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<document id="8BE8C2BFB02F0053E7EC13D67F0842AD" ID-DOI="10.11646/zootaxa.5531.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14023033" ID-ZooBank="27F9603C-D2AB-4133-AFB8-5A674ACCD110" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730453991549" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="024240545C5FFFC0FF3CFB218814C88C" docLanguage="en" docName="zootaxa.5531.1.1.pdf" docOrigin="Zootaxa 5531 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5531.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Asiageodromicus dolpoensis Shavrin 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="38" masterDocId="FE7B382C5C7CFFE5FFABFFD88913CA7D" masterDocTitle="Review of Hygrogeus Mulsant &amp; Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="36" updateTime="1730454835898" updateUser="ExternalLinkService">
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<mods:title id="CC72A130A62FC7F66C9FFBBA8E23C644">Review of Hygrogeus Mulsant &amp; Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae)</mods:title>
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<mods:date id="6CF2E658AE46099889CC836A8933C77D">2024</mods:date>
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<mods:classification id="C2E340EAC92A5208A555388386DADC9C">journal article</mods:classification>
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<treatment id="024240545C5FFFC0FF3CFB218814C88C" ID-DOI="http://doi.org/10.5281/zenodo.14023124" ID-Zenodo-Dep="14023124" LSID="urn:lsid:plazi:treatment:024240545C5FFFC0FF3CFB218814C88C" httpUri="http://treatment.plazi.org/id/024240545C5FFFC0FF3CFB218814C88C" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
<subSubSection id="C2F1A2C95C5FFFC6FF3CFB218A36CF69" box="[151,805,1273,1300]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8A54F1425C5FFFC6FF3CFB218A36CF69" blockId="35.[151,805,1273,1336]" box="[151,805,1273,1300]" pageId="35" pageNumber="36">
<heading id="D11C462E5C5FFFC6FF3CFB218A36CF69" bold="true" box="[151,805,1273,1300]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<emphasis id="B89F2D505C5FFFC6FF3CFB218A36CF69" bold="true" box="[151,805,1273,1300]" pageId="35" pageNumber="36">
<taxonomicName id="4DEB8AC15C5FFFC6FF3CFB218BA1CF6E" authority="(Coiffait, 1982)" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Coiffait" baseAuthorityYear="1982" box="[151,690,1273,1300]" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="dolpoensis" status="comb. nov.">
<emphasis id="B89F2D505C5FFFC6FF3CFB2188FDCF69" bold="true" box="[151,494,1273,1300]" italics="true" pageId="35" pageNumber="36">Asiageodromicus dolpoensis</emphasis>
(
<bibRefCitation id="EE7A8CB35C5FFFC6FE56FB218BB9CF6E" author="Coiffait, H." box="[509,682,1273,1299]" pageId="35" pageNumber="36" pagination="21 - 179" refId="ref43884" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A3AC902B5C5FFFC6FD11FB228A36CF69" box="[698,805,1274,1300]" pageId="35" pageNumber="36" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C5FFFC6FF3CFAC68868CF45" box="[151,379,1310,1336]" pageId="35" pageNumber="36" type="description">
<paragraph id="8A54F1425C5FFFC6FF3CFAC68868CF45" blockId="35.[151,805,1273,1336]" box="[151,379,1310,1336]" pageId="35" pageNumber="36">
(
<figureCitation id="12D0EDC75C5FFFC6FF34FAC689E5CF45" box="[159,246,1310,1336]" captionStart="FIGURES 66–67" captionStartId="30.[151,264,1791,1815]" captionTargetBox="[158,1429,491,1760]" captionTargetId="figure-97@30.[151,1436,484,1768]" captionTargetPageId="30" captionText="FIGURES 66–67. Habitus of Asiageodromicus: 66—A. dolpoensis (holotype), 67—A. weiperti (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023057" httpUri="https://zenodo.org/record/14023057/files/figure.png" pageId="35" pageNumber="36">Figs 66</figureCitation>
,
<figureCitation id="12D0EDC75C5FFFC6FEA9FAC68858CF45" box="[258,331,1310,1336]" captionStart="FIGURES 79–88" captionStartId="33.[151,264,1893,1917]" captionTargetBox="[158,1429,188,1863]" captionTargetId="figure-16@33.[151,1436,181,1869]" captionTargetPageId="33" captionText="FIGURES 79–88. Aedeagus (Figs 79–82) and abdominal segments (Figs 83–88) of Asiageodromicus (Figs 79–80: A. weiperti (holotype); Figs 81–82; A. dolpoensis (holotype); Figs 83–88: A. crassipalpis): 79, 81—aedeagus, parameral view, 80, 82— aedeagus, lateral view, 83—male tergite VIII, 84—male sternite VIII, 85—male genital segment, 86—female tergite VIII, 87—female sternite VIII, 88—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023065" httpUri="https://zenodo.org/record/14023065/files/figure.png" pageId="35" pageNumber="36">81–82</figureCitation>
,
<figureCitation id="12D0EDC75C5FFFC6FEFDFAC68860CF45" box="[342,371,1310,1336]" captionStart="FIGURE 89" captionStartId="36.[151,250,975,999]" captionTargetBox="[157,1429,187,945]" captionTargetId="figure-23@36.[151,1435,181,951]" captionTargetPageId="36" captionText="FIGURE 89. Distribution of Asiageodromicus in the Himalayan Region: A. crassipalpis (circles), A. dolpoensis (star), A. palpalis (Coiffait, 1977) (squares), A. spadiceus (triangles), A. weiperti sp. n. (diamonds)" figureDoi="http://doi.org/10.5281/zenodo.14023067" httpUri="https://zenodo.org/record/14023067/files/figure.png" pageId="35" pageNumber="36">89</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C5FFFC6FF3CFA848C2DCF09" box="[151,1342,1371,1396]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8A54F1425C5FFFC6FF3CFA848C2DCF09" blockId="35.[151,1342,1371,1396]" box="[151,1342,1371,1396]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AAFBD66C5C5FFFC6FF3CFA848C2DCF09" box="[151,1342,1371,1396]" pageId="35" pageNumber="36">
<treatmentCitation id="0B4AD7535C5FFFC6FF3CFA848B4FCF0E" author="Coiffait, H." box="[151,604,1371,1396]" page="155" pageId="35" pageNumber="36" year="1982">
<taxonomicName id="4DEB8AC15C5FFFC6FF3CFA848B4FCF0E" ID-CoL="3FPH4" authority="Coiffait, 1982: 155" authorityName="Coiffait" authorityPageNumber="155" authorityYear="1982" box="[151,604,1371,1396]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="dolpoensis">
<emphasis id="B89F2D505C5FFFC6FF3CFA848882CF09" box="[151,401,1372,1396]" italics="true" pageId="35" pageNumber="36">Geodromicus dolpoensis</emphasis>
<bibRefCitation id="EE7A8CB35C5FFFC6FE33FA848B4FCF0E" author="Coiffait, H." box="[408,604,1371,1396]" pageId="35" pageNumber="36" pagination="21 - 179" refId="ref43884" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982: 155</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0B4AD7535C5FFFC6FDCDFA848A35CF09" author="Herman, L. H." box="[614,806,1372,1396]" page="294" pageId="35" pageNumber="36" year="2001">
<bibRefCitation id="EE7A8CB35C5FFFC6FDCDFA848A35CF09" author="Herman, L. H." box="[614,806,1372,1396]" pageId="35" pageNumber="36" pagination="1 - 7" refId="ref44275" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001: 294</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0B4AD7535C5FFFC6FC9AFA848AE5CF09" author="Smetana, A." box="[817,1014,1372,1396]" page="243" pageId="35" pageNumber="36" year="2004">
<bibRefCitation id="EE7A8CB35C5FFFC6FC9AFA848AE5CF09" author="Smetana, A." box="[817,1014,1372,1396]" pageId="35" pageNumber="36" refId="ref45884" refString="Smetana, A. (2004) Subfamily Omaliinae MacLeay, 1825. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 237 - 368." type="book" year="2004">Smetana 2004: 243</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0B4AD7535C5FFFC6FBAAFA848C29CF09" author="Schulke, M. &amp; Smetana, A." box="[1025,1338,1371,1396]" page="313" pageId="35" pageNumber="36" year="2015">
<bibRefCitation id="EE7A8CB35C5FFFC6FBAAFA848C29CF09" author="Schulke, M. &amp; Smetana, A." box="[1025,1338,1371,1396]" pageId="35" pageNumber="36" refId="ref45143" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015: 313</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C5FFFC6FF3CFA438B40CC14" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8A54F1425C5FFFC6FF3CFA438B40CC14" blockId="35.[151,1437,1435,2038]" pageId="35" pageNumber="36">
<emphasis id="B89F2D505C5FFFC6FF3CFA4388D3CFC8" bold="true" box="[151,448,1435,1461]" pageId="35" pageNumber="36">Type material examined.</emphasis>
<typeStatus id="55504FE05C5FFFC6FE63FA438B23CFC8" box="[456,560,1435,1461]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
<specimenCount id="9CED3ACB5C5FFFC6FD93FA448B5ECFCB" box="[568,589,1436,1462]" count="1" pageId="35" pageNumber="36" type="male">♂</specimenCount>
(
<figureCitation id="12D0EDC75C5FFFC6FDF6FA438BA2CFC8" box="[605,689,1435,1461]" captionStart="FIGURES 66–67" captionStartId="30.[151,264,1791,1815]" captionTargetBox="[158,1429,491,1760]" captionTargetId="figure-97@30.[151,1436,484,1768]" captionTargetPageId="30" captionText="FIGURES 66–67. Habitus of Asiageodromicus: 66—A. dolpoensis (holotype), 67—A. weiperti (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023057" httpUri="https://zenodo.org/record/14023057/files/figure.png" pageId="35" pageNumber="36">Fig. 66</figureCitation>
; immature specimen; dissected): ‘Dolpo, Aufstieg | zum Namu La |
<quantity id="4D135CA75C5FFFC6FF0FFA678853CFA7" box="[164,320,1471,1498]" metricMagnitude="3" metricUnit="m" metricValue="4.375" metricValueMax="4.45" metricValueMin="4.3" pageId="35" pageNumber="36" unit="m" value="4375.0" valueMax="4450.0" valueMin="4300.0">4300–4450 m</quantity>
|
<date id="FE55D7825C5FFFC6FEFEFA6788F5CFA4" box="[341,486,1471,1498]" pageId="35" pageNumber="36" value="1973-06-17">17. VI. 1973</date>
’ &lt;photo label&gt;, ‘NEPAL-Expedition | Jochen Martens’ &lt;photo label&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘
<typeStatus id="55504FE05C5FFFC6FEA1FA3B8843CF80" box="[266,336,1507,1533]" pageId="35" pageNumber="36" type="holotype">Holo-</typeStatus>
| [handwritten in black] | Typus’ &lt;printed, with narrow black margin&gt;, ‘
<taxonomicName id="4DEB8AC15C5FFFC6FB2BFA3B8C0FCF80" authorityName="Redtenbacher" authorityYear="1857" box="[1152,1308,1507,1533]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| dolpensis [sic!] | H. COIFFAIT det. 19 [printed] 79’ &lt;handwritten&gt;, ‘SNG-Museum | Frankfurt/Main | Loan-000302’ &lt;printed&gt;, ‘Geodromius | dolpensis [sic!] Coiff.’ &lt;handwritten by blue pen&gt;, ‘
<taxonomicName id="4DEB8AC15C5FFFC6FC2FF9F38D5ECC38" authorityName="Shavrin" authorityYear="2024" box="[900,1101,1579,1605]" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="40" phylum="Arthropoda" rank="genus">Asiageodromicus</taxonomicName>
|
<taxonomicName id="4DEB8AC15C5FFFC6FBCAF9F38C83CC38" authority="(Coiffait, 1982)" baseAuthorityName="Coiffait" baseAuthorityYear="1982" box="[1121,1424,1579,1605]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="dolpoensis">
dolpoensis (
<bibRefCitation id="EE7A8CB35C5FFFC6FB4CF9F38C94CC38" author="Coiffait, H." box="[1255,1415,1579,1605]" pageId="35" pageNumber="36" pagination="21 - 179" refId="ref43884" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
)
</taxonomicName>
| Shavrin A.V. det. 2023’ &lt;printed&gt; (
<collectionCode id="ECFA69875C5FFFC6FD88F9978B54CC14" box="[547,583,1615,1641]" pageId="35" pageNumber="36">SF</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C5FFFC0FF6CF9AB8836C81C" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="description">
<paragraph id="8A54F1425C5FFFC6FF6CF9AB8851CCA8" blockId="35.[151,1437,1435,2038]" pageId="35" pageNumber="36">
<emphasis id="B89F2D505C5FFFC6FF6CF9AB8866CCF0" bold="true" box="[199,373,1651,1677]" pageId="35" pageNumber="36">Redescription.</emphasis>
Measurements: HW: 1.05; HL: 0.73; OL: 0.25; TL: 0.16; AL: 4.27; PL: 0.99; PWmax: 1.25; PWmin: 1.17; ESL: 1.95; EW: 1.88; MTbL: 1.75; MTrL: 0.62 (MTrL 1–4: 0.30; MTrL 5: 0.32); AW: 1.75; AedL: 1.10; BL: 4.90.
</paragraph>
<paragraph id="8A54F1425C5FFFC6FF6CF9078B4BCD88" blockId="35.[151,1437,1435,2038]" pageId="35" pageNumber="36">
Habitus as in
<figureCitation id="12D0EDC75C5FFFC6FEC1F90788D3CC84" box="[362,448,1759,1785]" captionStart="FIGURES 66–67" captionStartId="30.[151,264,1791,1815]" captionTargetBox="[158,1429,491,1760]" captionTargetId="figure-97@30.[151,1436,484,1768]" captionTargetPageId="30" captionText="FIGURES 66–67. Habitus of Asiageodromicus: 66—A. dolpoensis (holotype), 67—A. weiperti (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023057" httpUri="https://zenodo.org/record/14023057/files/figure.png" pageId="35" pageNumber="36">Fig. 66</figureCitation>
. Head, pronotum and abdomen brown; mouthparts, antennae, elytra and legs yellow-brown; apical segment of maxillary palpomeres and tarsi yellowish. Head with irregular and dense microsculpture: isodiametric in frontoclypeal portion, slightly coarser between antennal insertion, isodiametric in middle between eyes, finer in portions between anterior margin of eye and median part, coarser on infraorbital portions, subdiagonal in lateroposterior portion, coarser in mediobasal part in front of neck; neck with dense isodiametric microsculpture; pronotum with distinct dense and fine transverse microreticulation, finer in middle, mediobasal portion without meshes; scutellum with fine and moderately dense transverse meshes; elytra without microsculpture; abdomen with very dense transverse microreticulation.
</paragraph>
<caption id="DE94A1CA5C58FFC1FF3CFC178D32CE76" ID-DOI="http://doi.org/10.5281/zenodo.14023067" ID-Zenodo-Dep="14023067" httpUri="https://zenodo.org/record/14023067/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,975,999]" targetBox="[157,1429,187,945]" targetPageId="36" targetType="figure">
<paragraph id="8A54F1425C58FFC1FF3CFC178D32CE76" blockId="36.[151,1437,975,1035]" pageId="36" pageNumber="37">
<emphasis id="B89F2D505C58FFC1FF3CFC178837C99A" bold="true" box="[151,292,975,999]" pageId="36" pageNumber="37">FIGURE 89.</emphasis>
Distribution of
<taxonomicName id="4DEB8AC15C58FFC1FE7AFC178B93C99A" authorityName="Shavrin" authorityYear="2024" box="[465,640,975,999]" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B89F2D505C58FFC1FE7AFC178B93C99A" box="[465,640,975,999]" italics="true" pageId="36" pageNumber="37">Asiageodromicus</emphasis>
</taxonomicName>
in the Himalayan Region:
<taxonomicName id="4DEB8AC15C58FFC1FC08FC178D2EC99A" authorityName="Champion" authorityYear="1920" box="[931,1085,975,999]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crassipalpis">
<emphasis id="B89F2D505C58FFC1FC08FC178D2EC99A" box="[931,1085,975,999]" italics="true" pageId="36" pageNumber="37">A. crassipalpis</emphasis>
</taxonomicName>
(circles),
<taxonomicName id="4DEB8AC15C58FFC1FB00FC178C25C99A" baseAuthorityName="Coiffait" baseAuthorityYear="1982" box="[1195,1334,975,999]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dolpoensis">
<emphasis id="B89F2D505C58FFC1FB00FC178C25C99A" box="[1195,1334,975,999]" italics="true" pageId="36" pageNumber="37">A. dolpoensis</emphasis>
</taxonomicName>
(star),
<taxonomicName id="4DEB8AC15C58FFC1FA2DFC17889ECE76" authority="(Coiffait, 1977)" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Coiffait" baseAuthorityYear="1977" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="palpalis">
<emphasis id="B89F2D505C58FFC1FA2DFC1789FACE76" italics="true" pageId="36" pageNumber="37">A. palpalis</emphasis>
(
<bibRefCitation id="EE7A8CB35C58FFC1FF5DFC2B8894CE76" author="Coiffait, H." box="[246,391,1011,1035]" pageId="36" pageNumber="37" pagination="243 - 275" refId="ref43770" refString="Coiffait, H. (1977) Staphylinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977</bibRefCitation>
)
</taxonomicName>
(squares),
<taxonomicName id="4DEB8AC15C58FFC1FE55FC2B8B6DCE76" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Shavrin" baseAuthorityYear="2019" box="[510,638,1011,1035]" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="spadiceus">
<emphasis id="B89F2D505C58FFC1FE55FC2B8B6DCE76" box="[510,638,1011,1035]" italics="true" pageId="36" pageNumber="37">A. spadiceus</emphasis>
</taxonomicName>
(triangles),
<taxonomicName id="4DEB8AC15C58FFC1FD52FC2B8A7BCE76" authorityName="Shavrin" authorityYear="2024" box="[761,872,1011,1035]" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="weiperti" status="sp. n.">
<emphasis id="B89F2D505C58FFC1FD52FC2B8A7BCE76" box="[761,872,1011,1035]" italics="true" pageId="36" pageNumber="37">A. weiperti</emphasis>
</taxonomicName>
<emphasis id="B89F2D505C58FFC1FCC5FC2B8AB4CE76" bold="true" box="[878,935,1011,1035]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A3AC902B5C58FFC1FCC5FC2B8AB4CE76" box="[878,935,1011,1035]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(diamonds)
</paragraph>
</caption>
<paragraph id="8A54F1425C58FFC1FF6CFB858D28CC31" blockId="36.[151,1437,1117,2043]" pageId="36" pageNumber="37">Head 1.4 times as broad as long, elevated between anterior margins of eyes and infraorbital portions; frontal portion with strongly elevated supra-antennal prominences, with markedly wide and deep anteriomedian depression, gradually narrowed basad to about level of anterior margins of eyes; interocellar depression wide and moderately deep, slightly narrowed basad, separated from infraorbital portions by moderately deep and long anteocellar foveae convergent lateroanteriad and reaching level of middle length of eyes; temples convex, moderately long, distinctly more than 1/2 as long as longitudinal length of eyes (if see laterally). Eyes moderately large, convex. Ocelli small, distance between ocelli about as long as distance between ocellus and posterior margin of eyes. Frontoclypeal and middle portions without visible punctures; lateral parts with very small, sparse and irregular punctation; interocellar depression and neck without visible punctures. Maxillary palpomere 3 slightly shorter than preceding segment, significantly broadened apicad; last palpomere 1.7 times as long as preceding segment. Antennae very long, almost reaching apical margin of elytra reclined; basal antennomere wide, distinctly more than three times as long as broad, antennomere 2 about twice shorter and significantly narrower than basal antennomere, 3 very long, slightly less than twice and slightly narrower than 2, 4–5 slightly shorter than 3, 6–10 slightly longer than 5, apical antennomere about 1.3 times as long as 10, slightly broader in preapical part, with small rounded apex.</paragraph>
<paragraph id="8A54F1425C58FFC1FF6CF98E8A22CD35" blockId="36.[151,1437,1117,2043]" pageId="36" pageNumber="37">Pronotum 1.2 times as broad as long, widest in apical portion, with widely rounded apical angles strongly protruded anteriad, relatively strongly narrowed anteriad, laterobasal margins elongate, parallel-sided, hind angles obtuse; anterior margin widely rounded, distinctly shorter than slightly concave basal margin; medioanterior portion with wide and deep impression, narrowing toward middle, mediobasal part with distinct wide impression, deeper in front of basal margin; lateral portions impressed, wider and deeper in laterobasal parts, slightly reflexed laterobasally. Punctation fine and dense, finer and sparser in medioapical and distinctly sparser in mediobasal portions. Scutellum large, with widely rounded apex, without visible punctures.</paragraph>
<paragraph id="8A54F1425C58FFC1FF6CF8898801CD86" blockId="36.[151,1437,1117,2043]" pageId="36" pageNumber="37">Elytra slightly longer than broad, slightly less than twice as long as pronotum, distinctly broadened posteriad, reaching apical margin of abdominal tergite V; lateral margins narrowly flattened; hind margins widely rounded. Punctation dense, distinctly larger and deeper than that on pronotum, finer and slightly sparser in mediobasal portion, denser and larger in basal portion and around scutellum, slightly denser and larger along suture. Hind wings fully developed.</paragraph>
<paragraph id="8A54F1425C59FFC0FF6CFF4F8BD1CAA8" blockId="37.[151,1437,151,753]" pageId="37" pageNumber="38">Legs very long and slender; metatibia slightly curved in about middle and slightly broadened apically; metatarsus slightly less than three times as long as metatibia.</paragraph>
<paragraph id="8A54F1425C59FFC0FF6CFF078C52CB60" blockId="37.[151,1437,151,753]" pageId="37" pageNumber="38">Abdomen distinctly narrower than elytra, without visible tomentose spots in middle of abdominal tergites IV and V; apical margin of abdominal tergite VII with narrow palisade fringe. Punctation dense and very fine.</paragraph>
<paragraph id="8A54F1425C59FFC0FF6CFEFF8C37CBD0" blockId="37.[151,1437,151,753]" pageId="37" pageNumber="38">
Male. Posterior margin of abdominal tergite VIII straight. Posterior margin of abdominal sternite VIII slightly concave.Aedeagus with narrow basal part and moderately wide median lobe, from preapical part gradually narrowed toward rounded apex; parameres slightly less than apex of median lobe, with four moderately short apical setae; flagellum relatively long, spirally folded in basal part (
<figureCitation id="12D0EDC75C59FFC0FD57FE4B8A5CCBD0" box="[764,847,403,429]" captionStart="FIGURES 79–88" captionStartId="33.[151,264,1893,1917]" captionTargetBox="[158,1429,188,1863]" captionTargetId="figure-16@33.[151,1436,181,1869]" captionTargetPageId="33" captionText="FIGURES 79–88. Aedeagus (Figs 79–82) and abdominal segments (Figs 83–88) of Asiageodromicus (Figs 79–80: A. weiperti (holotype); Figs 81–82; A. dolpoensis (holotype); Figs 83–88: A. crassipalpis): 79, 81—aedeagus, parameral view, 80, 82— aedeagus, lateral view, 83—male tergite VIII, 84—male sternite VIII, 85—male genital segment, 86—female tergite VIII, 87—female sternite VIII, 88—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023065" httpUri="https://zenodo.org/record/14023065/files/figure.png" pageId="37" pageNumber="38">Fig. 81</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="12D0EDC75C59FFC0FB65FE4B8C33CBD0" box="[1230,1312,403,429]" captionStart="FIGURES 79–88" captionStartId="33.[151,264,1893,1917]" captionTargetBox="[158,1429,188,1863]" captionTargetId="figure-16@33.[151,1436,181,1869]" captionTargetPageId="33" captionText="FIGURES 79–88. Aedeagus (Figs 79–82) and abdominal segments (Figs 83–88) of Asiageodromicus (Figs 79–80: A. weiperti (holotype); Figs 81–82; A. dolpoensis (holotype); Figs 83–88: A. crassipalpis): 79, 81—aedeagus, parameral view, 80, 82— aedeagus, lateral view, 83—male tergite VIII, 84—male sternite VIII, 85—male genital segment, 86—female tergite VIII, 87—female sternite VIII, 88—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023065" httpUri="https://zenodo.org/record/14023065/files/figure.png" pageId="37" pageNumber="38">Fig. 82</figureCitation>
.
</paragraph>
<paragraph id="8A54F1425C59FFC0FF6CFE6F8882CBAC" blockId="37.[151,1437,151,753]" box="[199,401,439,465]" pageId="37" pageNumber="38">Female unknown.</paragraph>
<paragraph id="8A54F1425C59FFC0FF6CFE028836C81C" blockId="37.[151,1437,151,753]" pageId="37" pageNumber="38">
<emphasis id="B89F2D505C59FFC0FF6CFE0288A7CB88" bold="true" box="[199,436,474,501]" pageId="37" pageNumber="38">Comparative notes.</emphasis>
Based on the general shape of pronotum and aedeagus,
<taxonomicName id="4DEB8AC15C59FFC0FBE1FE038DF6CB88" baseAuthorityName="Coiffait" baseAuthorityYear="1982" box="[1098,1253,475,501]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="dolpoensis">
<emphasis id="B89F2D505C59FFC0FBE1FE038DF6CB88" box="[1098,1253,475,501]" italics="true" pageId="37" pageNumber="38">A. dolpoensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4DEB8AC15C59FFC0FA2FFE038832C864" authorityName="Shavrin" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="amplissimus">
<emphasis id="B89F2D505C59FFC0FA2FFE038832C864" italics="true" pageId="37" pageNumber="38">A. amplissimus</emphasis>
</taxonomicName>
,
<taxonomicName id="4DEB8AC15C59FFC0FE9BFE2788C5C864" authorityName="Champion" authorityYear="1920" box="[304,470,511,537]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="crassipalpis">
<emphasis id="B89F2D505C59FFC0FE9BFE2788C5C864" box="[304,470,511,537]" italics="true" pageId="37" pageNumber="38">A. crassipalpis</emphasis>
</taxonomicName>
,
<taxonomicName id="4DEB8AC15C59FFC0FE4EFE278B61C864" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Shavrin" baseAuthorityYear="2019" box="[485,626,511,537]" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="spadiceus">
<emphasis id="B89F2D505C59FFC0FE4EFE278B61C864" box="[485,626,511,537]" italics="true" pageId="37" pageNumber="38">A. spadiceus</emphasis>
</taxonomicName>
and
<taxonomicName id="4DEB8AC15C59FFC0FD01FE278A72C864" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Cheng &amp; Peng" baseAuthorityYear="2020" box="[682,865,511,537]" class="Insecta" family="Staphylinidae" genus="Asiageodromicus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="subquadratus">
<emphasis id="B89F2D505C59FFC0FD01FE278A72C864" box="[682,865,511,537]" italics="true" pageId="37" pageNumber="38">A. subquadratus</emphasis>
</taxonomicName>
, from which it can be distinguished by the smaller and paler body, narrower pronotum, shorter elytra and details of the external morphology of the aedeagus. See also a key above.
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C59FFC0FF6CFDB28814C88C" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8A54F1425C59FFC0FF6CFDB28859C8D4" blockId="37.[151,1437,151,753]" pageId="37" pageNumber="38">
<emphasis id="B89F2D505C59FFC0FF6CFDB28872C8F9" bold="true" box="[199,353,618,644]" pageId="37" pageNumber="38">Distribution.</emphasis>
<taxonomicName id="4DEB8AC15C59FFC0FEC2FDB38BB9C8F8" box="[361,682,619,645]" class="Insecta" family="Staphylinidae" genus="Asiogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="dolpoensis">
<emphasis id="B89F2D505C59FFC0FEC2FDB38BB9C8F8" box="[361,682,619,645]" italics="true" pageId="37" pageNumber="38">Asiogeodromicus dolpoensis</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="55504FE05C59FFC0FC6FFDB38AE7C8F8" box="[964,1012,619,645]" pageId="37" pageNumber="38">type</typeStatus>
locality in
<collectingRegion id="482F3FA05C59FFC0FBDDFDB38C2BC8F8" box="[1142,1336,619,645]" country="Nepal" name="Karnali" pageId="37" pageNumber="38">Karnali Province</collectingRegion>
, western
<collectingCountry id="F2FCB1D25C59FFC0FF3CFD5789C9C8D4" box="[151,218,655,681]" name="Nepal" pageId="37" pageNumber="38">Nepal</collectingCountry>
(
<figureCitation id="12D0EDC75C59FFC0FF42FD57882DC8D4" box="[233,318,655,681]" captionStart="FIGURE 89" captionStartId="36.[151,250,975,999]" captionTargetBox="[157,1429,187,945]" captionTargetId="figure-23@36.[151,1435,181,951]" captionTargetPageId="36" captionText="FIGURE 89. Distribution of Asiageodromicus in the Himalayan Region: A. crassipalpis (circles), A. dolpoensis (star), A. palpalis (Coiffait, 1977) (squares), A. spadiceus (triangles), A. weiperti sp. n. (diamonds)" figureDoi="http://doi.org/10.5281/zenodo.14023067" httpUri="https://zenodo.org/record/14023067/files/figure.png" pageId="37" pageNumber="38">Fig. 89</figureCitation>
).
</paragraph>
<paragraph id="8A54F1425C59FFC0FF6CFD6A8814C88C" blockId="37.[151,1437,151,753]" pageId="37" pageNumber="38">
<emphasis id="B89F2D505C59FFC0FF6CFD6A8858C8B1" bold="true" box="[199,331,690,716]" pageId="37" pageNumber="38">Bionomics.</emphasis>
The specimen was collected at elevations from
<quantity id="4D135CA75C59FFC0FCCDFD6B8D04C8B0" box="[870,1047,690,717]" metricMagnitude="3" metricUnit="m" metricValue="4.375" metricValueMax="4.45" metricValueMin="4.3" pageId="37" pageNumber="38" unit="m" value="4375.0" valueMax="4450.0" valueMin="4300.0">4300 to 4450 m</quantity>
a.s.l. Detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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