treatments-xml/data/19/BB/F8/19BBF8B07B5453979247DFD82C748143.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.881.34821" ID-GBIF-Dataset="818727b6-315f-4325-9d8d-98efc4d21f4f" ID-PMC="PMC6813177" ID-Pensoft-Pub="1313-2970-881-1" ID-Pensoft-UUID="3D35B64F5EFC52D4A6D044AD461A7339" ID-PubMed="31662608" ID-ZooBank="75F261B11A33426AB3624FABEE5429EE" ModsDocID="1313-2970-881-1" checkinTime="1571384729753" checkinUser="pensoft" docAuthor="Lu, Yuan-Yuan, Zorn, Carsten, Kral, David &amp; Bai, Ming" docDate="2019" docId="19BBF8B07B5453979247DFD82C748143" docLanguage="en" docName="ZooKeys 881: 1-11" docOrigin="ZooKeys 881" docSource="http://dx.doi.org/10.3897/zookeys.881.34821" docTitle="Callistethus hamus Lu &amp; Zorn, sp. nov." docType="treatment" docUuid="03158F0F-19B4-4940-8025-649989535595" docUuidSource="ZooBank" docVersion="7" id="3D35B64F5EFC52D4A6D044AD461A7339" lastPageNumber="6" masterDocId="3D35B64F5EFC52D4A6D044AD461A7339" masterDocTitle="Description of Callistethus hamus sp. nov. (Coleoptera, Scarabaeidae, Rutelinae) from continental Southeast Asia using synchrotron to illustrate the aedeagus" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1668167892700" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of Callistethus hamus sp. nov. (Coleoptera, Scarabaeidae, Rutelinae) from continental Southeast Asia using synchrotron to illustrate the aedeagus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lu, Yuan-Yuan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zorn, Carsten</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kral, David</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bai, Ming</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>881</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>11</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.881.34821</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.881.34821</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-881-1</mods:identifier>
<mods:identifier type="ZooBank">75F261B11A33426AB3624FABEE5429EE</mods:identifier>
<mods:identifier type="Pensoft-UUID">3D35B64F5EFC52D4A6D044AD461A7339</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159754688" LSID="urn:lsid:zoobank.org:act:03158F0F-19B4-4940-8025-649989535595" httpUri="http://treatment.plazi.org/id/19BBF8B07B5453979247DFD82C748143" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection pageId="2" pageNumber="3" type="nomenclature">
<paragraph pageId="2" pageNumber="3">
<taxonomicName LSID="19BBF8B0-7B54-5397-9247-DFD82C748143" authority="Lu &amp; Zorn" class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus hamus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hamus">Callistethus hamus Lu &amp; Zorn</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="3">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="2" pageNumber="3" tableDoi="10.3897/zookeys.881.34821.figures1-9">Figs 1-9</figureCitation>
, 
<figureCitation captionStart="Figures 19–23" captionStartId="F3" captionText="Figures 19 - 23. 3 D model for aedeagus of Callistethus hamus Lu &amp; Zorn, sp. nov. Paratype from China, Yunnan. These photos are screen shots of the 3 D model of the aedeagus." figureDoi="10.3897/zookeys.881.34821.figures19-23" httpUri="https://binary.pensoft.net/fig/347508" pageId="2" pageNumber="3" tableDoi="10.3897/zookeys.881.34821.figures19-23">19-23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="type locality">
<paragraph pageId="2" pageNumber="3">Type locality.</paragraph>
<paragraph pageId="2" pageNumber="3">
<materialsCitation ID-GBIF-Occurrence="2427924919" collectingDate="2019-01-01" collectingDateMax="2019-12-31" collectingDateMin="2019-01-01" country="China" county="Southern" elevation="730" location="Xishuangbanna" municipality="Nan Ban" specimenCount="730" stateProvince="Yunnan">
<collectingCountry name="China">China</collectingCountry>
, 
<collectingCounty>Southern</collectingCounty>
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
, 
<locationDeviation location="Jinghong">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.3" unit="km" value="23.0">23 km</quantity>
NW of Jinghong
</locationDeviation>
, vicinity of 
<collectingMunicipality>Nan Ban</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:2F71CFE952C9958EE431339B833569D8" country="China" county="Southern" municipality="Nan Ban" name="Xishuangbanna" stateProvince="Yunnan">Xishuangbanna</location>
[Prefecture], 22°09.49&quot;N, 100°39.92&quot;E, 
<specimenCount type="generic">
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">730 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="materials_examined">
<paragraph pageId="2" pageNumber="3">Material examined.</paragraph>
<paragraph pageId="2" pageNumber="3">
<materialsCitation ID-GBIF-Occurrence="2427924917" collectingDate="2008-05-20" collectionCode="NMEC" collectorName="A Weigel" country="China" county="Dai Autonomous Prefecture of Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong City" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis bold="true" pageId="2" pageNumber="3">
<typeStatus>Holotype</typeStatus>
(HT).
</emphasis>
<collectingCountry name="China">CHINA</collectingCountry>
• 
<specimenCount>♂</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Dai Autonomous Prefecture of Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong City</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:8EFBD177A47243A996A03A3B5348AE52" country="China" county="Dai Autonomous Prefecture of Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong City" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2008-05-20">
<collectingDate value="2008-05-20">20 May 2008</collectingDate>
</date>
; 
<collectorName>A Weigel</collectorName>
leg.; 
<collectionCode>NMEC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="paratypes">
<paragraph pageId="2" pageNumber="3">
<typeStatus>Paratypes</typeStatus>
.
</paragraph>
<paragraph lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<materialsCitation ID-GBIF-Occurrence="2427924923" collectingDate="2019-01-01" collectingDateMax="2019-12-31" collectingDateMin="2019-01-01" collectionCode="NMEC" collectorName="A Weigel" country="China" county="Dai Autonomous Prefecture of Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong City" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<collectingCountry name="China">CHINA</collectingCountry>
• 
<specimenCount type="female">1 ♀</specimenCount>
; same data as the holotype
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924930" collectingDate="2008-06-05" collectionCode="NMEC" collectorName="A. Weigel" country="China" county="Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:F266D90BD5D8DA3FE7100FE45650684A" country="China" county="Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2008-06-05">
<collectingDate value="2008-06-05">5 Jun. 2008</collectingDate>
</date>
; 
<collectorName>A. Weigel</collectorName>
leg.; 
<collectionCode>NMEC</collectionCode>
; [Micro-CT specimen]
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924918" collectingDate="2008-06-05" collectionCode="CZPC" collectorName="A. Weigel" country="China" county="Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:98F7214AC75AD0A85C06E2B0ADFB595A" country="China" county="Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2008-06-05">
<collectingDate value="2008-06-05">5 Jun. 2008</collectingDate>
</date>
; 
<collectorName>A. Weigel</collectorName>
leg.; 
<collectionCode>CZPC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924925" collectingDate="2008-06-15" collectionCode="NMEC" country="China" county="Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:6D07E50BB24AA6A919DD44F93B8A8DD8" country="China" county="Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2008-06-15">
<collectingDate value="2008-06-15">15 Jun. 2008</collectingDate>
</date>
; 
<collectionCode>NMEC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924921" collectingDate="2008-06-05" collectionCode="NMEC" collectorName="A. Weigel" country="China" county="Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:44E423FFBB0A516F507805B23E26B742" country="China" county="Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2008-06-05">
<collectingDate value="2008-06-05">5 Jun. 2008</collectingDate>
</date>
; 
<collectorName>A. Weigel</collectorName>
leg.; 
<collectionCode>NMEC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924928" collectingDate="2008-05-12" collectionCode="NMEC" collectorName="A. Weigel" country="China" county="Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:CB6E4BDA5DE4E7E85340156B71CC8FC4" country="China" county="Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2008-05-12">
<collectingDate value="2008-05-12">12 May 2008</collectingDate>
</date>
; 
<collectorName>A. Weigel</collectorName>
leg.; 
<collectionCode>NMEC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924929" collectingDate="2009-05-16" collectingMethod="Malaise trap" collectionCode="IZAS" collectorName="L. Z. Meng" country="China" county="Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" specimenCode="IOZ(E)1966878" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:D8DB4E9621820C4191769F301BB1592A" country="China" county="Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan Province">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2009-05-16">
<collectingDate value="2009-05-16">16 May 2009</collectingDate>
</date>
; 
<collectingMethod>Malaise trap</collectingMethod>
; 
<collectorName>L.Z. Meng</collectorName>
leg.; 
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
<specimenCode>IOZ(E)1966878</specimenCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924931" collectingDate="2009-05-26" collectingMethod="Malaise trap" collectionCode="IZAS" collectorName="L. Z. Meng" country="China" county="Xishuangbanna" latitude="22.130833" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66866" municipality="Jinghong" specimenCode="IOZ(E)1966879" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:4360947AC4A35DBAFB2E688BF2718E6C" country="China" county="Xishuangbanna" latitude="22.130833" longLatPrecision="12" longitude="100.66866" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan Province">
Na Ban River Watershed National 
<pageBreakToken pageId="3" pageNumber="4" start="start">Nature</pageBreakToken>
Reserve
</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="07.85" orientation="latitude" precision="9" value="22.130833">22°07.85'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="40.12" orientation="longitude" precision="9" value="100.66866">100°40.12'E</geoCoordinate>
; 
<date value="2009-05-26">
<collectingDate value="2009-05-26">26 May 2009</collectingDate>
</date>
; 
<collectingMethod>Malaise trap</collectingMethod>
; 
<collectorName>L.Z. Meng</collectorName>
leg.; 
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
<specimenCode>IOZ(E)1966879</specimenCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924932" collectingDate="2009-05-16" collectingMethod="flight interception traps" collectionCode="IZAS" collectorName="L. Z. Meng" country="China" county="Xishuangbanna" latitude="22.158167" location="Na Ban River Watershed National Nature Reserve" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" specimenCode="IOZ(E)1966880" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:9DDCCD70C2286205BFD77FE8E01870A0" country="China" county="Xishuangbanna" latitude="22.158167" longLatPrecision="12" longitude="100.66534" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan Province">Na Ban River Watershed National Nature Reserve</location>
; 
<geoCoordinate degrees="22" direction="north" minutes="09.49" orientation="latitude" precision="9" value="22.158167">22°09.49'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="39.92" orientation="longitude" precision="9" value="100.66534">100°39.92'E</geoCoordinate>
; 
<date value="2009-05-16">
<collectingDate value="2009-05-16">16 May 2009</collectingDate>
</date>
; 
<collectingMethod>flight interception traps</collectingMethod>
; 
<collectorName>L.Z. Meng</collectorName>
leg.; 
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
<specimenCode>IOZ(E)1966880</specimenCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924922" collectingDate="2013-07-31" collectionCode="FWPC" country="China" county="Xishuangbanna" location="Na Ban River Watershed National Nature Reserve" municipality="Jinghong" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Jinghong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:DFA80B38C06BD9C966BDD614765B95FE" country="China" county="Xishuangbanna" municipality="Jinghong" name="Na Ban River Watershed National Nature Reserve" stateProvince="Yunnan Province">Na Ban River Watershed National Nature Reserve</location>
; 
<date value="2013-07-31">
<collectingDate value="2013-07-31">31. Jul. 2013</collectingDate>
</date>
; 
<collectionCode>FWPC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924915" collectingDate="1958-06-04" collectionCode="IZAS" collectorName="L. Y. Zheng" country="China" latitude="21.841333" location="Menghun Town" longLatPrecision="12" longitude="100.38583" municipality="Menghai" specimenCode="IOZ(E)1966104" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingMunicipality>Menghai</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:227CBAF12440815C4DC9FBEB15A9792E" country="China" latitude="21.841333" longLatPrecision="12" longitude="100.38583" municipality="Menghai" name="Menghun Town" stateProvince="Yunnan Province">Menghun Town</location>
; 
<geoCoordinate degrees="21" direction="north" minutes="50.48" orientation="latitude" precision="9" value="21.841333">21°50.48'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="23.15" orientation="longitude" precision="9" value="100.38583">100°23.15'E</geoCoordinate>
; 
<date value="1958-06-04">
<collectingDate value="1958-06-04">4 Jun. 1958</collectingDate>
</date>
; 
<collectorName>L.Y. Zheng</collectorName>
leg.; 
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
<specimenCode>IOZ(E)1966104</specimenCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924926" collectingDate="1958-06-01" collectionCode="IZAS" collectorName="X. W. Meng" country="China" county="Xishuangbanna" latitude="21.841333" location="Menghun Town" longLatPrecision="12" longitude="100.38583" municipality="Menghai" specimenCode="IOZ(E)1966106" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Menghai</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:9F37F7749FC6D983991524277E3C5C6E" country="China" county="Xishuangbanna" latitude="21.841333" longLatPrecision="12" longitude="100.38583" municipality="Menghai" name="Menghun Town" stateProvince="Yunnan Province">Menghun Town</location>
; 
<geoCoordinate degrees="21" direction="north" minutes="50.48" orientation="latitude" precision="9" value="21.841333">21°50.48'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="23.15" orientation="longitude" precision="9" value="100.38583">100°23.15'E</geoCoordinate>
; 
<date value="1958-06-01">
<collectingDate value="1958-06-01">1 Jun. 1958</collectingDate>
</date>
; 
<collectorName>X.W. Meng</collectorName>
leg.; 
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
<specimenCode>IOZ(E)1966106</specimenCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924920" collectingDate="1958-06-04" collectionCode="IZAS" collectorName="L. Y. Zheng" country="China" county="Xishuangbanna" latitude="21.841333" location="Menghun Town" longLatPrecision="12" longitude="100.38583" municipality="Menghai" specimenCode="IOZ(E)1966107" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Menghai</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:E7D20966A3CFF9D77507E6A0039B8B99" country="China" county="Xishuangbanna" latitude="21.841333" longLatPrecision="12" longitude="100.38583" municipality="Menghai" name="Menghun Town" stateProvince="Yunnan Province">Menghun Town</location>
; 
<geoCoordinate degrees="21" direction="north" minutes="50.48" orientation="latitude" precision="9" value="21.841333">21°50.48'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="23.15" orientation="longitude" precision="9" value="100.38583">100°23.15'E</geoCoordinate>
; 
<date value="1958-06-04">
<collectingDate value="1958-06-04">4 Jun. 1958</collectingDate>
</date>
; 
<collectorName>L.Y. Zheng</collectorName>
leg.; 
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
<specimenCode>IOZ(E)1966107</specimenCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924916" collectingDate="1958-07-08" collectionCode="IZAS" collectorName="S. Y. Wang" country="China" county="Xishuangbanna" latitude="21.985834" location="Mengzhe Town" longLatPrecision="12" longitude="100.267" municipality="Menghai" specimenCode="IOZ(E)1966105" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
; 
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCounty>Xishuangbanna</collectingCounty>
, 
<collectingMunicipality>Menghai</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:21383E4ADF2E46EAB9FD11F60175D6F4" country="China" county="Xishuangbanna" latitude="21.985834" longLatPrecision="12" longitude="100.267" municipality="Menghai" name="Mengzhe Town" stateProvince="Yunnan Province">Mengzhe Town</location>
; 
<geoCoordinate degrees="21" direction="north" minutes="59.15" orientation="latitude" precision="9" value="21.985834">21°59.15'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="16.02" orientation="longitude" precision="9" value="100.267">100°16.02'E</geoCoordinate>
; 
<date value="1958-07-08">
<collectingDate value="1958-07-08">8 Jul. 1958</collectingDate>
</date>
; 
<collectorName>S.Y. Wang</collectorName>
leg.; 
<collectionCode country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" type="">IZAS</collectionCode>
<specimenCode>IOZ(E)1966105</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph pageId="3" pageNumber="4">
<materialsCitation ID-GBIF-Occurrence="2427924924" collectingDate="2005-05-04" collectingDateMax="2005-05-17" collectingDateMin="2005-05-04" collectionCode="CZPC" collectorName="P. Moravec" country="Laos" latitude="18.4025" location="Phou Khao Khoay" longLatPrecision="13" longitude="103.035835" specimenCount="1" specimenCount-male="1" stateProvince="Viangchan">
<collectingCountry name="Laos">LAOS</collectingCountry>
• 
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingRegion country="Laos" name="Viangchan">Vientiane Province</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:19BBF8B07B5453979247DFD82C748143:30944AB4AA36424D39460356AC61BF3A" country="Laos" latitude="18.4025" longLatPrecision="13" longitude="103.035835" name="Phou Khao Khoay" stateProvince="Viangchan">Phou Khao Khoay</location>
; 
<geoCoordinate degrees="18" direction="north" minutes="24.15" orientation="latitude" precision="9" value="18.4025">18°24.15'N</geoCoordinate>
, 
<geoCoordinate degrees="103" direction="east" minutes="02.15" orientation="longitude" precision="9" value="103.035835">103°02.15'E</geoCoordinate>
; 
<date value="2005-05-04" valueMax="2005-05-17" valueMin="2005-05-04">
<collectingDate value="2005-05-04" valueMax="2005-05-17" valueMin="2005-05-04">4-17 May 2005</collectingDate>
</date>
; 
<collectorName>P. Moravec</collectorName>
leg.; 
<collectionCode>CZPC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph pageId="3" pageNumber="4">
<materialsCitation ID-GBIF-Occurrence="2427924933" collectingDate="2017-05" collectionCode="CZPC" collectorName="Lam &amp; Prov. &amp; Dambri, B" country="Vietnam" latitude="11.640333" location="VIETNAM" longLatPrecision="13" longitude="107.742" specimenCount="6" specimenCount-female="2" specimenCount-male="4">
<collectingCountry name="Vietnam">VIETNAM</collectingCountry>
• 
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
; 
<normalizedToken originalValue="Lâm">
<collectorName>Lam</collectorName>
</normalizedToken>
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ồng 
<collectorName>Prov.</collectorName>
, 
<collectorName>Dambri, B</collectorName>
ảo 
<normalizedToken originalValue="Lâm">
<collectorName>Lam</collectorName>
</normalizedToken>
; 
<geoCoordinate degrees="11" direction="north" minutes="38.42" orientation="latitude" precision="9" value="11.640333">11°38.42'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="44.52" orientation="longitude" precision="9" value="107.742">107°44.52'E</geoCoordinate>
; 
<date value="2017-05">
<collectingDate value="2017-05">May 2017</collectingDate>
</date>
; local collector leg.; 
<collectionCode>CZPC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924913" collectingDate="2018-03" collectionCode="CZPC" collectorName="Lam &amp; Prov., B" country="Vietnam" latitude="11.548" longLatPrecision="13" longitude="107.80766" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 ♀</specimenCount>
; 
<normalizedToken originalValue="Lâm">
<collectorName>Lam</collectorName>
</normalizedToken>
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ồng 
<collectorName>Prov., B</collectorName>
ảo Lộc; 
<geoCoordinate degrees="11" direction="north" minutes="32.88" orientation="latitude" precision="9" value="11.548">11°32.88'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="48.46" orientation="longitude" precision="9" value="107.80766">107°48.46'E</geoCoordinate>
; 
<date value="2018-03">
<collectingDate value="2018-03">March 2018</collectingDate>
</date>
; local collector leg.; 
<collectionCode>CZPC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924927" collectingDate="2018-03" collectionCode="CZPC" collectorName="Lam &amp; Prov., B" country="Vietnam" latitude="11.548" longLatPrecision="13" longitude="107.80766" specimenCount="6" specimenCount-female="1" specimenCount-male="5">
<specimenCount type="male">5 ♂♂</specimenCount>
<specimenCount type="female">1 ♀♀</specimenCount>
; 
<normalizedToken originalValue="Lâm">
<collectorName>Lam</collectorName>
</normalizedToken>
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ồng 
<collectorName>Prov., B</collectorName>
ảo Lộc; 
<geoCoordinate degrees="11" direction="north" minutes="32.88" orientation="latitude" precision="9" value="11.548">11°32.88'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="48.46" orientation="longitude" precision="9" value="107.80766">107°48.46'E</geoCoordinate>
; 
<date value="2018-03">
<collectingDate value="2018-03">March 2018</collectingDate>
</date>
; local collector leg.; 
<collectionCode>CZPC</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="2427924914" collectingDate="2017-03" collectionCode="MSPC" collectorName="Lam &amp; Prov., B" country="Vietnam" latitude="11.548" longLatPrecision="13" longitude="107.80766" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
; 
<normalizedToken originalValue="Lâm">
<collectorName>Lam</collectorName>
</normalizedToken>
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ồng 
<collectorName>Prov., B</collectorName>
ảo Lộc; 
<geoCoordinate degrees="11" direction="north" minutes="32.88" orientation="latitude" precision="9" value="11.548">11°32.88'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="48.46" orientation="longitude" precision="9" value="107.80766">107°48.46'E</geoCoordinate>
; 
<date value="2017-03">
<collectingDate value="2017-03">March 2017</collectingDate>
</date>
; local collector leg.; 
<collectionCode country="Italy" httpUri="http://grbio.org/cool/t9g4-v7fn" name="Museo di Storia Naturale &amp;quot;Pietro Calderini&amp;quot;" type="">MSPC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph pageId="3" pageNumber="4">
<emphasis bold="true" pageId="3" pageNumber="4">
Description of 
<typeStatus>holotype</typeStatus>
(
<specimenCount>♂</specimenCount>
).
</emphasis>
Body shape elongate ovoid, convex.
</paragraph>
<paragraph pageId="3" pageNumber="4">
<emphasis bold="true" pageId="3" pageNumber="4">
<emphasis italics="true" pageId="3" pageNumber="4">Colour</emphasis>
.
</emphasis>
Head including antenna orange-brown, with two small black spots on vertex; pronotum orange-brown with two moderately large black spots in the middle of each side; an additional longitudinal stripe near the lateral margin (not reaching anterior and posterior margins) separating the yellow sides from the orange disc (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">Figs 1</figureCitation>
, 
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">4</figureCitation>
); elytra light reddish-brown with black markings as follows: an elongate spot on humeral callus, another smaller spot beneath shoulder at lateral margin, a zig-zag band of loosely connected spots crossing the middle of elytra, another zig-zag band shortly before the posterior margin (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 1</figureCitation>
); propygidium black with a transverse yellow spot at the outer posterior margin (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 5</figureCitation>
); pygidium black with a narrow yellow longitudinal middle line (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 5</figureCitation>
); underside yellow with black markings, with slight metallic shine (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 2</figureCitation>
); black macula in the middle and at extremities of femurs; various black markings present on all tibiae; pro- and mesotarsi brown, metatarsus black; major part of meso- and metaventrite black; meso-metaventral process yellow; major part of abdominal ventrites 1-3 yellow, abdominal ventrites 4-6 black with lateral yellow spot (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 2</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" start="Figures 1–9" startId="F1">
<paragraph pageId="3" pageNumber="4">
<emphasis bold="true" pageId="3" pageNumber="4">Figures 1-9.</emphasis>
<typeStatus>Holotype</typeStatus>
of 
<taxonomicName class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus hamus" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hamus">
<emphasis italics="true" pageId="3" pageNumber="4">Callistethus hamus</emphasis>
</taxonomicName>
Lu &amp; Zorn, sp. 
<date>
nov. 
<emphasis bold="true" pageId="3" pageNumber="4">1-3</emphasis>
</date>
habitus 
<emphasis bold="true" pageId="3" pageNumber="4">1</emphasis>
dorsal view 
<emphasis bold="true" pageId="3" pageNumber="4">2</emphasis>
ventral view 
<emphasis bold="true" pageId="3" pageNumber="4">3</emphasis>
lateral view from left 
<emphasis bold="true" pageId="3" pageNumber="4">4</emphasis>
head and pronotum 
<emphasis bold="true" pageId="3" pageNumber="4">5</emphasis>
propygidium and pygidium 
<emphasis bold="true" pageId="3" pageNumber="4">6-8</emphasis>
aedeagus 
<emphasis bold="true" pageId="3" pageNumber="4">6</emphasis>
dorsal view 
<emphasis bold="true" pageId="3" pageNumber="4">7</emphasis>
lateral view from right 
<emphasis bold="true" pageId="3" pageNumber="4">8</emphasis>
ventral view 
<emphasis bold="true" pageId="3" pageNumber="4">9</emphasis>
label. Key: Sc. 1, Scale bars: 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
(
<emphasis bold="true" pageId="3" pageNumber="4">1-3</emphasis>
); Sc. 2, 5 mm (
<emphasis bold="true" pageId="3" pageNumber="4">4, 5</emphasis>
); Sc. 3, 1 mm (
<emphasis bold="true" pageId="3" pageNumber="4">6-8</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="4">
<emphasis bold="true" italics="true" pageId="3" pageNumber="4">Head.</emphasis>
Clypeus subtrapezoidal, anterior angles rounded; anterior margin weakly reflexed, very densely, transversely rugo-punctate; frons rugo-punctate, with very shallow impression in the middle, confluently punctate at sides; vertex finely and sparsely punctate in the middle, more coarsely punctate laterally; ratio interocular width/width of head approximately 0.71; antennal club longer than antennomeres 1-6 combined (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">Figs 1</figureCitation>
, 
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="3" pageNumber="4" tableDoi="10.3897/zookeys.881.34821.figures1-9">4</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">
<pageBreakToken pageId="4" pageNumber="5" start="start">Pronotum</pageBreakToken>
</emphasis>
approximately 1.89 times wider than long; sparsely and finely punctate, punctures very fine on disc, becoming gradually larger toward the sides; surface with additional micropunctures; anterior angles sub-rectangular; posterior angles obtuse; broadest at base; sides evenly curved in the middle, straight and strongly converging before anterior angles; sides very slightly sinuate near posterior angles; basal marginal line only present near posterior angles; anterior marginal line indistinct in the middle (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 4</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Scutellum</emphasis>
subtriangular, slightly broader than long, finely and sparsely punctate (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 4</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Elytra</emphasis>
regularly striate; primary striae and secondary stria of subsutural interstice sulcate in posterior half, therefore intervals flat in anterior half and gradually more convex posteriad; strial punctation distinct, coarse; 2nd and 3rd interstices each with an indistinct row of punctures; elytral surface with additional sparse micropunctation; humeral umbone and apical protuberance rather prominent; epipleuron broad near humerus, ending approximately at the middle of elytron; posterior margins evenly, separately rounded; apico-sutural angle forming small dent (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 1</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Pygidium</emphasis>
convex; apex broadly rounded; moderately densely, coarsely punctate; apex with several long, erect brownish setae (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 5</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Ventral thoracic surface</emphasis>
smooth (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 2</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Meso-metaventral process</emphasis>
very long, reaching middle of procoxae; projecting upward in lateral view; apex acute (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Fig. 3</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Abdominal ventrites</emphasis>
with transverse row of rather sparse brown setae (usually only in lateral part); ventrites 1 and 2 carinate laterally (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Figs 2</figureCitation>
, 
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">3</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Legs.</emphasis>
Mesofemur with two bands of long brown setae: one along anterior margin; another transverse row of punctures parallel to posterior margin. Protibia bidentate, rather slender; proximal tooth short, situated close to the rather short, weakly outwards curved apical tooth; inner spur short, articulated in opposite to proximal tooth. Metatibia strongly fusiform; protarsomere 5 shorter than tarsomeres 1-4 combined in all legs; inner protarsal claw slightly widened and deeply incised apically, lower margin with obtuse angle basally, upper branch spiniform; outer mesotarsal claw long, curved, deeply incised at apex, upper branch spiniform; metatarsal claws somewhat unequal, outer claw distinctly broader and longer than inner (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Figs 1-3</figureCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Aedeagus.</emphasis>
Parameres short, with the hook-like apex in lateral view. See 
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures1-9">Figs 6-8</figureCitation>
.
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" pageId="4" pageNumber="5">Female.</emphasis>
Abdominal ventrites 4-6 with extensive yellow markings. Apical tooth of protibia long and somewhat spatulate; protarsus articulated slightly basally of level of proximal tooth; inner spur long; protarsomere 1 as long as combined length of protarsomeres 2-4; antennal club short, only slightly longer than antennomeres 2-6 combined.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="description">
<paragraph pageId="4" pageNumber="5">Measurements.</paragraph>
<paragraph pageId="4" pageNumber="5">
Total body length 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.44" metricValueMin="1.15" unit="mm" value="12.95" valueMax="14.4" valueMin="11.5">11.5-14.4 mm</quantity>
(HT 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.18" unit="mm" value="11.8">11.8 mm</quantity>
), total body width 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.45" metricValueMax="8.4" metricValueMin="6.5" unit="mm" value="7.45" valueMax="8.4" valueMin="6.5">6.5-8.4 mm</quantity>
(HT 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.5" unit="mm" value="7.5">7.5 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="morphological variability">
<paragraph pageId="4" pageNumber="5">Morphological variability.</paragraph>
<paragraph pageId="4" pageNumber="5">
Vertex with or without two black spots. Elytral spots vary slightly in shape and size. The extend of black markings of the ventral side variable. The secondary longitudinal rows of punctures in interstices 2 and 3 sometimes distinct until the posterior half. Shape of parameres very consistent (
<figureCitation captionStart="Figures 19–23" captionStartId="F3" captionText="Figures 19 - 23. 3 D model for aedeagus of Callistethus hamus Lu &amp; Zorn, sp. nov. Paratype from China, Yunnan. These photos are screen shots of the 3 D model of the aedeagus." figureDoi="10.3897/zookeys.881.34821.figures19-23" httpUri="https://binary.pensoft.net/fig/347508" pageId="4" pageNumber="5" tableDoi="10.3897/zookeys.881.34821.figures19-23">Figs 19-23</figureCitation>
, Appendix 1).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="differential diagnosis">
<paragraph pageId="5" pageNumber="6">
<pageBreakToken pageId="5" pageNumber="6" start="start">Differential</pageBreakToken>
diagnosis.
</paragraph>
<paragraph pageId="5" pageNumber="6">
<taxonomicName class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus hamus" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hamus">
<emphasis italics="true" pageId="5" pageNumber="6">Callistethus hamus</emphasis>
</taxonomicName>
sp. nov. resembles several other, similarly coloured South East Asian species of this genus. The reddish colour combined with black markings is also found in 
<taxonomicName lsidName="C. maculatus" pageId="5" pageNumber="6" rank="species" species="maculatus">
<emphasis italics="true" pageId="5" pageNumber="6">C. maculatus</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
, 1834), 
<taxonomicName lsidName="C. picturatus" pageId="5" pageNumber="6" rank="species" species="picturatus">
<emphasis italics="true" pageId="5" pageNumber="6">C. picturatus</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Candèze">Candeze</normalizedToken>
, 1869), 
<taxonomicName lsidName="C. rachelae" pageId="5" pageNumber="6" rank="species" species="rachelae">
<emphasis italics="true" pageId="5" pageNumber="6">C. rachelae</emphasis>
</taxonomicName>
(Arrow, 1917), 
<taxonomicName lsidName="C. spiniferus" pageId="5" pageNumber="6" rank="species" species="spiniferus">
<emphasis italics="true" pageId="5" pageNumber="6">C. spiniferus</emphasis>
</taxonomicName>
(Ohaus, 1915), and 
<taxonomicName lsidName="C. stolidopygus" pageId="5" pageNumber="6" rank="species" species="stolidopygus">
<emphasis italics="true" pageId="5" pageNumber="6">C. stolidopygus</emphasis>
</taxonomicName>
(Ohaus, 1915). The new species is most similar to 
<taxonomicName lsidName="C. rachelae" pageId="5" pageNumber="6" rank="species" species="rachelae">
<emphasis italics="true" pageId="5" pageNumber="6">C. rachelae</emphasis>
</taxonomicName>
, which also has only two black spots on the pronotum, not four as all other red species (
<figureCitation captionStart="Figures 10–18" captionStartId="F2" captionText="Figures 10 - 18. Holotype of Callistethus rachelae (Arrow, 1917). 10 - 12 habitus 10 dorsal view 11 ventral view 12 lateral view from left 13 head and pronotum 14 propygidium and pygidium 15 - 17 aedeagus 15 dorsal view 16 lateral view from right 17 ventral view 18 label. Key: Sc. 1. Scale bars: 5 mm (10 - 12); Sc. 2, 5 mm (13, 14); Sc. 3, 1 mm (15 - 17)." figureDoi="10.3897/zookeys.881.34821.figures10-18" httpUri="https://binary.pensoft.net/fig/347507" pageId="5" pageNumber="6" tableDoi="10.3897/zookeys.881.34821.figures10-18">Figs 10-18</figureCitation>
). 
<taxonomicName class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus hamus" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hamus">
<emphasis italics="true" pageId="5" pageNumber="6">Callistethus hamus</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="C. rachelae" pageId="5" pageNumber="6" rank="species" species="rachelae">
<emphasis italics="true" pageId="5" pageNumber="6">C. rachelae</emphasis>
</taxonomicName>
primarily in the shape of the aedeagus (
<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Holotype of Callistethus hamus Lu &amp; Zorn, sp. nov. 1 - 3 habitus 1 dorsal view 2 ventral view 3 lateral view from left 4 head and pronotum 5 propygidium and pygidium 6 - 8 aedeagus 6 dorsal view 7 lateral view from right 8 ventral view 9 label. Key: Sc. 1, Scale bars: 5 mm (1 - 3); Sc. 2, 5 mm (4, 5); Sc. 3, 1 mm (6 - 8)." figureDoi="10.3897/zookeys.881.34821.figures1-9" httpUri="https://binary.pensoft.net/fig/347506" pageId="5" pageNumber="6" tableDoi="10.3897/zookeys.881.34821.figures1-9">Figs 6-8</figureCitation>
, 
<figureCitation captionStart="Figures 10–18" captionStartId="F2" captionText="Figures 10 - 18. Holotype of Callistethus rachelae (Arrow, 1917). 10 - 12 habitus 10 dorsal view 11 ventral view 12 lateral view from left 13 head and pronotum 14 propygidium and pygidium 15 - 17 aedeagus 15 dorsal view 16 lateral view from right 17 ventral view 18 label. Key: Sc. 1. Scale bars: 5 mm (10 - 12); Sc. 2, 5 mm (13, 14); Sc. 3, 1 mm (15 - 17)." figureDoi="10.3897/zookeys.881.34821.figures10-18" httpUri="https://binary.pensoft.net/fig/347507" pageId="5" pageNumber="6" tableDoi="10.3897/zookeys.881.34821.figures10-18">15-17</figureCitation>
). The ventral margin of the parameres of 
<taxonomicName lsidName="C. rachelae" pageId="5" pageNumber="6" rank="species" species="rachelae">
<emphasis italics="true" pageId="5" pageNumber="6">C. rachelae</emphasis>
</taxonomicName>
is somewhat membranous and soft, but fully sclerotised in 
<taxonomicName lsidName="C. hamus" pageId="5" pageNumber="6" rank="species" species="hamus">
<emphasis italics="true" pageId="5" pageNumber="6">C. hamus</emphasis>
</taxonomicName>
sp. nov. The apices of the parameres in 
<taxonomicName lsidName="C. hamus" pageId="5" pageNumber="6" rank="species" species="hamus">
<emphasis italics="true" pageId="5" pageNumber="6">C. hamus</emphasis>
</taxonomicName>
sp. nov. are longer and more strongly curved compared to those of 
<taxonomicName lsidName="C. rachelae" pageId="5" pageNumber="6" rank="species" species="rachelae">
<emphasis italics="true" pageId="5" pageNumber="6">C. rachelae</emphasis>
</taxonomicName>
(
<figureCitation captionStart="Figures 10–18" captionStartId="F2" captionText="Figures 10 - 18. Holotype of Callistethus rachelae (Arrow, 1917). 10 - 12 habitus 10 dorsal view 11 ventral view 12 lateral view from left 13 head and pronotum 14 propygidium and pygidium 15 - 17 aedeagus 15 dorsal view 16 lateral view from right 17 ventral view 18 label. Key: Sc. 1. Scale bars: 5 mm (10 - 12); Sc. 2, 5 mm (13, 14); Sc. 3, 1 mm (15 - 17)." figureDoi="10.3897/zookeys.881.34821.figures10-18" httpUri="https://binary.pensoft.net/fig/347507" pageId="5" pageNumber="6" tableDoi="10.3897/zookeys.881.34821.figures10-18">Figs 15-17</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.881.34821.figures10-18" httpUri="https://binary.pensoft.net/fig/347507" pageId="5" pageNumber="6" start="Figures 10–18" startId="F2">
<paragraph pageId="5" pageNumber="6">
<emphasis bold="true" pageId="5" pageNumber="6">Figures 10-18.</emphasis>
<typeStatus>Holotype</typeStatus>
of 
<taxonomicName class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus rachelae" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rachelae">
<emphasis italics="true" pageId="5" pageNumber="6">Callistethus rachelae</emphasis>
</taxonomicName>
(Arrow, 1917). 
<emphasis bold="true" pageId="5" pageNumber="6">10-12</emphasis>
habitus 
<emphasis bold="true" pageId="5" pageNumber="6">10</emphasis>
dorsal view 
<emphasis bold="true" pageId="5" pageNumber="6">11</emphasis>
ventral view 
<emphasis bold="true" pageId="5" pageNumber="6">12</emphasis>
lateral view from left 
<emphasis bold="true" pageId="5" pageNumber="6">13</emphasis>
head and pronotum 
<emphasis bold="true" pageId="5" pageNumber="6">14</emphasis>
propygidium and pygidium 
<emphasis bold="true" pageId="5" pageNumber="6">15-17</emphasis>
aedeagus 
<emphasis bold="true" pageId="5" pageNumber="6">15</emphasis>
dorsal view 
<emphasis bold="true" pageId="5" pageNumber="6">16</emphasis>
lateral view from right 
<emphasis bold="true" pageId="5" pageNumber="6">17</emphasis>
ventral view 
<emphasis bold="true" pageId="5" pageNumber="6">18</emphasis>
label. Key: Sc. 1. Scale bars: 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
(
<emphasis bold="true" pageId="5" pageNumber="6">10-12</emphasis>
); Sc. 2, 5 mm (
<emphasis bold="true" pageId="5" pageNumber="6">13, 14</emphasis>
); Sc. 3, 1 mm (
<emphasis bold="true" pageId="5" pageNumber="6">15-17</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.881.34821.figures19-23" httpUri="https://binary.pensoft.net/fig/347508" pageId="5" pageNumber="6" start="Figures 19–23" startId="F3">
<paragraph pageId="5" pageNumber="6">
<emphasis bold="true" pageId="5" pageNumber="6">
Figures 
<date value="1919-03-23">19-23. 3</date>
</emphasis>
D model for aedeagus of 
<taxonomicName class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus hamus" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hamus">
<emphasis italics="true" pageId="5" pageNumber="6">Callistethus hamus</emphasis>
</taxonomicName>
Lu &amp; Zorn, sp. nov. 
<typeStatus>Paratype</typeStatus>
from 
<collectingCountry name="China">China</collectingCountry>
, 
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
. These photos are screen shots of the 3D model of the aedeagus.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="etymology">
<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="6">The specific epithet refers to the hook-like shape of the parameres of the new species.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="collecting data">
<paragraph pageId="5" pageNumber="6">Collecting data.</paragraph>
<paragraph pageId="5" pageNumber="6">Specimens collected in the Naban River Watershed National Nature Reserve by Lingzeng Meng and Andreas Weigel were collected with cross-window traps in the tree canopy and malaise trap.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="distribution">
<paragraph pageId="5" pageNumber="6">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="6">
<collectingCountry name="China">China</collectingCountry>
(
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
), 
<collectingCountry name="Laos">Laos</collectingCountry>
(
<collectingRegion country="Laos" name="Viangchan">Vientiane</collectingRegion>
), 
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
(
<normalizedToken originalValue="Lâm">Lam</normalizedToken>
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ồng) (
<figureCitation captionStart="Figure 24" captionStartId="F4" captionText="Figure 24. Map of Southeast Asia showing distributions of Callistethus hamus and Callistethus rachelae." figureDoi="10.3897/zookeys.881.34821.figure24" httpUri="https://binary.pensoft.net/fig/347509" pageId="5" pageNumber="6" tableDoi="10.3897/zookeys.881.34821.figure24">Fig. 24</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.881.34821.figure24" httpUri="https://binary.pensoft.net/fig/347509" pageId="5" pageNumber="6" start="Figure 24" startId="F4">
<paragraph pageId="5" pageNumber="6">
<emphasis bold="true" pageId="5" pageNumber="6">Figure 24.</emphasis>
Map of Southeast Asia showing distributions of 
<taxonomicName class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus hamus" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hamus">
<emphasis italics="true" pageId="5" pageNumber="6">Callistethus hamus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" family="Rutelidae" genus="Callistethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callistethus rachelae" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rachelae">
<emphasis italics="true" pageId="5" pageNumber="6">Callistethus rachelae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>