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<document ID-DOI="10.25221/fee.365.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7164252" approvalRequired="244" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="237" approvalRequired_for_treatments="1" checkinTime="1665241599349" checkinUser="felipe" docAuthor="Grichanov, I. Ya." docDate="2018" docId="382287BE1A4C6C09FF7A543FFBB96CCA" docLanguage="en" docName="FarEastEntomol.365.1-25.pdf" docOrigin="Far Eastern Entomologist 365" docSource="http://dx.doi.org/10.25221/fee.365.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Tenuopus soderlundi Grichanov 2018, sp. n." docType="treatment" docVersion="2" lastPageNumber="21" masterDocId="C41BFFC61A5E6C1DFF99575FFFE0683F" masterDocTitle="A new subfamily of Dolichopodidae (Diptera) for Tenuopus Curran, 1924 with description of new species from Tropical Africa" masterLastPageNumber="25" masterPageNumber="1" pageNumber="19" updateTime="1665252493061" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new subfamily of Dolichopodidae (Diptera) for Tenuopus Curran, 1924 with description of new species from Tropical Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grichanov, I. Ya.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-08-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>365</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>25</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.25221/fee.365.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.365.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7164252</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:382287BE1A4C6C09FF7A543FFBB96CCA" httpUri="http://treatment.plazi.org/id/382287BE1A4C6C09FF7A543FFBB96CCA" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection box="[227,662,863,888]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph blockId="18.[227,662,863,918]" box="[227,662,863,888]" pageId="18" pageNumber="19">
<heading box="[227,662,863,888]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="19" reason="2">
<emphasis bold="true" box="[227,662,863,888]" pageId="18" pageNumber="19">
<taxonomicName authority="Grichanov" authorityName="Grichanov" authorityYear="2018" box="[227,590,863,888]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="soderlundi" status="sp. n.">
<emphasis bold="true" box="[227,460,863,887]" italics="true" pageId="18" pageNumber="19">Tenuopus soderlundi</emphasis>
Grichanov
</taxonomicName>
,
<taxonomicNameLabel box="[602,662,864,888]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[227,346,894,918]" pageId="18" pageNumber="19" type="description">
<paragraph blockId="18.[227,662,863,918]" box="[227,346,894,918]" pageId="18" pageNumber="19">
<figureCitation box="[227,346,894,918]" captionStart="Figs 2328" captionStartId="19.[265,306,1571,1593]" captionTargetBox="[239,1114,411,1555]" captionTargetId="figure-42@19.[239,1114,411,1555]" captionTargetPageId="19" captionText="Figs 2328. Tenuopus spp. 2325 T. cognatus Parent; 2628 T. soderlundi sp. n.; 23," figureDoi="http://doi.org/10.5281/zenodo.7164268" httpUri="https://zenodo.org/record/7164268/files/figure.png" pageId="18" pageNumber="19">Figs 2628</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="materials_examined">
<paragraph blockId="18.[227,1127,953,1608]" box="[265,1127,953,978]" pageId="18" pageNumber="19">
<materialsCitation box="[265,1127,953,978]" collectionCode="TYPE, MATERIAL" country="South Africa" location="Cape Province" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<collectionCode box="[265,482,954,978]" pageId="18" pageNumber="19">TYPE MATERIAL</collectionCode>
.
<typeStatus box="[494,593,954,978]" pageId="18" pageNumber="19">Holotype</typeStatus>
<specimenCount box="[625,647,954,977]" pageId="18" pageNumber="19" type="male"></specimenCount>
,
<emphasis bold="true" box="[661,809,953,977]" pageId="18" pageNumber="19">
<collectingCountry box="[661,809,953,977]" name="South Africa" pageId="18" pageNumber="19">South Africa</collectingCountry>
</emphasis>
:
<collectingRegion box="[826,988,954,978]" country="South Africa" name="Western Cape" pageId="18" pageNumber="19">Cape Province</collectingRegion>
,
<quantity box="[1001,1071,954,978]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="km" value="10.0">10 km</quantity>
S of
</materialsCitation>
</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" pageId="18" pageNumber="19">
Citrusdal, Koomlandskloof,
<geoCoordinate box="[551,637,984,1008]" degrees="32" direction="south" minutes="40" orientation="latitude" pageId="18" pageNumber="19" precision="925" value="-32.666668">32°40'S</geoCoordinate>
,
<geoCoordinate box="[658,745,984,1008]" degrees="19" direction="east" minutes="01" orientation="longitude" pageId="18" pageNumber="19" precision="925" value="19.016666">19°01'E</geoCoordinate>
, Malaise-trap, at stream in low indigenous forest,
<date box="[429,560,1014,1038]" pageId="18" pageNumber="19" value="1994-10-05" valueMax="1994-10-08" valueMin="1994-10-05">58.X 1994</date>
, M. Söderlund [
<collectionCode box="[737,821,1014,1037]" country="Sweden" httpUri="http://grbio.org/cool/nzgn-b6sw" name="Lunds Universitet, Zoologiska Institutionen" pageId="18" pageNumber="19">ZMLU</collectionCode>
].
<typeStatus box="[840,938,1014,1038]" pageId="18" pageNumber="19">Paratype</typeStatus>
:
<specimenCount box="[950,985,1014,1038]" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
,
<collectingCountry box="[999,1126,1014,1038]" name="South Africa" pageId="18" pageNumber="19">Republic of</collectingCountry>
</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" box="[227,1126,1044,1068]" pageId="18" pageNumber="19">
<materialsCitation box="[227,1122,1044,1068]" latitude="-32.666668" location="Koomlandskloof" municipality="South Africa" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Western Cape">
<collectingMunicipality box="[227,374,1044,1068]" pageId="18" pageNumber="19">South Africa</collectingMunicipality>
:
<collectingRegion box="[390,556,1044,1068]" country="South Africa" name="Western Cape" pageId="18" pageNumber="19">Cape Province</collectingRegion>
,
<locationDeviation box="[572,821,1044,1068]" location="Citrusdal" pageId="18" pageNumber="19">
<quantity box="[572,645,1044,1068]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="km" value="10.0">10 km</quantity>
S of Citrusdal
</locationDeviation>
,
<location LSID="urn:lsid:plazi:treatment:382287BE1A4C6C09FF7A543FFBB96CCA:B55460731A4C6C0FFCDF534BFC1C6C13" box="[838,1020,1044,1068]" latitude="-32.666668" municipality="South Africa" name="Koomlandskloof" pageId="18" pageNumber="19" stateProvince="Western Cape">Koomlandskloof</location>
,
<geoCoordinate box="[1036,1122,1044,1068]" degrees="32" direction="south" minutes="40" orientation="latitude" pageId="18" pageNumber="19" precision="925" value="-32.666668">32°40'S</geoCoordinate>
</materialsCitation>
,
</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" box="[227,1127,1074,1098]" pageId="18" pageNumber="19">
<geoCoordinate box="[227,314,1074,1098]" degrees="19" direction="east" minutes="01" orientation="longitude" pageId="18" pageNumber="19" precision="925" value="19.016666">19°01'E</geoCoordinate>
, Malaise-trap, fynbos on mountain,
<date box="[753,891,1074,1098]" pageId="18" pageNumber="19" value="1994-10-05" valueMax="1994-10-07" valueMin="1994-10-05">57.X 1994</date>
, Michael Söderlund
</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" box="[227,328,1104,1127]" pageId="18" pageNumber="19">
[
<collectionCode box="[233,317,1104,1127]" country="Sweden" httpUri="http://grbio.org/cool/nzgn-b6sw" name="Lunds Universitet, Zoologiska Institutionen" pageId="18" pageNumber="19">ZMLU</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph blockId="18.[227,1127,953,1608]" pageId="18" pageNumber="19">
DESCRIPTION. Male (somewhat discolorated).
<emphasis box="[790,848,1134,1158]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
: frons black, shining; one pair of short postvertical setae; upper postocular setae black, slightly increasing in length upward; lateral and lower postoculars white; ventral postcranium covered with irregular white hairs; face silvery-white, 6 times as high as wide in middle,
</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" box="[227,1127,1254,1278]" pageId="18" pageNumber="19">nearly as wide as postpedicel; clypeus bulging; antennae as long as height of head,</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" pageId="18" pageNumber="19">
with yellow scape and pedicel and orange postpedicel; pedicel projected distally on inner side, with a crown of short black setae, 1-2 dorsal setae longer; postpedicel rounded, as long as high (1/1); arista-like stylus dorsal, with short hairs; length ratio of scape to pedicel to postpedicel to stylus (1
<superScript attach="left" box="[709,721,1370,1384]" fontSize="6" pageId="18" pageNumber="19">st</superScript>
and 2
<superScript attach="left" box="[787,804,1369,1383]" fontSize="6" pageId="18" pageNumber="19">nd</superScript>
segments), 5/6/8/6/39; palpus and proboscis short, yellow, covered with white hairs, proboscis also with a pair of black lateral setae.
</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" pageId="18" pageNumber="19">
<emphasis box="[265,342,1464,1488]" italics="true" pageId="18" pageNumber="19">Thorax</emphasis>
: pleura yellow; mesonotum orange, with median black stripe half as wide as surface between dorsocentral bristles, wider posteriorly; scutellum black dorsally; 6 dorsocentral setae with posterior pair shifted laterally; acrostichals absent; scutellum with 2 strong bristles and 2 short lateral hairs; proepisternum with
</paragraph>
<paragraph blockId="18.[227,1127,953,1608]" box="[227,441,1584,1608]" pageId="18" pageNumber="19">2 long yellow setae.</paragraph>
<paragraph blockId="19.[227,1127,264,380]" pageId="19" pageNumber="20">
<emphasis box="[265,316,265,288]" italics="true" pageId="19" pageNumber="20">Legs</emphasis>
including coxae yellow; tarsi brown from tip of basitarsus; fore and mid coxae with black hairs anteriorly and 4-7 black apical bristles of various length;
</paragraph>
<paragraph blockId="19.[227,1127,264,380]" pageId="19" pageNumber="20">hind coxa with one long black outer bristle above middle; fore leg without remarkable</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7164268" ID-Zenodo-Dep="7164268" box="[265,1127,1571,1593]" httpUri="https://zenodo.org/record/7164268/files/figure.png" pageId="19" pageNumber="20" startId="19.[265,306,1571,1593]" targetBox="[239,1114,411,1555]" targetPageId="19">
<paragraph blockId="19.[227,1127,1571,1648]" box="[265,1127,1571,1593]" pageId="19" pageNumber="20">
Figs 2328.
<taxonomicName box="[386,526,1572,1593]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[386,479,1572,1593]" italics="true" pageId="19" pageNumber="20">Tenuopus</emphasis>
spp.
</taxonomicName>
2325
<taxonomicName authority="Parent" authorityName="Parent" authorityYear="1934" box="[618,803,1572,1593]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis box="[618,731,1572,1593]" italics="true" pageId="19" pageNumber="20">T. cognatus</emphasis>
Parent
</taxonomicName>
; 2628
<taxonomicName authority="Grichanov, 2018" authorityName="Grichanov" authorityYear="2018" box="[898,1027,1571,1593]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="soderlundi" status="sp. n.">
<emphasis box="[898,1027,1571,1593]" italics="true" pageId="19" pageNumber="20">T. soderlundi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1034,1087,1572,1593]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
; 23,
</paragraph>
</caption>
<paragraph blockId="19.[227,1127,1571,1648]" pageId="19" pageNumber="20">26 head; 27 wing; 24, 28 hypopygium after maceration, right lateral view; 25 epandrial lobes and surstylus, right lateral view.</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">hairs and setae; fore tibia simple, with 1 short anterodorsal at base, 1-2 short apical setae; claws simple; mid femur with strong anterior preapical seta; mid tibia with 2-</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">3 anterior, 3 posterodorsal, 3-5 ventral, 4-5 apical setae; mid basitarsus with several short ventral setae; hind femur without long hairs, with 1 strong anterior preapical seta; hind tibia with 4-5 anterior, 2 anterodorsal, 2 posterodorsal, 3-4 apical setae.</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" box="[227,1126,417,441]" pageId="20" pageNumber="21">Femur, tibia and tarsomere (from first to fifth) length ratio: fore leg: 13/13/11/5/</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" box="[227,854,448,472]" pageId="20" pageNumber="21">3/2/2, mid leg: 16/21/14/6/4/2/2, hind leg: 19/27/8/9/5/3/2.</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" box="[265,1127,478,503]" pageId="20" pageNumber="21">
<emphasis box="[265,321,479,502]" italics="true" pageId="20" pageNumber="21">Wing</emphasis>
hyaline, veins yellow; subcosta very thin; ratio of part of costa between
</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">
R
<subScript attach="left" box="[245,272,520,534]" fontSize="6" pageId="20" pageNumber="21">2+3</subScript>
and R
<subScript attach="left" box="[342,369,520,534]" fontSize="6" pageId="20" pageNumber="21">4+5</subScript>
to that between R
<subScript attach="left" box="[564,591,520,534]" fontSize="6" pageId="20" pageNumber="21">4+5</subScript>
and M
<subScript attach="left" box="[667,676,520,534]" fontSize="6" pageId="20" pageNumber="21">1</subScript>
, 4/1; M
<subScript attach="left" box="[761,770,520,534]" fontSize="6" pageId="20" pageNumber="21">1</subScript>
with gentle arc to apex, reaching costa far before wing apex; M
<subScript attach="left" box="[578,587,551,565]" fontSize="6" pageId="20" pageNumber="21">2</subScript>
present as fold on membrane; crossvein
<emphasis box="[1066,1127,540,564]" italics="true" pageId="20" pageNumber="21">dm-m</emphasis>
</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">
straight; ratio of crossvein
<emphasis box="[521,582,570,594]" italics="true" pageId="20" pageNumber="21">dm-m</emphasis>
to apical part of M
<subScript attach="left" box="[797,824,582,596]" fontSize="6" pageId="20" pageNumber="21">1+2</subScript>
(fork-handle) to apical part of M
<subScript attach="left" box="[283,292,612,626]" fontSize="6" pageId="20" pageNumber="21">4</subScript>
, 2.5/6/11; anal vein foldlike, not reaching wing margin; anal angle obtuse;
</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">lower calypter yellow, with brown apex and pale setae; halter yellow, halter stem thin and long, with dorsal and ventral groups of short hairs distally.</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">
Abdomen mostly yellow, black setose; 1
<superScript attach="left" box="[727,739,689,703]" fontSize="6" pageId="20" pageNumber="21">st</superScript>
segment yellow, with white hairs ventrally; 5
<superScript attach="left" box="[352,365,719,733]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
segment mostly brown, 6
<superScript attach="left" box="[652,665,719,733]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
segment black; epandrium yellow-brown;
</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">surstylus and epandrial lobes brown; hypandrium relatively long, simple; phallus thin and simple; basoventral epandrial lobe reduced to 3 long apical setae at base of hypandrium; 1 long epandrial lobe distally with short setae at apex; 2-3 short setae between basoventral and distoventral lobes; surstylus short, about 1/3 as long as epandrium, strongly sclerotised, paw-shaped, with 4 apical processes; cercus yellow, half as long as epandrium, rounded, with short apical projection, covered with short hairs and several setae.</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">MEASUREMENTS (in mm). Body length 5-5.2; antenna length 1.1; wing length 4.2; wing width 1.4.</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" box="[265,470,1030,1054]" pageId="20" pageNumber="21">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="etymology">
<paragraph blockId="20.[227,1127,264,1269]" box="[265,1125,1061,1085]" pageId="20" pageNumber="21">ETYMOLOGY. The species is named after the Swedish entomologist, Dr. M.</paragraph>
<paragraph blockId="20.[227,1127,264,1269]" box="[227,828,1092,1116]" pageId="20" pageNumber="21">
<materialsCitation box="[227,825,1092,1116]" collectionCode="ZMLU" pageId="20" pageNumber="21" specimenCount="1">
Söderlund (
<collectionCode box="[351,432,1092,1116]" country="Sweden" httpUri="http://grbio.org/cool/nzgn-b6sw" name="Lunds Universitet, Zoologiska Institutionen" pageId="20" pageNumber="21">ZMLU</collectionCode>
), the collector of the
<typeStatus box="[657,703,1092,1116]" pageId="20" pageNumber="21">type</typeStatus>
specimens
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[265,610,1122,1146]" pageId="20" pageNumber="21" type="distribution">
<paragraph blockId="20.[227,1127,264,1269]" box="[265,610,1122,1146]" pageId="20" pageNumber="21">
DISTRIBUTION.
<collectingCountry box="[465,606,1122,1146]" name="South Africa" pageId="20" pageNumber="21">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="diagnosis">
<paragraph blockId="20.[227,1127,264,1269]" pageId="20" pageNumber="21">
DIAGNOSIS.
<taxonomicName authority="Grichanov, 2018" authorityName="Grichanov" authorityYear="2018" box="[422,565,1153,1177]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="soderlundi" status="sp. n.">
<emphasis box="[422,565,1153,1177]" italics="true" pageId="20" pageNumber="21">T. soderlundi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[572,630,1154,1177]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
is the smallest fly in the genus and belongs to a group of species with uniseriate acrostichals or without acrostichals, being the closest to
<taxonomicName authorityName="Parent" authorityYear="1934" box="[337,463,1215,1238]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis box="[337,463,1215,1238]" italics="true" pageId="20" pageNumber="21">T. cognatus</emphasis>
</taxonomicName>
, differing from the latter in wing vein M
<subScript attach="left" box="[913,922,1226,1240]" fontSize="6" pageId="20" pageNumber="21">4</subScript>
more than 4 times longer than
<emphasis box="[354,415,1245,1269]" italics="true" pageId="20" pageNumber="21">dm-m</emphasis>
; abdomen with 2
<superScript attach="none" box="[601,618,1240,1254]" fontSize="6" pageId="20" pageNumber="21">nd</superScript>
4
<superScript attach="left" box="[645,658,1240,1254]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
segments entirely yellow (see key above).
</paragraph>
</subSubSection>
</treatment>
</document>