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<document id="D9A68BBD5ACE6616E54D86D1EB3665E1" ID-DOI="10.11646/zootaxa.5531.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14023033" ID-ZooBank="27F9603C-D2AB-4133-AFB8-5A674ACCD110" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730453991549" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="024240545C67FFFEFF3CFF4F89C2C82B" docLanguage="en" docName="zootaxa.5531.1.1.pdf" docOrigin="Zootaxa 5531 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5531.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Trichodromeus pusillus" docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="FE7B382C5C7CFFE5FFABFFD88913CA7D" masterDocTitle="Review of Hygrogeus Mulsant &amp; Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="28" updateTime="1730454835898" updateUser="ExternalLinkService">
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<mods:title id="1E262C71C72964D15D0F16B8D4B537C7">Review of Hygrogeus Mulsant &amp; Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae)</mods:title>
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<mods:namePart id="83ECFF070D53A4F40C2FFCB28DA9B93B">Shavrin, Alexey V.</mods:namePart>
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<mods:classification id="085AE85D9DABF086CEBF66E6BD39422F">journal article</mods:classification>
<mods:identifier id="19105E321907DD481404CB94ED68E2AB" type="DOI">10.11646/zootaxa.5531.1.1</mods:identifier>
<mods:identifier id="35E994FDA5E652C04E027EFE603A453D" type="ISSN">1175-5334</mods:identifier>
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<treatment id="024240545C67FFFEFF3CFF4F89C2C82B" ID-DOI="http://doi.org/10.5281/zenodo.14023113" ID-Zenodo-Dep="14023113" LSID="urn:lsid:plazi:treatment:024240545C67FFFEFF3CFF4F89C2C82B" httpUri="http://treatment.plazi.org/id/024240545C67FFFEFF3CFF4F89C2C82B" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="C2F1A2C95C67FFFEFF3CFF4F8B69CACC" box="[151,634,151,178]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8A54F1425C67FFFEFF3CFF4F8B69CACC" blockId="27.[151,634,151,214]" box="[151,634,151,178]" pageId="27" pageNumber="28">
<heading id="D11C462E5C67FFFEFF3CFF4F8B69CACC" bold="true" box="[151,634,151,178]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<taxonomicName id="4DEB8AC15C67FFFEFF3CFF4F8B69CACC" ID-CoL="58CBQ" authority="(Coiffait, 1983)" baseAuthorityName="Coiffait" baseAuthorityYear="1983" box="[151,634,151,178]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B89F2D505C67FFFEFF3CFF4F8B69CACC" bold="true" box="[151,634,151,178]" pageId="27" pageNumber="28">
<emphasis id="B89F2D505C67FFFEFF3CFF4F88A5CACF" bold="true" box="[151,438,151,178]" italics="true" pageId="27" pageNumber="28">Trichodromeus pusillus</emphasis>
(
<bibRefCitation id="EE7A8CB35C67FFFEFE6EFF4F8B61CACC" author="Coiffait, H." box="[453,626,151,177]" pageId="27" pageNumber="28" pagination="273 - 289" refId="ref43923" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France, New Series, 19 (3), 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait, 1983</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C67FFFEFF3CFF6389EACAAB" box="[151,249,187,214]" pageId="27" pageNumber="28" type="description">
<paragraph id="8A54F1425C67FFFEFF3CFF6389EACAAB" blockId="27.[151,634,151,214]" box="[151,249,187,214]" pageId="27" pageNumber="28">
(
<figureCitation id="12D0EDC75C67FFFEFF34FF6389E2CAAB" box="[159,241,187,214]" captionStart="FIGURES 49–51" captionStartId="21.[151,264,830,854]" captionTargetBox="[179,1397,200,800]" captionTargetId="figure-16@21.[160,1428,178,806]" captionTargetPageId="21" captionText="FIGURES 49–51. Habitus of Trichodromeus: 49—T. penicillatus (Kazakhstan: Aksu-Dzhabagly), 50—T. pusillus (Kashmir), 51—T. telnovi (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023049" httpUri="https://zenodo.org/record/14023049/files/figure.png" pageId="27" pageNumber="28">Fig. 50</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C67FFFEFF3CFED88A99CB4B" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8A54F1425C67FFFEFF3CFED88ACDCB65" blockId="27.[151,990,256,310]" box="[151,990,256,280]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AAFBD66C5C67FFFEFF3CFED88ACDCB65" box="[151,990,256,280]" pageId="27" pageNumber="28">
<treatmentCitation id="0B4AD7535C67FFFEFF3CFED88B2DCB65" author="Coiffait, H." box="[151,574,256,280]" page="279" pageId="27" pageNumber="28" year="1983">
<taxonomicName id="4DEB8AC15C67FFFEFF3CFED88B2DCB65" ID-CoL="4NGRK" authority="Coiffait, 1983: 279" authorityName="Coiffait" authorityPageNumber="279" authorityYear="1983" box="[151,574,256,280]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B89F2D505C67FFFEFF3CFED88867CB65" box="[151,372,256,280]" italics="true" pageId="27" pageNumber="28">Geodromicus pusillus</emphasis>
<bibRefCitation id="EE7A8CB35C67FFFEFED1FED88B2DCB65" author="Coiffait, H." box="[378,574,256,280]" pageId="27" pageNumber="28" pagination="273 - 289" refId="ref43923" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France, New Series, 19 (3), 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait, 1983: 279</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0B4AD7535C67FFFEFDE2FED88A1ACB65" author="Herman, L. H." box="[585,777,256,280]" page="303" pageId="27" pageNumber="28" year="2001">
<bibRefCitation id="EE7A8CB35C67FFFEFDE2FED88A1ACB65" author="Herman, L. H." box="[585,777,256,280]" pageId="27" pageNumber="28" pagination="1 - 7" refId="ref44275" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001: 303</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0B4AD7535C67FFFEFCB8FED88AC9CB65" author="Smetana, A." box="[787,986,256,280]" page="244" pageId="27" pageNumber="28" year="2004">
<bibRefCitation id="EE7A8CB35C67FFFEFCB8FED88AC9CB65" author="Smetana, A." box="[787,986,256,280]" pageId="27" pageNumber="28" refId="ref45884" refString="Smetana, A. (2004) Subfamily Omaliinae MacLeay, 1825. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 237 - 368." type="book" year="2004">Smetana 2004: 244</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8A54F1425C67FFFEFF3CFEC68A99CB4B" blockId="27.[151,990,256,310]" box="[151,906,286,310]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AAFBD66C5C67FFFEFF3CFEC68A99CB4B" box="[151,906,286,310]" pageId="27" pageNumber="28">
<taxonomicName id="4DEB8AC15C67FFFEFF3CFEC68896CB4B" ID-CoL="58CBQ" baseAuthorityName="Coiffait" baseAuthorityYear="1983" box="[151,389,286,310]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B89F2D505C67FFFEFF3CFEC68896CB4B" box="[151,389,286,310]" italics="true" pageId="27" pageNumber="28">Trichodromeus pusillus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0B4AD7535C67FFFEFE39FEC68B50CB4B" ID-DOI="http://doi.org/10.5281/zenodo.6181902" ID-Zenodo-Dep="6181902" author="Shavrin, A. V." box="[402,579,286,310]" httpUri="http://treatment.plazi.org/id/82358795FF916446FF0DF9C4FA89FDB6" page="66" pageId="27" pageNumber="28" year="2012">
<bibRefCitation id="EE7A8CB35C67FFFEFE39FEC68B50CB4B" author="Shavrin, A. V." box="[402,579,286,310]" pageId="27" pageNumber="28" pagination="66 - 68" refId="ref45201" refString="Shavrin, A. V. (2012) On the taxonomic placement of Geodromicus pusillus Coiffait, 1983 (Coleoptera: Staphylinidae: Omaliinae). Zootaxa, 3552 (1), 66 - 68. https: // doi. org / 10.11646 / zootaxa. 3552.1.4" type="journal article" year="2012">Shavrin 2012: 66</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0B4AD7535C67FFFEFDE6FEC68A95CB4B" author="Schulke, M. &amp; Smetana, A." box="[589,902,286,310]" page="326" pageId="27" pageNumber="28" year="2015">
<bibRefCitation id="EE7A8CB35C67FFFEFDE6FEC68A95CB4B" author="Schulke, M. &amp; Smetana, A." box="[589,902,286,310]" pageId="27" pageNumber="28" refId="ref45143" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015: 326</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C67FFFEFF3CFEBC8A39CBBB" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8A54F1425C67FFFEFF3CFEBC8A39CBBB" blockId="27.[151,1436,356,598]" pageId="27" pageNumber="28">
<emphasis id="B89F2D505C67FFFEFF3CFEBC8892CB03" bold="true" box="[151,385,356,382]" pageId="27" pageNumber="28">Material examined.</emphasis>
<materialsCitation id="3A83FB1F5C67FFFEFE22FEBC8A35CBBB" collectingDate="1997-08-26" collectingDateMax="2003-10-07" collectingDateMin="1997-08-26" collectionCode="NSMT" collectorName="W. Heinz &amp; SW Garhi. &amp; NW Junkar." country="Pakistan" elevation="4050" location="Deosai" pageId="27" pageNumber="28" specimenCount="80" specimenCount-female="48" specimenCount-male="32">
<emphasis id="B89F2D505C67FFFEFE22FEBC8B9DCB03" bold="true" box="[393,654,356,382]" pageId="27" pageNumber="28">
<collectingCountry id="F2FCB1D25C67FFFEFE22FEBC8B09CB03" box="[393,538,356,382]" name="Pakistan" pageId="27" pageNumber="28">PAKISTAN</collectingCountry>
: SKARDU:
</emphasis>
<specimenCount id="9CED3ACB5C67FFFEFD3DFEBD8BD1CB02" box="[662,706,357,383]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9CED3ACB5C67FFFEFD64FEBD8BEFCB02" box="[719,764,357,383]" count="1" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
:
<location id="8F34A7995C67FFFEFCA2FEBC8A49CB03" LSID="urn:lsid:plazi:treatment:024240545C67FFFEFF3CFF4F89C2C82B:8F34A7995C67FFFEFCA2FEBC8A49CB03" box="[777,858,356,382]" country="Pakistan" name="Deosai" pageId="27" pageNumber="28">Deosai</location>
, Satpara.
<elevation id="01C616715C67FFFEFC63FEBD8DB3CB03" box="[968,1184,356,383]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="27" pageNumber="28" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">
<quantity id="4D135CA75C67FFFEFC63FEBD8D77CB02" box="[968,1124,357,383]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="27" pageNumber="28" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">4000–4100 m</quantity>
a.s.l.
</elevation>
<date id="FE55D7825C67FFFEFB0CFEBC8C3BCB03" box="[1191,1320,356,382]" pageId="27" pageNumber="28" value="1997-08-26">
<collectingDate id="EE112E6A5C67FFFEFB0CFEBC8C3BCB03" box="[1191,1320,356,382]" pageId="27" pageNumber="28" value="1997-08-26">26.08.1997</collectingDate>
</date>
.
<collectorName id="271E94945C67FFFEFA98FEBD8C8FCB03" box="[1331,1436,356,383]" pageId="27" pageNumber="28">W. Heinz</collectorName>
leg. (
<collectionCode id="ECFA69875C67FFFEFF7AFE518836CBDE" box="[209,293,393,419]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="27" pageNumber="28">NSMT</collectionCode>
);
<emphasis id="B89F2D505C67FFFEFE90FE508B35CBDF" bold="true" box="[315,550,392,418]" pageId="27" pageNumber="28">JAMMU &amp; KASHMIR:</emphasis>
<specimenCount id="9CED3ACB5C67FFFEFD84FE518B92CBDE" box="[559,641,393,419]" count="29" pageId="27" pageNumber="28" type="male">29 ♂♂</specimenCount>
,
<specimenCount id="9CED3ACB5C67FFFEFD24FE518BF3CBDE" box="[655,736,393,419]" count="47" pageId="27" pageNumber="28" type="female">47 ♀♀</specimenCount>
:
<collectorName id="271E94945C67FFFEFD44FE518A7ACBDF" box="[751,873,392,419]" pageId="27" pageNumber="28">SW Garhi.</collectorName>
<elevation id="01C616715C67FFFEFCDAFE508D15CBDF" box="[881,1030,392,419]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="28" unit="m" value="1500.0">
<quantity id="4D135CA75C67FFFEFCDAFE508ADBCBDE" box="[881,968,392,419]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="28" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
10–
<date id="FE55D7825C67FFFEFB9DFE518DA5CBDE" box="[1078,1206,393,419]" pageId="27" pageNumber="28" value="2003-07-20">
<collectingDate id="EE112E6A5C67FFFEFB9DFE518DA5CBDE" box="[1078,1206,393,419]" pageId="27" pageNumber="28" value="2003-07-20">20.07.2003</collectingDate>
</date>
(cSch, cSh);
<specimenCount id="9CED3ACB5C67FFFEFAF8FE518C8ACBDE" box="[1363,1433,393,419]" count="2" pageId="27" pageNumber="28" type="male">2 ♂♂</specimenCount>
: same data,
<collectorName id="271E94945C67FFFEFEBFFE758889CBBB" box="[276,410,428,455]" pageId="27" pageNumber="28">NW Junkar.</collectorName>
<elevation id="01C616715C67FFFEFE0AFE758B22CBBB" box="[417,561,428,455]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="4000.0">
<quantity id="4D135CA75C67FFFEFE0AFE7588E5CBBA" box="[417,502,429,455]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="4000.0">4000 m</quantity>
a.s.l.
</elevation>
01–
<date id="FE55D7825C67FFFEFDCBFE758BF3CBBA" box="[608,736,429,455]" pageId="27" pageNumber="28" value="2003-10-07">
<collectingDate id="EE112E6A5C67FFFEFDCBFE758BF3CBBA" box="[608,736,429,455]" pageId="27" pageNumber="28" value="2003-10-07">10.07.2003</collectingDate>
</date>
(cSh)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C67FFFEFF6CFE0989C2C82B" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8A54F1425C67FFFEFF6CFE0989C2C82B" blockId="27.[151,1436,356,598]" pageId="27" pageNumber="28">
<emphasis id="B89F2D505C67FFFEFF6CFE098828CB96" bold="true" box="[199,315,465,491]" pageId="27" pageNumber="28">Remarks.</emphasis>
<taxonomicName id="4DEB8AC15C67FFFEFEEAFE088B27CB97" authorityName="Coiffait" authorityYear="1983" box="[321,564,464,490]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B89F2D505C67FFFEFEEAFE088B27CB97" box="[321,564,464,490]" italics="true" pageId="27" pageNumber="28">Geodromicus pusillus</emphasis>
</taxonomicName>
was originally described from Kashmir,
<collectingCountry id="F2FCB1D25C67FFFEFC53FE088D26CB97" box="[1016,1077,464,490]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
.
<bibRefCitation id="EE7A8CB35C67FFFEFB94FE088DF4CB96" author="Shavrin, A. V." box="[1087,1255,464,491]" pageId="27" pageNumber="28" pagination="66 - 68" refId="ref45201" refString="Shavrin, A. V. (2012) On the taxonomic placement of Geodromicus pusillus Coiffait, 1983 (Coleoptera: Staphylinidae: Omaliinae). Zootaxa, 3552 (1), 66 - 68. https: // doi. org / 10.11646 / zootaxa. 3552.1.4" type="journal article" year="2012">Shavrin (2012)</bibRefCitation>
redescribed and transferred it to the genus
<taxonomicName id="4DEB8AC15C67FFFEFE69FE2C8B78C873" authorityName="Luze" authorityYear="1903" box="[450,619,500,526]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B89F2D505C67FFFEFE69FE2C8B78C873" box="[450,619,500,526]" italics="true" pageId="27" pageNumber="28">Trichodromeus</emphasis>
</taxonomicName>
, and provided additional records from Kashmir.
<taxonomicName id="4DEB8AC15C67FFFEFB3FFE2C8C8FC873" baseAuthorityName="Coiffait" baseAuthorityYear="1983" box="[1172,1436,500,526]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B89F2D505C67FFFEFB3FFE2C8C8FC873" box="[1172,1436,500,526]" italics="true" pageId="27" pageNumber="28">Trichodromeus pusillus</emphasis>
</taxonomicName>
(
<figureCitation id="12D0EDC75C67FFFEFF34FDC089E1C84E" box="[159,242,536,563]" captionStart="FIGURES 49–51" captionStartId="21.[151,264,830,854]" captionTargetBox="[179,1397,200,800]" captionTargetId="figure-16@21.[160,1428,178,806]" captionTargetPageId="21" captionText="FIGURES 49–51. Habitus of Trichodromeus: 49—T. penicillatus (Kazakhstan: Aksu-Dzhabagly), 50—T. pusillus (Kashmir), 51—T. telnovi (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023049" httpUri="https://zenodo.org/record/14023049/files/figure.png" pageId="27" pageNumber="28">Fig. 50</figureCitation>
) is known from north-eastern
<collectingCountry id="F2FCB1D25C67FFFEFDE5FDC08BBEC84F" box="[590,685,536,562]" name="Pakistan" pageId="27" pageNumber="28">Pakistan</collectingCountry>
and Kashmir,
<collectingCountry id="F2FCB1D25C67FFFEFCFEFDC08A81C84F" box="[853,914,536,562]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
. It is here recorded from
<collectingCountry id="F2FCB1D25C67FFFEFB1FFDC08C00C84F" box="[1204,1299,536,562]" name="Pakistan" pageId="27" pageNumber="28">Pakistan</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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