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<document id="3F8ACD5B1D72C6AD4FD6D5CFC349C1B0" ID-DOI="10.11646/zootaxa.5531.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14023033" ID-ZooBank="27F9603C-D2AB-4133-AFB8-5A674ACCD110" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730453991549" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="024240545C4CFFD4FF3CFB2A89E3CC74" docLanguage="en" docName="zootaxa.5531.1.1.pdf" docOrigin="Zootaxa 5531 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5531.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Himalodromeus grandis Shavrin, 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="50" masterDocId="FE7B382C5C7CFFE5FFABFFD88913CA7D" masterDocTitle="Review of Hygrogeus Mulsant &amp; Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="49" updateTime="1730454835898" updateUser="ExternalLinkService">
<mods:mods id="3C00CCD2B9BCBCD2CA1D1E276DA3A071" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3784784D5B1CA1A52CF9FDE9F5564693">
<mods:title id="42184D01E1EFEA646385A38A623B6C5B">Review of Hygrogeus Mulsant &amp; Rey, 1880 and Trichodromeus Luze, 1903, with descriptions of three new genera from the eastern Palaearctic Region (Coleoptera: Staphylinidae: Anthophagini: Omaliinae)</mods:title>
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<mods:namePart id="80296BEFDCCFEFA0CCA62C56106B978A">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="3AC667332DC8451911DADF64411E1330">Zootaxa</mods:title>
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<mods:date id="C5FA7BAF1FC38B766B4BE5347813B857">2024</mods:date>
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<mods:number id="58615BF91FF510BEEFF6C77AFAD4A790">2024-10-30</mods:number>
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<mods:number id="F73B8F41C9E1F007F83F52557B7AE61D">5531</mods:number>
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<mods:classification id="6E73921F1141C5206F4ADF9A95E57ED4">journal article</mods:classification>
<mods:identifier id="FB14C5ADBAD193530905F2869979ECAB" type="DOI">10.11646/zootaxa.5531.1.1</mods:identifier>
<mods:identifier id="2145DC7E9FD2F0907C0203EFE8E08A4E" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="552E560448E580FBF9258537C01F3EE6" type="Zenodo-Dep">14023033</mods:identifier>
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<treatment id="024240545C4CFFD4FF3CFB2A89E3CC74" ID-DOI="http://doi.org/10.5281/zenodo.14023145" ID-Zenodo-Dep="14023145" LSID="urn:lsid:plazi:treatment:024240545C4CFFD4FF3CFB2A89E3CC74" httpUri="http://treatment.plazi.org/id/024240545C4CFFD4FF3CFB2A89E3CC74" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<subSubSection id="C2F1A2C95C4CFFD5FF3CFB2A8B14CF70" box="[151,519,1266,1293]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="8A54F1425C4CFFD5FF3CFB2A8B14CF70" blockId="48.[151,519,1266,1329]" box="[151,519,1266,1293]" pageId="48" pageNumber="49">
<heading id="D11C462E5C4CFFD5FF3CFB2A8B14CF70" bold="true" box="[151,519,1266,1293]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="4">
<emphasis id="B89F2D505C4CFFD5FF3CFB2A8B14CF70" bold="true" box="[151,519,1266,1293]" pageId="48" pageNumber="49">
<taxonomicName id="4DEB8AC15C4CFFD5FF3CFB2A88AECF70" authorityName="Shavrin" authorityYear="2024" box="[151,445,1266,1293]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B89F2D505C4CFFD5FF3CFB2A88AECF70" bold="true" box="[151,445,1266,1293]" italics="true" pageId="48" pageNumber="49">Himalodromeus grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3AC902B5C4CFFD5FE6EFB2B8B14CF70" box="[453,519,1267,1293]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C4CFFD5FF3CFACF8B16CF4C" box="[151,517,1303,1329]" pageId="48" pageNumber="49" type="description">
<paragraph id="8A54F1425C4CFFD5FF3CFACF8B16CF4C" blockId="48.[151,519,1266,1329]" box="[151,517,1303,1329]" pageId="48" pageNumber="49">
(
<figureCitation id="12D0EDC75C4CFFD5FF34FACF8833CF4C" box="[159,288,1303,1329]" captionStart="FIGURES 90–97" captionStartId="40.[151,264,1707,1731]" captionTargetBox="[158,1429,188,1671]" captionTargetId="figure-23@40.[151,1436,181,1684]" captionTargetPageId="40" captionText="FIGURES 90–97. Habitus (Figs 90–94) and apical part of pronotum (Figs 95–97) of Himalodromeus: 90—H. cyanipennis (Uttarakhand), 91, 95—H. cyanipennis (paratype), 92—Hygrogeus cyanipennis var. atrocaeruleus (lectotype), 93, 96—H. grandis (holotype), 94, 97—H. reductus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023069" httpUri="https://zenodo.org/record/14023069/files/figure.png" pageId="48" pageNumber="49">Figs 93, 96</figureCitation>
,
<figureCitation id="12D0EDC75C4CFFD5FE87FACF889DCF4C" box="[300,398,1303,1329]" captionStart="FIGURES 110–115" captionStartId="43.[151,264,1849,1873]" captionTargetBox="[152,1429,188,1819]" captionTargetId="figure-16@43.[151,1436,181,1826]" captionTargetPageId="43" captionText="FIGURES 110–115. Aedeagus of Himalodromeus (Figs 110–111: H. cyanipennis (paralectotype), Figs 112–113: H. grandis (holotype), Figs 114–115: H. reductus (paratype): 110, 112, 114—parameral view, 111, 113, 115—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023075" httpUri="https://zenodo.org/record/14023075/files/figure.png" pageId="48" pageNumber="49">112–113</figureCitation>
,
<figureCitation id="12D0EDC75C4CFFD5FE31FACF88D5CF4C" box="[410,454,1303,1329]" captionStart="FIGURES 116–123" captionStartId="44.[151,264,1377,1401]" captionTargetBox="[158,1430,188,1346]" captionTargetId="figure-23@44.[151,1436,181,1353]" captionTargetPageId="44" captionText="FIGURES 116–123. Abdominal segments (ventral view) of Himalodromeus (Figs 116–121: H. cyanipennis; Fig. 122: H. grandis; Fig. 123: H. reductus): 116—male tergite VIII, 117, 122–123—male sternite VIII, 118—male genital segment, 119— female tergite VIII, 120—female sternite VIII, 121—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023077" httpUri="https://zenodo.org/record/14023077/files/figure.png" pageId="48" pageNumber="49">122</figureCitation>
,
<figureCitation id="12D0EDC75C4CFFD5FE79FACF88EECF4C" box="[466,509,1303,1329]" captionStart="FIGURE 124" captionStartId="45.[151,250,1548,1572]" captionTargetBox="[157,1429,567,1519]" captionTargetId="figure-187@45.[151,1435,561,1525]" captionTargetPageId="45" captionText="FIGURE 124. Distribution of Himalodromeus: H. cyanipennis (circles), H. grandis (square), H. reductus (triangles)." figureDoi="http://doi.org/10.5281/zenodo.14023079" httpUri="https://zenodo.org/record/14023079/files/figure.png" pageId="48" pageNumber="49">124</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C4CFFD5FF3CFA87887ECC50" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8A54F1425C4CFFD5FF3CFA878C8ACFBC" blockId="48.[151,1437,1374,2013]" pageId="48" pageNumber="49">
<emphasis id="B89F2D505C4CFFD5FF3CFA8788D2CF05" bold="true" box="[151,449,1374,1401]" pageId="48" pageNumber="49">Type material examined.</emphasis>
<typeStatus id="55504FE05C4CFFD5FE62FA878B22CF04" box="[457,561,1375,1401]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
<specimenCount id="9CED3ACB5C4CFFD5FD92FA878B5DCF04" box="[569,590,1375,1401]" count="1" pageId="48" pageNumber="49" type="male">♂</specimenCount>
(
<figureCitation id="12D0EDC75C4CFFD5FDF4FA878BA0CF04" box="[607,691,1375,1401]" captionStart="FIGURES 90–97" captionStartId="40.[151,264,1707,1731]" captionTargetBox="[158,1429,188,1671]" captionTargetId="figure-23@40.[151,1436,181,1684]" captionTargetPageId="40" captionText="FIGURES 90–97. Habitus (Figs 90–94) and apical part of pronotum (Figs 95–97) of Himalodromeus: 90—H. cyanipennis (Uttarakhand), 91, 95—H. cyanipennis (paratype), 92—Hygrogeus cyanipennis var. atrocaeruleus (lectotype), 93, 96—H. grandis (holotype), 94, 97—H. reductus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023069" httpUri="https://zenodo.org/record/14023069/files/figure.png" pageId="48" pageNumber="49">Fig. 93</figureCitation>
; dissected; left antennomeres 10–11 and 5–11 missing): ‘
<collectingCountry id="F2FCB1D25C4CFFD5FA93FA878C8FCF04" box="[1336,1436,1375,1401]" name="Pakistan" pageId="48" pageNumber="49">Pakistan</collectingCountry>
(
<collectionCode id="ECFA69875C4CFFD5FF34FA5B89E3CFE0" box="[159,240,1411,1437]" pageId="48" pageNumber="49">NWFP</collectionCode>
) | Balakot |
<date id="FE55D7825C4CFFD5FEC4FA5A88E0CFE0" box="[367,499,1410,1437]" pageId="48" pageNumber="49" value="1953-06-23">23-VI-1953</date>
| F.Schmid’ &lt;printed&gt;, ‘
<taxonomicName id="4DEB8AC15C4CFFD5FD52FA5B8A6ACFE0" authorityName="Mulsant and Rey" authorityYear="1880" box="[761,889,1411,1437]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Hygrogeus</taxonomicName>
| cyanipennis | Cam. | det. V. Puthz 19[printed]81’ &lt;handwritten&gt;, ‘
<typeStatus id="55504FE05C4CFFD5FEE4FA7F88FBCFBC" box="[335,488,1447,1473]" pageId="48" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4DEB8AC15C4CFFD5FE50FA7F8BBDCFBC" authorityName="Shavrin" authorityYear="2024" box="[507,686,1447,1473]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B89F2D505C4CFFD5FE50FA7F8BBDCFBC" box="[507,686,1447,1473]" italics="true" pageId="48" pageNumber="49">Himalodromeus</emphasis>
</taxonomicName>
|
<taxonomicName id="4DEB8AC15C4CFFD5FD69FA7F8A0BCFBC" authorityName="Shavrin" authorityYear="2024" box="[706,792,1447,1473]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B89F2D505C4CFFD5FD69FA7F8A0BCFBC" box="[706,792,1447,1473]" italics="true" pageId="48" pageNumber="49">grandis</emphasis>
</taxonomicName>
<emphasis id="B89F2D505C4CFFD5FCB4FA7F8A4DCFBC" bold="true" box="[799,862,1447,1473]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A3AC902B5C4CFFD5FCB4FA7F8A4DCFBC" box="[799,862,1447,1473]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="ECFA69875C4CFFD5FA9AFA7F8C9ECFBC" box="[1329,1421,1447,1473]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="48" pageNumber="49" type="Museum">MHNG</collectionCode>
).
</paragraph>
<paragraph id="8A54F1425C4CFFD5FF6CFA13887ECC50" blockId="48.[151,1437,1374,2013]" pageId="48" pageNumber="49">
<typeStatus id="55504FE05C4CFFD5FF6CFA13883ACF98" box="[199,297,1483,1509]" pageId="48" pageNumber="49" type="paratype">Paratype</typeStatus>
<specimenCount id="9CED3ACB5C4CFFD5FE9AFA138855CF98" box="[305,326,1483,1509]" count="1" pageId="48" pageNumber="49" type="female">♀</specimenCount>
(left antenna and right antennomeres 2–11, and right fore leg missing): same data as the holotype, with additional red printed label: ‘
<typeStatus id="55504FE05C4CFFD5FDB7FA378BBFCC74" box="[540,684,1519,1545]" pageId="48" pageNumber="49" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4DEB8AC15C4CFFD5FD6EFA378A6BCC74" authorityName="Shavrin" authorityYear="2024" box="[709,888,1519,1545]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B89F2D505C4CFFD5FD6EFA378A6BCC74" box="[709,888,1519,1545]" italics="true" pageId="48" pageNumber="49">Himalodromeus</emphasis>
</taxonomicName>
|
<taxonomicName id="4DEB8AC15C4CFFD5FC3BFA378AF5CC74" authorityName="Shavrin" authorityYear="2024" box="[912,998,1519,1545]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B89F2D505C4CFFD5FC3BFA378AF5CC74" box="[912,998,1519,1545]" italics="true" pageId="48" pageNumber="49">grandis</emphasis>
</taxonomicName>
<emphasis id="B89F2D505C4CFFD5FC44FA378D22CC74" bold="true" box="[1007,1073,1519,1545]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A3AC902B5C4CFFD5FC44FA378D22CC74" box="[1007,1073,1519,1545]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="ECFA69875C4CFFD5FEAEF9CB8872CC50" box="[261,353,1555,1581]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="48" pageNumber="49" type="Museum">MHNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C4CFFD4FF6CF9EE8A01CF28" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
<paragraph id="8A54F1425C4CFFD5FF6CF9EE8D6FCCE4" blockId="48.[151,1437,1374,2013]" pageId="48" pageNumber="49">
<emphasis id="B89F2D505C4CFFD5FF6CF9EE884ACC2D" bold="true" box="[199,345,1590,1616]" pageId="48" pageNumber="49">Description.</emphasis>
Measurements (n=2): HW: 1.60–1.64; HL: 1.15; OL: 0.32–0.38; TL: 0.30; PL: 1.70–2.25; PWmax: 2.30–2.60; PWmin: 1.71–1.85; ESL: 2.65–2.75; EW: 3.10–3.45; MTbL (
<typeStatus id="55504FE05C4CFFD5FC71F9838D50CC08" box="[986,1091,1627,1653]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
): 3.25; MTrL (
<typeStatus id="55504FE05C4CFFD5FB46F9838C45CC08" box="[1261,1366,1627,1653]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
): 1.28 (MTrL 1–4: 0.68; MTrL 5: 0.60); AW: 2.90–3.35; AedL: 1.60; BL: 8.25–9.45 (
<typeStatus id="55504FE05C4CFFD5FBACF9A78D63CCE4" box="[1031,1136,1663,1689]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8A54F1425C4CFFD5FF6CF97B8ADCCD54" blockId="48.[151,1437,1374,2013]" pageId="48" pageNumber="49">
Habitus as in
<figureCitation id="12D0EDC75C4CFFD5FEF4F97B88A2CCC0" box="[351,433,1699,1725]" captionStart="FIGURES 90–97" captionStartId="40.[151,264,1707,1731]" captionTargetBox="[158,1429,188,1671]" captionTargetId="figure-23@40.[151,1436,181,1684]" captionTargetPageId="40" captionText="FIGURES 90–97. Habitus (Figs 90–94) and apical part of pronotum (Figs 95–97) of Himalodromeus: 90—H. cyanipennis (Uttarakhand), 91, 95—H. cyanipennis (paratype), 92—Hygrogeus cyanipennis var. atrocaeruleus (lectotype), 93, 96—H. grandis (holotype), 94, 97—H. reductus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023069" httpUri="https://zenodo.org/record/14023069/files/figure.png" pageId="48" pageNumber="49">Fig. 94</figureCitation>
. Body dark-brown, with bluish elytra; legs brown; antennae, mouthparts and tarsi yellow-brown (apical maxillary palpomeres distinctly paler). Head with dense isodiametric microsculpture, coarser between ocelli and on neck; pronotum with dense isodiametric microsculpture, finer in mediobasal and medioapical portions; scutellum with dense and moderately coarse transverse microreticulation.
</paragraph>
<paragraph id="8A54F1425C4CFFD4FF6CF8EB8821CB60" blockId="48.[151,1437,1374,2013]" lastBlockId="49.[151,1437,151,1545]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">Head 1.3–1.4 times as broad as long, with slightly elevated infraorbital portions; anterio-median depression connected with slightly depressed subrectangular interocellar depression; anteocellar foveae reaching about level of middle of eyes; basal part of interocellar depression in front of ocelli with widely rounded elevation; temples slightly shorter than longitudinal length of eyes. Ocelli moderately large; distance between ocelli about as long as distance between ocellus and posterior margin of eyes. Punctation dense and large, sparser and finer in middle; mediobasal part between ocelli and neck with fine and sparse pucntation. Apical maxillary palpomere about 1.4 times as long as penultimate segment, distinctly broadened in preapical portion, with rounded apex. Antennomere 2 shorter and narrower than basal antennomere, 3 distinctly longer than 2, 4–6 slightly longer than 3 and 7–9 slightly longer than 6.</paragraph>
<paragraph id="8A54F1425C4DFFD4FF6CFEFF8BC6C81C" blockId="49.[151,1437,151,1545]" pageId="49" pageNumber="50">
Pronotum 1.1–1.3 times as broad as long, broader slightly in front of middle, from widest part gradually narrowed anteriad and slightly more sharply narrowed posteriad; latero-apical margin distinctly concave; frontal part strongly protruded anteriad, with widely concave apical margin (
<figureCitation id="12D0EDC75C4DFFD4FC3CFEB78AF8CBF4" box="[919,1003,367,393]" captionStart="FIGURES 90–97" captionStartId="40.[151,264,1707,1731]" captionTargetBox="[158,1429,188,1671]" captionTargetId="figure-23@40.[151,1436,181,1684]" captionTargetPageId="40" captionText="FIGURES 90–97. Habitus (Figs 90–94) and apical part of pronotum (Figs 95–97) of Himalodromeus: 90—H. cyanipennis (Uttarakhand), 91, 95—H. cyanipennis (paratype), 92—Hygrogeus cyanipennis var. atrocaeruleus (lectotype), 93, 96—H. grandis (holotype), 94, 97—H. reductus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023069" httpUri="https://zenodo.org/record/14023069/files/figure.png" pageId="49" pageNumber="50">Fig. 96</figureCitation>
); laterobasal margins in front of obtuse hind angles moderately short, subparallel; posterior margin slightly concave; middle portion of disc with shallow and narrow longitudinal depression; mediobasal portion in front of basal margin indistinctly and widely impressed; latero-apical and lateral margins narrowly bordered, reflexed latero-apically (
<typeStatus id="55504FE05C4DFFD4FBAEFE038D7ECB88" box="[1029,1133,475,501]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
), with smooth crenulation, stronger in
<typeStatus id="55504FE05C4DFFD4FEB3FE27886DC864" box="[280,382,511,537]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
. Punctation about as that on head, but significantly denser, with interstices between punctures in middle about as long as diameter of nearest punctures, sparser in medioapical and mediobasal portions. Scutellum with distinct dense and moderately fine punctation.
</paragraph>
<paragraph id="8A54F1425C4DFFD4FF6CFDB38AA1C88C" blockId="49.[151,1437,151,1545]" pageId="49" pageNumber="50">
Elytra 1.1–1.2 times as broad as long, 1.2–1.5 times as long as pronotum; disc of each elytron with distinct four narrow and elongate elevations in middle, from about mediobasal portion subdiagonally stretching posteriad toward medioapical part; lateral potion widely impressed and slightly reflexed. Punctation about as that on pronotum, but sparser, finer and denser around scutellum and along suture (
<typeStatus id="55504FE05C4DFFD4FC94FD0F8AB5C88C" box="[831,934,727,753]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8A54F1425C4DFFD4FF6CFD238B91C968" blockId="49.[151,1437,151,1545]" box="[199,642,763,789]" pageId="49" pageNumber="50">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="8A54F1425C4DFFD4FF6CFCC78B9DCE24" blockId="49.[151,1437,151,1545]" pageId="49" pageNumber="50">
Male. Pronotum distinctly broader and more convex. Profemora and protarsomeres 1–4 wide. Posterior margin of abdominal tergite VIII straight. Posterior margin of abdominal sternite VIII with significant deep emargination, with dense and very long setation on each latero-apical portion (
<figureCitation id="12D0EDC75C4DFFD4FCF2FCBF8AA9C9FC" box="[857,954,871,897]" captionStart="FIGURES 116–123" captionStartId="44.[151,264,1377,1401]" captionTargetBox="[158,1430,188,1346]" captionTargetId="figure-23@44.[151,1436,181,1353]" captionTargetPageId="44" captionText="FIGURES 116–123. Abdominal segments (ventral view) of Himalodromeus (Figs 116–121: H. cyanipennis; Fig. 122: H. grandis; Fig. 123: H. reductus): 116—male tergite VIII, 117, 122–123—male sternite VIII, 118—male genital segment, 119— female tergite VIII, 120—female sternite VIII, 121—female genital segment. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023077" httpUri="https://zenodo.org/record/14023077/files/figure.png" pageId="49" pageNumber="50">Fig. 122</figureCitation>
). Aedeagus with broad basal part, gradually narrowed toward moderately broad median lobe, from preapical portion strongly narrowed toward elongate and acute apex; basal portions of median lobe with long and narrow lateral sclerotized lobes, curved apically about at level of apical third of median lobe; parameres shortened, not reaching middle of median lobe, with four moderately short apical setae; internal sac without spines, with narrow elongated structures in preapical and middle portions, with long and narrow flagellum, spirally folded in basal portion and significantly broadened in basal part (
<figureCitation id="12D0EDC75C4DFFD4FA84FBC38C83CE48" box="[1327,1424,1051,1077]" captionStart="FIGURES 110–115" captionStartId="43.[151,264,1849,1873]" captionTargetBox="[152,1429,188,1819]" captionTargetId="figure-16@43.[151,1436,181,1826]" captionTargetPageId="43" captionText="FIGURES 110–115. Aedeagus of Himalodromeus (Figs 110–111: H. cyanipennis (paralectotype), Figs 112–113: H. grandis (holotype), Figs 114–115: H. reductus (paratype): 110, 112, 114—parameral view, 111, 113, 115—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023075" httpUri="https://zenodo.org/record/14023075/files/figure.png" pageId="49" pageNumber="50">Fig. 112</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="12D0EDC75C4DFFD4FD80FBE78B99CE24" box="[555,650,1087,1113]" captionStart="FIGURES 110–115" captionStartId="43.[151,264,1849,1873]" captionTargetBox="[152,1429,188,1819]" captionTargetId="figure-16@43.[151,1436,181,1826]" captionTargetPageId="43" captionText="FIGURES 110–115. Aedeagus of Himalodromeus (Figs 110–111: H. cyanipennis (paralectotype), Figs 112–113: H. grandis (holotype), Figs 114–115: H. reductus (paratype): 110, 112, 114—parameral view, 111, 113, 115—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023075" httpUri="https://zenodo.org/record/14023075/files/figure.png" pageId="49" pageNumber="50">Fig. 113</figureCitation>
.
</paragraph>
<paragraph id="8A54F1425C4DFFD4FF6CFBBB8A3BCEDC" blockId="49.[151,1437,151,1545]" pageId="49" pageNumber="50">Female. Pronotum narrower, less convex. Profemuri and protarsomeres 1–4 moderately narrow. Posterior margin of abdominal tergite VIII and sternite VIII straight.</paragraph>
<paragraph id="8A54F1425C4DFFD4FF6CFB728A01CF28" blockId="49.[151,1437,151,1545]" pageId="49" pageNumber="50">
<emphasis id="B89F2D505C4DFFD4FF6CFB7288A7CEB8" bold="true" box="[199,436,1194,1221]" pageId="49" pageNumber="50">Comparative notes.</emphasis>
Based on the coloration of the body and the bluish elytra, general shape of pronotum and aedeagus,
<taxonomicName id="4DEB8AC15C4DFFD4FE94FB1788AACE94" authorityName="Shavrin" authorityYear="2024" box="[319,441,1231,1257]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="grandis" status="sp. n.">
<emphasis id="B89F2D505C4DFFD4FE94FB178840CE95" box="[319,339,1231,1256]" italics="true" pageId="49" pageNumber="50">H</emphasis>
.
<emphasis id="B89F2D505C4DFFD4FEC8FB1788AACE94" box="[355,441,1231,1257]" italics="true" pageId="49" pageNumber="50">grandis</emphasis>
</taxonomicName>
<emphasis id="B89F2D505C4DFFD4FE69FB178B10CE94" bold="true" box="[450,515,1231,1257]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A3AC902B5C4DFFD4FE69FB178B10CE94" box="[450,515,1231,1257]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is more similar to
<taxonomicName id="4DEB8AC15C4DFFD4FD74FB178A9ACE95" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[735,905,1231,1256]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cyanipennis">
<emphasis id="B89F2D505C4DFFD4FD74FB178A9ACE95" box="[735,905,1231,1256]" italics="true" pageId="49" pageNumber="50">H. cyanipennis</emphasis>
</taxonomicName>
, from which it differs by the larger pronotum, broader slightly in front of middle, with frontal part more protruded anteriad, denser punctation of the pronotum, the presence of distinct longitudinal elevations on each elytron, the broader apical part of the aedeagus, and other details of the internal and external morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C4DFFD4FF6CFA868A18CFBC" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8A54F1425C4DFFD4FF6CFA86881DCFE0" blockId="49.[151,1437,151,1545]" pageId="49" pageNumber="50">
<emphasis id="B89F2D505C4DFFD4FF6CFA868872CF05" bold="true" box="[199,353,1374,1400]" pageId="49" pageNumber="50">Distribution.</emphasis>
<taxonomicName id="4DEB8AC15C4DFFD4FEC2FA878B6FCF04" authorityName="Shavrin" authorityYear="2024" box="[361,636,1375,1401]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B89F2D505C4DFFD4FEC2FA878B6FCF04" box="[361,636,1375,1401]" italics="true" pageId="49" pageNumber="50">Himalodromeus grandis</emphasis>
</taxonomicName>
<emphasis id="B89F2D505C4DFFD4FD2FFA878BD5CF04" bold="true" box="[644,710,1375,1401]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A3AC902B5C4DFFD4FD2FFA878BD5CF04" box="[644,710,1375,1401]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="55504FE05C4DFFD4FC4FFA878D07CF04" box="[996,1044,1375,1401]" pageId="49" pageNumber="50">type</typeStatus>
locality in north-western
<collectingCountry id="F2FCB1D25C4DFFD4FA96FA878C8FCF04" box="[1341,1436,1375,1401]" name="Pakistan" pageId="49" pageNumber="50">Pakistan</collectingCountry>
(
<figureCitation id="12D0EDC75C4DFFD4FF34FA5B8811CFE0" box="[159,258,1411,1437]" captionStart="FIGURE 124" captionStartId="45.[151,250,1548,1572]" captionTargetBox="[157,1429,567,1519]" captionTargetId="figure-187@45.[151,1435,561,1525]" captionTargetPageId="45" captionText="FIGURE 124. Distribution of Himalodromeus: H. cyanipennis (circles), H. grandis (square), H. reductus (triangles)." figureDoi="http://doi.org/10.5281/zenodo.14023079" httpUri="https://zenodo.org/record/14023079/files/figure.png" pageId="49" pageNumber="50">Fig. 124</figureCitation>
).
</paragraph>
<paragraph id="8A54F1425C4DFFD4FF6CFA7E8A18CFBC" blockId="49.[151,1437,151,1545]" box="[199,779,1446,1473]" pageId="49" pageNumber="50">
<emphasis id="B89F2D505C4DFFD4FF6CFA7E8858CFBD" bold="true" box="[199,331,1446,1472]" pageId="49" pageNumber="50">Bionomics.</emphasis>
Detailed bionomical data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C2F1A2C95C4DFFD4FF6CFA1389E3CC74" pageId="49" pageNumber="50" type="etymology">
<paragraph id="8A54F1425C4DFFD4FF6CFA1389E3CC74" blockId="49.[151,1437,151,1545]" pageId="49" pageNumber="50">
<emphasis id="B89F2D505C4DFFD4FF6CFA13885DCF98" bold="true" box="[199,334,1483,1509]" pageId="49" pageNumber="50">Etymology.</emphasis>
The specific epithet is the Latin adjective
<taxonomicName id="4DEB8AC15C4DFFD4FCE9FA138A8BCF98" authorityName="Shavrin" authorityYear="2024" box="[834,920,1483,1509]" class="Insecta" family="Staphylinidae" genus="Himalodromeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B89F2D505C4DFFD4FCE9FA138A8BCF98" box="[834,920,1483,1509]" italics="true" pageId="49" pageNumber="50">grandis</emphasis>
</taxonomicName>
,
<emphasis id="B89F2D505C4DFFD4FC02FA138AADCF99" box="[937,958,1483,1508]" italics="true" pageId="49" pageNumber="50">-e</emphasis>
(large). It refers to the large body of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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