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<mods:title id="4B49BF3DFB97DA0E44FA35F2560C904B">Review of Hygrogeus Mulsant &amp; Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae)</mods:title>
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<treatment id="024240545C73FFEAFF3CFEFE8B5AC99F" ID-DOI="http://doi.org/10.5281/zenodo.14023089" ID-Zenodo-Dep="14023089" LSID="urn:lsid:plazi:treatment:024240545C73FFEAFF3CFEFE8B5AC99F" httpUri="http://treatment.plazi.org/id/024240545C73FFEAFF3CFEFE8B5AC99F" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="C2F1A2C95C73FFEAFF3CFEFE8B80CB3D" box="[151,659,294,321]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8A54F1425C73FFEAFF3CFEFE8B80CB3D" blockId="15.[151,659,294,357]" box="[151,659,294,321]" pageId="15" pageNumber="16">
<heading id="D11C462E5C73FFEAFF3CFEFE8B80CB3D" bold="true" box="[151,659,294,321]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="4">
<taxonomicName id="4DEB8AC15C73FFEAFF3CFEFE8B80CB3D" ID-CoL="58CBH" authority="(Cameron, 1941)" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[151,659,294,321]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B89F2D505C73FFEAFF3CFEFE8B80CB3D" bold="true" box="[151,659,294,321]" pageId="15" pageNumber="16">
<emphasis id="B89F2D505C73FFEAFF3CFEFE88A9CB3C" bold="true" box="[151,442,294,321]" italics="true" pageId="15" pageNumber="16">Trichodromeus armatus</emphasis>
(
<bibRefCitation id="EE7A8CB35C73FFEAFE61FEFE8B98CB3D" author="Cameron, M." box="[458,651,294,320]" pageId="15" pageNumber="16" pagination="142 - 147" refId="ref43534" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). The Proceedings of the Royal Entomological Society of London (B), 10, 142 - 147. https: // doi. org / 10.1111 / j. 1365 - 3113.1941. tb 00709. x" type="journal article" year="1941">Cameron, 1941</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C73FFEAFF3CFE93884CCB18" box="[151,351,331,357]" pageId="15" pageNumber="16" type="description">
<paragraph id="8A54F1425C73FFEAFF3CFE93884CCB18" blockId="15.[151,659,294,357]" box="[151,351,331,357]" pageId="15" pageNumber="16">
(
<figureCitation id="12D0EDC75C73FFEAFF34FE938817CB18" box="[159,260,331,357]" captionStart="FIGURES 16" captionStartId="7.[151,264,1732,1756]" captionTargetBox="[157,1429,187,1702]" captionTargetId="figure-16@7.[151,1435,181,1708]" captionTargetPageId="7" captionText="FIGURES 16. Habitus of Hygrogeus (Fig. 1) and Trichodromeus (Figs 26): 1—H. aemulus (Tirol), 2—T. armatus (Kashmir: Baltal), 3—T. armatus (Kashmir: Kolahoi Glacier), 4—T. ketmeniensis (Kyrgyzstan: Chatkal Mts.), 5—T. ovchinnikovi (holotype), 6—T. kumaonensis (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023035" httpUri="https://zenodo.org/record/14023035/files/figure.png" pageId="15" pageNumber="16">Figs 23</figureCitation>
,
<figureCitation id="12D0EDC75C73FFEAFEBBFE938844CB18" box="[272,343,331,357]" captionStart="FIGURES 3946" captionStartId="16.[151,264,1817,1841]" captionTargetBox="[158,1428,188,1786]" captionTargetId="figure-23@16.[151,1436,181,1793]" captionTargetPageId="16" captionText="FIGURES 3946. Aedeagus of Trichodromeus (Figs 3940: T. armatus (Kashmir: Kolahoi Glacier), Figs 4142: T. jaegeri (Nepal: Gandaki), Figs 4346: T. ketmeniensis (Figs 4344: Kazakhstan, Kulbastau), Fig. 45: Kyrgyzstan, Sary-Chelek, Fig. 46: Kazakhstan, Dolaity): 39, 42, 43, 4546—parameral view, 40, 42, 44—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023043" httpUri="https://zenodo.org/record/14023043/files/figure.png" pageId="15" pageNumber="16">3940</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C73FFEAFF3CFE488C3DCBBB" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8A54F1425C73FFEAFF3CFE488B6FCBDA" blockId="15.[151,1326,399,454]" box="[151,636,399,423]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AAFBD66C5C73FFEAFF3CFE488B6FCBDA" box="[151,636,399,423]" pageId="15" pageNumber="16">
<treatmentCitation id="0B4AD7535C73FFEAFF3CFE488B6BCBDA" author="Cameron, M." box="[151,632,399,423]" page="144" pageId="15" pageNumber="16" year="1941">
<taxonomicName id="4DEB8AC15C73FFEAFF3CFE488B6BCBDA" ID-CoL="3NBJT" authority="Cameron, 1941: 144" authorityName="Cameron" authorityPageNumber="144" authorityYear="1941" box="[151,632,399,423]" class="Insecta" family="Staphylinidae" genus="Hygrogoeus" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B89F2D505C73FFEAFF3CFE488801CBDA" box="[151,274,400,423]" italics="true" pageId="15" pageNumber="16">Hygrogoeus</emphasis>
[sic]
<emphasis id="B89F2D505C73FFEAFEE7FE48888CCBDA" box="[332,415,400,423]" italics="true" pageId="15" pageNumber="16">armatus</emphasis>
<bibRefCitation id="EE7A8CB35C73FFEAFE0EFE578B6BCBDA" author="Cameron, M." box="[421,632,399,423]" pageId="15" pageNumber="16" pagination="142 - 147" refId="ref43534" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). The Proceedings of the Royal Entomological Society of London (B), 10, 142 - 147. https: // doi. org / 10.1111 / j. 1365 - 3113.1941. tb 00709. x" type="journal article" year="1941">Cameron, 1941: 144</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8A54F1425C73FFEAFF3CFE768C3DCBBB" blockId="15.[151,1326,399,454]" box="[151,1326,430,454]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AAFBD66C5C73FFEAFF3CFE768C3DCBBB" box="[151,1326,430,454]" pageId="15" pageNumber="16">
<taxonomicName id="4DEB8AC15C73FFEAFF3CFE76889ACBBB" ID-CoL="58CBH" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[151,393,430,454]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B89F2D505C73FFEAFF3CFE76889ACBBB" box="[151,393,430,454]" italics="true" pageId="15" pageNumber="16">Trichodromeus armatus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0B4AD7535C73FFEAFE3DFE768B5FCBBB" author="Zerche, L." box="[406,588,430,454]" page="134" pageId="15" pageNumber="16" year="1992">
<bibRefCitation id="EE7A8CB35C73FFEAFE3DFE768B5FCBBB" author="Zerche, L." box="[406,588,430,454]" pageId="15" pageNumber="16" pagination="105 - 142" refId="ref46115" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992: 134</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0B4AD7535C73FFEAFDFCFE768A05CBBB" author="Herman, L. H." box="[599,790,430,454]" page="373" pageId="15" pageNumber="16" year="2001">
<bibRefCitation id="EE7A8CB35C73FFEAFDFCFE768A05CBBB" author="Herman, L. H." box="[599,790,430,454]" pageId="15" pageNumber="16" pagination="1 - 7" refId="ref44275" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001: 373</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0B4AD7535C73FFEAFC8AFE768AF4CBBB" author="Smetana, A." box="[801,999,430,454]" page="251" pageId="15" pageNumber="16" year="2004">
<bibRefCitation id="EE7A8CB35C73FFEAFC8AFE768AF4CBBB" author="Smetana, A." box="[801,999,430,454]" pageId="15" pageNumber="16" refId="ref45884" refString="Smetana, A. (2004) Subfamily Omaliinae MacLeay, 1825. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 237 - 368." type="book" year="2004">Smetana 2004: 251</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0B4AD7535C73FFEAFC5AFE768C39CBBB" author="Schulke, M. &amp; Smetana, A." box="[1009,1322,430,454]" page="326" pageId="15" pageNumber="16" year="2015">
<bibRefCitation id="EE7A8CB35C73FFEAFC5AFE768C39CBBB" author="Schulke, M. &amp; Smetana, A." box="[1009,1322,430,454]" pageId="15" pageNumber="16" refId="ref45143" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015: 326</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C73FFEAFF3CFE2C8BAFC977" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8A54F1425C73FFEAFF3CFE2C8D9BC807" blockId="15.[151,1437,500,995]" pageId="15" pageNumber="16">
<emphasis id="B89F2D505C73FFEAFF3CFE2C88AEC873" bold="true" box="[151,445,500,526]" pageId="15" pageNumber="16">Type material examined.</emphasis>
<typeStatus id="55504FE05C73FFEAFE68FE2C8B38C873" box="[451,555,500,526]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
<specimenCount id="9CED3ACB5C73FFEAFD9AFE2D8B5AC872" box="[561,585,501,527]" count="1" pageId="15" pageNumber="16" type="male"></specimenCount>
:
<typeStatus id="55504FE05C73FFEAFDF3FE2C8B8DC873" box="[600,670,500,526]" pageId="15" pageNumber="16" type="holotype">Holo-</typeStatus>
| type &lt;round label with red margin&gt;, Kashmir | Aphar Wat |
<quantity id="4D135CA75C73FFEAFAE4FE2C89A3C84F" metricMagnitude="3" metricUnit="m" metricValue="3.9624" pageId="15" pageNumber="16" unit="ft" value="13000.0">13,000 ft.</quantity>
| Dr.Cameron &lt;printed&gt;, H. | armatus | TYPE [red, underlined]
<collectingRegion id="482F3FA05C73FFEAFC30FDC08AC5C84F" box="[923,982,536,562]" country="Myanmar" name="Chin" pageId="15" pageNumber="16">Cam</collectingRegion>
. &lt;handwritten in black&gt;, M.Cameron | Bequest. | B.M. 1955-147. &lt;printed&gt;, UNIQUE TYPE | H.K.Kenward [printed] | det. 197[printed]7 &lt;handwritten in black&gt;,
<taxonomicName id="4DEB8AC15C73FFEAFEB9FDB888D2C807" authorityName="Luze" authorityYear="1903" box="[274,449,608,634]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Trichodromeus</taxonomicName>
|
<taxonomicName id="4DEB8AC15C73FFEAFE7EFDB98B90C807" authority="(Cam.)" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[469,643,608,635]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="armatus">armatus (Cam.)</taxonomicName>
| Shavrin A.V. det. 2016 &lt;printed&gt; (
<collectionCode id="ECFA69875C73FFEAFB89FDB88D6FC807" box="[1058,1148,608,634]" pageId="15" pageNumber="16">BMNH</collectionCode>
).
</paragraph>
<paragraph id="8A54F1425C73FFEAFF6CFD5C8BAFC977" blockId="15.[151,1437,500,995]" pageId="15" pageNumber="16">
<emphasis id="B89F2D505C73FFEAFF6CFD5C8A3EC8E3" bold="true" box="[199,813,644,670]" pageId="15" pageNumber="16">
Additional material examined.
<collectingCountry id="F2FCB1D25C73FFEAFDE5FD5C8BB6C8E3" box="[590,677,644,670]" name="India" pageId="15" pageNumber="16">INDIA</collectingCountry>
: KASHMIR:
</emphasis>
<materialsCitation id="3A83FB1F5C73FFEAFC90FD5C8B47C8BF" collectionCode="BMNH" collectorName="Hygrogeus'" country="India" elevation="3962" location="Kashmir Aphar Wat" municipality="Dr. Cameron'" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1">
<specimenCount id="9CED3ACB5C73FFEAFC90FD5C8A7DC8E2" box="[827,878,644,671]" count="1" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
:
<location id="8F34A7995C73FFEAFC2EFD5C8D66C8E2" LSID="urn:lsid:plazi:treatment:024240545C73FFEAFF3CFEFE8B5AC99F:8F34A7995C73FFEAFC2EFD5C8D66C8E2" box="[901,1141,644,671]" country="India" municipality="Dr. Cameron'" name="Kashmir Aphar Wat" pageId="15" pageNumber="16">Kashmir Aphar Wat</location>
<quantity id="4D135CA75C73FFEAFB28FD5C8DE6C8E3" box="[1155,1269,644,670]" metricMagnitude="3" metricUnit="m" metricValue="3.9624" pageId="15" pageNumber="16" unit="ft" value="13000.0">
<elevation id="01C616715C73FFEAFB28FD5C8DE6C8E3" box="[1155,1269,644,670]" metricMagnitude="3" metricUnit="m" metricValue="3.9624" pageId="15" pageNumber="16" unit="ft" value="13000.0">13,000 ft.</elevation>
</quantity>
<collectingMunicipality id="6A306B385C73FFEAFAA9FD5C8C8BC8E3" box="[1282,1432,644,670]" pageId="15" pageNumber="16">Dr.Cameron</collectingMunicipality>
,
<collectorName id="271E94945C73FFEAFF37FD70883BC8BF" box="[156,296,680,706]" pageId="15" pageNumber="16">Hygrogeus</collectorName>
,
<taxonomicName id="4DEB8AC15C73FFEAFE9FFD7088CEC8BF" authority="Cam." authorityName="Cam." box="[308,477,680,706]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="armatus">armatus Cam.</taxonomicName>
(
<collectionCode id="ECFA69875C73FFEAFE59FD708B5FC8BF" box="[498,588,680,706]" pageId="15" pageNumber="16">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3A83FB1F5C73FFEAFDCAFD708B20C89B" collectionCode="BMNH" collectorName="M. Cameron Bequest." country="India" elevation="3962" location="Kashmir Aphar Wat" municipality="Dr. Cameron'" pageId="15" pageNumber="16" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="9CED3ACB5C73FFEAFDCAFD708BB0C8BE" box="[609,675,680,707]" count="3" pageId="15" pageNumber="16" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9CED3ACB5C73FFEAFD1BFD708BE0C8BE" box="[688,755,680,707]" count="3" pageId="15" pageNumber="16" type="female">3 ♀♀</specimenCount>
:
<location id="8F34A7995C73FFEAFCAEFD708AF8C8BE" LSID="urn:lsid:plazi:treatment:024240545C73FFEAFF3CFEFE8B5AC99F:8F34A7995C73FFEAFCAEFD708AF8C8BE" box="[773,1003,680,707]" country="India" municipality="Dr. Cameron'" name="Kashmir Aphar Wat" pageId="15" pageNumber="16">Kashmir Aphar Wat</location>
<quantity id="4D135CA75C73FFEAFC5FFD708D71C8BF" box="[1012,1122,680,706]" metricMagnitude="3" metricUnit="m" metricValue="3.9624" pageId="15" pageNumber="16" unit="ft" value="13000.0">
<elevation id="01C616715C73FFEAFC5FFD708D71C8BF" box="[1012,1122,680,706]" metricMagnitude="3" metricUnit="m" metricValue="3.9624" pageId="15" pageNumber="16" unit="ft" value="13000.0">13,000 ft.</elevation>
</quantity>
<collectingMunicipality id="6A306B385C73FFEAFBC1FD708C13C8BF" box="[1130,1280,680,706]" pageId="15" pageNumber="16">Dr.Cameron</collectingMunicipality>
,
<collectorName id="271E94945C73FFEAFABAFD7089EAC89A" pageId="15" pageNumber="16">M.Cameron Bequest.</collectorName>
B.M. 1955-147. (
<collectionCode id="ECFA69875C73FFEAFE7AFD148B38C89B" box="[465,555,716,742]" pageId="15" pageNumber="16">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3A83FB1F5C73FFEAFDEAFD148D4BC89B" box="[577,1112,716,743]" collectingDate="1981-07-13" collectorName="G. Ledoux" country="India" location="Baltal." pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1">
<specimenCount id="9CED3ACB5C73FFEAFDEAFD148B63C89A" box="[577,624,716,743]" count="1" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
:
<location id="8F34A7995C73FFEAFDD4FD148BDAC89B" LSID="urn:lsid:plazi:treatment:024240545C73FFEAFF3CFEFE8B5AC99F:8F34A7995C73FFEAFDD4FD148BDAC89B" box="[639,713,716,742]" country="India" name="Baltal." pageId="15" pageNumber="16">Baltal.</location>
<date id="FE55D7825C73FFEAFD78FD148A47C89B" box="[723,852,716,742]" pageId="15" pageNumber="16" value="1981-07-13">
<collectingDate id="EE112E6A5C73FFEAFD78FD148A47C89B" box="[723,852,716,742]" pageId="15" pageNumber="16" value="1981-07-13">13.07.1981</collectingDate>
</date>
.
<collectorName id="271E94945C73FFEAFCCAFD148ACFC89B" box="[865,988,716,742]" pageId="15" pageNumber="16">G. Ledoux</collectorName>
leg. (cSh)
</materialsCitation>
;
<materialsCitation id="3A83FB1F5C73FFEAFBCDFD148BAAC977" collectingDate="2007-11-08" collectorName="NE Srinagar &amp; C. Reuter" country="India" elevation="3500" location="Kolahoi Glacier." pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1">
<specimenCount id="9CED3ACB5C73FFEAFBCDFD148D85C89A" box="[1126,1174,716,743]" count="1" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
:
<collectorName id="271E94945C73FFEAFB0EFD158C26C89B" box="[1189,1333,716,743]" pageId="15" pageNumber="16">NE Srinagar</collectorName>
,
<location id="8F34A7995C73FFEAFAE9FD1489E3C977" LSID="urn:lsid:plazi:treatment:024240545C73FFEAFF3CFEFE8B5AC99F:8F34A7995C73FFEAFAE9FD1489E3C977" country="India" name="Kolahoi Glacier." pageId="15" pageNumber="16">Kolahoi Glacier.</location>
<quantity id="4D135CA75C73FFEAFF5DFD28885DC976" box="[246,334,752,779]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="m" value="3500.0">
<elevation id="01C616715C73FFEAFF5DFD28885DC976" box="[246,334,752,779]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="m" value="3500.0">3500 m</elevation>
</quantity>
.
<date id="FE55D7825C73FFEAFEF2FD2888CAC977" box="[345,473,752,778]" pageId="15" pageNumber="16" value="2007-11-08">
<collectingDate id="EE112E6A5C73FFEAFEF2FD2888CAC977" box="[345,473,752,778]" pageId="15" pageNumber="16" value="2007-11-08">11.08.2007</collectingDate>
</date>
.
<collectorName id="271E94945C73FFEAFE4FFD288B5CC976" box="[484,591,752,779]" pageId="15" pageNumber="16">C. Reuter</collectorName>
leg. (cF)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C73FFEAFF6CFCCC8B5AC99F" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8A54F1425C73FFEAFF6CFCCC8B5AC99F" blockId="15.[151,1437,500,995]" pageId="15" pageNumber="16">
<emphasis id="B89F2D505C73FFEAFF6CFCCC8828C953" bold="true" box="[199,315,788,814]" pageId="15" pageNumber="16">Remarks.</emphasis>
<taxonomicName id="4DEB8AC15C73FFEAFEEFFCCD8B22C953" authorityName="Cameron" authorityYear="1941" box="[324,561,789,814]" class="Insecta" family="Staphylinidae" genus="Hygrogoeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B89F2D505C73FFEAFEEFFCCD8B22C953" box="[324,561,789,814]" italics="true" pageId="15" pageNumber="16">Hygrogoeus armatus</emphasis>
</taxonomicName>
was originally described based on the
<typeStatus id="55504FE05C73FFEAFC58FCCC8D46C953" box="[1011,1109,788,814]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
from “Kashmir: Aphar Wat, altitude
<quantity id="4D135CA75C73FFEAFF58FCE0886BC92F" box="[243,376,824,850]" metricMagnitude="3" metricUnit="m" metricValue="3.9624" pageId="15" pageNumber="16" unit="ft" value="13000.0">13,000 feet</quantity>
”.
<bibRefCitation id="EE7A8CB35C73FFEAFE26FCE08B3EC92F" author="Zerche, L." box="[397,557,824,850]" pageId="15" pageNumber="16" pagination="105 - 142" refId="ref46115" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche (1992)</bibRefCitation>
transferred it to
<taxonomicName id="4DEB8AC15C73FFEAFD40FCE08A87C92F" authorityName="Luze" authorityYear="1903" box="[747,916,824,850]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B89F2D505C73FFEAFD40FCE08A87C92F" box="[747,916,824,850]" italics="true" pageId="15" pageNumber="16">Trichodromeus</emphasis>
</taxonomicName>
and recorded from
<collectingCountry id="F2FCB1D25C73FFEAFBD8FCE08DC5C92F" box="[1139,1238,824,850]" name="Pakistan" pageId="15" pageNumber="16">Pakistan</collectingCountry>
. The shape of the body is variable (
<figureCitation id="12D0EDC75C73FFEAFEFCFC8488AEC90B" box="[343,445,860,886]" captionStart="FIGURES 16" captionStartId="7.[151,264,1732,1756]" captionTargetBox="[157,1429,187,1702]" captionTargetId="figure-16@7.[151,1435,181,1708]" captionTargetPageId="7" captionText="FIGURES 16. Habitus of Hygrogeus (Fig. 1) and Trichodromeus (Figs 26): 1—H. aemulus (Tirol), 2—T. armatus (Kashmir: Baltal), 3—T. armatus (Kashmir: Kolahoi Glacier), 4—T. ketmeniensis (Kyrgyzstan: Chatkal Mts.), 5—T. ovchinnikovi (holotype), 6—T. kumaonensis (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023035" httpUri="https://zenodo.org/record/14023035/files/figure.png" pageId="15" pageNumber="16">Figs 23</figureCitation>
). The studied specimen from Kolahoi Glacier has a narrower body and shorter elytra, but the shape of the aedeagus (
<figureCitation id="12D0EDC75C73FFEAFE68FC588B57C9E7" box="[451,580,896,922]" captionStart="FIGURES 3946" captionStartId="16.[151,264,1817,1841]" captionTargetBox="[158,1428,188,1786]" captionTargetId="figure-23@16.[151,1436,181,1793]" captionTargetPageId="16" captionText="FIGURES 3946. Aedeagus of Trichodromeus (Figs 3940: T. armatus (Kashmir: Kolahoi Glacier), Figs 4142: T. jaegeri (Nepal: Gandaki), Figs 4346: T. ketmeniensis (Figs 4344: Kazakhstan, Kulbastau), Fig. 45: Kyrgyzstan, Sary-Chelek, Fig. 46: Kazakhstan, Dolaity): 39, 42, 43, 4546—parameral view, 40, 42, 44—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023043" httpUri="https://zenodo.org/record/14023043/files/figure.png" pageId="15" pageNumber="16">Figs 3940</figureCitation>
) is identical with that in other studied specimens. From all known species of the genus
<taxonomicName id="4DEB8AC15C73FFEAFEA0FC7D8890C9C3" authorityName="Cameron" authorityYear="1941" box="[267,387,933,958]" class="Insecta" family="Staphylinidae" genus="Hygrogoeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B89F2D505C73FFEAFEA0FC7D8890C9C3" box="[267,387,933,958]" italics="true" pageId="15" pageNumber="16">T. armatus</emphasis>
</taxonomicName>
can be easily recognized by the unusual modification in the middle of the inner side of each metatibia with a long and strong tooth.
</paragraph>
</subSubSection>
</treatment>
</document>