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<document ID-DOI="10.25221/fee.365.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7164252" approvalRequired="244" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="237" approvalRequired_for_treatments="1" checkinTime="1665241599349" checkinUser="felipe" docAuthor="Grichanov, I. Ya." docDate="2018" docId="382287BE1A526C13FF7A534AFCAD6C57" docLanguage="en" docName="FarEastEntomol.365.1-25.pdf" docOrigin="Far Eastern Entomologist 365" docSource="http://dx.doi.org/10.25221/fee.365.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Tenuopus kylei Grichanov 2018, sp. n." docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="C41BFFC61A5E6C1DFF99575FFFE0683F" masterDocTitle="A new subfamily of Dolichopodidae (Diptera) for Tenuopus Curran, 1924 with description of new species from Tropical Africa" masterLastPageNumber="25" masterPageNumber="1" pageNumber="13" updateTime="1665268663009" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new subfamily of Dolichopodidae (Diptera) for Tenuopus Curran, 1924 with description of new species from Tropical Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grichanov, I. Ya.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-08-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>365</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>25</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.25221/fee.365.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.365.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7164252</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:382287BE1A526C13FF7A534AFCAD6C57" httpUri="http://treatment.plazi.org/id/382287BE1A526C13FF7A534AFCAD6C57" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection box="[227,596,1044,1068]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph blockId="12.[227,596,1044,1098]" box="[227,596,1044,1068]" pageId="12" pageNumber="13">
<heading box="[227,596,1044,1068]" centered="true" fontSize="9" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis bold="true" box="[227,596,1044,1068]" pageId="12" pageNumber="13">
<taxonomicName authority="Grichanov" authorityName="Grichanov" authorityYear="2018" box="[227,525,1044,1068]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kylei" status="sp. n.">
<emphasis bold="true" box="[227,395,1044,1068]" italics="true" pageId="12" pageNumber="13">Tenuopus kylei</emphasis>
Grichanov
</taxonomicName>
,
<taxonomicNameLabel box="[537,596,1044,1068]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[227,373,1074,1098]" pageId="12" pageNumber="13" type="description">
<paragraph blockId="12.[227,596,1044,1098]" box="[227,373,1074,1098]" pageId="12" pageNumber="13">
<figureCitation box="[227,296,1074,1098]" captionStart="Figs 1–6" captionStartId="6.[265,306,1584,1606]" captionTargetBox="[229,1124,257,1519]" captionTargetId="figure-0@6.[229,1124,257,1520]" captionTargetPageId="6" captionText="Figs 1–6. Tenuopus spp., head: 1 – T. bururiensis sp. n.; 2 – T. gorongosaensis sp. n.; 3 – T. kirkspriggsi sp. n.; 4 – T. kylei sp. n.; 5 – T. lomholdti sp. n.; 6 – T. unicolor (Becker)." figureDoi="http://doi.org/10.5281/zenodo.7166072" httpUri="https://zenodo.org/record/7166072/files/figure.png" pageId="12" pageNumber="13">Figs 4</figureCitation>
,
<figureCitation box="[306,334,1074,1098]" captionStart="Figs 7–14" captionStartId="10.[265,306,1503,1525]" captionTargetBox="[229,1124,257,1481]" captionTargetId="figure-0@10.[229,1124,257,1482]" captionTargetPageId="10" captionText="Figs 7–14. Tenuopus spp. 7 – T. shcherbakovi Grichanov, head in ethanol with everted" figureDoi="http://doi.org/10.5281/zenodo.7166074" httpUri="https://zenodo.org/record/7166074/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
,
<figureCitation box="[346,373,1074,1098]" captionStart="Figs 19–22" captionStartId="17.[265,306,1423,1445]" captionTargetBox="[229,1124,717,1388]" captionTargetId="figure-199@17.[229,1124,717,1389]" captionTargetPageId="17" captionText="Figs 19–22. Tenuopus spp., epandrial lobes and surstylus, right lateral view: 19 – T. kylei" figureDoi="http://doi.org/10.5281/zenodo.7164266" httpUri="https://zenodo.org/record/7164266/files/figure.png" pageId="12" pageNumber="13">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="materials_examined">
<paragraph blockId="12.[227,1127,1134,1608]" box="[265,1126,1134,1158]" pageId="12" pageNumber="13">
<materialsCitation box="[265,1126,1134,1158]" collectionCode="TYPE, MATERIAL" country="South Africa" county="Kosi" location="Kwazulu" municipality="Madlangula" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode box="[265,481,1134,1158]" pageId="12" pageNumber="13">TYPE MATERIAL</collectionCode>
.
<typeStatus box="[493,592,1134,1158]" pageId="12" pageNumber="13">Holotype</typeStatus>
–
<specimenCount box="[623,645,1134,1157]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
,
<emphasis bold="true" box="[658,806,1134,1158]" pageId="12" pageNumber="13">
<collectingCountry box="[658,806,1134,1158]" name="South Africa" pageId="12" pageNumber="13">South Africa</collectingCountry>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:382287BE1A526C13FF7A534AFCAD6C57:B55460731A526C11FCAF5331FC796CB9" box="[822,921,1134,1158]" country="South Africa" county="Kosi" municipality="Madlangula" name="Kwazulu" pageId="12" pageNumber="13">Kwazulu</location>
,
<collectingMunicipality box="[934,1064,1134,1158]" pageId="12" pageNumber="13">Madlangula</collectingMunicipality>
,
<collectingCounty box="[1076,1126,1134,1158]" pageId="12" pageNumber="13">Kosi</collectingCounty>
</materialsCitation>
</paragraph>
<paragraph blockId="12.[227,1127,1134,1608]" box="[227,1037,1164,1189]" pageId="12" pageNumber="13">
<materialsCitation box="[227,484,1164,1189]" collectingDate="1985-01-21" collectingDateMax="1985-02-06" collectingDateMin="1985-01-21" collectionCode="R" pageId="12" pageNumber="13" specimenCount="1">
Bay,
<date box="[283,449,1164,1188]" pageId="12" pageNumber="13" value="1985-01-21" valueMax="1985-02-06" valueMin="1985-01-21">
<collectingDate box="[283,449,1164,1188]" pageId="12" pageNumber="13" value="1985-01-21" valueMax="1985-02-06" valueMin="1985-01-21">21.I – 6.II 1985</collectingDate>
</date>
,
<collectionCode box="[463,484,1165,1188]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
</materialsCitation>
.
<materialsCitation box="[494,803,1164,1189]" collectionCode="NMSA" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
Kyle [
<collectionCode box="[559,644,1164,1188]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="12" pageNumber="13" type="Museum">NMSA</collectionCode>
].
<typeStatus box="[661,758,1165,1189]" pageId="12" pageNumber="13">Paratype</typeStatus>
:
<specimenCount box="[768,803,1164,1188]" pageId="12" pageNumber="13" type="male">1♂</specimenCount>
</materialsCitation>
, same label [
<collectionCode box="[941,1026,1164,1188]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="12" pageNumber="13" type="Museum">NMSA</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph blockId="12.[227,1127,1134,1608]" pageId="12" pageNumber="13">
DESCRIPTION. Male.
<emphasis box="[526,584,1194,1218]" italics="true" pageId="12" pageNumber="13">Head</emphasis>
: frons black, whitish pollinose; one pair of short postvertical setae, as long as uppermost postocular seta; upper postocular setae black, increasing in length upward; lateral and lower postoculars white; ventral postcranium covered with irregular white hairs; face silvery-white, 7 times as high as wide in middle, nearly as wide as postpedicel; clypeus slightly bulging; antennae as long as height of head, with yellow scape and pedicel and yellow-orange postpedicel; pedicel projected distally on inner side, with a crown of short black setae,
</paragraph>
<paragraph blockId="12.[227,1127,1134,1608]" box="[227,1126,1404,1428]" pageId="12" pageNumber="13">one of dorsal setae as long as pedicel; postpedicel rounded, as long as high at base</paragraph>
<paragraph blockId="12.[227,1127,1134,1608]" pageId="12" pageNumber="13">
(1/1); arista-like stylus dorsal, with short hairs; length ratio of scape to pedicel to postpedicel to stylus (1
<superScript attach="left" box="[478,490,1460,1474]" fontSize="6" pageId="12" pageNumber="13">st</superScript>
and 2
<superScript attach="left" box="[558,575,1459,1473]" fontSize="6" pageId="12" pageNumber="13">nd</superScript>
segments), 5/7/9/3/40; palpus and proboscis short,
</paragraph>
<paragraph blockId="12.[227,1127,1134,1608]" box="[227,1096,1494,1518]" pageId="12" pageNumber="13">yellow, covered with white hairs, proboscis also with a pair of black lateral setae.</paragraph>
<paragraph blockId="12.[227,1127,1134,1608]" lastBlockId="13.[227,1127,258,677]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis box="[265,342,1524,1548]" italics="true" pageId="12" pageNumber="13">Thorax</emphasis>
: pleura yellow; mesonotum mostly orange-yellow, with median greenish blue stripe half as wide as surface between dorsocentral bristles, wider posteriorly and black on scutellum dorsally; 6 dorsocentral setae with posterior pair shifted laterally; acrostichals biseriate, strong, nearly reaching 5
<superScript attach="left" box="[847,860,258,272]" fontSize="6" pageId="13" pageNumber="14">th</superScript>
dorsocentrals; scutellum with 2 strong bristles and 2 short lateral hairs; proepisternum with 2 yellow setae.
</paragraph>
<paragraph blockId="13.[227,1127,258,677]" pageId="13" pageNumber="14">
<emphasis box="[265,316,323,346]" italics="true" pageId="13" pageNumber="14">Legs</emphasis>
yellow; last segments of hind tarsus brown; fore and mid coxae with black hairs anteriorly and 5-7 black apical bristles of various length; hind coxa with one long black outer bristle above middle; fore femur with light and dark fine erect ventral hairs, at most half as long as diameter of femur, and 1-2 black posteroventral cilia; fore tibia simple, with 1 anterodorsal and 1 posterodorsal at base, 1 dorsal in middle, 1-2 apical setae; basitarsus with 1 dorsal; other tarsomeres simple; claws simple; mid femur with very small anterior preapical seta, practically glabrous ventrally, with few dark ventral hairs; mid tibia with 3 anterior, 3 posterodorsal, 3
</paragraph>
<paragraph blockId="13.[227,1127,258,677]" pageId="13" pageNumber="14">ventral, 3-4 apical seta; hind femur without long hairs, with 1 strong anterior preapical seta; hind tibia with 3 anterior, 2 dorsal, 3 posterodorsal, 2-3 apical setae. Femur,</paragraph>
<paragraph blockId="13.[227,1127,258,677]" pageId="13" pageNumber="14">tibia and tarsomere (from first to fifth) length ratio: fore leg: 9/12/15/8/4/2/1, mid leg: 10/15/12/5/4/2/1, hind leg: 12/20/6/6/4/2/1.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166076" ID-Zenodo-Dep="7166076" box="[265,1127,1531,1553]" httpUri="https://zenodo.org/record/7166076/files/figure.png" pageId="13" pageNumber="14" startId="13.[265,306,1531,1553]" targetBox="[229,1124,708,1495]" targetPageId="13">
<paragraph blockId="13.[227,1127,1531,1608]" box="[265,1127,1531,1553]" pageId="13" pageNumber="14">
Figs 15–18.
<taxonomicName box="[384,526,1532,1553]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[384,477,1532,1553]" italics="true" pageId="13" pageNumber="14">Tenuopus</emphasis>
spp.
</taxonomicName>
, epandrial lobes and surstylus, right (15, 18) and left (16) lateral
</paragraph>
</caption>
<paragraph blockId="13.[227,1127,1531,1608]" box="[227,1127,1559,1581]" pageId="13" pageNumber="14">
view, and cercus, dorsal view (17): 15 –
<taxonomicName authority="Grichanov, 2018" authorityName="Grichanov" authorityYear="2018" box="[623,758,1559,1581]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bururiensis" status="sp. n.">
<emphasis box="[623,758,1559,1581]" italics="true" pageId="13" pageNumber="14">T. bururiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[765,819,1559,1580]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
; 16–17 –
<taxonomicName authority="Grichanov, 2018" authorityName="Grichanov" authorityYear="2018" box="[916,1093,1559,1581]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gorongosaensis" status="sp.">
<emphasis box="[916,1093,1559,1581]" italics="true" pageId="13" pageNumber="14">T. gorongosaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1099,1127,1559,1580]" pageId="13" pageNumber="14">sp.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="13.[227,1127,1531,1608]" box="[227,504,1586,1608]" pageId="13" pageNumber="14">
n.; 18 –
<taxonomicName authority="Grichanov, 2018" authorityName="Grichanov" authorityYear="2018" box="[306,445,1586,1608]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kirkspriggsi" status="sp. n.">
<emphasis box="[306,445,1586,1608]" italics="true" pageId="13" pageNumber="14">T. kirkspriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[452,504,1587,1608]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" pageId="14" pageNumber="15">
<emphasis box="[265,321,263,286]" italics="true" pageId="14" pageNumber="15">Wing</emphasis>
greyish, almost hyaline, veins brown; subcosta very thin; ratio of part of costa between R
<subScript attach="left" box="[403,430,303,317]" fontSize="6" pageId="14" pageNumber="15">2+3</subScript>
and R
<subScript attach="left" box="[501,528,303,317]" fontSize="6" pageId="14" pageNumber="15">4+5</subScript>
to that between R
<subScript attach="left" box="[724,751,303,317]" fontSize="6" pageId="14" pageNumber="15">4+5</subScript>
and M
<subScript attach="left" box="[828,837,303,317]" fontSize="6" pageId="14" pageNumber="15">1</subScript>
, 7/1; M
<subScript attach="left" box="[923,932,303,317]" fontSize="6" pageId="14" pageNumber="15">1</subScript>
with gentle arc to apex, reaching costa right before wing apex; M
<subScript attach="left" box="[724,733,332,346]" fontSize="6" pageId="14" pageNumber="15">2</subScript>
present as fold on membrane; crossvein
<emphasis box="[281,342,350,374]" italics="true" pageId="14" pageNumber="15">dm-m</emphasis>
straight; ratio of crossvein
<emphasis box="[644,705,350,374]" italics="true" pageId="14" pageNumber="15">dm-m</emphasis>
to apical part of M
<subScript attach="left" box="[920,947,361,375]" fontSize="6" pageId="14" pageNumber="15">1+2</subScript>
(fork-handle) to apical part of M
<subScript attach="left" box="[399,408,390,404]" fontSize="6" pageId="14" pageNumber="15">4</subScript>
, 5/16/13; anal vein foldlike, not reaching wing margin; anal angle obtuse; lower calypter yellow, with black apex and pale setae; halter yellow with orange knob, halter stem thin and long, with dorsal and ventral groups of short hairs distally.
</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" pageId="14" pageNumber="15">
<emphasis box="[265,366,495,519]" italics="true" pageId="14" pageNumber="15">Abdomen</emphasis>
mostly yellow-orange, black setose; 2
<superScript attach="right" box="[770,787,491,505]" fontSize="6" pageId="14" pageNumber="15">nd</superScript>
–6
<superScript attach="left" box="[814,827,491,505]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
yellow, with narrow brown or black anterior and posterior edging; 8
<superScript attach="left" box="[668,681,519,533]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
segment and epandrium entirely yellow;
</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" pageId="14" pageNumber="15">
8
<superScript attach="left" box="[240,253,548,562]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
segment with yellow hairs; cerci and surstyli yellow; surstylus and epandrial lobe black at apex; cercus covered with dark-yellow hairs and setae; hypandrium small, simple; phallus thin and simple; epandrial lobe prominent, finger-like,
</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" pageId="14" pageNumber="15">narrow at apex, with 2 ventral setae; 3 pedunculate epandrial setae of different length and 2 epandrial processes between hypandrium and base of epandrial lobe,</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" pageId="14" pageNumber="15">including 1 process with rather long apical seta and 1 slightly longer process with microscopic apical seta; surstylus about as long as cercus, with apical incision, with thin dorsal and thick ventral lobes; dorsal lobe of surstylus with 1 small apical seta;</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" pageId="14" pageNumber="15">ventral lobe of surstylus with 2 small apical setae; cercus nearly as long as epandrium, subtriangular, covered with hairs dorsally and setae distally.</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" box="[265,1127,842,866]" pageId="14" pageNumber="15">MEASUREMENTS (in mm). Body length 6.0; antenna length 1.4; wing length</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" box="[227,442,871,895]" pageId="14" pageNumber="15">5.1; wing width 1.6.</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" box="[265,470,901,925]" pageId="14" pageNumber="15">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="etymology">
<paragraph blockId="14.[227,1127,263,1129]" box="[265,1127,930,954]" pageId="14" pageNumber="15">ETYMOLOGY. The species is named after the South African zoologist, Dr. R.</paragraph>
<paragraph blockId="14.[227,1127,263,1129]" box="[227,1069,959,983]" pageId="14" pageNumber="15">
Kyle (Ezemvelo KwaZulu Natal Wildlife), the collector of the
<typeStatus box="[898,944,959,983]" pageId="14" pageNumber="15">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
<subSubSection box="[265,610,988,1012]" pageId="14" pageNumber="15" type="distribution">
<paragraph blockId="14.[227,1127,263,1129]" box="[265,610,988,1012]" pageId="14" pageNumber="15">
DISTRIBUTION.
<collectingCountry box="[465,606,988,1012]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="diagnosis">
<paragraph blockId="14.[227,1127,263,1129]" pageId="14" pageNumber="15">
DIAGNOSIS.
<taxonomicName authority="Grichanov, 2018" authorityName="Grichanov" authorityYear="2018" box="[430,516,1017,1041]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kylei" status="sp. n.">
<emphasis box="[430,516,1017,1041]" italics="true" pageId="14" pageNumber="15">T. kylei</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[531,596,1018,1041]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
belongs to a group of species with biseriate acrostichals, being the closest to
<taxonomicName authorityName="Grichanov" authorityYear="1996" box="[597,752,1046,1070]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kononenkoi">
<emphasis box="[597,752,1046,1070]" italics="true" pageId="14" pageNumber="15">T. kononenkoi</emphasis>
</taxonomicName>
, differing from the latter in male fore basitarsus longer, about 1.3 times as long as fore tibia; and male surstylus bifurcated only at apex, with broad lobes (see key above).
</paragraph>
</subSubSection>
</treatment>
</document>
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