treatments-xml/data/D4/20/87/D42087ADFF8E88782A2DE1B1C0A4FD23.xml

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<mods:title>Phylogenetic classification of the family Terebridae (Neogastropoda: Conoidea)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fedosov, Alexander E</mods:namePart>
<mods:affiliation>A. N. Severtsov Institute of Ecology and Evolution of Russian Academy of Sciences, Leninskiy Prospect, 33, Moscow 119071, Russia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Malcolm, Gavin</mods:namePart>
<mods:affiliation>Bird Hill, Barnes Lane, Milford on Sea, Hampshire, UK</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Terryn, Yves</mods:namePart>
<mods:affiliation>Kapiteinstraat 27, 9000 Gent, Belgium</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gorson, Juliette</mods:namePart>
<mods:affiliation>Department of Chemistry, Hunter College Belfer Research Center, New York, NY 10021, USA; 5 Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA; 6 Program in Biology, Graduate Center, City University of New York, New York, NY 10016, USA;</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Modica, Maria Vittoria</mods:namePart>
<mods:affiliation>Stazione Zoologica Anton Dohrn, Naples, Italy</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Holford, Mandë</mods:namePart>
<mods:affiliation>Department of Chemistry, Hunter College Belfer Research Center, New York, NY 10021, USA; 5 Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA; 6 Program in Biology, Graduate Center, City University of New York, New York, NY 10016, USA; Programs in Biology, Chemistry &amp; Biochemistry, Graduate Center, City University of New York, New York, NY 10016, USA; 9 Department of Biochemistry, Weill Cornell Medicine, Cornell University, New York, NY 10021, USA.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Puillandre, Nicolas</mods:namePart>
<mods:affiliation>Institut Systématique Evolution Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, 57 rue Cuvier, CP 26, 75005 Paris, France</mods:affiliation>
<mods:nameIdentifier type="email">nicolaspuillandre@gmail.com</mods:nameIdentifier>
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<mods:title>Journal of Molluscan Studies The Malacological Society of London</mods:title>
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<mods:date>2019</mods:date>
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<mods:number>2020-01-09</mods:number>
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<mods:number>85</mods:number>
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<subSubSection box="[258,706,670,692]" pageId="8" pageNumber="9" type="nomenclature">
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<heading box="[258,706,670,692]" level="4" pageId="8" pageNumber="9">
<emphasis bold="true" box="[258,706,670,692]" pageId="8" pageNumber="9">
Genus 
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<emphasis bold="true" box="[336,455,671,692]" italics="true" pageId="8" pageNumber="9">Partecosta</emphasis>
Dance &amp; Eames, 1966
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</emphasis>
</heading>
</paragraph>
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<subSubSection box="[438,548,697,719]" pageId="8" pageNumber="9" type="description">
<paragraph blockId="8.[258,706,670,719]" box="[438,548,697,719]" pageId="8" pageNumber="9">
(
<figureCitation box="[445,495,697,719]" captionStart="Figure 3" captionStartId="7.[158,223,1668,1687]" captionTargetBox="[161,1475,180,1639]" captionTargetId="figure-6@7.[161,1476,179,1639]" captionTargetPageId="7" captionText="Figure 3. Subfamily Pervicaciinae A. Duplicaria duplicata, MNHN-IM-2009-29454, WESTERN AUSTRALIA 2011 Stn WB32, 33◦33'S, 115◦04'E, 5–15 m, 26.7 mm. B. Duplicaria tricincta, MNHN-IM-2013-5638, PAPUA NIUGINI Stn PD67, 05◦15.5'S, 145◦46.8'E, 2–6 m, 6.9 mm. C. Duplicaria tristis lectotype, NHMUK 1979115, ‘Seas of Japan’, 17.7 mm. D. Duplicaria brevicula, MNHN-IM-2013-66140, off Namibe, S Angola, 40–60 m, 12.9 mm. E. Duplicaria bernardi radula, MNHN-IM-2009-10908, Australia, 26◦56'607''S; 153◦23'813''E, shell broken. F. Terebra fuscobasis lectotype, NHMUK 1873.7.5.8/1, Persian Gulf, 11.3 mm. G. Terebra nassoides lectotype, NHMUK 1968251/1, Red Sea, 13.5 mm. H. Partecosta varia, MNHN-IM-2013-52342, ATIMO VATAE Stn TM27, 24◦56.4'S, 47◦06.9'E, 0–1 m, 10.4 mm. I. Partecosta sandrinae, MNHN-IM-2013-52359, INHACA 2011 Stn MM7 PL5, 26◦03.7'S, 32◦54.1'E, 0–1 m, 8.8 mm. J. Partecosta n. sp. aff fuscolutea radula, MNHN-IM-2009-10133, ATIMO VATAE Stn BS06, 25◦26.8'S, 44◦54.9'E, 0–27 m, 6.9 mm. K. Partecosta bozzettii n. sp. paratype, MNHN-IM-2009-10162, ATIMO VATAE Stn TP29, 25◦03.7–03.8'S, 46◦57.7'E, 3–4 m, 12.4 mm. L. Partecosta trilineata, MNHN-IM-2009-10164, ATIMO VATAE Stn TP24, 25◦03.7–03.8'S, 46◦57.6–57.7'E, 2–7 m, 8.2 mm. M. Partecosta macleani, MNHN-IM-2009-10115 ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 8.9 mm. N, O. Partecosta macleani radula, MNHN-IM- 2009-10111, ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 12 mm." figureDoi="http://doi.org/10.5281/zenodo.4469850" httpUri="https://zenodo.org/record/4469850/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
F–O)
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="type_taxon">
<paragraph blockId="8.[151,789,750,799]" pageId="8" pageNumber="9">
<emphasis box="[151,259,751,772]" italics="true" pageId="8" pageNumber="9">
<typeStatus box="[151,191,751,772]" pageId="8" pageNumber="9">Type</typeStatus>
species:
</emphasis>
<treatmentCitation author="Dance &amp; Eames" box="[269,686,750,772]" pageId="8" pageNumber="9" year="1966">
<taxonomicName authority="Dance &amp; Eames, 1966" authorityName="Dance &amp; Eames" authorityYear="1966" box="[269,686,750,772]" class="Gastropoda" family="Terebridae" genus="Strioterebrum" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="wilkinsi">
<treatmentCitationGroup box="[269,451,750,771]" pageId="8" pageNumber="9">
<emphasis box="[269,451,750,771]" italics="true" pageId="8" pageNumber="9">Strioterebrum wilkinsi</emphasis>
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Dance &amp; Eames, 1966
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(= 
<treatmentCitation author="E.A. Smith" pageId="8" pageNumber="9" year="1877">
<taxonomicName authority="E. A. Smith, 1877" authorityName="E. A. Smith" authorityYear="1877" class="Gastropoda" family="Terebridae" genus="Terebra" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fuscobasis">
<emphasis italics="true" pageId="8" pageNumber="9">Terebra fuscobasis</emphasis>
E.A. Smith, 1877
</taxonomicName>
</treatmentCitation>
); OD.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="discussion">
<paragraph blockId="8.[151,790,830,959]" pageId="8" pageNumber="9">
<emphasis box="[151,237,830,851]" italics="true" pageId="8" pageNumber="9">Definition:</emphasis>
Includes species with combination of conchological, anatomical and distributional characteristics closely comparable with 
<taxonomicName authorityName="Dance &amp; Eames" authorityYear="1966" box="[207,394,883,904]" class="Gastropoda" family="Terebridae" genus="Strioterebrum" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="wilkinsi">
<emphasis box="[207,394,883,904]" italics="true" pageId="8" pageNumber="9">Strioterebrum wilkinsi</emphasis>
</taxonomicName>
, including all species in clade F2 of 
<bibRefCitation author="MODICA, M. V. &amp; GORSON, J. &amp; FEDOSOV, A. E. &amp; MALCOLM, G. &amp; TERRYN, Y. &amp; PUILLANDRE, N. &amp; HOLFORD, M." box="[151,337,910,932]" journalOrPublisher="Systematic Biology" pageId="8" pageNumber="9" refId="ref29363" refString="MODICA, M. V., GORSON, J., FEDOSOV, A. E., MALCOLM, G., TERRYN, Y., PUILLANDRE, N. &amp; HOLFORD, M. 2019. Macroevolutionary analyses suggest environmental factors, not venom apparatus, play key role in Terebridae marine snail diversification. Systematic Biology. DOI: https: // doi. org / 10.1093 / sysbio / syz 059." title="Macroevolutionary analyses suggest environmental factors, not venom apparatus, play key role in Terebridae marine snail diversification" type="journal article" year="2019">
Modica 
<emphasis box="[232,273,910,932]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2019)
</bibRefCitation>
and any species proven to be a member of this clade.
</paragraph>
</subSubSection>
<subSubSection box="[151,698,990,1012]" pageId="8" pageNumber="9" type="diagnosis">
<paragraph blockId="8.[151,698,990,1012]" box="[151,698,990,1012]" pageId="8" pageNumber="9">
<emphasis box="[151,232,991,1012]" italics="true" pageId="8" pageNumber="9">Diagnosis</emphasis>
: Diagnostic nucleotide combinations in 
<tableCitation box="[619,692,990,1012]" captionStart="Table 2" captionStartId="8.[153,209,151,170]" captionTargetBox="[153,784,233,426]" captionText="Table 2. Genera Duplicaria and Patecosta: diagnostic combinations of nucleotides in COI alignment." httpUri="http://table.plazi.org/id/08F66633FF8E88782BB6E3B8C5B2FF49" pageId="8" pageNumber="9" tableUuid="08F66633FF8E88782BB6E3B8C5B2FF49">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="description">
<paragraph blockId="8.[151,790,1043,1252]" pageId="8" pageNumber="9">
<emphasis box="[151,191,1043,1064]" italics="true" pageId="8" pageNumber="9">Shell</emphasis>
: Small, 
<quantity box="[276,386,1043,1065]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="mm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 mm</quantity>
, pale or tan, often with spiral bands and dark basal area. Protoconch 1.5–3 whorls. Predominant sculpture of axial ribs spanning whorl height, resulting in even, slightly convex whorl outline, typically interrupted by depression or narrow groove forming subsutural row of nodules. Spiral sculpture indistinct or of fine regular striae. Siphonal canal very short, stout, indistinctly or shallowly notched, with strong fasciole. Aperture wide, pyriform.
</paragraph>
<paragraph blockId="8.[151,789,1283,1385]" pageId="8" pageNumber="9">
<emphasis box="[151,224,1283,1304]" italics="true" pageId="8" pageNumber="9">Anatomy</emphasis>
: Rhynchodeal introvert small; salivary glands and radular sac present; radula of solid curved marginal teeth attached to strong membrane. Venom gland, proboscis and accessory proboscis structure absent.
</paragraph>
</subSubSection>
<subSubSection box="[151,705,1416,1438]" pageId="8" pageNumber="9" type="distribution">
<paragraph blockId="8.[151,705,1416,1438]" box="[151,705,1416,1438]" pageId="8" pageNumber="9">
<emphasis box="[151,256,1416,1437]" italics="true" pageId="8" pageNumber="9">Distribution:</emphasis>
Indian Ocean; intertidal and shallow subtidal.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="discussion">
<paragraph blockId="8.[151,790,1470,1961]" pageId="8" pageNumber="9">
<emphasis box="[151,222,1470,1491]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
: Three clusters with varying sculpture patterns can be recognized:
</paragraph>
<paragraph blockId="8.[151,790,1470,1961]" pageId="8" pageNumber="9">
<taxonomicName baseAuthorityName="E. A. Smith" baseAuthorityYear="1877" box="[175,349,1523,1545]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fuscobasis">
<emphasis box="[175,349,1523,1545]" italics="true" pageId="8" pageNumber="9">Partecosta fuscobasis</emphasis>
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group: sculpture of strong ribs and clearly delineated subsutural band forming distinct nodules at intersections with ribs (
<figureCitation box="[310,365,1576,1598]" captionStart="Figure 3" captionStartId="7.[158,223,1668,1687]" captionTargetBox="[161,1475,180,1639]" captionTargetId="figure-6@7.[161,1476,179,1639]" captionTargetPageId="7" captionText="Figure 3. Subfamily Pervicaciinae A. Duplicaria duplicata, MNHN-IM-2009-29454, WESTERN AUSTRALIA 2011 Stn WB32, 33◦33'S, 115◦04'E, 5–15 m, 26.7 mm. B. Duplicaria tricincta, MNHN-IM-2013-5638, PAPUA NIUGINI Stn PD67, 05◦15.5'S, 145◦46.8'E, 2–6 m, 6.9 mm. C. Duplicaria tristis lectotype, NHMUK 1979115, ‘Seas of Japan’, 17.7 mm. D. Duplicaria brevicula, MNHN-IM-2013-66140, off Namibe, S Angola, 40–60 m, 12.9 mm. E. Duplicaria bernardi radula, MNHN-IM-2009-10908, Australia, 26◦56'607''S; 153◦23'813''E, shell broken. F. Terebra fuscobasis lectotype, NHMUK 1873.7.5.8/1, Persian Gulf, 11.3 mm. G. Terebra nassoides lectotype, NHMUK 1968251/1, Red Sea, 13.5 mm. H. Partecosta varia, MNHN-IM-2013-52342, ATIMO VATAE Stn TM27, 24◦56.4'S, 47◦06.9'E, 0–1 m, 10.4 mm. I. Partecosta sandrinae, MNHN-IM-2013-52359, INHACA 2011 Stn MM7 PL5, 26◦03.7'S, 32◦54.1'E, 0–1 m, 8.8 mm. J. Partecosta n. sp. aff fuscolutea radula, MNHN-IM-2009-10133, ATIMO VATAE Stn BS06, 25◦26.8'S, 44◦54.9'E, 0–27 m, 6.9 mm. K. Partecosta bozzettii n. sp. paratype, MNHN-IM-2009-10162, ATIMO VATAE Stn TP29, 25◦03.7–03.8'S, 46◦57.7'E, 3–4 m, 12.4 mm. L. Partecosta trilineata, MNHN-IM-2009-10164, ATIMO VATAE Stn TP24, 25◦03.7–03.8'S, 46◦57.6–57.7'E, 2–7 m, 8.2 mm. M. Partecosta macleani, MNHN-IM-2009-10115 ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 8.9 mm. N, O. Partecosta macleani radula, MNHN-IM- 2009-10111, ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 12 mm." figureDoi="http://doi.org/10.5281/zenodo.4469850" httpUri="https://zenodo.org/record/4469850/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
F–I). Closely comparable with New World genus 
<taxonomicName authorityName="Fedosov &amp; Malcolm &amp; Terryn &amp; Gorson &amp; Modica &amp; Holford &amp; Puillandre" authorityYear="2019" box="[216,301,1603,1624]" class="Gastropoda" family="Terebridae" genus="Neoterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis box="[216,301,1603,1624]" italics="true" pageId="8" pageNumber="9">Neoterebra</emphasis>
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, but can be differentiated by smaller size, wider aperture, usually with wide anterior notch, and solid curved radular teeth.
</paragraph>
<paragraph blockId="8.[151,790,1470,1961]" pageId="8" pageNumber="9">
<taxonomicName baseAuthorityName="Bozzetti" baseAuthorityYear="2008" box="[185,346,1691,1713]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="trilineata">
<emphasis box="[185,346,1691,1713]" italics="true" pageId="8" pageNumber="9">Partecosta trilineata</emphasis>
</taxonomicName>
group: glossy shells sculptured by strong continuous ribs; no distinct spiral sculpture or subsutural band (
<figureCitation box="[158,266,1744,1766]" captionStart="Figure 3" captionStartId="7.[158,223,1668,1687]" captionTargetBox="[161,1475,180,1639]" captionTargetId="figure-6@7.[161,1476,179,1639]" captionTargetPageId="7" captionText="Figure 3. Subfamily Pervicaciinae A. Duplicaria duplicata, MNHN-IM-2009-29454, WESTERN AUSTRALIA 2011 Stn WB32, 33◦33'S, 115◦04'E, 5–15 m, 26.7 mm. B. Duplicaria tricincta, MNHN-IM-2013-5638, PAPUA NIUGINI Stn PD67, 05◦15.5'S, 145◦46.8'E, 2–6 m, 6.9 mm. C. Duplicaria tristis lectotype, NHMUK 1979115, ‘Seas of Japan’, 17.7 mm. D. Duplicaria brevicula, MNHN-IM-2013-66140, off Namibe, S Angola, 40–60 m, 12.9 mm. E. Duplicaria bernardi radula, MNHN-IM-2009-10908, Australia, 26◦56'607''S; 153◦23'813''E, shell broken. F. Terebra fuscobasis lectotype, NHMUK 1873.7.5.8/1, Persian Gulf, 11.3 mm. G. Terebra nassoides lectotype, NHMUK 1968251/1, Red Sea, 13.5 mm. H. Partecosta varia, MNHN-IM-2013-52342, ATIMO VATAE Stn TM27, 24◦56.4'S, 47◦06.9'E, 0–1 m, 10.4 mm. I. Partecosta sandrinae, MNHN-IM-2013-52359, INHACA 2011 Stn MM7 PL5, 26◦03.7'S, 32◦54.1'E, 0–1 m, 8.8 mm. J. Partecosta n. sp. aff fuscolutea radula, MNHN-IM-2009-10133, ATIMO VATAE Stn BS06, 25◦26.8'S, 44◦54.9'E, 0–27 m, 6.9 mm. K. Partecosta bozzettii n. sp. paratype, MNHN-IM-2009-10162, ATIMO VATAE Stn TP29, 25◦03.7–03.8'S, 46◦57.7'E, 3–4 m, 12.4 mm. L. Partecosta trilineata, MNHN-IM-2009-10164, ATIMO VATAE Stn TP24, 25◦03.7–03.8'S, 46◦57.6–57.7'E, 2–7 m, 8.2 mm. M. Partecosta macleani, MNHN-IM-2009-10115 ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 8.9 mm. N, O. Partecosta macleani radula, MNHN-IM- 2009-10111, ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 12 mm." figureDoi="http://doi.org/10.5281/zenodo.4469850" httpUri="https://zenodo.org/record/4469850/files/figure.png" pageId="8" pageNumber="9">Fig. 3K, L</figureCitation>
). Very similar sympatric forms are known in genus 
<taxonomicName authorityName="Bartsch" authorityYear="1923" box="[151,261,1771,1792]" class="Gastropoda" family="Terebridae" genus="Punctoterebra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis box="[151,261,1771,1792]" italics="true" pageId="8" pageNumber="9">Punctoterebra</emphasis>
</taxonomicName>
(i.e. 
<taxonomicName authority="Bozzetti, 2015" authorityName="Bozzetti" authorityYear="2015" box="[317,666,1771,1793]" class="Gastropoda" family="Terebridae" genus="Punctoterebra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="solangeae">
<emphasis box="[317,512,1771,1792]" italics="true" pageId="8" pageNumber="9">Punctoterebra solangeae</emphasis>
Bozzetti, 2015
</taxonomicName>
group) that can only be reliably differentiated by anatomical characters (radula lacking in 
<taxonomicName authorityName="Bozzetti" authorityYear="2015" box="[286,388,1824,1846]" class="Gastropoda" family="Terebridae" genus="Punctoterebra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="solangeae">
<emphasis box="[286,299,1825,1846]" italics="true" pageId="8" pageNumber="9">P</emphasis>
. 
<emphasis box="[313,388,1824,1845]" italics="true" pageId="8" pageNumber="9">solangeae</emphasis>
</taxonomicName>
; of solid curved teeth in 
<taxonomicName authorityName="Dance &amp; Eames" authorityYear="1966" box="[629,711,1825,1846]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis box="[629,711,1825,1846]" italics="true" pageId="8" pageNumber="9">Partecosta</emphasis>
</taxonomicName>
spp.).
</paragraph>
<paragraph blockId="8.[151,790,1470,1961]" lastBlockId="8.[837,1474,152,201]" pageId="8" pageNumber="9">
<taxonomicName baseAuthorityName="Bratcher" baseAuthorityYear="1988" box="[178,335,1859,1881]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="macleani">
<emphasis box="[178,335,1859,1881]" italics="true" pageId="8" pageNumber="9">Partecosta macleani</emphasis>
</taxonomicName>
group: flattened whorl outline, very short robust siphonal canal; very weak sculpture of rounded, indistinct subsutural nodules extended into even less distinct axial folds on whorl periphery (
<figureCitation box="[257,332,1939,1961]" captionStart="Figure 3" captionStartId="7.[158,223,1668,1687]" captionTargetBox="[161,1475,180,1639]" captionTargetId="figure-6@7.[161,1476,179,1639]" captionTargetPageId="7" captionText="Figure 3. Subfamily Pervicaciinae A. Duplicaria duplicata, MNHN-IM-2009-29454, WESTERN AUSTRALIA 2011 Stn WB32, 33◦33'S, 115◦04'E, 5–15 m, 26.7 mm. B. Duplicaria tricincta, MNHN-IM-2013-5638, PAPUA NIUGINI Stn PD67, 05◦15.5'S, 145◦46.8'E, 2–6 m, 6.9 mm. C. Duplicaria tristis lectotype, NHMUK 1979115, ‘Seas of Japan’, 17.7 mm. D. Duplicaria brevicula, MNHN-IM-2013-66140, off Namibe, S Angola, 40–60 m, 12.9 mm. E. Duplicaria bernardi radula, MNHN-IM-2009-10908, Australia, 26◦56'607''S; 153◦23'813''E, shell broken. F. Terebra fuscobasis lectotype, NHMUK 1873.7.5.8/1, Persian Gulf, 11.3 mm. G. Terebra nassoides lectotype, NHMUK 1968251/1, Red Sea, 13.5 mm. H. Partecosta varia, MNHN-IM-2013-52342, ATIMO VATAE Stn TM27, 24◦56.4'S, 47◦06.9'E, 0–1 m, 10.4 mm. I. Partecosta sandrinae, MNHN-IM-2013-52359, INHACA 2011 Stn MM7 PL5, 26◦03.7'S, 32◦54.1'E, 0–1 m, 8.8 mm. J. Partecosta n. sp. aff fuscolutea radula, MNHN-IM-2009-10133, ATIMO VATAE Stn BS06, 25◦26.8'S, 44◦54.9'E, 0–27 m, 6.9 mm. K. Partecosta bozzettii n. sp. paratype, MNHN-IM-2009-10162, ATIMO VATAE Stn TP29, 25◦03.7–03.8'S, 46◦57.7'E, 3–4 m, 12.4 mm. L. Partecosta trilineata, MNHN-IM-2009-10164, ATIMO VATAE Stn TP24, 25◦03.7–03.8'S, 46◦57.6–57.7'E, 2–7 m, 8.2 mm. M. Partecosta macleani, MNHN-IM-2009-10115 ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 8.9 mm. N, O. Partecosta macleani radula, MNHN-IM- 2009-10111, ATIMO VATAE Stn TP19, 25◦04.4–04.7'S, 46◦55.3–56.3'E, 16–26 m, 12 mm." figureDoi="http://doi.org/10.5281/zenodo.4469850" httpUri="https://zenodo.org/record/4469850/files/figure.png" pageId="8" pageNumber="9">Fig. 3M</figureCitation>
). Some species of 
<taxonomicName authorityName="H. Adams &amp; A. Adams" authorityYear="1858" box="[506,573,1939,1960]" class="Gastropoda" family="Terebridae" genus="Hastula" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis box="[506,573,1939,1960]" italics="true" pageId="8" pageNumber="9">Hastula</emphasis>
</taxonomicName>
are close to 
<taxonomicName baseAuthorityName="Bratcher" baseAuthorityYear="1988" box="[691,782,1939,1961]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="macleani">
<emphasis box="[691,782,1939,1961]" italics="true" pageId="8" pageNumber="9">P. macleani</emphasis>
</taxonomicName>
, but are commonly larger, have polished surface and retain distinct ribbing at least on adapical portions of whorls.
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[837,970,232,254]" pageId="8" pageNumber="9">
<emphasis box="[837,970,232,254]" italics="true" pageId="8" pageNumber="9">Included species:</emphasis>
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1300,257,281]" pageId="8" pageNumber="9">
<taxonomicName authority="(Bozzetti, 2008)" baseAuthorityName="Bozzetti" baseAuthorityYear="2008" box="[861,1195,259,281]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="albofuscata">
<emphasis box="[861,1041,259,280]" italics="true" pageId="8" pageNumber="9">Partecosta albofuscata</emphasis>
(Bozzetti, 2008)
</taxonomicName>
<specimenCount box="[1195,1229,257,280]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1195,1202,257,269]" fontSize="5" pageId="8" pageNumber="9">1</superScript>
<taxonomicNameLabel box="[1209,1292,259,281]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1297,284,307]" pageId="8" pageNumber="9">
<taxonomicName authority="Alexander E Fedosov &amp; Gavin Malcolm &amp; Yves Terryn &amp; Juliette Gorson &amp; Maria Vittoria Modica &amp; Mandë Holford &amp; Nicolas Puillandre, 2019" authorityName="Fedosov &amp; Malcolm &amp; Terryn &amp; Gorson &amp; Modica &amp; Holford &amp; Puillandre" authorityYear="2019" box="[861,949,285,307]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bozzettii" status="n. sp.">
<emphasis box="[861,949,285,307]" italics="true" pageId="8" pageNumber="9">P. bozzettii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[956,1007,286,307]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
Malcolm, Terryn &amp; Fedosov 
<superScript attach="left" box="[1283,1290,284,296]" fontSize="5" pageId="8" pageNumber="9">1</superScript>
;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1208,311,334]" pageId="8" pageNumber="9">
<taxonomicName authority="Bozzetti, 20082" authorityName="Bozzetti" authorityYear="2008" box="[861,1111,311,334]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fuscolutea">
<emphasis box="[861,960,312,334]" italics="true" pageId="8" pageNumber="9">P. fuscolutea</emphasis>
Bozzetti, 
<specimenCount box="[1056,1111,311,333]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1056,1111,311,333]" fontSize="5" pageId="8" pageNumber="9">20082</superScript>
</specimenCount>
</taxonomicName>
<taxonomicNameLabel box="[1118,1200,312,334]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1216,337,361]" pageId="8" pageNumber="9">
<taxonomicName authority="(Bratcher, 1988)" baseAuthorityName="Bratcher" baseAuthorityYear="1988" box="[861,1112,339,361]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="macleani">
<emphasis box="[861,952,339,360]" italics="true" pageId="8" pageNumber="9">P. macleani</emphasis>
(Bratcher, 1988)
</taxonomicName>
<specimenCount box="[1112,1145,337,360]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1112,1119,337,349]" fontSize="5" pageId="8" pageNumber="9">1</superScript>
<taxonomicNameLabel box="[1126,1208,339,361]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1196,364,388]" pageId="8" pageNumber="9">
<taxonomicName authority="(Hinds, 1844)" baseAuthorityName="Hinds" baseAuthorityYear="1844" box="[861,1091,365,388]" class="Gastropoda" family="Terebridae" genus="Terebra" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="nassoides">
<emphasis box="[861,956,365,387]" italics="true" pageId="8" pageNumber="9">P. nassoides</emphasis>
(Hinds, 1844)
</taxonomicName>
<specimenCount box="[1091,1125,364,387]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1091,1098,364,376]" fontSize="5" pageId="8" pageNumber="9">1</superScript>
<taxonomicNameLabel box="[1105,1188,365,387]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1199,391,414]" pageId="8" pageNumber="9">
<taxonomicName authority="(Aubry, 2008)" baseAuthorityName="Aubry" baseAuthorityYear="2008" box="[861,1094,392,414]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="sandrinae">
<emphasis box="[861,959,392,414]" italics="true" pageId="8" pageNumber="9">P. sandrinae</emphasis>
(Aubry, 2008)
</taxonomicName>
<specimenCount box="[1095,1128,391,414]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1095,1102,391,403]" fontSize="5" pageId="8" pageNumber="9">1</superScript>
<taxonomicNameLabel box="[1109,1191,392,414]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1213,418,441]" pageId="8" pageNumber="9">
<taxonomicName authority="(Bozzetti, 2008)" baseAuthorityName="Bozzetti" baseAuthorityYear="2008" box="[861,1109,419,441]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="trilineata">
<emphasis box="[861,954,419,440]" italics="true" pageId="8" pageNumber="9">P. trilineata</emphasis>
(Bozzetti, 2008)
</taxonomicName>
<specimenCount box="[1109,1142,418,440]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1109,1116,418,430]" fontSize="5" pageId="8" pageNumber="9">1</superScript>
<taxonomicNameLabel box="[1123,1206,419,441]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1181,444,468]" pageId="8" pageNumber="9">
<taxonomicName authority="(Bozzetti, 2008)" baseAuthorityName="Bozzetti" baseAuthorityYear="2008" box="[861,1077,445,468]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="varia">
<emphasis box="[861,922,446,467]" italics="true" pageId="8" pageNumber="9">P. varia</emphasis>
(Bozzetti, 2008)
</taxonomicName>
<specimenCount box="[1077,1110,444,467]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1077,1084,444,456]" fontSize="5" pageId="8" pageNumber="9">1</superScript>
<taxonomicNameLabel box="[1091,1174,445,467]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1251,471,494]" pageId="8" pageNumber="9">
<taxonomicName authority="(E. A. Smith, 1877)" baseAuthorityName="E. A. Smith" baseAuthorityYear="1877" box="[861,1148,472,494]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fuscobasis">
<emphasis box="[861,962,472,494]" italics="true" pageId="8" pageNumber="9">P. fuscobasis</emphasis>
(E.A. Smith, 1877)
</taxonomicName>
<specimenCount box="[1147,1180,471,494]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1147,1154,471,483]" fontSize="5" pageId="8" pageNumber="9">2</superScript>
<taxonomicNameLabel box="[1161,1244,472,494]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1257,497,521]" pageId="8" pageNumber="9">
<taxonomicName authority="(E. A. Smith, 1877)" baseAuthorityName="E. A. Smith" baseAuthorityYear="1877" box="[861,1154,499,521]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fuscocincta">
<emphasis box="[861,968,499,520]" italics="true" pageId="8" pageNumber="9">P. fuscocincta</emphasis>
(E.A. Smith, 1877)
</taxonomicName>
<specimenCount box="[1153,1186,497,520]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1153,1160,497,509]" fontSize="5" pageId="8" pageNumber="9">2</superScript>
<taxonomicNameLabel box="[1167,1250,499,521]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1288,524,548]" pageId="8" pageNumber="9">
<taxonomicName authority="(Terryn &amp; Rosado, 2011)" baseAuthorityName="Terryn &amp; Rosado" baseAuthorityYear="2011" box="[861,1183,525,548]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="herosae">
<emphasis box="[861,939,525,547]" italics="true" pageId="8" pageNumber="9">P. herosae</emphasis>
(Terryn &amp; Rosado, 2011)
</taxonomicName>
<specimenCount box="[1183,1217,524,547]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1183,1190,524,536]" fontSize="5" pageId="8" pageNumber="9">2</superScript>
<taxonomicNameLabel box="[1197,1280,525,547]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1275,551,574]" pageId="8" pageNumber="9">
<taxonomicName authority="(E. A. Smith, 1877)" baseAuthorityName="E. A. Smith" baseAuthorityYear="1877" box="[861,1171,552,574]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="macandrewii">
<emphasis box="[861,985,552,574]" italics="true" pageId="8" pageNumber="9">P. macandrewii</emphasis>
(E.A. Smith, 1877)
</taxonomicName>
<specimenCount box="[1171,1204,551,574]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1171,1178,551,563]" fontSize="5" pageId="8" pageNumber="9">2</superScript>
<taxonomicNameLabel box="[1185,1268,552,574]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1222,577,601]" pageId="8" pageNumber="9">
<taxonomicName authority="(Thiele, 1925)" baseAuthorityName="Thiele" baseAuthorityYear="1925" box="[861,1118,579,601]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="padangensis">
<emphasis box="[861,978,579,600]" italics="true" pageId="8" pageNumber="9">P. padangensis</emphasis>
(Thiele, 1925)
</taxonomicName>
<specimenCount box="[1118,1151,577,600]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1118,1125,577,589]" fontSize="5" pageId="8" pageNumber="9">2</superScript>
<taxonomicNameLabel box="[1132,1214,579,601]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1229,604,628]" pageId="8" pageNumber="9">
<taxonomicName authority="(E. A. Smith, 1873)" baseAuthorityName="E. A. Smith" baseAuthorityYear="1873" box="[861,1125,605,628]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="tantilla">
<emphasis box="[861,940,605,627]" italics="true" pageId="8" pageNumber="9">P. tantilla</emphasis>
(E.A. Smith, 1873)
</taxonomicName>
<specimenCount box="[1125,1158,604,627]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1125,1132,604,616]" fontSize="5" pageId="8" pageNumber="9">2</superScript>
<taxonomicNameLabel box="[1139,1222,605,627]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1163,631,654]" pageId="8" pageNumber="9">
<taxonomicName authority="(Hinds, 1844)" baseAuthorityName="Hinds" baseAuthorityYear="1844" box="[861,1064,632,654]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="tenera">
<emphasis box="[861,928,633,654]" italics="true" pageId="8" pageNumber="9">P. tenera</emphasis>
(Hinds, 1844)
</taxonomicName>
<specimenCount box="[1064,1097,631,654]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1064,1071,631,643]" fontSize="5" pageId="8" pageNumber="9">3</superScript>
<taxonomicNameLabel box="[1078,1159,632,654]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
;
</paragraph>
<paragraph blockId="8.[837,1300,232,681]" box="[861,1160,658,681]" pageId="8" pageNumber="9">
<taxonomicName authority="(Aubry, 1991)" baseAuthorityName="Aubry" baseAuthorityYear="1991" box="[861,1062,659,681]" class="Gastropoda" family="Terebridae" genus="Partecosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="veliae">
<emphasis box="[861,926,659,680]" italics="true" pageId="8" pageNumber="9">P. veliae</emphasis>
(Aubry, 1991)
</taxonomicName>
<specimenCount box="[1061,1094,658,680]" pageId="8" pageNumber="9" type="generic">
<superScript attach="left" box="[1061,1068,658,670]" fontSize="5" pageId="8" pageNumber="9">3</superScript>
<taxonomicNameLabel box="[1075,1156,659,681]" pageId="8" pageNumber="9">n. comb</taxonomicNameLabel>
</specimenCount>
.
</paragraph>
</subSubSection>
</treatment>
</document>