360,1387]" pageId="4" pageNumber="329">
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<heading id="B2180314FF98FF8DFF3AFAE1FDC2A10B" bold="true" box="[151,524,1360,1387]" fontSize="11" level="1" pageId="4" pageNumber="329" reason="1">
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||
<emphasis id="DB9B686AFF98FF8DFF3AFAE1FDC2A10B" bold="true" box="[151,524,1360,1387]" pageId="4" pageNumber="329">
|
||
<taxonomicName id="2EEFCFFBFF98FF8DFF3AFAE1FE69A10A" ID-CoL="72DKZ" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[151,423,1360,1386]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="329" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF98FF8DFF3AFAE1FE69A10A" bold="true" box="[151,423,1360,1386]" italics="true" pageId="4" pageNumber="329">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C0A8D511FF98FF8DFE03FAE0FDC2A10B" box="[430,524,1361,1387]" pageId="4" pageNumber="329" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1F5E7F3FF98FF8DFF3AFA26FA4BA2B0" pageId="4" pageNumber="329" type="materials_examined">
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||
<paragraph id="E950B478FF98FF8DFF3AFA26FB1CA198" blockId="4.[151,1436,1431,2032]" pageId="4" pageNumber="329">
|
||
<materialsCitation id="5987BE25FF98FF8DFF3AFA26FB1CA198" ID-GBIF-Occurrence="1805284686" collectingDate="2010-08-03" collectionCode="NCSM, BLS" collectorName="Bryan L. Stuart & Niane Sivongxay & Sengvilay Seateun" country="Laos" elevation="1235" latitude="15.06237" location="Bolaven Plateau" longLatPrecision="1" longitude="106.21075" municipality="Pakxong District" pageId="4" pageNumber="329" specimenCode="NCSM 77788, BLS 13957" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" stateProvince="Champasak Province" typeStatus="holotype">
|
||
<emphasis id="DB9B686AFF98FF8DFF3AFA26FEC3A1D0" bold="true" box="[151,269,1431,1456]" pageId="4" pageNumber="329">
|
||
<typeStatus id="36540ADAFF98FF8DFF3AFA26FEC6A1D0" box="[151,264,1431,1456]" pageId="4" pageNumber="329" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="B9491C03FF98FF8DFEB9FA29FE7EA1D0" box="[276,432,1432,1456]" pageId="4" pageNumber="329">NCSM 77788</specimenCode>
|
||
(field tag
|
||
<specimenCode id="B9491C03FF98FF8DFD8EFA29FD69A1D0" box="[547,679,1431,1456]" pageId="4" pageNumber="329">BLS 13957</specimenCode>
|
||
),
|
||
<specimenCount id="FFE97FF1FF98FF8DFD16FA26FD3DA1D0" box="[699,755,1431,1456]" pageId="4" pageNumber="329" type="adult">adult</specimenCount>
|
||
<specimenCount id="FFE97FF1FF98FF8DFD57FA26FCFEA1D0" box="[762,816,1431,1456]" pageId="4" pageNumber="329" type="male">male</specimenCount>
|
||
(
|
||
<figureCitation id="71D4A8FDFF98FF8DFC92FA26FC4DA1D0" box="[831,899,1431,1456]" captionStart="FIGURE 2" captionStartId="5.[151,250,1218,1240]" captionTargetBox="[156,1429,343,1189]" captionTargetId="figure@5.[150,1436,337,1196]" captionTargetPageId="5" captionText="FIGURE 2. Holotype male of Limnonectes coffeatus sp. nov. (NCSM 77788, field tag BLS 13957) in dorsal (A), ventral (B), profile (C), and posterior (D)views immediately prior to preservation." httpUri="https://zenodo.org/record/1158874/files/figure.png" pageId="4" pageNumber="329">Fig. 2</figureCitation>
|
||
),
|
||
<collectingCountry id="91F8F4E8FF98FF8DFC34FA29FC1CA1CF" box="[921,978,1432,1455]" name="Laos" pageId="4" pageNumber="329">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="2B2B7A9AFF98FF8DFC70FA26FB00A1D0" box="[989,1230,1431,1456]" country="Laos" name="Champasak" pageId="4" pageNumber="329">Champasak Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="09342E02FF98FF8DFB75FA26FA56A1D0" box="[1240,1432,1431,1456]" pageId="4" pageNumber="329">Pakxong District</collectingMunicipality>
|
||
,
|
||
<location id="EC30E2A3FF98FF8DFF3AFA0AFE9AA1B4" LSID="urn:lsid:plazi:treatment:6146056EFF98FF82FF3AFAE1FE77A78C:EC30E2A3FF98FF8DFF3AFA0AFE9AA1B4" box="[151,340,1467,1492]" country="Laos" latitude="15.06237" longLatPrecision="1" longitude="106.21075" municipality="Pakxong District" name="Bolaven Plateau" pageId="4" pageNumber="329" stateProvince="Champasak Province">Bolaven Plateau</location>
|
||
,
|
||
<location id="EC30E2A3FF98FF8DFEC9FA0DFCF8A1B4" LSID="urn:lsid:plazi:treatment:6146056EFF98FF82FF3AFAE1FE77A78C:EC30E2A3FF98FF8DFEC9FA0DFCF8A1B4" box="[356,822,1467,1492]" country="Laos" latitude="15.06237" longLatPrecision="1" longitude="106.21075" municipality="Pakxong District" name="Dong Hua Sao National Protected Area" pageId="4" pageNumber="329" stateProvince="Champasak Province">Dong Hua Sao National Protected Area</location>
|
||
,
|
||
<location id="EC30E2A3FF98FF8DFCEBFA0DFBAAA1B4" LSID="urn:lsid:plazi:treatment:6146056EFF98FF82FF3AFAE1FE77A78C:EC30E2A3FF98FF8DFCEBFA0DFBAAA1B4" box="[838,1124,1467,1492]" country="Laos" latitude="15.06237" longLatPrecision="1" longitude="106.21075" municipality="Pakxong District" name="Houay Tad Seua Stream" pageId="4" pageNumber="329" stateProvince="Champasak Province">Houay Tad Seua Stream</location>
|
||
,
|
||
<geoCoordinate id="8CDBD2BFFF98FF8DFBDEFA0AFB34A1B4" box="[1139,1274,1467,1492]" direction="north" orientation="latitude" pageId="4" pageNumber="329" precision="1" value="15.06237">15.06237°N</geoCoordinate>
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||
<geoCoordinate id="8CDBD2BFFF98FF8DFAA8FA0AFA56A1B4" box="[1285,1432,1467,1492]" direction="east" orientation="longitude" pageId="4" pageNumber="329" precision="1" value="106.21075">106.21075°E</geoCoordinate>
|
||
,
|
||
<elevation id="62C2534BFF98FF8DFF3AFA6EFF3DA197" box="[151,243,1503,1528]" metricMagnitude="3" metricUnit="m" metricValue="1.2349999999999999" pageId="4" pageNumber="329" unit="m" value="1235.0">
|
||
<quantity id="2E17199DFF98FF8DFF3AFA6EFF3DA197" box="[151,243,1503,1528]" metricMagnitude="3" metricUnit="m" metricValue="1.2349999999999999" pageId="4" pageNumber="329" unit="m" value="1235.0">1,235 m</quantity>
|
||
</elevation>
|
||
elev., coll.
|
||
<collectingDate id="8D156B50FF98FF8DFED9FA51FDD6A198" box="[372,536,1504,1528]" pageId="4" pageNumber="329" value="2010-08-03">
|
||
<date id="9D5192B8FF98FF8DFED9FA51FDD6A198" box="[372,536,1504,1528]" pageId="4" pageNumber="329" value="2010-08-03">3 August 2010</date>
|
||
</collectingDate>
|
||
by
|
||
<collectorName id="441AD1AEFF98FF8DFDEFFA51FD3CA198" box="[578,754,1504,1528]" pageId="4" pageNumber="329">Bryan L. Stuart</collectorName>
|
||
,
|
||
<collectorName id="441AD1AEFF98FF8DFD50FA6EFC0EA198" box="[765,960,1503,1528]" pageId="4" pageNumber="329">Niane Sivongxay</collectorName>
|
||
, and
|
||
<collectorName id="441AD1AEFF98FF8DFC56FA6EFB00A198" box="[1019,1230,1503,1528]" pageId="4" pageNumber="329">Sengvilay Seateun</collectorName>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E950B478FF98FF8DFF6AF9B2FBA6A2CC" blockId="4.[151,1436,1431,2032]" pageId="4" pageNumber="329">
|
||
<emphasis id="DB9B686AFF98FF8DFF6AF9B2FE86A27C" bold="true" box="[199,328,1539,1564]" pageId="4" pageNumber="329">
|
||
<typeStatus id="36540ADAFF98FF8DFF6AF9B2FE8DA27C" box="[199,323,1539,1564]" pageId="4" pageNumber="329" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectionCode id="8FFE2CBDFF98FF8DFEFEF9B5FE6CA27C" box="[339,418,1540,1564]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="4" pageNumber="329">NCSM</collectionCode>
|
||
77786, adult female, same data as
|
||
<typeStatus id="36540ADAFF98FF8DFCE7F9B2FC62A27C" box="[842,940,1539,1564]" pageId="4" pageNumber="329" type="holotype">holotype</typeStatus>
|
||
except coll.
|
||
<date id="9D5192B8FF98FF8DFBE4F9B5FB28A27C" box="[1097,1254,1539,1564]" pageId="4" pageNumber="329" value="2010-07-30">30 July 2010</date>
|
||
.
|
||
<collectionCode id="8FFE2CBDFF98FF8DFB59F9B5FA8DA27C" box="[1268,1347,1540,1564]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="4" pageNumber="329">NCSM</collectionCode>
|
||
77785,
|
||
<collectionCode id="8FFE2CBDFF98FF8DFF3AF999FF28A220" box="[151,230,1576,1600]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="4" pageNumber="329">NCSM</collectionCode>
|
||
77787 (
|
||
<figureCitation id="71D4A8FDFF98FF8DFEEAF996FE64A220" box="[327,426,1575,1600]" captionStart-0="FIGURE 3" captionStart-1="FIGURE4" captionStartId-0="6.[151,250,1287,1309]" captionStartId-1="8.[151,250,1050,1072]" captionTargetBox-0="[156,1429,414,1260]" captionTargetBox-1="[409,1177,194,1029]" captionTargetId-0="figure@6.[151,1435,409,1266]" captionTargetId-1="figure@8.[409,1177,194,1029]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURE 3. Paratype female of Limnonectes coffeatus sp. nov. (NCSM 77787, field tag BLS 13934) in lateral (A) and dorsal (B) views in life, and in ventral (C) and lateral (D) views immediately prior to preservation." captionText-1="FIGURE4. Paratypefemale (NCSM 77787, left)and holotype male (NCSM77788,right)of Limnonectes coffeatussp.nov. in dorsal (A) and ventral (B) views in preservative." httpUri-0="https://zenodo.org/record/1158876/files/figure.png" httpUri-1="https://zenodo.org/record/1158878/files/figure.png" pageId="4" pageNumber="329">Fig. 3–4</figureCitation>
|
||
), two adult females;
|
||
<collectionCode id="8FFE2CBDFF98FF8DFD30F999FD25A220" box="[669,747,1576,1600]" country="Laos" httpUri="http://grbio.org/cool/i4me-fta1" name="National University of Laos" pageId="4" pageNumber="329">NUOL</collectionCode>
|
||
0 0 0 60, one immature female; same data as
|
||
<typeStatus id="36540ADAFF98FF8DFB45F996FA84A220" box="[1256,1354,1575,1600]" pageId="4" pageNumber="329" type="holotype">holotype</typeStatus>
|
||
except coll.
|
||
<geoCoordinate id="8CDBD2BFFF98FF8DFF62F9FAFE98A204" box="[207,342,1611,1636]" direction="north" orientation="latitude" pageId="4" pageNumber="329" precision="1" value="15.06515">15.06515°N</geoCoordinate>
|
||
<geoCoordinate id="8CDBD2BFFF98FF8DFEF1F9FAFE3EA204" box="[348,496,1611,1636]" direction="east" orientation="longitude" pageId="4" pageNumber="329" precision="1" value="106.21394">106.21394°E</geoCoordinate>
|
||
,
|
||
<quantity id="2E17199DFF98FF8DFE56F9FAFD96A203" box="[507,600,1611,1636]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" pageId="4" pageNumber="329" unit="m" value="1245.0">1,245 m</quantity>
|
||
elev.,
|
||
<date id="9D5192B8FF98FF8DFD0CF9FDFC76A204" box="[673,952,1611,1636]" pageId="4" pageNumber="329" value="2010-07-28" valueMax="2010-08-01" valueMin="2010-07-28">28 July–01 August 2010</date>
|
||
.
|
||
<collectionCode id="8FFE2CBDFF98FF8DFC6EF9FDFBDAA203" box="[963,1044,1612,1635]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="329">FMNH</collectionCode>
|
||
258440, immature female;
|
||
<collectionCode id="8FFE2CBDFF98FF8DFAE6F9FDFA52A203" box="[1355,1436,1612,1635]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="329">FMNH</collectionCode>
|
||
258441,
|
||
<collectionCode id="8FFE2CBDFF98FF8DFF57F9C1FE85A2E7" box="[250,331,1648,1671]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="329">FMNH</collectionCode>
|
||
258530, two immature males, same data as
|
||
<typeStatus id="36540ADAFF98FF8DFCE9F9DEFC68A2E8" box="[836,934,1647,1672]" pageId="4" pageNumber="329" type="holotype">holotype</typeStatus>
|
||
except coll. near
|
||
<geoCoordinate id="8CDBD2BFFF98FF8DFBDCF9DEFB34A2E8" box="[1137,1274,1647,1672]" direction="north" orientation="latitude" pageId="4" pageNumber="329" precision="1" value="15.06528">15.06528°N</geoCoordinate>
|
||
<geoCoordinate id="8CDBD2BFFF98FF8DFAAFF9DEFA57A2E8" box="[1282,1433,1647,1672]" direction="east" orientation="longitude" pageId="4" pageNumber="329" precision="1" value="106.2175">106.21750°E</geoCoordinate>
|
||
,
|
||
<quantity id="2E17199DFF98FF8DFF3AF925FF3DA2CB" box="[151,243,1684,1708]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="329" unit="m" value="1200.0">1,200 m</quantity>
|
||
elev., coll.
|
||
<date id="9D5192B8FF98FF8DFED9F925FDB9A2CC" box="[372,631,1683,1708]" pageId="4" pageNumber="329" value="1999-09-22" valueMax="1999-09-23" valueMin="1999-09-22">22–23 September 1999</date>
|
||
by Bryan L. Stuart and Harold F. Heatwole.
|
||
</paragraph>
|
||
<paragraph id="E950B478FF98FF8DFF6AF906FA4BA2B0" blockId="4.[151,1436,1431,2032]" box="[199,1413,1719,1744]" pageId="4" pageNumber="329">
|
||
<emphasis id="DB9B686AFF98FF8DFF6AF906FE0EA2B0" bold="true" box="[199,448,1719,1744]" pageId="4" pageNumber="329">Referred Specimens:</emphasis>
|
||
<collectionCode id="8FFE2CBDFF98FF8DFE6AF909FDD8A2B0" box="[455,534,1720,1744]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="4" pageNumber="329">NCSM</collectionCode>
|
||
77944, larvae (
|
||
<emphasis id="DB9B686AFF98FF8DFD6BF909FD1AA2AF" box="[710,724,1720,1743]" italics="true" pageId="4" pageNumber="329">n</emphasis>
|
||
=13;
|
||
<figureCitation id="71D4A8FDFF98FF8DFCA3F906FC9FA2B0" box="[782,849,1719,1744]" captionStart="FIGURE 5" captionStartId="8.[151,250,1980,2002]" captionTargetBox="[187,1406,1147,1957]" captionTargetId="figure@8.[180,1415,1137,1963]" captionTargetPageId="8" captionText="FIGURE 5. Larvae of Limnonectes coffeatus sp. nov. (NCSM 77944, field tag BLS 13928) immediately prior to preservation (A), and inlateral (B), ventral (C) and oral (D) views in preservative." httpUri="https://zenodo.org/record/1158880/files/figure.png" pageId="4" pageNumber="329">Fig. 5</figureCitation>
|
||
), same data as
|
||
<typeStatus id="36540ADAFF98FF8DFC51F906FB90A2B0" box="[1020,1118,1719,1744]" pageId="4" pageNumber="329" type="holotype">holotype</typeStatus>
|
||
except coll.
|
||
<date id="9D5192B8FF98FF8DFB40F909FA4FA2B0" box="[1261,1409,1719,1744]" pageId="4" pageNumber="329" value="2010-07-30">30 July 2010</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1F5E7F3FF98FF8DFF6AF96AFE7AA35B" pageId="4" pageNumber="329" type="etymology">
|
||
<paragraph id="E950B478FF98FF8DFF6AF96AFE7AA35B" blockId="4.[151,1436,1431,2032]" pageId="4" pageNumber="329">
|
||
<emphasis id="DB9B686AFF98FF8DFF6AF96AFE9FA294" bold="true" box="[199,337,1755,1780]" pageId="4" pageNumber="329">Etymology:</emphasis>
|
||
The specific epithet refers to the coffee plant genus
|
||
<taxonomicName id="2EEFCFFBFF98FF8DFC1DF96DFC35A294" box="[944,1019,1756,1780]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="329" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="DB9B686AFF98FF8DFC1DF96DFC35A294" box="[944,1019,1756,1780]" italics="true" pageId="4" pageNumber="329">Coffea</emphasis>
|
||
</taxonomicName>
|
||
, in reference to the extensive coffee plantations that are encroaching into the natural habitat of the new species at upper elevations on the Bolaven Plateau of southern
|
||
<collectingCountry id="91F8F4E8FF98FF8DFEDBF895FE7EA35B" box="[374,432,1828,1851]" name="Laos" pageId="4" pageNumber="329">Laos</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1F5E7F3FF98FF8CFF6AF8F6FBF8A57D" lastPageId="5" lastPageNumber="330" pageId="4" pageNumber="329" type="diagnosis">
|
||
<paragraph id="E950B478FF98FF8CFF6AF8F6FBF8A57D" blockId="4.[151,1436,1431,2032]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="330" pageId="4" pageNumber="329">
|
||
<emphasis id="DB9B686AFF98FF8DFF6AF8F6FE8DA300" bold="true" box="[199,323,1863,1888]" pageId="4" pageNumber="329">Diagnosis:</emphasis>
|
||
Assigned to the genus
|
||
<taxonomicName id="2EEFCFFBFF98FF8DFDE2F8F9FD12A33F" authorityName="Fitzinger" authorityYear="1843" box="[591,732,1864,1887]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="329" phylum="Chordata" rank="genus">
|
||
<emphasis id="DB9B686AFF98FF8DFDE2F8F9FD12A33F" box="[591,732,1864,1887]" italics="true" pageId="4" pageNumber="329">Limnonectes</emphasis>
|
||
</taxonomicName>
|
||
on the basis of its inferred phylogenetic position (
|
||
<figureCitation id="71D4A8FDFF98FF8DFAB0F8F6FAADA300" box="[1309,1379,1863,1888]" captionStart="FIGURE 6" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[166,1414,530,1812]" captionTargetId="figure@9.[151,1436,517,1829]" captionTargetPageId="9" captionText="FIGURE 6. Fifty percent majority-rule consensus phylogram resulting from Bayesian analysis of 1,633 aligned characters of the mitochondrial 16S gene from frogs of the dicroglossid genera Fejervarya, Quasipaa, and Limnonectes. Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95, and no circle at nodes indicate Bayesian posterior probabilities <0.95. Numbers at terminal tips are GenBank accession numbers." httpUri="https://zenodo.org/record/1158882/files/figure.png" pageId="4" pageNumber="329">Fig. 6</figureCitation>
|
||
), the presence of fang-like odontoid processes on the lower jaw (
|
||
<bibRefCitation id="8D7EC989FF98FF8DFC98F8DDFBD2A3E4" author="Emerson" box="[821,1052,1900,1924]" pageId="4" pageNumber="329" refString="Emerson, S. B., Inger, R. F. & Iskandar, D. (2000) Molecular systematics and biogeography of the fanged frogs of Southeast Asia. Molecular Phylogenetics and Evolution, 16, 131 - 142. https: // doi. org / 10.1006 / mpev. 2000.0778" type="journal article" year="2000">
|
||
Emerson
|
||
<emphasis id="DB9B686AFF98FF8DFC0CF8DDFC14A3E4" box="[929,986,1900,1924]" italics="true" pageId="4" pageNumber="329">et al.</emphasis>
|
||
2000
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D7EC989FF98FF8DFB85F8DAFAD9A3E4" author="Lambertz" box="[1064,1303,1899,1924]" pageId="4" pageNumber="329" refString="Lambertz, M., Hartmann, T., Walsh, S., Geissler, P. & McLeod, D. S. (2014) Anatomy, histology, and systematic implications of the head ornamentation in the males of four species of Limnonectes (Anura: Dicroglossidae). Zoological Journal of the Linnean Society, 172, 117 - 132. https: // doi. org / 10.1111 / zoj. 12171" type="journal article" year="2014">
|
||
Lambertz
|
||
<emphasis id="DB9B686AFF98FF8DFB36F8DDFB1AA3E4" box="[1179,1236,1900,1924]" italics="true" pageId="4" pageNumber="329">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
), and males with hypertrophied heads (
|
||
<bibRefCitation id="8D7EC989FF98FF8DFE6EF83EFD7BA3C8" author="Lambertz" box="[451,693,1935,1960]" pageId="4" pageNumber="329" refString="Lambertz, M., Hartmann, T., Walsh, S., Geissler, P. & McLeod, D. S. (2014) Anatomy, histology, and systematic implications of the head ornamentation in the males of four species of Limnonectes (Anura: Dicroglossidae). Zoological Journal of the Linnean Society, 172, 117 - 132. https: // doi. org / 10.1111 / zoj. 12171" type="journal article" year="2014">
|
||
Lambertz
|
||
<emphasis id="DB9B686AFF98FF8DFD95F821FDBFA3C7" box="[568,625,1935,1959]" italics="true" pageId="4" pageNumber="329">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). A small-sized
|
||
<taxonomicName id="2EEFCFFBFF98FF8DFCC6F821FC36A3C7" authorityName="Fitzinger" authorityYear="1843" box="[875,1016,1936,1959]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="329" phylum="Chordata" rank="genus">
|
||
<emphasis id="DB9B686AFF98FF8DFCC6F821FC36A3C7" box="[875,1016,1936,1959]" italics="true" pageId="4" pageNumber="329">Limnonectes</emphasis>
|
||
</taxonomicName>
|
||
having the combination of the single available adult male with SVL 37.9, adult females with SVL 38.1–42.1; male lacking postorbital caruncle; male with hypertrophied head; odontoid processes on anterior margin of lower jaw, larger in male than in females; male with horizontal diameter of tympanum greater than that of eye, females with horizontal diameter of tympanum less than or equal to that of eye; tubercles on dorsum enlarged and rounded, but not arranged in rows; interdigital webbing of foot present; throat uniformly gray in male; tympanum uniformly colored, without distinct dark marking on upper half; flank without distinct spots; and ova with pigmented poles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="BD90E4F0FF99FF8CFF3AFB73FCAEA096" httpUri="https://zenodo.org/record/1158874/files/figure.png" pageId="5" pageNumber="330" targetBox="[156,1429,343,1189]" targetPageId="5">
|
||
<paragraph id="E950B478FF99FF8CFF3AFB73FCAEA096" blockId="5.[151,1436,1218,1270]" pageId="5" pageNumber="330">
|
||
<emphasis id="DB9B686AFF99FF8CFF3AFB73FEDDA0B7" bold="true" box="[151,275,1218,1240]" pageId="5" pageNumber="330">FIGURE 2.</emphasis>
|
||
Holotype male of
|
||
<taxonomicName id="2EEFCFFBFF99FF8CFE7EFB72FD78A0B8" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[467,694,1218,1240]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="330" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF99FF8CFE7EFB72FD78A0B8" box="[467,694,1218,1240]" italics="true" pageId="5" pageNumber="330">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF99FF8CFD11FB73FCC3A0B7" bold="true" box="[700,781,1218,1239]" pageId="5" pageNumber="330">
|
||
<taxonomicNameLabel id="C0A8D511FF99FF8CFD11FB73FCC3A0B7" box="[700,781,1218,1239]" pageId="5" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NCSM 77788, field tag BLS 13957) in dorsal (A), ventral (B), profile (C), and posterior (D) views immediately prior to preservation.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A1F5E7F3FF99FF80FF6AFAACFF26A5B0" lastPageId="9" lastPageNumber="334" pageId="5" pageNumber="330" type="description">
|
||
<paragraph id="E950B478FF99FF8CFF6AFAACFE55A21A" blockId="5.[151,1437,1309,2018]" pageId="5" pageNumber="330">
|
||
<emphasis id="DB9B686AFF99FF8CFF6AFAACFE29A156" bold="true" box="[199,487,1309,1334]" pageId="5" pageNumber="330">
|
||
Description of
|
||
<typeStatus id="36540ADAFF99FF8CFEDAFAACFE2CA156" box="[375,482,1309,1334]" pageId="5" pageNumber="330" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Habitus moderately stocky; body tapering to groin. Head broad and depressed; head width greater than head length. Snout obtusely pointed in dorsal view, round in profile, projecting well beyond lower jaw in profile; nostril dorsolateral, much closer to tip of snout than to eye, below canthus; internarial distance greater than interorbital distance; canthus rostralis indistinct, rounded, slightly constricted behind nostrils; lores concave, oblique; eye diameter 55% of snout length, interorbital distance greater than upper eyelid width; pineal ocellus visible; tympanum round, not elevated from side of head, annulus visible, tympanum diameter approximately 117% eye diameter and greater than distance between tympanum and eye; small, slit-like vocal sac openings on floor of mouth near lateral margin of tongue; vomerine teeth on two oblique ridges, closer to choanae than to each other; two large odontoid processes at front of mandible; median triangular protuberance at mandibular symphysis.
|
||
</paragraph>
|
||
<paragraph id="E950B478FF99FF8CFF6AF934FEA4A36A" blockId="5.[151,1437,1309,2018]" pageId="5" pageNumber="330">Forelimbs moderately robust. Fingers relatively slender, without webbing or skin flaps; tip of fingers rounded, weakly expanded into discs; relative finger lengths II = IV <I <III; distinct subarticular tubercles, one on Fingers I–II, two on Fingers III–IV; distinct thenar tubercle; two palmar tubercles in contact at base of Fingers II–IV; nuptial pad absent.</paragraph>
|
||
<paragraph id="E950B478FF99FF8FFF6AF8A4FB5BA4B5" blockId="5.[151,1437,1309,2018]" lastBlockId="6.[151,1436,151,357]" lastPageId="6" lastPageNumber="331" pageId="5" pageNumber="330">Hindlimbs moderately robust. Toes relatively slender; tips of toes rounded, expanded into small discs; relative toe lengths I <II <V <III <IV; webbing on Toe I to base of disc, on preaxial side of Toe II to distal margin of subarticular tubercle and continuing as a fringe to base of tip, on postaxial side of Toe II to base of tip, on preaxial side of Toe III to level of distal subarticular tubercle continuing as a fringe to base of tip, on postaxial side of Toe III to base of tip, on preaxial and postaxial sides of Toe IV to level of penultimate subarticular tubercle and continuing as a fringe to base of tip, and on Toe V to base of tip; moveable flap of skin on outer margins of Toes I and V; distinct fold on distal two-thirds of tarsus; distinct, elongate, oval, inner metatarsal tubercle, length approximately 59% distance between tip of toe I and tubercle; no outer metatarsal tubercle.</paragraph>
|
||
<paragraph id="E950B478FF9AFF8FFF6AFF6EFCE1A505" blockId="6.[151,1436,151,357]" pageId="6" pageNumber="331">Skin on dorsum shagreened with large round tubercles, not elongated or arranged in rows; tubercular ridges from posterior margin of eyelid to occiput forming “W”-shape; caruncle absent from interorbital region; distinct supratympanic ridge from posterior corner of eye to axilla; rictal gland absent; dorsolateral fold absent; skin on throat wrinkled, that on remaining ventral surfaces smooth.</paragraph>
|
||
<caption id="BD90E4F0FF9AFF8FFF3AFAB6FBF6A15C" httpUri="https://zenodo.org/record/1158876/files/figure.png" pageId="6" pageNumber="331" targetBox="[156,1429,414,1260]" targetPageId="6">
|
||
<paragraph id="E950B478FF9AFF8FFF3AFAB6FBF6A15C" blockId="6.[151,1436,1287,1340]" pageId="6" pageNumber="331">
|
||
<emphasis id="DB9B686AFF9AFF8FFF3AFAB6FEDDA17C" bold="true" box="[151,275,1287,1309]" pageId="6" pageNumber="331">FIGURE 3.</emphasis>
|
||
Paratype female of
|
||
<taxonomicName id="2EEFCFFBFF9AFF8FFE4FFAB8FD0AA17D" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[482,708,1287,1310]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="331" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF9AFF8FFE4FFAB8FD0AA17D" box="[482,708,1287,1310]" italics="true" pageId="6" pageNumber="331">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF9AFF8FFD66FAB9FCD2A17D" bold="true" box="[715,796,1288,1309]" pageId="6" pageNumber="331">
|
||
<taxonomicNameLabel id="C0A8D511FF9AFF8FFD66FAB9FCD2A17D" box="[715,796,1288,1309]" pageId="6" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NCSM 77787, field tag BLS 13934) in lateral (A) and dorsal (B) views in life, and in ventral (C) and lateral (D) views immediately prior to preservation.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E950B478FF9AFF8FFF6AFA33FAD8A24B" blockId="6.[151,1437,1410,2011]" pageId="6" pageNumber="331">
|
||
<emphasis id="DB9B686AFF9AFF8FFF6AFA33FE24A1FB" bold="true" box="[199,490,1410,1435]" pageId="6" pageNumber="331">
|
||
Color of
|
||
<typeStatus id="36540ADAFF9AFF8FFE9DFA33FE59A1FB" box="[304,407,1410,1435]" pageId="6" pageNumber="331" type="holotype">holotype</typeStatus>
|
||
in life:
|
||
</emphasis>
|
||
Dorsum brown with ochre wash; larger tubercles on dorsal surface of head and back orange-red with black margins, those on occiput forming “W”-shape; dark brown triangular marking between eyes with yellow anterior margin; dark brown alternating white-gray bands on upper and lower lips; dorsal surfaces of limbs with indistinct gray-brown bands; dorsal surfaces of thighs with yellowish-wash. Chin and throat gray; ventral surfaces of chest, belly and limbs yellow; ventral surfaces of hands and feet brown; iris bronze.
|
||
</paragraph>
|
||
<paragraph id="E950B478FF9AFF8FFF6AF987FA53A3BB" blockId="6.[151,1437,1410,2011]" pageId="6" pageNumber="331">
|
||
<emphasis id="DB9B686AFF9AFF8FFF6AF987FDBEA22F" bold="true" box="[199,624,1590,1615]" pageId="6" pageNumber="331">Description of larvae NCSM 77944:</emphasis>
|
||
Based on Gosner stage 28 individual in series of thirteen larvae (
|
||
<figureCitation id="71D4A8FDFF9AFF8FFAE4F987FA40A22F" box="[1353,1422,1590,1615]" captionStart="FIGURE 5" captionStartId="8.[151,250,1980,2002]" captionTargetBox="[187,1406,1147,1957]" captionTargetId="figure@8.[180,1415,1137,1963]" captionTargetPageId="8" captionText="FIGURE 5. Larvae of Limnonectes coffeatus sp. nov. (NCSM 77944, field tag BLS 13928) immediately prior to preservation (A), and inlateral (B), ventral (C) and oral (D) views in preservative." httpUri="https://zenodo.org/record/1158880/files/figure.png" pageId="6" pageNumber="331">Fig. 5</figureCitation>
|
||
). Body oval in dorsal view, slightly compressed dorsoventrally, maximum body width slightly anterior to level of spiracle. Nares dorsal, without raised rim. Eyes dorsolateral, not visible from below. Spiracular tube single, sinistral on left side, angled slightly dorsally, aperture near midline of side and projecting posteriorly, approximately midway between eye and end of body. Tail slender, tapering gradually in distal one-third to rounded tip, origins of dorsal and ventral fins at end of body, dorsal fin only slightly deeper than ventral fin. Oral disk ventral, subterminal, width about 36% maximum width of body. Anterior labium with single row of papillae on lateral margins; posterior labium with two staggered rows of papillae on lateral and posterior margins; papillae homogenous in length. Labial tooth row formula 2(2)/3(1). All posterior tooth rows subequal in length. Upper and lower jaw sheaths black with serrated margins, upper sheath without median convexity. No distinct cluster of glandules posterior to eye on dorsal surface of body, subequal to interorbital distance. Color in life brown with gold flecking on belly and distinct black spots on dorsal surface of body, caudal muscle, and dorsal and ventral tail fins.
|
||
</paragraph>
|
||
<paragraph id="E950B478FF9BFF8EFF3AFF29FDDCA4B4" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="332">
|
||
In preservative, brown faded to light brown, black faded to dark brown, gold flecking absent. Measurements of larvae are summarized in
|
||
<tableCitation id="A46D81C3FF9BFF8EFE15FF0DFDC3A4B4" box="[440,525,188,213]" captionStart="TABLE 3" captionStartId="7.[151,239,1472,1495]" captionText="TABLE 3. Measurements (mm) of 12larvae of Limnonectes coffeatus sp. nov. (NCSM 77944). Abbreviations defined in the text." pageId="7" pageNumber="332">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E950B478FF9BFF8EFF6AFF6EFD17A5B0" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFF6AFF6EFE8FA498" bold="true" box="[199,321,223,248]" pageId="7" pageNumber="332">Variation:</emphasis>
|
||
Females have narrower heads in dorsal view than
|
||
<typeStatus id="36540ADAFF9BFF8EFC2FFF6EFC2AA498" box="[898,996,223,248]" pageId="7" pageNumber="332" type="holotype">holotype</typeStatus>
|
||
male (
|
||
<figureCitation id="71D4A8FDFF9BFF8EFB9EFF6EFBB4A498" box="[1075,1146,223,248]" captionStart="FIGURE4" captionStartId="8.[151,250,1050,1072]" captionTargetBox="[409,1177,194,1029]" captionTargetId="figure@8.[409,1177,194,1029]" captionTargetPageId="8" captionText="FIGURE4. Paratypefemale (NCSM 77787, left)and holotype male (NCSM77788,right)of Limnonectes coffeatussp.nov. in dorsal (A) and ventral (B) views in preservative." httpUri="https://zenodo.org/record/1158878/files/figure.png" pageId="7" pageNumber="332">Fig. 4</figureCitation>
|
||
). Females have relatively smaller tympana than
|
||
<typeStatus id="36540ADAFF9BFF8EFE23FEB5FE3EA57D" box="[398,496,260,285]" pageId="7" pageNumber="332" type="holotype">holotype</typeStatus>
|
||
male, with the horizontal diameter of the tympanum less than or equal to that of the eye. Females have smaller odontoid processes at front of mandible than
|
||
<typeStatus id="36540ADAFF9BFF8EFC64FE96FBE5A520" box="[969,1067,295,320]" pageId="7" pageNumber="332" type="holotype">holotype</typeStatus>
|
||
male. Female NCSM 77787 has wider black margins on larger tubercles on dorsal surface of head and back than
|
||
<typeStatus id="36540ADAFF9BFF8EFB9AFEFDFB52A505" box="[1079,1180,332,357]" pageId="7" pageNumber="332" type="holotype">holotype</typeStatus>
|
||
, and with black rather than brown or gray-brown bands on interorbital region, lips, and dorsal surfaces of limbs. Females lack wrinkled skin and have paler coloration on throat than
|
||
<typeStatus id="36540ADAFF9BFF8EFD7FFE25FCFAA5CD" box="[722,820,404,429]" pageId="7" pageNumber="332" type="holotype">holotype</typeStatus>
|
||
male. Females contain ova with pigmented poles. Measurements of adults are summarized in
|
||
<tableCitation id="A46D81C3FF9BFF8EFDD2FE06FD1BA5B0" box="[639,725,439,464]" captionStart="TABLE 2" captionStartId="7.[151,239,508,531]" captionText="TABLE 2. Measurements (mm) of types of Limnonectes coffeatus sp. nov. and L. kohchangae. Abbreviations defined in the text." pageId="7" pageNumber="332">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="BD90E4F0FF9BFF8EFF3AFE4DFF23A653" pageId="7" pageNumber="332">
|
||
<paragraph id="E950B478FF9BFF8EFF3AFE4DFF23A653" blockId="7.[151,1436,507,563]" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFF3AFE4DFEC7A673" bold="true" box="[151,265,508,531]" pageId="7" pageNumber="332">TABLE 2.</emphasis>
|
||
Measurements (mm) of types of
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFDC1FE4DFC97A673" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[620,857,507,531]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF9BFF8EFDC1FE4DFC97A673" box="[620,857,507,531]" italics="true" pageId="7" pageNumber="332">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF9BFF8EFCCDFE4DFC78A673" bold="true" box="[864,950,508,531]" pageId="7" pageNumber="332">
|
||
<taxonomicNameLabel id="C0A8D511FF9BFF8EFCCDFE4DFC78A673" box="[864,950,508,531]" pageId="7" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="DB9B686AFF9BFF8EFC44FE4DFB43A673" box="[1001,1165,507,531]" italics="true" pageId="7" pageNumber="332">
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFC44FE4DFB47A673" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[1001,1161,507,531]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="kohchangae">L. kohchangae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Abbreviations defined in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<table id="9BEF46D8FF9B0076FF32FDFFFA46A11F" box="[159,1416,590,1407]" gridcols="6" gridrows="21" pageId="7" pageNumber="332">
|
||
<paragraph id="E950B478FF9BFF8EFF32FDFEFABFA6A0" pageId="7" pageNumber="332">
|
||
<table id="9BEF46D8FF9B0076FF32FDFFFABFA6A0" box="[159,1393,590,704]" gridcols="6" gridrows="4" pageId="7" pageNumber="332">
|
||
<tr id="57DFB63AFF9B0076FF32FDFFFABFA605" box="[159,1393,590,613]" gridrow="0" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FDFFFEE7A605" box="[159,297,590,613]" gridcol="0" gridrow="0" pageId="7" pageNumber="332">Measurement</th>
|
||
<th id="140EDF46FF9B0076FEECFDFFFE17A605" box="[321,473,590,613]" gridcol="1" gridrow="0" pageId="7" pageNumber="332">
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFEECFDE1FE79A604" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[321,439,590,613]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="coffeatus">
|
||
<emphasis id="DB9B686AFF9BFF8EFEECFDE1FE79A604" box="[321,439,590,613]" italics="true" pageId="7" pageNumber="332">L. coffeatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="140EDF46FF9B0076FE59FDFFFD75A605" box="[500,699,590,613]" gridcol="2" gridrow="0" pageId="7" pageNumber="332">
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFE59FDE1FDA4A604" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[500,618,590,613]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF9BFF8EFE59FDE1FDA4A604" box="[500,618,590,613]" italics="true" pageId="7" pageNumber="332">L. coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF9BFF8EFDDDFDFEFD75A604" bold="true" box="[624,699,591,612]" pageId="7" pageNumber="332">
|
||
<taxonomicNameLabel id="C0A8D511FF9BFF8EFDDDFDFEFD75A604" box="[624,699,591,612]" pageId="7" pageNumber="332" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="140EDF46FF9B0076FD52FDFFFC5AA605" box="[767,916,590,613]" gridcol="3" gridrow="0" pageId="7" pageNumber="332">
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFD52FDE1FC5AA604" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[767,916,590,613]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF9BFF8EFD52FDE1FC5AA604" box="[767,916,590,613]" italics="true" pageId="7" pageNumber="332">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="140EDF46FF9B0076FC06FDFFFBA3A605" box="[939,1133,590,613]" gridcol="4" gridrow="0" pageId="7" pageNumber="332">
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFC06FDE1FBF1A604" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[939,1087,590,613]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF9BFF8EFC06FDE1FBF1A604" box="[939,1087,590,613]" italics="true" pageId="7" pageNumber="332">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="140EDF46FF9B0076FB02FDFFFABFA605" box="[1199,1393,590,613]" gridcol="5" gridrow="0" pageId="7" pageNumber="332">
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFB02FDE1FA8AA604" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[1199,1348,590,613]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF9BFF8EFB02FDE1FA8AA604" box="[1199,1348,590,613]" italics="true" pageId="7" pageNumber="332">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FDDCFABFA6E3" box="[159,1393,621,643]" gridrow="1" pageId="7" pageNumber="332" rowspan-0="1">
|
||
<td id="140EDF46FF9B0076FEECFDDCFE17A6E3" box="[321,473,621,643]" gridcol="1" gridrow="1" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFEECFDDCFE48A6E2" bold="true" box="[321,390,621,642]" pageId="7" pageNumber="332">sp.nov</emphasis>
|
||
</td>
|
||
<td id="140EDF46FF9B0076FE59FDDCFD75A6E3" box="[500,699,621,643]" gridcol="2" gridrow="1" pageId="7" pageNumber="332">Paratype Females</td>
|
||
<td id="140EDF46FF9B0076FD52FDDCFC5AA6E3" box="[767,916,621,643]" gridcol="3" gridrow="1" pageId="7" pageNumber="332">Lectotype</td>
|
||
<td id="140EDF46FF9B0076FC06FDDCFBA3A6E3" box="[939,1133,621,643]" gridcol="4" gridrow="1" pageId="7" pageNumber="332">Other Males</td>
|
||
<td id="140EDF46FF9B0076FB02FDDCFABFA6E3" box="[1199,1393,621,643]" gridcol="5" gridrow="1" pageId="7" pageNumber="332">Females</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FD3DFABFA6C2" box="[159,1393,652,674]" gridrow="2" pageId="7" pageNumber="332" rowspan-0="1">
|
||
<td id="140EDF46FF9B0076FEECFD3DFE17A6C2" box="[321,473,652,674]" gridcol="1" gridrow="2" pageId="7" pageNumber="332">Holotype Male</td>
|
||
<td id="140EDF46FF9B0076FE59FD3DFD75A6C2" box="[500,699,652,674]" gridcol="2" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFE59FD3CFDCEA6C2" box="[500,512,653,674]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=3
|
||
</td>
|
||
<td id="140EDF46FF9B0076FD52FD3DFC5AA6C2" box="[767,916,652,674]" gridcol="3" gridrow="2" pageId="7" pageNumber="332">Male BMNH</td>
|
||
<td id="140EDF46FF9B0076FC06FD3DFBA3A6C2" box="[939,1133,652,674]" gridcol="4" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFC06FD3CFC79A6C2" box="[939,951,653,674]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=8
|
||
</td>
|
||
<td id="140EDF46FF9B0076FB02FD3DFABFA6C2" box="[1199,1393,652,674]" gridcol="5" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFB02FD3CFB72A6C2" box="[1199,1212,653,674]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=4
|
||
</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FD1BFABFA6A0" box="[159,1393,682,704]" gridrow="3" pageId="7" pageNumber="332" rowspan-0="1">
|
||
<td id="140EDF46FF9B0076FEECFD1BFE17A6A0" box="[321,473,682,704]" gridcol="1" gridrow="3" pageId="7" pageNumber="332">NCSM 77788</td>
|
||
<td id="140EDF46FF9B0076FE59FD1BFD75A6A0" box="[500,699,682,704]" gridcol="2" gridrow="3" pageId="7" pageNumber="332">Range; Mean ± SD</td>
|
||
<td id="140EDF46FF9B0076FD52FD1BFC5AA6A0" box="[767,916,682,704]" gridcol="3" gridrow="3" pageId="7" pageNumber="332">1947.2.3.60</td>
|
||
<td id="140EDF46FF9B0076FC06FD1BFBA3A6A0" box="[939,1133,682,704]" gridcol="4" gridrow="3" pageId="7" pageNumber="332">Range; Mean ± SD</td>
|
||
<td id="140EDF46FF9B0076FB02FD1BFABFA6A0" box="[1199,1393,682,704]" gridcol="5" gridrow="3" pageId="7" pageNumber="332">Range; Mean ± SD</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="E950B478FF9BFF8EFF32FD65FAACA11F" pageId="7" pageNumber="332">
|
||
<tr id="57DFB63AFF9B0076FF32FD65FA46A68A" box="[159,1416,724,746]" gridrow="4" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FD65FEE7A68A" box="[159,297,724,746]" gridcol="0" gridrow="4" pageId="7" pageNumber="332">SVL</th>
|
||
<td id="140EDF46FF9B0076FEECFD65FE17A68A" box="[321,473,724,746]" gridcol="1" gridrow="4" pageId="7" pageNumber="332">37.9</td>
|
||
<td id="140EDF46FF9B0076FE59FD65FD03A68A" box="[500,717,724,746]" gridcol="2" gridrow="4" pageId="7" pageNumber="332">38.1–42.1; 40.5 ± 2.1</td>
|
||
<td id="140EDF46FF9B0076FD52FD65FC5AA68A" box="[767,916,724,746]" gridcol="3" gridrow="4" pageId="7" pageNumber="332">41.1</td>
|
||
<td id="140EDF46FF9B0076FC06FD65FB4AA68A" box="[939,1156,724,746]" gridcol="4" gridrow="4" pageId="7" pageNumber="332">37.6–46.4; 40.9 ± 2.9</td>
|
||
<td id="140EDF46FF9B0076FB02FD65FA46A68A" box="[1199,1416,724,746]" gridcol="5" gridrow="4" pageId="7" pageNumber="332">36.0–41.2; 38.1 ± 2.2</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FD4CFA46A773" box="[159,1416,765,787]" gridrow="5" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FD4CFEE7A773" box="[159,297,765,787]" gridcol="0" gridrow="5" pageId="7" pageNumber="332">HDL</th>
|
||
<td id="140EDF46FF9B0076FEECFD4CFE17A773" box="[321,473,765,787]" gridcol="1" gridrow="5" pageId="7" pageNumber="332">17.9</td>
|
||
<td id="140EDF46FF9B0076FE59FD4CFD03A773" box="[500,717,765,787]" gridcol="2" gridrow="5" pageId="7" pageNumber="332">15.8–17.7; 16.9 ± 1.0</td>
|
||
<td id="140EDF46FF9B0076FD52FD4CFC5AA773" box="[767,916,765,787]" gridcol="3" gridrow="5" pageId="7" pageNumber="332">19.8</td>
|
||
<td id="140EDF46FF9B0076FC06FD4CFB4AA773" box="[939,1156,765,787]" gridcol="4" gridrow="5" pageId="7" pageNumber="332">16.0–21.4; 18.9 ± 1.6</td>
|
||
<td id="140EDF46FF9B0076FB02FD4CFA46A773" box="[1199,1416,765,787]" gridcol="5" gridrow="5" pageId="7" pageNumber="332">14.4–16.8; 15.7 ± 0.9</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FC97FA46A75C" box="[159,1416,806,828]" gridrow="6" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FC97FEE7A75C" box="[159,297,806,828]" gridcol="0" gridrow="6" pageId="7" pageNumber="332">HDW</th>
|
||
<td id="140EDF46FF9B0076FEECFC97FE17A75C" box="[321,473,806,828]" gridcol="1" gridrow="6" pageId="7" pageNumber="332">18.6</td>
|
||
<td id="140EDF46FF9B0076FE59FC97FD03A75C" box="[500,717,806,828]" gridcol="2" gridrow="6" pageId="7" pageNumber="332">15.2–17.3; 16.6 ± 1.2</td>
|
||
<td id="140EDF46FF9B0076FD52FC97FC5AA75C" box="[767,916,806,828]" gridcol="3" gridrow="6" pageId="7" pageNumber="332">18.0</td>
|
||
<td id="140EDF46FF9B0076FC06FC97FB4AA75C" box="[939,1156,806,828]" gridcol="4" gridrow="6" pageId="7" pageNumber="332">17.6–23.2; 20.0 ± 1.8</td>
|
||
<td id="140EDF46FF9B0076FB02FC97FA46A75C" box="[1199,1416,806,828]" gridcol="5" gridrow="6" pageId="7" pageNumber="332">13.1–17.7; 15.9 ± 1.7</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FCE1FA46A706" box="[159,1416,848,870]" gridrow="7" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FCE1FEE7A706" box="[159,297,848,870]" gridcol="0" gridrow="7" pageId="7" pageNumber="332">SNT</th>
|
||
<td id="140EDF46FF9B0076FEECFCE1FE17A706" box="[321,473,848,870]" gridcol="1" gridrow="7" pageId="7" pageNumber="332">7.6</td>
|
||
<td id="140EDF46FF9B0076FE59FCE1FD03A706" box="[500,717,848,870]" gridcol="2" gridrow="7" pageId="7" pageNumber="332">5.9–7.3; 6.8 ± 0.8</td>
|
||
<td id="140EDF46FF9B0076FD52FCE1FC5AA706" box="[767,916,848,870]" gridcol="3" gridrow="7" pageId="7" pageNumber="332">6.7</td>
|
||
<td id="140EDF46FF9B0076FC06FCE1FB4AA706" box="[939,1156,848,870]" gridcol="4" gridrow="7" pageId="7" pageNumber="332">6.1–9.0; 7.4 ± 0.8</td>
|
||
<td id="140EDF46FF9B0076FB02FCE1FA46A706" box="[1199,1416,848,870]" gridcol="5" gridrow="7" pageId="7" pageNumber="332">5.6–6.9; 6.5 ± 0.5</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FCC8FA46A7EF" box="[159,1416,889,911]" gridrow="8" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FCC8FEE7A7EF" box="[159,297,889,911]" gridcol="0" gridrow="8" pageId="7" pageNumber="332">EYE</th>
|
||
<td id="140EDF46FF9B0076FEECFCC8FE17A7EF" box="[321,473,889,911]" gridcol="1" gridrow="8" pageId="7" pageNumber="332">4.2</td>
|
||
<td id="140EDF46FF9B0076FE59FCC8FD03A7EF" box="[500,717,889,911]" gridcol="2" gridrow="8" pageId="7" pageNumber="332">3.6–4.4; 4.1 ± 0.4</td>
|
||
<td id="140EDF46FF9B0076FD52FCC8FC5AA7EF" box="[767,916,889,911]" gridcol="3" gridrow="8" pageId="7" pageNumber="332">5.5</td>
|
||
<td id="140EDF46FF9B0076FC06FCC8FB4AA7EF" box="[939,1156,889,911]" gridcol="4" gridrow="8" pageId="7" pageNumber="332">4.3–5.7; 5.0 ± 0.4</td>
|
||
<td id="140EDF46FF9B0076FB02FCC8FA46A7EF" box="[1199,1416,889,911]" gridcol="5" gridrow="8" pageId="7" pageNumber="332">4.2–5.1; 4.7 ± 0.3</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FC13FA46A7D8" box="[159,1416,930,952]" gridrow="9" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FC13FEE7A7D8" box="[159,297,930,952]" gridcol="0" gridrow="9" pageId="7" pageNumber="332">IOD</th>
|
||
<td id="140EDF46FF9B0076FEECFC13FE17A7D8" box="[321,473,930,952]" gridcol="1" gridrow="9" pageId="7" pageNumber="332">3.8</td>
|
||
<td id="140EDF46FF9B0076FE59FC13FD03A7D8" box="[500,717,930,952]" gridcol="2" gridrow="9" pageId="7" pageNumber="332">3.2–3.3; 3.3 ± 0.1</td>
|
||
<td id="140EDF46FF9B0076FD52FC13FC5AA7D8" box="[767,916,930,952]" gridcol="3" gridrow="9" pageId="7" pageNumber="332">4.4</td>
|
||
<td id="140EDF46FF9B0076FC06FC13FB4AA7D8" box="[939,1156,930,952]" gridcol="4" gridrow="9" pageId="7" pageNumber="332">4.0–6.2; 5.1 ± 0.7</td>
|
||
<td id="140EDF46FF9B0076FB02FC13FA46A7D8" box="[1199,1416,930,952]" gridcol="5" gridrow="9" pageId="7" pageNumber="332">2.6–4.6; 3.7 ± 0.7</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FC7DFA46A782" box="[159,1416,972,994]" gridrow="10" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FC7DFEE7A782" box="[159,297,972,994]" gridcol="0" gridrow="10" pageId="7" pageNumber="332">TMP</th>
|
||
<td id="140EDF46FF9B0076FEECFC7DFE17A782" box="[321,473,972,994]" gridcol="1" gridrow="10" pageId="7" pageNumber="332">4.9</td>
|
||
<td id="140EDF46FF9B0076FE59FC7DFD03A782" box="[500,717,972,994]" gridcol="2" gridrow="10" pageId="7" pageNumber="332">3.5–4.3; 3.9 ± 0.4</td>
|
||
<td id="140EDF46FF9B0076FD52FC7DFC5AA782" box="[767,916,972,994]" gridcol="3" gridrow="10" pageId="7" pageNumber="332">5.3</td>
|
||
<td id="140EDF46FF9B0076FC06FC7DFB4AA782" box="[939,1156,972,994]" gridcol="4" gridrow="10" pageId="7" pageNumber="332">4.4–7.3; 5.6 ± 0.9</td>
|
||
<td id="140EDF46FF9B0076FB02FC7DFA46A782" box="[1199,1416,972,994]" gridcol="5" gridrow="10" pageId="7" pageNumber="332">2.8–3.8; 3.4 ± 0.4</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FC44FA46A06B" box="[159,1416,1013,1035]" gridrow="11" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FC44FEE7A06B" box="[159,297,1013,1035]" gridcol="0" gridrow="11" pageId="7" pageNumber="332">IND</th>
|
||
<td id="140EDF46FF9B0076FEECFC44FE17A06B" box="[321,473,1013,1035]" gridcol="1" gridrow="11" pageId="7" pageNumber="332">4.6</td>
|
||
<td id="140EDF46FF9B0076FE59FC44FD03A06B" box="[500,717,1013,1035]" gridcol="2" gridrow="11" pageId="7" pageNumber="332">4.3–4.6; 4.5 ± 0.2</td>
|
||
<td id="140EDF46FF9B0076FD52FC44FC5AA06B" box="[767,916,1013,1035]" gridcol="3" gridrow="11" pageId="7" pageNumber="332">4.8</td>
|
||
<td id="140EDF46FF9B0076FC06FC44FB4AA06B" box="[939,1156,1013,1035]" gridcol="4" gridrow="11" pageId="7" pageNumber="332">4.3–6.2; 5.3 ± 0.6</td>
|
||
<td id="140EDF46FF9B0076FB02FC44FA46A06B" box="[1199,1416,1013,1035]" gridcol="5" gridrow="11" pageId="7" pageNumber="332">3.6–5.2; 4.6 ± 0.6</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FBAFFA46A054" box="[159,1416,1054,1076]" gridrow="12" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FBAFFEE7A054" box="[159,297,1054,1076]" gridcol="0" gridrow="12" pageId="7" pageNumber="332">SHK</th>
|
||
<td id="140EDF46FF9B0076FEECFBAFFE17A054" box="[321,473,1054,1076]" gridcol="1" gridrow="12" pageId="7" pageNumber="332">20.2</td>
|
||
<td id="140EDF46FF9B0076FE59FBAFFD03A054" box="[500,717,1054,1076]" gridcol="2" gridrow="12" pageId="7" pageNumber="332">19.4–20.0; 19.7 ± 0.3</td>
|
||
<td id="140EDF46FF9B0076FD52FBAFFC5AA054" box="[767,916,1054,1076]" gridcol="3" gridrow="12" pageId="7" pageNumber="332">20.5</td>
|
||
<td id="140EDF46FF9B0076FC06FBAFFB4AA054" box="[939,1156,1054,1076]" gridcol="4" gridrow="12" pageId="7" pageNumber="332">17.6–22.1; 20.4 ± 1.3</td>
|
||
<td id="140EDF46FF9B0076FB02FBAFFA46A054" box="[1199,1416,1054,1076]" gridcol="5" gridrow="12" pageId="7" pageNumber="332">17.6–20.5; 19.5 ± 1.2</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FBF9FA46A03E" box="[159,1416,1096,1118]" gridrow="13" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FBF9FEE7A03E" box="[159,297,1096,1118]" gridcol="0" gridrow="13" pageId="7" pageNumber="332">TGH</th>
|
||
<td id="140EDF46FF9B0076FEECFBF9FE17A03E" box="[321,473,1096,1118]" gridcol="1" gridrow="13" pageId="7" pageNumber="332">20.3</td>
|
||
<td id="140EDF46FF9B0076FE59FBF9FD03A03E" box="[500,717,1096,1118]" gridcol="2" gridrow="13" pageId="7" pageNumber="332">19.4–21.0; 20.5 ± 0.9</td>
|
||
<td id="140EDF46FF9B0076FD52FBF9FC5AA03E" box="[767,916,1096,1118]" gridcol="3" gridrow="13" pageId="7" pageNumber="332">18.0</td>
|
||
<td id="140EDF46FF9B0076FC06FBF9FB4AA03E" box="[939,1156,1096,1118]" gridcol="4" gridrow="13" pageId="7" pageNumber="332">18.9–22.9; 20.9 ± 1.4</td>
|
||
<td id="140EDF46FF9B0076FB02FBF9FA46A03E" box="[1199,1416,1096,1118]" gridcol="5" gridrow="13" pageId="7" pageNumber="332">18.5–20.7; 20.0 ± 0.9</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FBC0FA46A0E7" box="[159,1416,1137,1159]" gridrow="14" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FBC0FEE7A0E7" box="[159,297,1137,1159]" gridcol="0" gridrow="14" pageId="7" pageNumber="332">LAL</th>
|
||
<td id="140EDF46FF9B0076FEECFBC0FE17A0E7" box="[321,473,1137,1159]" gridcol="1" gridrow="14" pageId="7" pageNumber="332">7.2</td>
|
||
<td id="140EDF46FF9B0076FE59FBC0FD03A0E7" box="[500,717,1137,1159]" gridcol="2" gridrow="14" pageId="7" pageNumber="332">7.9–8.4; 8.2 ± 0.3</td>
|
||
<td id="140EDF46FF9B0076FD52FBC0FC5AA0E7" box="[767,916,1137,1159]" gridcol="3" gridrow="14" pageId="7" pageNumber="332">n/a</td>
|
||
<td id="140EDF46FF9B0076FC06FBC0FB4AA0E7" box="[939,1156,1137,1159]" gridcol="4" gridrow="14" pageId="7" pageNumber="332">7.5–8.7; 8.0 ± 0.4</td>
|
||
<td id="140EDF46FF9B0076FB02FBC0FA46A0E7" box="[1199,1416,1137,1159]" gridcol="5" gridrow="14" pageId="7" pageNumber="332">6.7–8.0; 7.6 ± 0.6</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FB2BFA46A0D0" box="[159,1416,1178,1200]" gridrow="15" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FB2BFEE7A0D0" box="[159,297,1178,1200]" gridcol="0" gridrow="15" pageId="7" pageNumber="332">HND</th>
|
||
<td id="140EDF46FF9B0076FEECFB2BFE17A0D0" box="[321,473,1178,1200]" gridcol="1" gridrow="15" pageId="7" pageNumber="332">10.5</td>
|
||
<td id="140EDF46FF9B0076FE59FB2BFD03A0D0" box="[500,717,1178,1200]" gridcol="2" gridrow="15" pageId="7" pageNumber="332">10.0–11.0; 10.4 ± 0.5</td>
|
||
<td id="140EDF46FF9B0076FD52FB2BFC5AA0D0" box="[767,916,1178,1200]" gridcol="3" gridrow="15" pageId="7" pageNumber="332">n/a</td>
|
||
<td id="140EDF46FF9B0076FC06FB2BFB4AA0D0" box="[939,1156,1178,1200]" gridcol="4" gridrow="15" pageId="7" pageNumber="332">9.1–12.0; 10.6 ± 0.8</td>
|
||
<td id="140EDF46FF9B0076FB02FB2BFA46A0D0" box="[1199,1416,1178,1200]" gridcol="5" gridrow="15" pageId="7" pageNumber="332">8.5–10.9; 10.0 ± 0.9</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FB75FA46A0BA" box="[159,1416,1220,1242]" gridrow="16" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FB75FEE7A0BA" box="[159,297,1220,1242]" gridcol="0" gridrow="16" pageId="7" pageNumber="332">FTL</th>
|
||
<td id="140EDF46FF9B0076FEECFB75FE17A0BA" box="[321,473,1220,1242]" gridcol="1" gridrow="16" pageId="7" pageNumber="332">19.3</td>
|
||
<td id="140EDF46FF9B0076FE59FB75FD03A0BA" box="[500,717,1220,1242]" gridcol="2" gridrow="16" pageId="7" pageNumber="332">18.7–20.7; 19.9 ± 1.0</td>
|
||
<td id="140EDF46FF9B0076FD52FB75FC5AA0BA" box="[767,916,1220,1242]" gridcol="3" gridrow="16" pageId="7" pageNumber="332">n/a</td>
|
||
<td id="140EDF46FF9B0076FC06FB75FB4AA0BA" box="[939,1156,1220,1242]" gridcol="4" gridrow="16" pageId="7" pageNumber="332">17.5–22.4; 20.3 ± 1.5</td>
|
||
<td id="140EDF46FF9B0076FB02FB75FA46A0BA" box="[1199,1416,1220,1242]" gridcol="5" gridrow="16" pageId="7" pageNumber="332">16.7–20.1; 19.3 ± 1.4</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FB5CFA46A163" box="[159,1416,1261,1283]" gridrow="17" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FB5CFEE7A163" box="[159,297,1261,1283]" gridcol="0" gridrow="17" pageId="7" pageNumber="332">IML</th>
|
||
<td id="140EDF46FF9B0076FEECFB5CFE17A163" box="[321,473,1261,1283]" gridcol="1" gridrow="17" pageId="7" pageNumber="332">3.0</td>
|
||
<td id="140EDF46FF9B0076FE59FB5CFD03A163" box="[500,717,1261,1283]" gridcol="2" gridrow="17" pageId="7" pageNumber="332">2.5–2.6; 2.5 ± 0.0</td>
|
||
<td id="140EDF46FF9B0076FD52FB5CFC5AA163" box="[767,916,1261,1283]" gridcol="3" gridrow="17" pageId="7" pageNumber="332">n/a</td>
|
||
<td id="140EDF46FF9B0076FC06FB5CFB4AA163" box="[939,1156,1261,1283]" gridcol="4" gridrow="17" pageId="7" pageNumber="332">1.8–2.7; 2.3 ± 0.3</td>
|
||
<td id="140EDF46FF9B0076FB02FB5CFA46A163" box="[1199,1416,1261,1283]" gridcol="5" gridrow="17" pageId="7" pageNumber="332">1.8–3.1; 2.5 ± 0.4</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FAA7FA46A14C" box="[159,1416,1302,1324]" gridrow="18" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FAA7FEE7A14C" box="[159,297,1302,1324]" gridcol="0" gridrow="18" pageId="7" pageNumber="332">IMW</th>
|
||
<td id="140EDF46FF9B0076FEECFAA7FE17A14C" box="[321,473,1302,1324]" gridcol="1" gridrow="18" pageId="7" pageNumber="332">1.2</td>
|
||
<td id="140EDF46FF9B0076FE59FAA7FD03A14C" box="[500,717,1302,1324]" gridcol="2" gridrow="18" pageId="7" pageNumber="332">0.9–1.2; 1.1 ± 0.1</td>
|
||
<td id="140EDF46FF9B0076FD52FAA7FC5AA14C" box="[767,916,1302,1324]" gridcol="3" gridrow="18" pageId="7" pageNumber="332">n/a</td>
|
||
<td id="140EDF46FF9B0076FC06FAA7FB4AA14C" box="[939,1156,1302,1324]" gridcol="4" gridrow="18" pageId="7" pageNumber="332">0.8–1.1; 1.0 ± 0.1</td>
|
||
<td id="140EDF46FF9B0076FB02FAA7FA46A14C" box="[1199,1416,1302,1324]" gridcol="5" gridrow="18" pageId="7" pageNumber="332">0.7–1.2; 0.9 ± 0.2</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FAF1FA46A136" box="[159,1416,1344,1366]" gridrow="19" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FAF1FEE7A136" box="[159,297,1344,1366]" gridcol="0" gridrow="19" pageId="7" pageNumber="332">TMP:EYE</th>
|
||
<td id="140EDF46FF9B0076FEECFAF1FE17A136" box="[321,473,1344,1366]" gridcol="1" gridrow="19" pageId="7" pageNumber="332">1.2</td>
|
||
<td id="140EDF46FF9B0076FE59FAF1FD03A136" box="[500,717,1344,1366]" gridcol="2" gridrow="19" pageId="7" pageNumber="332">0.9–1.0; 1.0 ± 0.0</td>
|
||
<td id="140EDF46FF9B0076FD52FAF1FC5AA136" box="[767,916,1344,1366]" gridcol="3" gridrow="19" pageId="7" pageNumber="332">1.0</td>
|
||
<td id="140EDF46FF9B0076FC06FAF1FB4AA136" box="[939,1156,1344,1366]" gridcol="4" gridrow="19" pageId="7" pageNumber="332">1.0–1.4; 1.1 ± 0.2</td>
|
||
<td id="140EDF46FF9B0076FB02FAF1FA46A136" box="[1199,1416,1344,1366]" gridcol="5" gridrow="19" pageId="7" pageNumber="332">0.5–0.8; 0.7 ± 0.1</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32FAD8FA46A11F" box="[159,1416,1385,1407]" gridrow="20" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32FAD8FEE7A11F" box="[159,297,1385,1407]" gridcol="0" gridrow="20" pageId="7" pageNumber="332">TMP:SVL</th>
|
||
<td id="140EDF46FF9B0076FEECFAD8FE17A11F" box="[321,473,1385,1407]" gridcol="1" gridrow="20" pageId="7" pageNumber="332">0.1</td>
|
||
<td id="140EDF46FF9B0076FE59FAD8FD03A11F" box="[500,717,1385,1407]" gridcol="2" gridrow="20" pageId="7" pageNumber="332">0.1–0.1; 0.1 ± 0.0</td>
|
||
<td id="140EDF46FF9B0076FD52FAD8FC5AA11F" box="[767,916,1385,1407]" gridcol="3" gridrow="20" pageId="7" pageNumber="332">0.1</td>
|
||
<td id="140EDF46FF9B0076FC06FAD8FB4AA11F" box="[939,1156,1385,1407]" gridcol="4" gridrow="20" pageId="7" pageNumber="332">0.1–0.2; 0.1 ± 0.0</td>
|
||
<td id="140EDF46FF9B0076FB02FAD8FA46A11F" box="[1199,1416,1385,1407]" gridcol="5" gridrow="20" pageId="7" pageNumber="332">0.1–0.1; 0.1 ± 0.0</td>
|
||
</tr>
|
||
</paragraph>
|
||
</table>
|
||
<caption id="BD90E4F0FF9BFF8EFF3AFA71FF23A197" pageId="7" pageNumber="332">
|
||
<paragraph id="E950B478FF9BFF8EFF3AFA71FF23A197" blockId="7.[151,1436,1471,1527]" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFF3AFA71FEC6A1B7" bold="true" box="[151,264,1472,1495]" pageId="7" pageNumber="332">TABLE 3.</emphasis>
|
||
Measurements (mm) of 12 larvae of
|
||
<taxonomicName id="2EEFCFFBFF9BFF8EFD3EFA71FC4EA1B7" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[659,896,1471,1495]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="332" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF9BFF8EFD3EFA71FC4EA1B7" box="[659,896,1471,1495]" italics="true" pageId="7" pageNumber="332">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF9BFF8EFC2BFA71FC24A1B7" bold="true" box="[902,1002,1472,1495]" pageId="7" pageNumber="332">
|
||
<taxonomicNameLabel id="C0A8D511FF9BFF8EFC2BFA71FC15A1B7" box="[902,987,1472,1495]" pageId="7" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
</emphasis>
|
||
NCSM 77944). Abbreviations defined in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E950B478FF9BFF8EFF32F9A2FA9DA3DB" pageId="7" pageNumber="332">
|
||
<table id="9BEF46D8FF9B0076FF32F9A3FA91A3DB" box="[159,1375,1554,1979]" gridcols="9" gridrows="10" pageId="7" pageNumber="332">
|
||
<tr id="57DFB63AFF9B0076FF32F9A3FA91A248" box="[159,1375,1554,1576]" gridrow="0" pageId="7" pageNumber="332" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="140EDF46FF9B0076FF32F9A3FEE7A248" box="[159,297,1554,1576]" gridcol="0" gridrow="0" pageId="7" pageNumber="332">Measurement</th>
|
||
<th id="140EDF46FF9B0076FEF6F9A3FDFEA248" box="[347,560,1554,1576]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="7" pageNumber="332">Gosner stage</th>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F98DFA91A232" box="[159,1375,1596,1618]" gridrow="1" pageId="7" pageNumber="332" rowspan-0="1">
|
||
<td id="140EDF46FF9B0076FEF6F98DFE46A232" box="[347,392,1596,1618]" gridcol="1" gridrow="1" pageId="7" pageNumber="332">28</td>
|
||
<td id="140EDF46FF9B0076FE67F98DFDFEA232" box="[458,560,1596,1618]" gridcol="2" gridrow="1" pageId="7" pageNumber="332">31</td>
|
||
<td id="140EDF46FF9B0076FDE2F98DFDB5A232" box="[591,635,1596,1618]" gridcol="3" gridrow="1" pageId="7" pageNumber="332">34</td>
|
||
<td id="140EDF46FF9B0076FD16F98DFC5BA232" box="[699,917,1596,1618]" gridcol="4" gridrow="1" pageId="7" pageNumber="332">35</td>
|
||
<td id="140EDF46FF9B0076FC16F98DFBEFA232" box="[955,1057,1596,1618]" gridcol="5" gridrow="1" pageId="7" pageNumber="332">36</td>
|
||
<td id="140EDF46FF9B0076FBFCF98DFBB3A232" box="[1105,1149,1596,1618]" gridcol="6" gridrow="1" pageId="7" pageNumber="332">38</td>
|
||
<td id="140EDF46FF9B0076FB6FF98DFB21A232" box="[1218,1263,1596,1618]" gridcol="7" gridrow="1" pageId="7" pageNumber="332">39</td>
|
||
<td id="140EDF46FF9B0076FA9EF98DFA91A232" box="[1331,1375,1596,1618]" gridcol="8" gridrow="1" pageId="7" pageNumber="332">40</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F9D4FA91A2FA" box="[159,1375,1637,1690]" gridrow="2" pageId="7" pageNumber="332" rowspan-0="1">
|
||
<td id="140EDF46FF9B0076FEF6F9D4FE46A2FA" box="[347,392,1637,1690]" gridcol="1" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFEF1F9D7FEA6A21B" box="[348,360,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=1
|
||
</td>
|
||
<td id="140EDF46FF9B0076FE67F9D4FDFEA2FA" box="[458,560,1637,1690]" gridcol="2" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFE67F9D7FE19A21B" box="[458,471,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=2 Range
|
||
</td>
|
||
<td id="140EDF46FF9B0076FDE2F9D4FDB5A2FA" box="[591,635,1637,1690]" gridcol="3" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFDE2F9D7FD95A21B" box="[591,603,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=1
|
||
</td>
|
||
<td id="140EDF46FF9B0076FD16F9D4FC5BA2FA" box="[699,917,1637,1690]" gridcol="4" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFD11F9D7FD06A21B" box="[700,712,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=3 Range; Mean ± SD
|
||
</td>
|
||
<td id="140EDF46FF9B0076FC16F9D4FBEFA2FA" box="[955,1057,1637,1690]" gridcol="5" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFC11F9D7FC06A21B" box="[956,968,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=2 Range
|
||
</td>
|
||
<td id="140EDF46FF9B0076FBFCF9D4FBB3A2FA" box="[1105,1149,1637,1690]" gridcol="6" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFBFCF9D7FB93A21B" box="[1105,1117,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=1
|
||
</td>
|
||
<td id="140EDF46FF9B0076FB6FF9D4FB21A2FA" box="[1218,1263,1637,1690]" gridcol="7" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFB6EF9D7FB01A21B" box="[1219,1231,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=1
|
||
</td>
|
||
<td id="140EDF46FF9B0076FA9EF9D4FA91A2FA" box="[1331,1375,1637,1690]" gridcol="8" gridrow="2" pageId="7" pageNumber="332">
|
||
<emphasis id="DB9B686AFF9BFF8EFA99F9D7FA8FA21B" box="[1332,1345,1638,1659]" italics="true" pageId="7" pageNumber="332">n</emphasis>
|
||
=1
|
||
</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F91CFA91A2A3" box="[159,1375,1709,1731]" gridrow="3" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32F91CFEE7A2A3" box="[159,297,1709,1731]" gridcol="0" gridrow="3" pageId="7" pageNumber="332">BL</th>
|
||
<td id="140EDF46FF9B0076FEF6F91CFE46A2A3" box="[347,392,1709,1731]" gridcol="1" gridrow="3" pageId="7" pageNumber="332">12.1</td>
|
||
<td id="140EDF46FF9B0076FE67F91CFDFEA2A3" box="[458,560,1709,1731]" gridcol="2" gridrow="3" pageId="7" pageNumber="332">12.6–13.1</td>
|
||
<td id="140EDF46FF9B0076FDE2F91CFDB5A2A3" box="[591,635,1709,1731]" gridcol="3" gridrow="3" pageId="7" pageNumber="332">15.2</td>
|
||
<td id="140EDF46FF9B0076FD16F91CFC5BA2A3" box="[699,917,1709,1731]" gridcol="4" gridrow="3" pageId="7" pageNumber="332">11.9–12.3; 12.1 ± 0.2</td>
|
||
<td id="140EDF46FF9B0076FC16F91CFBEFA2A3" box="[955,1057,1709,1731]" gridcol="5" gridrow="3" pageId="7" pageNumber="332">12.4</td>
|
||
<td id="140EDF46FF9B0076FBFCF91CFBB3A2A3" box="[1105,1149,1709,1731]" gridcol="6" gridrow="3" pageId="7" pageNumber="332">13.0</td>
|
||
<td id="140EDF46FF9B0076FB6FF91CFB21A2A3" box="[1218,1263,1709,1731]" gridcol="7" gridrow="3" pageId="7" pageNumber="332">13.8</td>
|
||
<td id="140EDF46FF9B0076FA9EF91CFA91A2A3" box="[1331,1375,1709,1731]" gridcol="8" gridrow="3" pageId="7" pageNumber="332">12.9</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F967FA91A28C" box="[159,1375,1750,1772]" gridrow="4" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32F967FEE7A28C" box="[159,297,1750,1772]" gridcol="0" gridrow="4" pageId="7" pageNumber="332">TAL</th>
|
||
<td id="140EDF46FF9B0076FEF6F967FE46A28C" box="[347,392,1750,1772]" gridcol="1" gridrow="4" pageId="7" pageNumber="332">22.3</td>
|
||
<td id="140EDF46FF9B0076FE67F967FDFEA28C" box="[458,560,1750,1772]" gridcol="2" gridrow="4" pageId="7" pageNumber="332">24.5–25.0</td>
|
||
<td id="140EDF46FF9B0076FDE2F967FDB5A28C" box="[591,635,1750,1772]" gridcol="3" gridrow="4" pageId="7" pageNumber="332">28.9</td>
|
||
<td id="140EDF46FF9B0076FD16F967FC5BA28C" box="[699,917,1750,1772]" gridcol="4" gridrow="4" pageId="7" pageNumber="332">21.3–22.5; 21.9 ± 0.2</td>
|
||
<td id="140EDF46FF9B0076FC16F967FBEFA28C" box="[955,1057,1750,1772]" gridcol="5" gridrow="4" pageId="7" pageNumber="332">21.7–21.8</td>
|
||
<td id="140EDF46FF9B0076FBFCF967FBB3A28C" box="[1105,1149,1750,1772]" gridcol="6" gridrow="4" pageId="7" pageNumber="332">22.9</td>
|
||
<td id="140EDF46FF9B0076FB6FF967FB21A28C" box="[1218,1263,1750,1772]" gridcol="7" gridrow="4" pageId="7" pageNumber="332">24.6</td>
|
||
<td id="140EDF46FF9B0076FA9EF967FA91A28C" box="[1331,1375,1750,1772]" gridcol="8" gridrow="4" pageId="7" pageNumber="332">25.0</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F8B1FA91A376" box="[159,1375,1792,1814]" gridrow="5" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32F8B1FEE7A376" box="[159,297,1792,1814]" gridcol="0" gridrow="5" pageId="7" pageNumber="332">TMH</th>
|
||
<td id="140EDF46FF9B0076FEF6F8B1FE46A376" box="[347,392,1792,1814]" gridcol="1" gridrow="5" pageId="7" pageNumber="332">3.8</td>
|
||
<td id="140EDF46FF9B0076FE67F8B1FDFEA376" box="[458,560,1792,1814]" gridcol="2" gridrow="5" pageId="7" pageNumber="332">3.8–4.7</td>
|
||
<td id="140EDF46FF9B0076FDE2F8B1FDB5A376" box="[591,635,1792,1814]" gridcol="3" gridrow="5" pageId="7" pageNumber="332">6.8</td>
|
||
<td id="140EDF46FF9B0076FD16F8B1FC5BA376" box="[699,917,1792,1814]" gridcol="4" gridrow="5" pageId="7" pageNumber="332">3.6–3.8; 3.7 ± 0.1</td>
|
||
<td id="140EDF46FF9B0076FC16F8B1FBEFA376" box="[955,1057,1792,1814]" gridcol="5" gridrow="5" pageId="7" pageNumber="332">3.2–3.3</td>
|
||
<td id="140EDF46FF9B0076FBFCF8B1FBB3A376" box="[1105,1149,1792,1814]" gridcol="6" gridrow="5" pageId="7" pageNumber="332">3.8</td>
|
||
<td id="140EDF46FF9B0076FB6FF8B1FB21A376" box="[1218,1263,1792,1814]" gridcol="7" gridrow="5" pageId="7" pageNumber="332">5.4</td>
|
||
<td id="140EDF46FF9B0076FA9EF8B1FA91A376" box="[1331,1375,1792,1814]" gridcol="8" gridrow="5" pageId="7" pageNumber="332">4.9</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F898FA91A35F" box="[159,1375,1833,1855]" gridrow="6" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32F898FEE7A35F" box="[159,297,1833,1855]" gridcol="0" gridrow="6" pageId="7" pageNumber="332">MTH</th>
|
||
<td id="140EDF46FF9B0076FEF6F898FE46A35F" box="[347,392,1833,1855]" gridcol="1" gridrow="6" pageId="7" pageNumber="332">6.5</td>
|
||
<td id="140EDF46FF9B0076FE67F898FDFEA35F" box="[458,560,1833,1855]" gridcol="2" gridrow="6" pageId="7" pageNumber="332">6.5–7.8</td>
|
||
<td id="140EDF46FF9B0076FDE2F898FDB5A35F" box="[591,635,1833,1855]" gridcol="3" gridrow="6" pageId="7" pageNumber="332">8.7</td>
|
||
<td id="140EDF46FF9B0076FD16F898FC5BA35F" box="[699,917,1833,1855]" gridcol="4" gridrow="6" pageId="7" pageNumber="332">5.6–5.9; 5.7 ± 0.2</td>
|
||
<td id="140EDF46FF9B0076FC16F898FBEFA35F" box="[955,1057,1833,1855]" gridcol="5" gridrow="6" pageId="7" pageNumber="332">5.3–5.9</td>
|
||
<td id="140EDF46FF9B0076FBFCF898FBB3A35F" box="[1105,1149,1833,1855]" gridcol="6" gridrow="6" pageId="7" pageNumber="332">6.9</td>
|
||
<td id="140EDF46FF9B0076FB6FF898FB21A35F" box="[1218,1263,1833,1855]" gridcol="7" gridrow="6" pageId="7" pageNumber="332">5.9</td>
|
||
<td id="140EDF46FF9B0076FA9EF898FA91A35F" box="[1331,1375,1833,1855]" gridcol="8" gridrow="6" pageId="7" pageNumber="332">6.7</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F8E3FA91A308" box="[159,1375,1874,1896]" gridrow="7" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32F8E3FEE7A308" box="[159,297,1874,1896]" gridcol="0" gridrow="7" pageId="7" pageNumber="332">TMW</th>
|
||
<td id="140EDF46FF9B0076FEF6F8E3FE46A308" box="[347,392,1874,1896]" gridcol="1" gridrow="7" pageId="7" pageNumber="332">3.0</td>
|
||
<td id="140EDF46FF9B0076FE67F8E3FDFEA308" box="[458,560,1874,1896]" gridcol="2" gridrow="7" pageId="7" pageNumber="332">2.7–3.4</td>
|
||
<td id="140EDF46FF9B0076FDE2F8E3FDB5A308" box="[591,635,1874,1896]" gridcol="3" gridrow="7" pageId="7" pageNumber="332">4.0</td>
|
||
<td id="140EDF46FF9B0076FD16F8E3FC5BA308" box="[699,917,1874,1896]" gridcol="4" gridrow="7" pageId="7" pageNumber="332">2.5–2.7; 2.6 ± 0.1</td>
|
||
<td id="140EDF46FF9B0076FC16F8E3FBEFA308" box="[955,1057,1874,1896]" gridcol="5" gridrow="7" pageId="7" pageNumber="332">2.6–2.7</td>
|
||
<td id="140EDF46FF9B0076FBFCF8E3FBB3A308" box="[1105,1149,1874,1896]" gridcol="6" gridrow="7" pageId="7" pageNumber="332">3.0</td>
|
||
<td id="140EDF46FF9B0076FB6FF8E3FB21A308" box="[1218,1263,1874,1896]" gridcol="7" gridrow="7" pageId="7" pageNumber="332">2.9</td>
|
||
<td id="140EDF46FF9B0076FA9EF8E3FA91A308" box="[1331,1375,1874,1896]" gridcol="8" gridrow="7" pageId="7" pageNumber="332">3.1</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F8CDFA91A3F2" box="[159,1375,1916,1938]" gridrow="8" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32F8CDFEE7A3F2" box="[159,297,1916,1938]" gridcol="0" gridrow="8" pageId="7" pageNumber="332">IOD</th>
|
||
<td id="140EDF46FF9B0076FEF6F8CDFE46A3F2" box="[347,392,1916,1938]" gridcol="1" gridrow="8" pageId="7" pageNumber="332">4.4</td>
|
||
<td id="140EDF46FF9B0076FE67F8CDFDFEA3F2" box="[458,560,1916,1938]" gridcol="2" gridrow="8" pageId="7" pageNumber="332">4.5–4.8</td>
|
||
<td id="140EDF46FF9B0076FDE2F8CDFDB5A3F2" box="[591,635,1916,1938]" gridcol="3" gridrow="8" pageId="7" pageNumber="332">5.7</td>
|
||
<td id="140EDF46FF9B0076FD16F8CDFC5BA3F2" box="[699,917,1916,1938]" gridcol="4" gridrow="8" pageId="7" pageNumber="332">3.5–3.7; 3.6 ± 0.1</td>
|
||
<td id="140EDF46FF9B0076FC16F8CDFBEFA3F2" box="[955,1057,1916,1938]" gridcol="5" gridrow="8" pageId="7" pageNumber="332">3.4–3.9</td>
|
||
<td id="140EDF46FF9B0076FBFCF8CDFBB3A3F2" box="[1105,1149,1916,1938]" gridcol="6" gridrow="8" pageId="7" pageNumber="332">4.1</td>
|
||
<td id="140EDF46FF9B0076FB6FF8CDFB21A3F2" box="[1218,1263,1916,1938]" gridcol="7" gridrow="8" pageId="7" pageNumber="332">4.1</td>
|
||
<td id="140EDF46FF9B0076FA9EF8CDFA91A3F2" box="[1331,1375,1916,1938]" gridcol="8" gridrow="8" pageId="7" pageNumber="332">3.7</td>
|
||
</tr>
|
||
<tr id="57DFB63AFF9B0076FF32F814FA91A3DB" box="[159,1375,1957,1979]" gridrow="9" pageId="7" pageNumber="332">
|
||
<th id="140EDF46FF9B0076FF32F814FEE7A3DB" box="[159,297,1957,1979]" gridcol="0" gridrow="9" pageId="7" pageNumber="332">IND</th>
|
||
<td id="140EDF46FF9B0076FEF6F814FE46A3DB" box="[347,392,1957,1979]" gridcol="1" gridrow="9" pageId="7" pageNumber="332">2.6</td>
|
||
<td id="140EDF46FF9B0076FE67F814FDFEA3DB" box="[458,560,1957,1979]" gridcol="2" gridrow="9" pageId="7" pageNumber="332">2.9–3.0</td>
|
||
<td id="140EDF46FF9B0076FDE2F814FDB5A3DB" box="[591,635,1957,1979]" gridcol="3" gridrow="9" pageId="7" pageNumber="332">3.2</td>
|
||
<td id="140EDF46FF9B0076FD16F814FC5BA3DB" box="[699,917,1957,1979]" gridcol="4" gridrow="9" pageId="7" pageNumber="332">2.2–2.8; 2.4 ± 0.3</td>
|
||
<td id="140EDF46FF9B0076FC16F814FBEFA3DB" box="[955,1057,1957,1979]" gridcol="5" gridrow="9" pageId="7" pageNumber="332">2.2–2.3</td>
|
||
<td id="140EDF46FF9B0076FBFCF814FBB3A3DB" box="[1105,1149,1957,1979]" gridcol="6" gridrow="9" pageId="7" pageNumber="332">2.3</td>
|
||
<td id="140EDF46FF9B0076FB6FF814FB21A3DB" box="[1218,1263,1957,1979]" gridcol="7" gridrow="9" pageId="7" pageNumber="332">1.9</td>
|
||
<td id="140EDF46FF9B0076FA9EF814FA91A3DB" box="[1331,1375,1957,1979]" gridcol="8" gridrow="9" pageId="7" pageNumber="332">2.2</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="BD90E4F0FF94FF81FF3AFBABFD4CA02F" httpUri="https://zenodo.org/record/1158878/files/figure.png" pageId="8" pageNumber="333" targetBox="[409,1177,194,1029]" targetPageId="8">
|
||
<paragraph id="E950B478FF94FF81FF3AFBABFD4CA02F" blockId="8.[151,1436,1050,1103]" pageId="8" pageNumber="333">
|
||
<emphasis id="DB9B686AFF94FF81FF3AFBABFEDCA04F" bold="true" box="[151,274,1050,1072]" pageId="8" pageNumber="333">FIGURE 4.</emphasis>
|
||
Paratype female (NCSM 77787, left) and holotype male (NCSM 77788, right) of
|
||
<taxonomicName id="2EEFCFFBFF94FF81FBE6FBAAFAE3A050" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[1099,1325,1050,1072]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="333" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF94FF81FBE6FBAAFAE3A050" box="[1099,1325,1050,1072]" italics="true" pageId="8" pageNumber="333">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF94FF81FA9FFBAAFA4CA050" bold="true" box="[1330,1410,1051,1072]" pageId="8" pageNumber="333">
|
||
<taxonomicNameLabel id="C0A8D511FF94FF81FA9FFBAAFA4CA050" box="[1330,1410,1051,1072]" pageId="8" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in dorsal (A) and ventral (B) views in preservative.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BD90E4F0FF94FF81FF3AF80DFC9AA391" httpUri="https://zenodo.org/record/1158880/files/figure.png" pageId="8" pageNumber="333" targetBox="[187,1406,1147,1957]" targetPageId="8">
|
||
<paragraph id="E950B478FF94FF81FF3AF80DFC9AA391" blockId="8.[151,1436,1980,2033]" pageId="8" pageNumber="333">
|
||
<emphasis id="DB9B686AFF94FF81FF3AF80DFEDDA3B2" bold="true" box="[151,275,1980,2002]" pageId="8" pageNumber="333">FIGURE 5.</emphasis>
|
||
Larvae of
|
||
<taxonomicName id="2EEFCFFBFF94FF81FE2EF80CFDABA3B2" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[387,613,1980,2002]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="333" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF94FF81FE2EF80CFDABA3B2" box="[387,613,1980,2002]" italics="true" pageId="8" pageNumber="333">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF94FF81FDC1F80CFD73A3B2" bold="true" box="[620,701,1981,2002]" pageId="8" pageNumber="333">
|
||
<taxonomicNameLabel id="C0A8D511FF94FF81FDC1F80CFD73A3B2" box="[620,701,1981,2002]" pageId="8" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NCSM 77944, field tag BLS 13928) immediately prior to preservation (A), and in lateral (B), ventral (C) and oral (D) views in preservative.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E950B478FF95FF80FF6AFF26FF26A5B0" blockId="9.[151,1436,151,464]" pageId="9" pageNumber="334">
|
||
<emphasis id="DB9B686AFF95FF80FF6AFF26FE86A4D0" bold="true" box="[199,328,151,176]" pageId="9" pageNumber="334">Molecules:</emphasis>
|
||
The aligned dataset contained 1,633 characters. The standard deviation of split frequencies was 0.003805 among the four Bayesian runs, and all parameters had an Estimated Sample Size (ESS) ≥267. The new species was recovered as a member of a clade containing
|
||
<emphasis id="DB9B686AFF95FF80FC8CFF50FAA2A498" box="[801,1388,224,248]" italics="true" pageId="9" pageNumber="334">
|
||
<taxonomicName id="2EEFCFFBFF95FF80FC8CFF50FC45A498" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[801,907,224,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="doriae">L. doriae</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EEFCFFBFF95FF80FC3AFF50FBE2A498" baseAuthorityName="Stoliczka" baseAuthorityYear="1873" box="[919,1068,224,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="plicatellus">L. plicatellus</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EEFCFFBFF95FF80FB95FF50FB15A498" baseAuthorityName="Stoliczka" baseAuthorityYear="1870" box="[1080,1243,224,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="hascheanus">L. hascheanus</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EEFCFFBFF95FF80FB45FF50FAA6A498" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[1256,1384,224,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="limborgi">L. limborgi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="2EEFCFFBFF95FF80FF3AFEB4FE9DA57C" baseAuthorityName="Boulenger" baseAuthorityYear="1917" box="[151,339,260,284]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="macrognathus">
|
||
<emphasis id="DB9B686AFF95FF80FF3AFEB4FE9DA57C" box="[151,339,260,284]" italics="true" pageId="9" pageNumber="334">L. macrognathus</emphasis>
|
||
</taxonomicName>
|
||
, and this clade was sister to
|
||
<taxonomicName id="2EEFCFFBFF95FF80FD3EFEB4FCF9A57C" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[659,823,260,284]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF95FF80FD3EFEB4FCF9A57C" box="[659,823,260,284]" italics="true" pageId="9" pageNumber="334">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="71D4A8FDFF95FF80FCEBFEB5FC44A57D" box="[838,906,260,285]" captionStart="FIGURE 6" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[166,1414,530,1812]" captionTargetId="figure@9.[151,1436,517,1829]" captionTargetPageId="9" captionText="FIGURE 6. Fifty percent majority-rule consensus phylogram resulting from Bayesian analysis of 1,633 aligned characters of the mitochondrial 16S gene from frogs of the dicroglossid genera Fejervarya, Quasipaa, and Limnonectes. Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95, and no circle at nodes indicate Bayesian posterior probabilities <0.95. Numbers at terminal tips are GenBank accession numbers." httpUri="https://zenodo.org/record/1158882/files/figure.png" pageId="9" pageNumber="334">Fig. 6</figureCitation>
|
||
). The exact sister taxon relationship of the new species was unresolved, but the new species was not recovered to be the sister species to
|
||
<taxonomicName id="2EEFCFFBFF95FF80FB3CFE98FAF9A520" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[1169,1335,296,320]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF95FF80FB3CFE98FAF9A520" box="[1169,1335,296,320]" italics="true" pageId="9" pageNumber="334">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="71D4A8FDFF95FF80FAE5FE96FA43A520" box="[1352,1421,295,320]" captionStart="FIGURE 6" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[166,1414,530,1812]" captionTargetId="figure@9.[151,1436,517,1829]" captionTargetPageId="9" captionText="FIGURE 6. Fifty percent majority-rule consensus phylogram resulting from Bayesian analysis of 1,633 aligned characters of the mitochondrial 16S gene from frogs of the dicroglossid genera Fejervarya, Quasipaa, and Limnonectes. Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95, and no circle at nodes indicate Bayesian posterior probabilities <0.95. Numbers at terminal tips are GenBank accession numbers." httpUri="https://zenodo.org/record/1158882/files/figure.png" pageId="9" pageNumber="334">Fig. 6</figureCitation>
|
||
). The
|
||
<typeStatus id="36540ADAFF95FF80FF61FEFDFEFFA505" box="[204,305,332,357]" pageId="9" pageNumber="334" type="holotype">holotype</typeStatus>
|
||
, six
|
||
<typeStatus id="36540ADAFF95FF80FEC7FEFCFE16A505" box="[362,472,333,357]" pageId="9" pageNumber="334" type="paratype">paratypes</typeStatus>
|
||
, and referred larvae of
|
||
<taxonomicName id="2EEFCFFBFF95FF80FD47FEFCFCBEA504" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[746,880,332,356]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF95FF80FD47FEFCFCBEA504" box="[746,880,332,356]" italics="true" pageId="9" pageNumber="334">L. coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF95FF80FCD7FEFDFC18A505" bold="true" box="[890,982,332,357]" pageId="9" pageNumber="334">
|
||
<taxonomicNameLabel id="C0A8D511FF95FF80FCD7FEFDFC18A505" box="[890,982,332,357]" pageId="9" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are identical in the 16S gene fragment (differing only by a single insertion-deletion event), but have an uncorrected pairwise divergence of 10.13–11.72% from
|
||
<taxonomicName id="2EEFCFFBFF95FF80FF79FE24FEB9A5CC" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[212,375,404,428]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF95FF80FF79FE24FEB9A5CC" box="[212,375,404,428]" italics="true" pageId="9" pageNumber="334">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="DB9B686AFF95FF80FE2AFE24FE5BA5CC" box="[391,405,405,428]" italics="true" pageId="9" pageNumber="334">n</emphasis>
|
||
=11), the species to which it is phenotypically most similar (but not phylogenetically related;
|
||
<figureCitation id="71D4A8FDFF95FF80FF3AFE06FF14A5B0" box="[151,218,439,464]" captionStart="FIGURE 6" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[166,1414,530,1812]" captionTargetId="figure@9.[151,1436,517,1829]" captionTargetPageId="9" captionText="FIGURE 6. Fifty percent majority-rule consensus phylogram resulting from Bayesian analysis of 1,633 aligned characters of the mitochondrial 16S gene from frogs of the dicroglossid genera Fejervarya, Quasipaa, and Limnonectes. Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95, and no circle at nodes indicate Bayesian posterior probabilities <0.95. Numbers at terminal tips are GenBank accession numbers." httpUri="https://zenodo.org/record/1158882/files/figure.png" pageId="9" pageNumber="334">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="BD90E4F0FF95FF80FF3AF88AFF3BA3AB" httpUri="https://zenodo.org/record/1158882/files/figure.png" pageId="9" pageNumber="334" targetBox="[166,1414,530,1812]" targetPageId="9">
|
||
<paragraph id="E950B478FF95FF80FF3AF88AFF3BA3AB" blockId="9.[151,1436,1851,1995]" pageId="9" pageNumber="334">
|
||
<emphasis id="DB9B686AFF95FF80FF3AF88AFEDDA330" bold="true" box="[151,275,1851,1873]" pageId="9" pageNumber="334">FIGURE 6.</emphasis>
|
||
Fifty percent majority-rule consensus phylogram resulting from Bayesian analysis of 1,633 aligned characters of the mitochondrial 16S gene from frogs of the dicroglossid genera
|
||
<taxonomicName id="2EEFCFFBFF95FF80FCE1F8EBFC73A30F" authorityName="Bolkay" authorityYear="1915" box="[844,957,1882,1903]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="genus">
|
||
<emphasis id="DB9B686AFF95FF80FCE1F8EBFC73A30F" box="[844,957,1882,1903]" italics="true" pageId="9" pageNumber="334">Fejervarya</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EEFCFFBFF95FF80FC61F8EBFBE1A30F" authorityName="Dubois" authorityYear="1992" box="[972,1071,1882,1903]" class="Amphibia" family="Dicroglossidae" genus="Quasipaa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="genus">
|
||
<emphasis id="DB9B686AFF95FF80FC61F8EBFBE1A30F" box="[972,1071,1882,1903]" italics="true" pageId="9" pageNumber="334">Quasipaa</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2EEFCFFBFF95FF80FBC1F8EBFB22A30F" authorityName="Fitzinger" authorityYear="1843" box="[1132,1260,1882,1903]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="334" phylum="Chordata" rank="genus">
|
||
<emphasis id="DB9B686AFF95FF80FBC1F8EBFB22A30F" box="[1132,1260,1882,1903]" italics="true" pageId="9" pageNumber="334">Limnonectes</emphasis>
|
||
</taxonomicName>
|
||
. Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95, and no circle at nodes indicate Bayesian posterior probabilities <0.95. Numbers at terminal tips are GenBank accession numbers.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BD90E4F0FF96FF83FF3AFB64FDA4A16A" httpUri="https://zenodo.org/record/1158884/files/figure.png" pageId="10" pageNumber="335" targetBox="[411,1178,187,1207]" targetPageId="10">
|
||
<paragraph id="E950B478FF96FF83FF3AFB64FDA4A16A" blockId="10.[151,1436,1237,1290]" pageId="10" pageNumber="335">
|
||
<emphasis id="DB9B686AFF96FF83FF3AFB64FEDAA08B" bold="true" box="[151,276,1237,1259]" pageId="10" pageNumber="335">FIGURE 7.</emphasis>
|
||
<taxonomicName id="2EEFCFFBFF96FF83FEB0FB67FDCFA08B" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[285,513,1237,1259]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="335" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF96FF83FEB0FB67FDCFA08B" box="[285,513,1237,1259]" italics="true" pageId="10" pageNumber="335">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF96FF83FDA4FB67FD95A08B" bold="true" box="[521,603,1238,1259]" pageId="10" pageNumber="335">
|
||
<taxonomicNameLabel id="C0A8D511FF96FF83FDA4FB67FD95A08B" box="[521,603,1238,1259]" pageId="10" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
habitat of adults (A) and larvae (B) at the type locality on the Bolaven Plateau, Pakxong District, Champasak Province, Laos.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A1F5E7F3FF96FF82FF6AFA80FAB2A57D" lastPageId="11" lastPageNumber="336" pageId="10" pageNumber="335" type="distribution">
|
||
<paragraph id="E950B478FF96FF83FF6AFA80FB04A20A" blockId="10.[151,1437,1329,2038]" pageId="10" pageNumber="335">
|
||
<emphasis id="DB9B686AFF96FF83FF6AFA80FCC4A12A" bold="true" box="[199,778,1329,1354]" pageId="10" pageNumber="335">Distribution, natural history, and conservation:</emphasis>
|
||
<taxonomicName id="2EEFCFFBFF96FF83FCB9FA83FBDCA129" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[788,1042,1329,1353]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="335" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF96FF83FCB9FA83FBDCA129" box="[788,1042,1329,1353]" italics="true" pageId="10" pageNumber="335">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF96FF83FBB1FA80FBB7A12A" bold="true" box="[1052,1145,1329,1354]" pageId="10" pageNumber="335">
|
||
<taxonomicNameLabel id="C0A8D511FF96FF83FBB1FA80FBB7A12A" box="[1052,1145,1329,1354]" pageId="10" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is only known from wet evergreen forest (
|
||
<figureCitation id="71D4A8FDFF96FF83FEC9FAE4FE62A10E" box="[356,428,1365,1390]" captionStart="FIGURE 7" captionStartId="10.[151,250,1237,1259]" captionTargetBox="[411,1178,187,1207]" captionTargetId="figure@10.[399,1188,177,1216]" captionTargetPageId="10" captionText="FIGURE 7. Limnonectes coffeatus sp. nov. habitat of adults (A) and larvae (B) at the type locality on the Bolaven Plateau, Pakxong District, Champasak Province, Laos." httpUri="https://zenodo.org/record/1158884/files/figure.png" pageId="10" pageNumber="335">Fig. 7</figureCitation>
|
||
) near to the top (≥
|
||
<quantity id="2E17199DFF96FF83FD3FFAE7FD3AA10D" box="[658,756,1366,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="335" unit="m" value="1200.0">1,200 m</quantity>
|
||
elev.) of the Bolaven Plateau in Dong Hua Sao National Protected Area, Pakxong District,
|
||
<collectingRegion id="2B2B7A9AFF96FF83FDB9FAC8FCCAA1F2" box="[532,772,1401,1426]" country="Laos" name="Champasak" pageId="10" pageNumber="335">Champasak Province</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="91F8F4E8FF96FF83FCD8FACBFC65A1F1" box="[885,939,1402,1425]" name="Laos" pageId="10" pageNumber="335">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="71D4A8FDFF96FF83FC17FAC8FC35A1F2" box="[954,1019,1401,1426]" captionStart="FIGURE 1" captionStartId="4.[151,250,1127,1149]" captionTargetBox="[153,1435,195,1102]" captionTargetId="figure@4.[151,1435,195,1103]" captionTargetPageId="4" captionText="FIGURE 1. Map illustrating the type locality (white star) of Limnonectes coffeatus sp. nov. in Champasak Province, Laos; the type locality (black star) of L. kohchangae in Trat Province, Koh Chang Island, Thailand; and localities of studied specimens (black circles) of L.kohchangae in Pursat and Kampong Speu Provinces,Cambodia." httpUri="https://zenodo.org/record/1158872/files/figure.png" pageId="10" pageNumber="335">Fig. 1</figureCitation>
|
||
). Most of the known specimens were collected at night on the forest floor within
|
||
<quantity id="2E17199DFF96FF83FD38FA2FFD0DA1D5" box="[661,707,1438,1462]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="335" unit="m" value="3.0">3 m</quantity>
|
||
of the bank, or in shallow water, of small, rocky streams. The immature female (NUOL 00060) was found at night (2215 h) during heavy rain on a forest trail, and one adult female (NCSM 77787) was found during the morning (1030 h) on leaf litter on the forest floor
|
||
<quantity id="2E17199DFF96FF83FB16FA57FB3AA19D" box="[1211,1268,1510,1534]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="335" unit="m" value="20.0">20 m</quantity>
|
||
from a stream. Larvae (NCSM 77944) were collected at night (2130 h) in a
|
||
<quantity id="2E17199DFF96FF83FC94F9B8FCB7A241" box="[825,889,1545,1570]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="335" unit="m" value="1.5">1.5 m</quantity>
|
||
wide slow-moving stream, approximately
|
||
<quantity id="2E17199DFF96FF83FAFAF9BBFA52A241" box="[1367,1436,1546,1570]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="335" unit="cm" value="20.0">20 cm</quantity>
|
||
deep, with a substrate of sand and algae-covered rocks (
|
||
<figureCitation id="71D4A8FDFF96FF83FCBEF99CFC96A226" box="[787,856,1581,1606]" captionStart="FIGURE 7" captionStartId="10.[151,250,1237,1259]" captionTargetBox="[411,1178,187,1207]" captionTargetId="figure@10.[399,1188,177,1216]" captionTargetPageId="10" captionText="FIGURE 7. Limnonectes coffeatus sp. nov. habitat of adults (A) and larvae (B) at the type locality on the Bolaven Plateau, Pakxong District, Champasak Province, Laos." httpUri="https://zenodo.org/record/1158884/files/figure.png" pageId="10" pageNumber="335">Fig. 7</figureCitation>
|
||
). An adult female (NCSM 77786) was collected in shallow water at the same time and place as the larvae. Calls and oviposition sites are unknown.
|
||
</paragraph>
|
||
<paragraph id="E950B478FF96FF82FF6AF9C4FAB2A57D" blockId="10.[151,1437,1329,2038]" lastBlockId="11.[151,1436,151,1005]" lastPageId="11" lastPageNumber="336" pageId="10" pageNumber="335">
|
||
Coffee (
|
||
<taxonomicName id="2EEFCFFBFF96FF83FE88F9C4FEBEA2ED" box="[293,368,1653,1677]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="335" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="DB9B686AFF96FF83FE88F9C4FEBEA2ED" box="[293,368,1653,1677]" italics="true" pageId="10" pageNumber="335">Coffea</emphasis>
|
||
</taxonomicName>
|
||
spp.) is the major agricultural export from
|
||
<collectingCountry id="91F8F4E8FF96FF83FCC7F9C7FC6EA2ED" box="[874,928,1654,1677]" name="Laos" pageId="10" pageNumber="335">Laos</collectingCountry>
|
||
(for example, it was the largest agricultural export from the country in 2010), and 95% of Lao coffee is grown on the Bolaven Plateau (85% from Pakxong District alone) (
|
||
<bibRefCitation id="8D7EC989FF96FF83FEEAF90CFD53A2B6" author="Schonweger" box="[327,669,1725,1750]" pageId="10" pageNumber="335" refString="Schonweger, O. & Messerli, P. (2015) Land acquisition, investment, and development in the Lao coffee sector: successes and failures. Critical Asian Studies, 47, 94 - 122. https: // doi. org / 10.1080 / 14672715.2015.997095" type="journal article" year="2015">Schönweger & Messerli 2015</bibRefCitation>
|
||
). Rainfall, temperature and physical properties of the soil render the upper elevations of the Bolaven Plateau highly favorable for growing coffee (
|
||
<bibRefCitation id="8D7EC989FF96FF83FC5AF950FB24A29A" author="Trelo-Ges" box="[1015,1258,1761,1786]" pageId="10" pageNumber="335" refString="Trelo-Ges, V., Nilavong, K. & Polthanee, A. (2010) Soil quality assessment for coffee production in Pakxong District, Champasak Province of Lao People's Democratic Republic. International Journal of Environmental and Rural Development, 1 - 2, 88 - 93." type="book chapter" year="2010">
|
||
Trelo-Ges
|
||
<emphasis id="DB9B686AFF96FF83FBDDF953FB66A299" box="[1136,1192,1761,1785]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D7EC989FF96FF83FB58F950FEF4A37E" author="Schonweger" pageId="10" pageNumber="335" refString="Schonweger, O. & Messerli, P. (2015) Land acquisition, investment, and development in the Lao coffee sector: successes and failures. Critical Asian Studies, 47, 94 - 122. https: // doi. org / 10.1080 / 14672715.2015.997095" type="journal article" year="2015">Schönweger & Messerli 2015</bibRefCitation>
|
||
), and a system of small landholder coffee production has been in place on the Bolaven Plateau since French colonial times (
|
||
<bibRefCitation id="8D7EC989FF96FF83FE05F898FD40A322" author="Delang" box="[424,654,1833,1858]" pageId="10" pageNumber="335" refString="Delang, C. O., Toro, M. & Charlet-Phommachanh, M. (2012) Coffee, mines and dams: conflicts over land in the Bolaven Plateau, southern Lao PDR. The Geographical Journal, 2012, 1 - 15." type="journal article" year="2012">
|
||
Delang
|
||
<emphasis id="DB9B686AFF96FF83FDAAF89BFD8BA321" box="[519,581,1833,1857]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). Today, approximately 15,000 households (69% of farms) on the Bolaven Plateau depend on coffee production as their primary source of income (
|
||
<bibRefCitation id="8D7EC989FF96FF83FB82F8FCFACBA306" author="Delang" box="[1071,1285,1869,1894]" pageId="10" pageNumber="335" refString="Delang, C. O., Toro, M. & Charlet-Phommachanh, M. (2012) Coffee, mines and dams: conflicts over land in the Bolaven Plateau, southern Lao PDR. The Geographical Journal, 2012, 1 - 15." type="journal article" year="2012">
|
||
Delang
|
||
<emphasis id="DB9B686AFF96FF83FB24F8FFFB0CA305" box="[1161,1218,1869,1893]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D7EC989FF96FF83FABFF8FCFE92A3EA" author="Schonweger" pageId="10" pageNumber="335" refString="Schonweger, O. & Messerli, P. (2015) Land acquisition, investment, and development in the Lao coffee sector: successes and failures. Critical Asian Studies, 47, 94 - 122. https: // doi. org / 10.1080 / 14672715.2015.997095" type="journal article" year="2015">Schönweger & Messerli 2015</bibRefCitation>
|
||
). Large-scale coffee plantations have also been established in recent years (
|
||
<bibRefCitation id="8D7EC989FF96FF83FB13F8C0FA56A3EA" author="Delang" box="[1214,1432,1905,1930]" pageId="10" pageNumber="335" refString="Delang, C. O., Toro, M. & Charlet-Phommachanh, M. (2012) Coffee, mines and dams: conflicts over land in the Bolaven Plateau, southern Lao PDR. The Geographical Journal, 2012, 1 - 15." type="journal article" year="2012">
|
||
Delang
|
||
<emphasis id="DB9B686AFF96FF83FAB4F8C3FA9AA3E9" box="[1305,1364,1905,1929]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D7EC989FF96FF83FF3AF824FE3FA3CE" author="Schonweger" box="[151,497,1941,1966]" pageId="10" pageNumber="335" refString="Schonweger, O. & Messerli, P. (2015) Land acquisition, investment, and development in the Lao coffee sector: successes and failures. Critical Asian Studies, 47, 94 - 122. https: // doi. org / 10.1080 / 14672715.2015.997095" type="journal article" year="2015">Schönweger & Messerli 2015</bibRefCitation>
|
||
). As such, forest at upper elevations on the Bolaven Plateau has been extensively converted into coffee plantations (
|
||
<bibRefCitation id="8D7EC989FF96FF83FDB3F808FCCFA3B2" author="Thewlis" box="[542,769,1977,2002]" pageId="10" pageNumber="335" refString="Thewlis, R. M., Timmins, R. J., Evans, T. D. & Duckworth, J. W. (1998) The conservation status of birds in Laos: a review of key species. Bird Conservation International, 8 (Supplement), 1 - 159. https: // doi. org / 10.1017 / S 0959270900002197" type="journal article" year="1998">
|
||
Thewlis
|
||
<emphasis id="DB9B686AFF96FF83FD2EF80BFD70A3B1" box="[643,702,1977,2001]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
), with human settlements (and accompanying plantations) encroaching even inside the boundaries of Dong Hua Sao National Protected Area (
|
||
<bibRefCitation id="8D7EC989FF96FF83FBFBF86CFAFFA396" author="Delang" box="[1110,1329,2013,2038]" pageId="10" pageNumber="335" refString="Delang, C. O., Toro, M. & Charlet-Phommachanh, M. (2012) Coffee, mines and dams: conflicts over land in the Bolaven Plateau, southern Lao PDR. The Geographical Journal, 2012, 1 - 15." type="journal article" year="2012">
|
||
Delang
|
||
<emphasis id="DB9B686AFF96FF83FB1CF86FFB25A395" box="[1201,1259,2013,2037]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). Bauxite mining, rubber plantations, and hydroelectric dam construction place additional pressures on forested areas of the Bolaven Plateau (
|
||
<bibRefCitation id="8D7EC989FF97FF82FEF1FF0DFDFCA4B4" author="Delang" box="[348,562,188,213]" pageId="11" pageNumber="336" refString="Delang, C. O., Toro, M. & Charlet-Phommachanh, M. (2012) Coffee, mines and dams: conflicts over land in the Bolaven Plateau, southern Lao PDR. The Geographical Journal, 2012, 1 - 15." type="journal article" year="2012">
|
||
Delang
|
||
<emphasis id="DB9B686AFF97FF82FE1BFF0CFE20A4B4" box="[438,494,188,212]" italics="true" pageId="11" pageNumber="336">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). All known records of the new species were obtained in closed-canopy forest (and not in coffee plantations), and so we expect that the continued conversion of forest into coffee plantations at upper elevations of the Bolaven Plateau likely presents a significant threat to the persistence of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1F5E7F3FF97FF82FF6AFE96FC6BA5CC" pageId="11" pageNumber="336" type="discussion">
|
||
<paragraph id="E950B478FF97FF82FF6AFE96FC6BA5CC" blockId="11.[151,1436,151,1005]" pageId="11" pageNumber="336">
|
||
<emphasis id="DB9B686AFF97FF82FF6AFE96FEA0A520" bold="true" box="[199,366,295,320]" pageId="11" pageNumber="336">Comparisons:</emphasis>
|
||
Four other species of
|
||
<taxonomicName id="2EEFCFFBFF97FF82FDCAFE99FD3AA55F" authorityName="Fitzinger" authorityYear="1843" box="[615,756,296,319]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="genus">
|
||
<emphasis id="DB9B686AFF97FF82FDCAFE99FD3AA55F" box="[615,756,296,319]" italics="true" pageId="11" pageNumber="336">Limnonectes</emphasis>
|
||
</taxonomicName>
|
||
in the region (
|
||
<collectingCountry id="91F8F4E8FF97FF82FC35FE99FC1CA520" box="[920,978,296,320]" name="Laos" pageId="11" pageNumber="336">Laos</collectingCountry>
|
||
,
|
||
<collectingCountry id="91F8F4E8FF97FF82FC70FE96FBF1A520" box="[989,1087,295,320]" name="Vietnam" pageId="11" pageNumber="336">Vietnam</collectingCountry>
|
||
,
|
||
<collectingCountry id="91F8F4E8FF97FF82FBE7FE96FB73A520" box="[1098,1213,295,320]" name="Cambodia" pageId="11" pageNumber="336">Cambodia</collectingCountry>
|
||
and
|
||
<collectingCountry id="91F8F4E8FF97FF82FB5EFE96FA97A520" box="[1267,1369,295,320]" name="Thailand" pageId="11" pageNumber="336">Thailand</collectingCountry>
|
||
) have the combination of males lacking a cephalic caruncle (swollen or cap-like structure;
|
||
<emphasis id="DB9B686AFF97FF82FBFAFEFCFB5BA504" box="[1111,1173,333,356]" italics="true" pageId="11" pageNumber="336">sensu</emphasis>
|
||
<bibRefCitation id="8D7EC989FF97FF82FB33FEFDFA5EA504" author="Lambertz" box="[1182,1424,332,357]" pageId="11" pageNumber="336" refString="Lambertz, M., Hartmann, T., Walsh, S., Geissler, P. & McLeod, D. S. (2014) Anatomy, histology, and systematic implications of the head ornamentation in the males of four species of Limnonectes (Anura: Dicroglossidae). Zoological Journal of the Linnean Society, 172, 117 - 132. https: // doi. org / 10.1111 / zoj. 12171" type="journal article" year="2014">
|
||
Lambertz
|
||
<emphasis id="DB9B686AFF97FF82FABCFEFCFA85A504" box="[1297,1355,332,356]" italics="true" pageId="11" pageNumber="336">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
); adult SVL <
|
||
<quantity id="2E17199DFF97FF82FE86FEDEFEB3A5E7" box="[299,381,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="336" unit="mm" value="60.0">60 mm</quantity>
|
||
; and reduced interdigital foot webbing:
|
||
<taxonomicName id="2EEFCFFBFF97FF82FCECFEC0FBA1A5E8" authority="(Sclater, 1892)" authorityName="Sclater" authorityYear="1892" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[833,1135,367,392]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="limborgi">
|
||
<emphasis id="DB9B686AFF97FF82FCECFEC0FC70A5E8" box="[833,958,368,392]" italics="true" pageId="11" pageNumber="336">L. limborgi</emphasis>
|
||
(
|
||
<bibRefCitation id="8D7EC989FF97FF82FC63FEDEFBA9A5E8" author="Sclater" box="[974,1127,367,392]" pageId="11" pageNumber="336" refString="Sclater, W. L. (1892) On some specimens of frogs in the Indian Museum, Calcutta, with descriptions of several new species. Proceedings of the Zoological Society of London, 1892, 341 - 348." type="proceedings paper" year="1892">Sclater, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EEFCFFBFF97FF82FBD6FEC0FF14A5CC" authority="(Stoliczka, 1870)" authorityName="Stoliczka" authorityYear="1870" baseAuthorityName="Stoliczka" baseAuthorityYear="1870" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="hascheanus">
|
||
<emphasis id="DB9B686AFF97FF82FBD6FEC0FAD3A5E8" box="[1147,1309,368,392]" italics="true" pageId="11" pageNumber="336">L. hascheanus</emphasis>
|
||
(
|
||
<bibRefCitation id="8D7EC989FF97FF82FA81FEDEFF1DA5CC" author="Stoliczka" pageId="11" pageNumber="336" refString="Stoliczka, F. (1870) Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal, 39, 134 - 228. https: // doi. org / 10.1080 / 00222937008696209" type="journal article" year="1870">Stoliczka, 1870</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2EEFCFFBFF97FF82FF4BFE24FDEEA5CC" authority="(Boulenger, 1887)" authorityName="Boulenger" authorityYear="1887" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[230,544,404,429]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="doriae">
|
||
<emphasis id="DB9B686AFF97FF82FF4BFE24FE83A5CC" box="[230,333,404,428]" italics="true" pageId="11" pageNumber="336">L. doriae</emphasis>
|
||
(
|
||
<bibRefCitation id="8D7EC989FF97FF82FEF6FE25FDD6A5CC" author="Boulenger" box="[347,536,404,429]" pageId="11" pageNumber="336" refString="Boulenger, G. A. (1887) An account of the reptiles and batrachians obtained in Tenasserim by M. L. Fea of the Genoa Civic Museum. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 25, 474 - 486." type="journal article" year="1887">Boulenger, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2EEFCFFBFF97FF82FDFBFE24FC6FA5CC" authority="(Smith, 1922)" authorityName="Smith" authorityYear="1922" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[598,929,404,429]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF97FF82FDFBFE24FD35A5CC" box="[598,763,404,428]" italics="true" pageId="11" pageNumber="336">L. kohchangae</emphasis>
|
||
(
|
||
<bibRefCitation id="8D7EC989FF97FF82FCA7FE25FC56A5CC" author="Smith" box="[778,920,404,429]" pageId="11" pageNumber="336" refString="Smith, M. A. (1922) The frogs allied to Rana doriae. The Journal of the Natural History Society of Siam, 4, 215 - 229." type="journal article" year="1922">Smith, 1922</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1F5E7F3FF97FF82FF6AFE09FE77A78C" pageId="11" pageNumber="336" type="diagnosis">
|
||
<paragraph id="E950B478FF97FF82FF6AFE09FA52A678" blockId="11.[151,1436,151,1005]" pageId="11" pageNumber="336">
|
||
<taxonomicName id="2EEFCFFBFF97FF82FF6AFE09FE0AA5B0" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[199,452,440,464]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF97FF82FF6AFE09FE0AA5B0" box="[199,452,440,464]" italics="true" pageId="11" pageNumber="336">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF97FF82FE63FE06FDE5A5B0" bold="true" box="[462,555,439,464]" pageId="11" pageNumber="336">
|
||
<taxonomicNameLabel id="C0A8D511FF97FF82FE63FE06FDE5A5B0" box="[462,555,439,464]" pageId="11" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="2EEFCFFBFF97FF82FD67FE08FC84A5B0" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[714,842,440,464]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="limborgi">
|
||
<emphasis id="DB9B686AFF97FF82FD67FE08FC84A5B0" box="[714,842,440,464]" italics="true" pageId="11" pageNumber="336">L. limborgi</emphasis>
|
||
</taxonomicName>
|
||
by having webbing on Toe V reaching base of tip (webbing more reduced in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FE6BFE6CFD8DA594" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[454,579,476,500]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="limborgi">
|
||
<emphasis id="DB9B686AFF97FF82FE6BFE6CFD8DA594" box="[454,579,476,500]" italics="true" pageId="11" pageNumber="336">L. limborgi</emphasis>
|
||
</taxonomicName>
|
||
); dorsum with rounded tubercles, not arranged in rows (if present, tubercles elongated and arranged in rows in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FDB5FDB0FD5AA678" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[536,660,512,536]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="limborgi">
|
||
<emphasis id="DB9B686AFF97FF82FDB5FDB0FD5AA678" box="[536,660,512,536]" italics="true" pageId="11" pageNumber="336">L. limborgi</emphasis>
|
||
</taxonomicName>
|
||
), and having a uniformly gray throat in males (absent in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FAA2FDB0FA45A678" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[1295,1419,512,536]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="limborgi">
|
||
<emphasis id="DB9B686AFF97FF82FAA2FDB0FA45A678" box="[1295,1419,512,536]" italics="true" pageId="11" pageNumber="336">L. limborgi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E950B478FF97FF82FF6AFD94FACEA6E4" blockId="11.[151,1436,151,1005]" pageId="11" pageNumber="336">
|
||
<taxonomicName id="2EEFCFFBFF97FF82FF6AFD94FE0EA65C" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[199,448,548,572]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF97FF82FF6AFD94FE0EA65C" box="[199,448,548,572]" italics="true" pageId="11" pageNumber="336">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF97FF82FE6AFD95FDEEA65D" bold="true" box="[455,544,548,573]" pageId="11" pageNumber="336">
|
||
<taxonomicNameLabel id="C0A8D511FF97FF82FE6AFD95FDEEA65D" box="[455,544,548,573]" pageId="11" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="2EEFCFFBFF97FF82FD1EFD94FC9DA65C" baseAuthorityName="Stoliczka" baseAuthorityYear="1870" box="[691,851,548,572]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="hascheanus">
|
||
<emphasis id="DB9B686AFF97FF82FD1EFD94FC9DA65C" box="[691,851,548,572]" italics="true" pageId="11" pageNumber="336">L. hascheanus</emphasis>
|
||
</taxonomicName>
|
||
by having adults with SVL> 26 (adults with SVL <
|
||
<quantity id="2E17199DFF97FF82FF3AFDF6FF19A600" box="[151,215,583,608]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="11" pageNumber="336" unit="in" value="26.0">26 in</quantity>
|
||
<taxonomicName id="2EEFCFFBFF97FF82FF48FDF8FE43A600" baseAuthorityName="Stoliczka" baseAuthorityYear="1870" box="[229,397,584,608]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="hascheanus">
|
||
<emphasis id="DB9B686AFF97FF82FF48FDF8FE43A600" box="[229,397,584,608]" italics="true" pageId="11" pageNumber="336">L. hascheanus</emphasis>
|
||
</taxonomicName>
|
||
); having males with two large odontoid processes at front of mandible (absent in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FA28FDF8FED4A6E4" baseAuthorityName="Stoliczka" baseAuthorityYear="1870" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="hascheanus">
|
||
<emphasis id="DB9B686AFF97FF82FA28FDF8FED4A6E4" italics="true" pageId="11" pageNumber="336">L. hascheanus</emphasis>
|
||
</taxonomicName>
|
||
); and having ova with pigmented poles (enlarged, non-pigmented ova in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FBE2FDDCFB3EA6E4" baseAuthorityName="Stoliczka" baseAuthorityYear="1870" box="[1103,1264,620,644]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="hascheanus">
|
||
<emphasis id="DB9B686AFF97FF82FBE2FDDCFB3EA6E4" box="[1103,1264,620,644]" italics="true" pageId="11" pageNumber="336">L. hascheanus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E950B478FF97FF82FF6AFD21FDC8A690" blockId="11.[151,1436,151,1005]" pageId="11" pageNumber="336">
|
||
<taxonomicName id="2EEFCFFBFF97FF82FF6AFD21FE0CA6C8" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[199,450,656,680]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF97FF82FF6AFD21FE0CA6C8" box="[199,450,656,680]" italics="true" pageId="11" pageNumber="336">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF97FF82FE61FD3EFDE9A6C8" bold="true" box="[460,551,655,680]" pageId="11" pageNumber="336">
|
||
<taxonomicNameLabel id="C0A8D511FF97FF82FE61FD3EFDE9A6C8" box="[460,551,655,680]" pageId="11" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="2EEFCFFBFF97FF82FD6CFD20FCE4A6C8" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[705,810,656,680]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="doriae">
|
||
<emphasis id="DB9B686AFF97FF82FD6CFD20FCE4A6C8" box="[705,810,656,680]" italics="true" pageId="11" pageNumber="336">L. doriae</emphasis>
|
||
</taxonomicName>
|
||
by having males with two large odontoid processes at front of mandible (absent in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FE79FD04FDF4A6AC" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[468,570,692,716]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="doriae">
|
||
<emphasis id="DB9B686AFF97FF82FE79FD04FDF4A6AC" box="[468,570,692,716]" italics="true" pageId="11" pageNumber="336">L. doriae</emphasis>
|
||
</taxonomicName>
|
||
); having adults with SVL <43 (SVL
|
||
<quantity id="2E17199DFF97FF82FC4DFD05FBD7A6AD" box="[992,1049,692,717]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="11" pageNumber="336" unit="in" value="56.0">56 in</quantity>
|
||
<taxonomicName id="2EEFCFFBFF97FF82FBB2FD04FB4BA6AC" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[1055,1157,692,716]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="doriae">
|
||
<emphasis id="DB9B686AFF97FF82FBB2FD04FB4BA6AC" box="[1055,1157,692,716]" italics="true" pageId="11" pageNumber="336">L. doriae</emphasis>
|
||
</taxonomicName>
|
||
); and males with TMP> EYE (TMP <EYE in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FE22FD68FE38A690" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[399,502,728,752]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="doriae">
|
||
<emphasis id="DB9B686AFF97FF82FE22FD68FE38A690" box="[399,502,728,752]" italics="true" pageId="11" pageNumber="336">L. doriae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E950B478FF97FF82FF6AFD4CFE77A78C" blockId="11.[151,1436,151,1005]" pageId="11" pageNumber="336">
|
||
<taxonomicName id="2EEFCFFBFF97FF82FF6AFD4CFE0CA774" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[199,450,764,788]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF97FF82FF6AFD4CFE0CA774" box="[199,450,764,788]" italics="true" pageId="11" pageNumber="336">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DB9B686AFF97FF82FE66FD4DFDE8A775" bold="true" box="[459,550,764,789]" pageId="11" pageNumber="336">
|
||
<taxonomicNameLabel id="C0A8D511FF97FF82FE66FD4DFDE8A775" box="[459,550,764,789]" pageId="11" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is phenotypically most similar (but not phylogenetically related;
|
||
<figureCitation id="71D4A8FDFF97FF82FABFFD4DFA98A775" box="[1298,1366,764,789]" captionStart="FIGURE 6" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[166,1414,530,1812]" captionTargetId="figure@9.[151,1436,517,1829]" captionTargetPageId="9" captionText="FIGURE 6. Fifty percent majority-rule consensus phylogram resulting from Bayesian analysis of 1,633 aligned characters of the mitochondrial 16S gene from frogs of the dicroglossid genera Fejervarya, Quasipaa, and Limnonectes. Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95, and no circle at nodes indicate Bayesian posterior probabilities <0.95. Numbers at terminal tips are GenBank accession numbers." httpUri="https://zenodo.org/record/1158882/files/figure.png" pageId="11" pageNumber="336">Fig. 6</figureCitation>
|
||
) to
|
||
<taxonomicName id="2EEFCFFBFF97FF82FA28FD4CFED0A758" baseAuthorityName="Smith" baseAuthorityYear="1922" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF97FF82FA28FD4CFED0A758" italics="true" pageId="11" pageNumber="336">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="71D4A8FDFF97FF82FE82FCAEFEBAA758" box="[303,372,799,824]" captionStart="FIGURE 8" captionStartId="11.[151,250,1943,1965]" captionTargetBox="[154,1431,1065,1912]" captionTargetId="figure@11.[150,1436,1060,1921]" captionTargetPageId="11" captionText="FIGURE 8. Adult males of Limnonectes kohchangae in life at the type locality of Ko Chang Island, Ko Chang District, Trat Province, Thailand (A; ZMKU AM 01158) and at Phnom Aural, Kampong Speu Province, Aural District, Cambodia (B; CBC 01907).Lectotype male (BMNH 1947.2.3.60) of L. kohchangae in dorsal (C) and ventral (D) views." httpUri="https://zenodo.org/record/1158886/files/figure.png" pageId="11" pageNumber="336">Fig. 8</figureCitation>
|
||
).
|
||
<taxonomicName id="2EEFCFFBFF97FF82FE21FC91FD49A758" authority="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart, 2018" authorityName="Phimmachak & Sivongxay & Seateun & Yodthong & Rujirawan & Neang & Aowphol & Stuart" authorityYear="2018" box="[396,647,800,824]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="coffeatus" status="sp. nov.">
|
||
<emphasis id="DB9B686AFF97FF82FE21FC91FD49A758" box="[396,647,800,824]" italics="true" pageId="11" pageNumber="336">Limnonectes coffeatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C0A8D511FF97FF82FD3DFC91FD26A757" box="[656,744,800,823]" pageId="11" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="2EEFCFFBFF97FF82FC2DFC90FBE8A758" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[896,1062,800,824]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF97FF82FC2DFC90FBE8A758" box="[896,1062,800,824]" italics="true" pageId="11" pageNumber="336">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
by having dorsum with rounded tubercles, not arranged in rows (distinctly elongated and arranged in rows in
|
||
<emphasis id="DB9B686AFF97FF82FC52FCF4FB60A73C" box="[1023,1198,836,860]" italics="true" pageId="11" pageNumber="336">
|
||
<taxonomicName id="2EEFCFFBFF97FF82FC52FCF4FB68A73C" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[1023,1190,836,860]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="kohchangae">L. kohchangae</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<figureCitation id="71D4A8FDFF97FF82FB1BFCF5FB34A73C" box="[1206,1274,836,861]" captionStart="FIGURE 9" captionStartId="12.[151,250,1142,1164]" captionTargetBox="[409,1177,193,1120]" captionTargetId="figure@12.[409,1177,193,1120]" captionTargetPageId="12" captionText="FIGURE 9. Dorsal views in preservative of males (A) of L. kohchangae (left, NCSM 79541) and L. coffeatus sp. nov. (right, holotype NCSM 77788) and females (B) of L. kohchangae (left, NCSM 79542) and L. coffeatus sp. nov. (right, paratype NCSM 77787)." httpUri="https://zenodo.org/record/1158888/files/figure.png" pageId="11" pageNumber="336">Fig. 9</figureCitation>
|
||
); by lacking a distinctly bi-colored tympanum, with the upper half heavily pigmented (present in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FBCEFCD8FAC5A7E0" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[1123,1291,872,896]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF97FF82FBCEFCD8FAC5A7E0" box="[1123,1291,872,896]" italics="true" pageId="11" pageNumber="336">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="71D4A8FDFF97FF82FAB3FCD6FAABA7E0" box="[1310,1381,871,896]" captionStart="FIGURE 8" captionStartId="11.[151,250,1943,1965]" captionTargetBox="[154,1431,1065,1912]" captionTargetId="figure@11.[150,1436,1060,1921]" captionTargetPageId="11" captionText="FIGURE 8. Adult males of Limnonectes kohchangae in life at the type locality of Ko Chang Island, Ko Chang District, Trat Province, Thailand (A; ZMKU AM 01158) and at Phnom Aural, Kampong Speu Province, Aural District, Cambodia (B; CBC 01907).Lectotype male (BMNH 1947.2.3.60) of L. kohchangae in dorsal (C) and ventral (D) views." httpUri="https://zenodo.org/record/1158886/files/figure.png" pageId="11" pageNumber="336">Fig. 8</figureCitation>
|
||
); by lacking dark spots on flank (present in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FDE5FC3CFD22A7C4" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[584,748,908,932]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF97FF82FDE5FC3CFD22A7C4" box="[584,748,908,932]" italics="true" pageId="11" pageNumber="336">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="71D4A8FDFF97FF82FD56FC3DFCF2A7C4" box="[763,828,908,933]" captionStart="FIGURE 8" captionStartId="11.[151,250,1943,1965]" captionTargetBox="[154,1431,1065,1912]" captionTargetId="figure@11.[150,1436,1060,1921]" captionTargetPageId="11" captionText="FIGURE 8. Adult males of Limnonectes kohchangae in life at the type locality of Ko Chang Island, Ko Chang District, Trat Province, Thailand (A; ZMKU AM 01158) and at Phnom Aural, Kampong Speu Province, Aural District, Cambodia (B; CBC 01907).Lectotype male (BMNH 1947.2.3.60) of L. kohchangae in dorsal (C) and ventral (D) views." httpUri="https://zenodo.org/record/1158886/files/figure.png" pageId="11" pageNumber="336">Fig. 8</figureCitation>
|
||
); and having narrower limb bands, with the maximum shank band width <50% horizontal diameter of eye (maximum shank band width>50% horizontal diameter of eye in
|
||
<taxonomicName id="2EEFCFFBFF97FF82FF19FC64FE96A78C" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[180,344,980,1004]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="336" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="DB9B686AFF97FF82FF19FC64FE96A78C" box="[180,344,980,1004]" italics="true" pageId="11" pageNumber="336">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="71D4A8FDFF97FF82FECAFC65FE62A78C" box="[359,428,980,1005]" captionStart="FIGURE 9" captionStartId="12.[151,250,1142,1164]" captionTargetBox="[409,1177,193,1120]" captionTargetId="figure@12.[409,1177,193,1120]" captionTargetPageId="12" captionText="FIGURE 9. Dorsal views in preservative of males (A) of L. kohchangae (left, NCSM 79541) and L. coffeatus sp. nov. (right, holotype NCSM 77788) and females (B) of L. kohchangae (left, NCSM 79542) and L. coffeatus sp. nov. (right, paratype NCSM 77787)." httpUri="https://zenodo.org/record/1158888/files/figure.png" pageId="11" pageNumber="336">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |