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<mods:title id="1CDFCFF5359D52E6B389AA4069257264">A new genus of speleophriid copepod (Copepoda: Misophrioida) from a cenote in the Yucatan, Mexico with a phylogenetic analysis at the species level</mods:title>
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<mods:namePart id="85E61DAD3E893BD37FB788843A0D9B6A">Boxshall, Geoff A.</mods:namePart>
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<mods:namePart id="71510A8B51093417B2462B9BA8B80FC5">Jaume, Damià</mods:namePart>
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<mods:namePart id="F9B2596A27C26B95BDA2A7940BEB2C1F">Iliffe, Thomas M.</mods:namePart>
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<treatment id="8A3384428735FFD847FAFD1D1B480438" ID-DOI="http://doi.org/10.5281/zenodo.5667157" ID-GBIF-Taxon="119508048" ID-Zenodo-Dep="5667157" LSID="urn:lsid:zoobank.org:act:7B91FBD1-2ECB-4C0F-83CF-D1B59242282B" httpUri="http://treatment.plazi.org/id/8A3384428735FFD847FAFD1D1B480438" lastPageId="4" lastPageNumber="325" pageId="3" pageNumber="324">
<subSubSection id="4A8066DF8735FFDF47FAFD1D1BE40494" box="[151,423,665,692]" pageId="3" pageNumber="324" type="nomenclature">
<paragraph id="022535548735FFDF47FAFD1D1BE40494" blockId="3.[151,423,665,692]" box="[151,423,665,692]" pageId="3" pageNumber="324">
<heading id="596D82388735FFDF47FAFD1D1BE40494" bold="true" box="[151,423,665,692]" fontSize="11" level="1" pageId="3" pageNumber="324" reason="1">
<emphasis id="30EEE9468735FFDF47FAFD1D1BE40494" bold="true" box="[151,423,665,692]" pageId="3" pageNumber="324">
<taxonomicName id="C59A4ED78735FFDF47FAFD1D1B710493" box="[151,306,665,691]" class="Maxillopoda" family="Speleophriidae" genus="Mexicophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="30EEE9468735FFDF47FAFD1D1B710493" bold="true" box="[151,306,665,691]" italics="true" pageId="3" pageNumber="324">Mexicophria</emphasis>
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<taxonomicNameLabel id="2BDD543D8735FFDF4654FD1E1BE40494" box="[313,423,666,692]" pageId="3" pageNumber="324" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="4A8066DF8735FFDF47FAFD641A960319" pageId="3" pageNumber="324" type="diagnosis">
<paragraph id="022535548735FFDF47FAFD641A960319" blockId="3.[151,1437,736,2021]" pageId="3" pageNumber="324">
<emphasis id="30EEE9468735FFDF47FAFD641B4904D9" bold="true" box="[151,266,736,761]" pageId="3" pageNumber="324">Diagnosis</emphasis>
. Cephalosome not produced posteriorly into carapace-like extension. First pedigerous somite free. Female urosome 5-segmented, with genital and first abdominal somites fused to form genital double-somite; single copulatory pore opening mid-ventrally. Free abdominal somites separated by extensive arthrodial membrane allowing telescoping of adjacent somites. Caudal rami armed with 6 setae. Antennules 27-segmented in female, with compound apical segment derived from ancestral segments XXVIIXXVIII. Proximal segment swollen dorsolaterally, forming bulbous lobe: segments IIX and XIXII not completely separated along posterior margin; elongate aesthetascs on segments XI and XVI. Antennule 23-segmented in male; geniculate between segments 19 (XIXXX) and 20 (XXIXXIII); segment 21 (XXIV) free, segment 22 (XXVXXVI) and apical segment (XXVIIXXVIII) double. Antenna with 1 seta on basis; exopod 7-segmented bearing 1, 1, 1, 1, 1, 1, 4 setae; endopod indistinctly 3-segmented bearing 2, 5, 7 setae. Mandible with 2 setae on basis, endopod with 4, 7 setae; exopod with 1, 1, 1, 3 setae. Maxillulary basal exite absent. Allobasis of maxilla with armature formula 4, 2. Maxilliped syncoxa lacking praecoxal seta; endopod 6-segmented, setal formula 2, 2, 2, 2, 2+1, 4. Swimming legs 14 biramous with 3-segmented rami except for 2-segmented endopod of leg 1; spine and seta formula as below for
<typeStatus id="DD218BF68735FFDF47FAFB311A8402ED" box="[151,199,1205,1229]" pageId="3" pageNumber="324">type</typeStatus>
species. Fifth legs uniramous in both sexes, located immediately adjacent to ventral midline, lacking intercoxal sclerite; segmentation sexually dimorphic, 2-segmented, with unsegmented protopodal segment and single exopodal segment in female, 3-segmented, with separate coxa and basis plus single exopodal segment in male.
</paragraph>
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<paragraph id="022535548735FFDF47AAFAC01E02037D" blockId="3.[151,1437,736,2021]" box="[199,1089,1348,1373]" pageId="3" pageNumber="324">
<emphasis id="30EEE9468735FFDF47AAFAC01B21037D" bold="true" box="[199,354,1348,1373]" pageId="3" pageNumber="324">
<typeStatus id="DD218BF68735FFDF47AAFAC01B41037D" box="[199,258,1348,1373]" pageId="3" pageNumber="324">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="C59A4ED78735FFDF4604FAC01830037D" box="[361,627,1348,1373]" class="Maxillopoda" family="Speleophriidae" genus="Mexicophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="cenoticola">
<emphasis id="30EEE9468735FFDF4604FAC01830037D" box="[361,627,1348,1373]" italics="true" pageId="3" pageNumber="324">Mexicophria cenoticola</emphasis>
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<emphasis id="30EEE9468735FFDF4516FAC01973037D" bold="true" box="[635,816,1348,1373]" pageId="3" pageNumber="324">gen. et sp. nov.,</emphasis>
by original designation.
</paragraph>
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<subSubSection id="4A8066DF8735FFDF47AAFAEC19250385" pageId="3" pageNumber="324" type="etymology">
<paragraph id="022535548735FFDF47AAFAEC19250385" blockId="3.[151,1437,736,2021]" pageId="3" pageNumber="324">
<emphasis id="30EEE9468735FFDF47AAFAEC1B0D03A1" bold="true" box="[199,334,1384,1409]" pageId="3" pageNumber="324">Etymology.</emphasis>
The generic name is based on the name of the country where it is found,
<collectingCountry id="7A8D75C48735FFDF43FDFAEC1EAB03A1" box="[1168,1256,1384,1409]" name="Mexico" pageId="3" pageNumber="324">Mexico</collectingCountry>
, combined with the ending
<emphasis id="30EEE9468735FFDF4645FA081B300385" box="[296,371,1420,1445]" italics="true" pageId="3" pageNumber="324">ophria</emphasis>
, which is commonly used within the family.
</paragraph>
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<paragraph id="022535548735FFDF47AAFA341FDE0159" blockId="3.[151,1437,736,2021]" pageId="3" pageNumber="324">
<emphasis id="30EEE9468735FFDF47AAFA341B7803E9" bold="true" box="[199,315,1456,1481]" pageId="3" pageNumber="324">Remarks.</emphasis>
The fifth legs of the new genus are located immediately adjacent to the ventral midline of the somite and lack an intercoxal sclerite, as in several other speleophriid genera, but they are more strongly reduced than in most other genera.
<emphasis id="30EEE9468735FFDF4617FA7C1E300030" box="[378,1139,1528,1553]" italics="true" pageId="3" pageNumber="324">
<taxonomicName id="C59A4ED78735FFDF4617FA7C18730031" box="[378,560,1528,1553]" class="Maxillopoda" family="Speleophriidae" genus="Speleophriopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">Speleophriopsis</taxonomicName>
,
<taxonomicName id="C59A4ED78735FFDF4552FA7C188A0031" box="[575,713,1528,1553]" class="Maxillopoda" family="Speleophriidae" genus="Speleophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">Speleophria</taxonomicName>
,
<taxonomicName id="C59A4ED78735FFDF45B4FA7C19170031" box="[729,852,1528,1553]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">Boxshallia</taxonomicName>
,
<taxonomicName id="C59A4ED78735FFDF440EFA7C1E560031" box="[867,1045,1528,1553]" class="Maxillopoda" family="Speleophriidae" genus="Archimisophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">Archimisophria</taxonomicName>
,
<taxonomicName id="C59A4ED78735FFDF4349FA7D1E300030" box="[1060,1139,1529,1552]" class="Maxillopoda" family="Speleophriidae" genus="Huysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">Huysia</taxonomicName>
</emphasis>
and
<taxonomicName id="C59A4ED78735FFDF43DFFA7C1F360031" box="[1202,1397,1528,1553]" class="Maxillopoda" family="Speleophriidae" genus="Protospeleophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468735FFDF43DFFA7C1F360031" box="[1202,1397,1528,1553]" italics="true" pageId="3" pageNumber="324">Protospeleophria</emphasis>
</taxonomicName>
all have a fifth leg in the female consisting of at least three segments. The enigmatic genus
<taxonomicName id="C59A4ED78735FFDF43CBF9981F700015" box="[1190,1331,1564,1589]" class="Maxillopoda" family="Misophriidae" genus="Dimisophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468735FFDF43CBF9981F700015" box="[1190,1331,1564,1589]" italics="true" pageId="3" pageNumber="324">Dimisophria</emphasis>
</taxonomicName>
has a 2- segmented fifth leg (
<bibRefCitation id="660B48A58735FFDF46EDF9C418C60079" author="Boxshall" box="[384,645,1600,1625]" pageId="3" pageNumber="324" refString="Boxshall, G. A. &amp; Iliffe, T. M. (1987) Three new genera and five new species of Misophrioid copepods (Crustacea) from anchialine caves on Indo-West Pacific and North Atlantic Islands. Zoological Journal of the Linnean Society, 91, 223 - 252. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1987. tb 01510. x" type="journal article" year="1987">Boxshall &amp; Iliffe 1987</bibRefCitation>
), but this genus can be distinguished from
<taxonomicName id="C59A4ED78735FFDF4306F9C41EB90079" box="[1131,1274,1600,1625]" class="Maxillopoda" family="Speleophriidae" genus="Mexicophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="30EEE9468735FFDF4306F9C41EB90079" box="[1131,1274,1600,1625]" italics="true" pageId="3" pageNumber="324">Mexicophria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BDD543D8735FFDF426FF9C51F0D0079" box="[1282,1358,1601,1625]" pageId="3" pageNumber="324" rank="genus">n. gen.</taxonomicNameLabel>
by the lack of a bulbous lobe on the proximal segment of the antennule, and by mouthpart characters, such as the reduced setation on the mandibular endopod and the maxillulary exopod.
<taxonomicName id="C59A4ED78735FFDF44E0F90C1E690081" box="[909,1066,1672,1697]" class="Maxillopoda" family="Speleophriidae" genus="Expansophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468735FFDF44E0F90C1E690081" box="[909,1066,1672,1697]" italics="true" pageId="3" pageNumber="324">Expansophria</emphasis>
</taxonomicName>
species vary in the form of the female fifth leg. In female
<taxonomicName id="C59A4ED78735FFDF46B0F92A19D300E5" authority="Boxshall &amp; Iliffe, 1987" authorityName="Boxshall &amp; Iliffe" authorityYear="1987" box="[477,912,1708,1733]" class="Maxillopoda" family="Speleophriidae" genus="Expansophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="dimorpha">
<emphasis id="30EEE9468735FFDF46B0F92A182C00E5" box="[477,623,1708,1733]" italics="true" pageId="3" pageNumber="324">E. dimorpha</emphasis>
<bibRefCitation id="660B48A58735FFDF4516F92819D300E5" author="Boxshall" box="[635,912,1708,1733]" pageId="3" pageNumber="324" refString="Boxshall, G. A. &amp; Iliffe, T. M. (1987) Three new genera and five new species of Misophrioid copepods (Crustacea) from anchialine caves on Indo-West Pacific and North Atlantic Islands. Zoological Journal of the Linnean Society, 91, 223 - 252. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1987. tb 01510. x" type="journal article" year="1987">Boxshall &amp; Iliffe, 1987</bibRefCitation>
</taxonomicName>
the fifth leg is 3-segmented, comprising an unarmed coxa, the basis with a single outer seta and a 1-segmented exopod with 2 apical setae but in
<taxonomicName id="C59A4ED78735FFDF4259F9551BDD012D" authority="Boxshall &amp; Iliffe, 1987" authorityName="Boxshall &amp; Iliffe" authorityYear="1987" class="Maxillopoda" family="Speleophriidae" genus="Expansophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="apoda">
<emphasis id="30EEE9468735FFDF4259F9551FDF00C9" box="[1332,1436,1744,1769]" italics="true" pageId="3" pageNumber="324">E. apoda</emphasis>
<bibRefCitation id="660B48A58735FFDF47FAF9701BDD012D" author="Boxshall" box="[151,414,1780,1805]" pageId="3" pageNumber="324" refString="Boxshall, G. A. &amp; Iliffe, T. M. (1987) Three new genera and five new species of Misophrioid copepods (Crustacea) from anchialine caves on Indo-West Pacific and North Atlantic Islands. Zoological Journal of the Linnean Society, 91, 223 - 252. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1987. tb 01510. x" type="journal article" year="1987">Boxshall &amp; Iliffe, 1987</bibRefCitation>
</taxonomicName>
, the female fifth leg is lacking. Within the family, this 3-segmented state is the most similar to that found in the new genus. However, the male fifth legs differ markedly between the new genus and
<taxonomicName id="C59A4ED78735FFDF47FAF8B81B770175" box="[151,308,1852,1877]" class="Maxillopoda" family="Speleophriidae" genus="Expansophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468735FFDF47FAF8B81B770175" box="[151,308,1852,1877]" italics="true" pageId="3" pageNumber="324">Expansophria</emphasis>
</taxonomicName>
: in the former they are 3-segmented with 1-segmented exopod whereas known males of the latter genus have a 5-segmented fifth leg with a 3-segmented exopod (
<bibRefCitation id="660B48A58735FFDF4408F8E41E250159" author="Boxshall" box="[869,1126,1888,1913]" pageId="3" pageNumber="324" refString="Boxshall, G. A. &amp; Iliffe, T. M. (1987) Three new genera and five new species of Misophrioid copepods (Crustacea) from anchialine caves on Indo-West Pacific and North Atlantic Islands. Zoological Journal of the Linnean Society, 91, 223 - 252. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1987. tb 01510. x" type="journal article" year="1987">Boxshall &amp; Iliffe 1987</bibRefCitation>
,
<bibRefCitation id="660B48A58735FFDF431CF8E51FD30159" author="Jaume" box="[1137,1424,1888,1913]" pageId="3" pageNumber="324" refString="Jaume, D. &amp; Boxshall, G. A. (1996 b) The persistence of an ancient marine Fauna in Mediterranean waters: new evidence from misophrioid copepods living in anchihaline caves. Journal of Natural History, 30, 1583 - 1595. http: // dx. doi. org / 10.1080 / 00222939600770921" type="journal article" year="1996" yearSuffix="b">Jaume &amp; Boxshall 1996b</bibRefCitation>
).
</paragraph>
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The new genus exhibits reduced leg setation. The second endopodal segment of legs 2 to 4 bears only 1 inner seta and this is a unique character state for the family. The third exopodal segment of legs 1 to 4 carries 3, 4, 4, 3 inner margin setae, respectively, as in
<taxonomicName id="C59A4ED78735FFDF4520F848188601C5" box="[589,709,1996,2021]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468735FFDF4520F848188601C5" box="[589,709,1996,2021]" italics="true" pageId="3" pageNumber="324">Boxshallia</emphasis>
</taxonomicName>
,
<taxonomicName id="C59A4ED78735FFDF45B8F84A196701C5" box="[725,804,1998,2021]" class="Maxillopoda" family="Speleophriidae" genus="Huysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468735FFDF45B8F84A196701C5" box="[725,804,1998,2021]" italics="true" pageId="3" pageNumber="324">Huysia</emphasis>
</taxonomicName>
, and
<taxonomicName id="C59A4ED78735FFDF440BF8481E6A01C5" box="[870,1065,1996,2021]" class="Maxillopoda" family="Speleophriidae" genus="Protospeleophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="3" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468735FFDF440BF8481E6A01C5" box="[870,1065,1996,2021]" italics="true" pageId="3" pageNumber="324">Protospeleophria</emphasis>
</taxonomicName>
(
<bibRefCitation id="660B48A58735FFDF4354F8491EF901C5" author="Huys" box="[1081,1210,1997,2021]" pageId="3" pageNumber="324" refString="Huys, R. (1988) Boxshallia bulbantennulata gen. et sp. nov. (Copepoda: Misophrioida) from an anchialine lava pool on Lanzarote, Canary Islands. Stygologia, 4, 138 - 154." type="journal article" year="1988">Huys 1988</bibRefCitation>
,
<bibRefCitation id="660B48A58735FFDF43AAF8491FD701C5" author="Jaume" box="[1223,1428,1996,2021]" pageId="3" pageNumber="324" refString="Jaume, D., Boxshall, G. A. &amp; Iliffe, T. M. (1998) Two new genera of Misophrioid copepods (Crustacea) from an anchihaline cave in the Bahamas. Journal of Natural History, 32, 661 - 681. http: // dx. doi. org / 10.1080 / 00222939800770351" type="journal article" year="1998">
Jaume
<emphasis id="30EEE9468735FFDF4275F84A1F0801C5" box="[1304,1355,1996,2021]" italics="true" pageId="3" pageNumber="324">et al</emphasis>
. 1998
</bibRefCitation>
) and 3, 4, 4, 3 outer and distal spines. These other genera do not share the latter formula: in
<taxonomicName id="C59A4ED78732FFD8439DFF131F2B0690" box="[1264,1384,151,176]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD8439DFF131F2B0690" box="[1264,1384,151,176]" italics="true" pageId="4" pageNumber="325">Boxshallia</emphasis>
</taxonomicName>
and
<taxonomicName id="C59A4ED78732FFD847FAFF381B1906F5" box="[151,346,188,213]" class="Maxillopoda" family="Speleophriidae" genus="Protospeleophria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD847FAFF381B1906F5" box="[151,346,188,213]" italics="true" pageId="4" pageNumber="325">Protospeleophria</emphasis>
</taxonomicName>
it is 4, 4, 4, 4, while in
<taxonomicName id="C59A4ED78732FFD8450BFF3918F606F4" box="[614,693,189,212]" class="Maxillopoda" family="Speleophriidae" genus="Huysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD8450BFF3918F606F4" box="[614,693,189,212]" italics="true" pageId="4" pageNumber="325">Huysia</emphasis>
</taxonomicName>
it is 3, 4, 4, 4. The presence of flagellated tips on the outer spines of exopodal segments 2 and 3 of leg 1 is a fine scale similarity shared only with
<taxonomicName id="C59A4ED78732FFD84322FF5B1E8406D8" box="[1103,1223,223,248]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD84322FF5B1E8406D8" box="[1103,1223,223,248]" italics="true" pageId="4" pageNumber="325">Boxshallia</emphasis>
</taxonomicName>
out of these three genera.
<taxonomicName id="C59A4ED78732FFD8479CFE801B2A073D" box="[241,361,260,285]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD8479CFE801B2A073D" box="[241,361,260,285]" italics="true" pageId="4" pageNumber="325">Boxshallia</emphasis>
</taxonomicName>
and the new genus also share the possession of the conspicuous bulbous lobe on the proximal segment of the antennule.
</paragraph>
<paragraph id="022535548732FFD847AAFEC81B480438" blockId="4.[151,1437,151,536]" pageId="4" pageNumber="325">
The new genus can be distinguished from
<taxonomicName id="C59A4ED78732FFD845C6FEC819600745" box="[683,803,332,357]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD845C6FEC819600745" box="[683,803,332,357]" italics="true" pageId="4" pageNumber="325">Boxshallia</emphasis>
</taxonomicName>
by its derived 2-segmented endopod of leg 1 since this ramus retains a 3-segmented state in
<taxonomicName id="C59A4ED78732FFD8452FFEEB18F907A8" box="[578,698,367,392]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD8452FFEEB18F907A8" box="[578,698,367,392]" italics="true" pageId="4" pageNumber="325">Boxshallia</emphasis>
</taxonomicName>
. Another major difference between these two genera is the fifth leg. In
<taxonomicName id="C59A4ED78732FFD84788FE101B1E078D" box="[229,349,404,429]" class="Maxillopoda" family="Speleophriidae" genus="Boxshallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="30EEE9468732FFD84788FE101B1E078D" box="[229,349,404,429]" italics="true" pageId="4" pageNumber="325">Boxshallia</emphasis>
</taxonomicName>
the intercoxal sclerite is retained, the coxa and basis are separate in both sexes and the exopod is 2-segmented, plus the endopod is represented by an inner seta on the basis, so the leg is biramous (
<bibRefCitation id="660B48A58732FFD8439CFE3C1F3207F0" author="Huys" box="[1265,1393,440,464]" pageId="4" pageNumber="325" refString="Huys, R. (1988) Boxshallia bulbantennulata gen. et sp. nov. (Copepoda: Misophrioida) from an anchialine lava pool on Lanzarote, Canary Islands. Stygologia, 4, 138 - 154." type="journal article" year="1988">Huys 1988</bibRefCitation>
). In contrast, in the new genus the leg is uniramous, the exopod is 1-segmented in both sexes and the intercoxal sclerite is lacking.
</paragraph>
</subSubSection>
</treatment>
</document>