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<mods:title id="7417FA18EB37AE777B5336669ADD9B26">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<heading id="2A0581ADFFB7951AFF49FC01D6C458A5" bold="true" box="[151,1192,906,932]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
<emphasis id="4386EAD3FFB7951AFF49FC01D6C458A5" bold="true" box="[151,1192,906,932]" pageId="2" pageNumber="153">
Key to genera of
<taxonomicName id="B6F24D42FFB7951AFEB2FC01D02F58A5" authorityName="Crowson" authorityYear="1952" box="[364,579,906,932]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
(modified from
<bibRefCitation id="15634B30FFB7951AFCD1FC01D1D458A5" author="Leschen, R. A. B." box="[783,952,906,932]" pageId="2" pageNumber="153" refId="ref19416" refString="Leschen, R. A. B. (2003) Fauna of New Zealand 47. Erotylidae (Insecta: Coleoptera: Cucujoidea): Phylogeny and Review. Manaaki Whenua Press, Lincoln, 108 pp." type="book" year="2003">Leschen 2003</bibRefCitation>
and
<bibRefCitation id="15634B30FFB7951AFC2BFC01D6CC58A5" author="Xu, G. &amp; Tang, W. &amp; Skelley, P. &amp; Liu, N. &amp; Rich, S." box="[1013,1184,906,932]" pageId="2" pageNumber="153" pagination="251 - 278" refId="ref20834" refString="Xu, G., Tang, W., Skelley, P., Liu, N. &amp; Rich, S. (2015) Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis (Cycadaceae) in Asia. Zootaxa, 3986 (3), 251 - 278. https: // doi. org / 10.11646 / zootaxa. 3986.3.1" type="journal article" year="2015">
Xu
<emphasis id="4386EAD3FFB7951AFBFCFC01D63158A5" bold="true" box="[1058,1117,906,932]" italics="true" pageId="2" pageNumber="153">et al.</emphasis>
2015
</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
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<paragraph id="714D36C1FFB7951AFF49FC5BD7F758E7" blockId="2.[151,1436,976,1732]" box="[151,1435,976,998]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FC5BD7F758E7" box="[151,1435,976,998]" pageId="2" pageNumber="153">
1 Antennal club of 2 antennomeres..........................................................
<taxonomicName id="B6F24D42FFB7951AFB00FC5BD7F758E7" authority="Jacobson" authorityName="Jacobson" authorityYear="1915" box="[1246,1435,976,998]" class="Insecta" family="Erotylidae" genus="Setariola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="4386EAD3FFB7951AFB00FC5BD7F758E7" bold="true" box="[1246,1435,976,998]" pageId="2" pageNumber="153">
<emphasis id="4386EAD3FFB7951AFB00FC5BD75B58E7" bold="true" box="[1246,1335,976,998]" italics="true" pageId="2" pageNumber="153">Setariola</emphasis>
Jacobson
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<keyLead id="CA0395F4FFB7951AFF49FC65D7F05F02" box="[151,1436,1005,1027]" pageId="2" pageNumber="153">
- Antennal club of 3 antennomeres.........................................................................
<emphasis id="4386EAD3FFB7951AFA4EFC65D7F05F02" bold="true" box="[1424,1436,1006,1027]" pageId="2" pageNumber="153">2</emphasis>
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<paragraph id="714D36C1FFB7951AFF49FB80D7F75F20" blockId="2.[151,1436,976,1732]" box="[151,1435,1035,1057]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FB80D7F75F20" box="[151,1435,1035,1057]" pageId="2" pageNumber="153">
2 Pronotal callosities present; vestiture of scale-like setae................................
<taxonomicName id="B6F24D42FFB7951AFBA0FB80D7F75F20" authority="Rosenhauer" authorityName="Rosenhauer" authorityYear="1856" box="[1150,1435,1035,1057]" class="Insecta" family="Erotylidae" genus="Leucohimatium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="4386EAD3FFB7951AFBA0FB80D7F75F20" bold="true" box="[1150,1435,1035,1057]" pageId="2" pageNumber="153">
<emphasis id="4386EAD3FFB7951AFBA0FB80D7775F20" bold="true" box="[1150,1307,1035,1057]" italics="true" pageId="2" pageNumber="153">Leucohimatium</emphasis>
Rosenhauer
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<keyLead id="CA0395F4FFB7951AFF49FBA2D7F05F3F" box="[151,1436,1064,1086]" pageId="2" pageNumber="153">
- Pronotal callosities absent; vestiture of simple setae..........................................................
<emphasis id="4386EAD3FFB7951AFA4EFBA2D7F05F3F" bold="true" box="[1424,1436,1065,1086]" pageId="2" pageNumber="153">3</emphasis>
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<paragraph id="714D36C1FFB7951AFF49FBCED7F05F78" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FBCED7F05F78" pageId="2" pageNumber="153">
3 Eyes small, ventrally not encroaching upon gular area, distance between eye and lateral corner of mentum about equal to length of eye in ventral view; not associated with cycads......................................................
<emphasis id="4386EAD3FFB7951AFA4EFBEFD7F05F78" bold="true" box="[1424,1436,1124,1145]" pageId="2" pageNumber="153">4</emphasis>
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<keyLead id="CA0395F4FFB7951AFF49FB0AD7F05FB2" pageId="2" pageNumber="153">
- Eyes large, ventrally encroaching upon gular area, distance between eye and lateral corner of mentum &lt;1/2 eye length in ventral view; most associated with cycads.....................................................................
<emphasis id="4386EAD3FFB7951AFA4EFB15D7F05FB2" bold="true" box="[1424,1436,1182,1203]" pageId="2" pageNumber="153">5</emphasis>
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<paragraph id="714D36C1FFB7951AFF49FB30D7F05FD0" blockId="2.[151,1436,976,1732]" box="[151,1436,1211,1233]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FB30D7F05FD0" box="[151,1436,1211,1233]" pageId="2" pageNumber="153">
4 Dorsal punctation strong and impressed....................................................
<taxonomicName id="B6F24D42FFB7951AFB11FB30D7F05FD0" authority="Reitter" authorityName="Reitter" authorityYear="1878" box="[1231,1436,1211,1233]" class="Insecta" family="Erotylidae" genus="Henoticonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="4386EAD3FFB7951AFB11FB30D7F05FD0" bold="true" box="[1231,1436,1211,1233]" pageId="2" pageNumber="153">
<emphasis id="4386EAD3FFB7951AFB11FB30D7215FD0" bold="true" box="[1231,1357,1211,1233]" italics="true" pageId="2" pageNumber="153">Henoticonus</emphasis>
Reitter
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="714D36C1FFB7951AFF49FB52D7F05FEF" blockId="2.[151,1436,976,1732]" box="[151,1436,1240,1262]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FB52D7F05FEF" box="[151,1436,1240,1262]" pageId="2" pageNumber="153">
- Dorsal punctation weak and not strongly impressed.........................................
<taxonomicName id="B6F24D42FFB7951AFB62FB53D7F05FEF" authority="Reitter" authorityName="Reitter" authorityYear="1876" box="[1212,1436,1240,1262]" class="Insecta" family="Erotylidae" genus="Loberogosmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="4386EAD3FFB7951AFB62FB53D7F05FEF" bold="true" box="[1212,1436,1240,1262]" pageId="2" pageNumber="153">
<emphasis id="4386EAD3FFB7951AFB62FB53D7215FEF" bold="true" box="[1212,1357,1240,1262]" italics="true" pageId="2" pageNumber="153">Loberogosmus</emphasis>
Reitter
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA062E64FFB7951AFF49FB7ED7F05E9F" pageId="2" pageNumber="153">
<paragraph id="714D36C1FFB7951AFF49FB7ED7F05E47" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FB7ED7F05E47" pageId="2" pageNumber="153">
5 Submental-gular suture visible as depressed darkened area, often with row of punctures and setae; anterior pronotal margin of most with fine, distinct, complete marginal line; male genitalia with median lobe and tegmen laterally flattened and slightly to strongly twisted (at least basally); spiculum gastrale asymmetrical; Asia...............
<taxonomicName id="B6F24D42FFB7951AFB8DFABBD7F05E47" authority="Xu, Tang &amp; Skelley" authorityName="Xu, Tang &amp; Skelley" box="[1107,1436,1328,1350]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="4386EAD3FFB7951AFB8DFABBD7F05E47" bold="true" box="[1107,1436,1328,1350]" pageId="2" pageNumber="153">
<emphasis id="4386EAD3FFB7951AFB8DFABBD6A35E47" bold="true" box="[1107,1231,1328,1350]" italics="true" pageId="2" pageNumber="153">Cycadophila</emphasis>
Xu, Tang &amp; Skelley
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="714D36C1FFB7951AFF49FAC5D7F05E9F" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FAC5D7F05E9F" pageId="2" pageNumber="153">
- Submental-gular suture apparently lacking, surface flattened; anterior pronotal margin lacking marginal line, some with row of fine punctures; male genitalia with median lobe cylindrical and tegmen dorsoventrally or laterally flattened, but usually not twisted; spiculum gastrale symmetrical; Americas............................................................
<emphasis id="4386EAD3FFB7951AFA4EFA02D7F05E9F" bold="true" box="[1424,1436,1417,1438]" pageId="2" pageNumber="153">6</emphasis>
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<paragraph id="714D36C1FFB7951AFF49FA2ED7F05D30" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49FA2ED7F05D30" pageId="2" pageNumber="153">
6 Lateral pronotal carina thicker anteriorly in lateral view (
<figureCitation id="E9C92A44FFB7951AFCCBFA2ED1395EBA" box="[789,853,1445,1467]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Figs. 1</figureCitation>
DE); head with supraocular lines and elytra lacking basal bead; stridulatory file at base of head separated by distance ~ 0.5× width of the scutellar shield (
<figureCitation id="E9C92A44FFB7951AFBA4FA48D6A95ED8" box="[1146,1221,1475,1497]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Fig. 1B</figureCitation>
); tarsomeres IIII distinctly widened (
<figureCitation id="E9C92A44FFB7951AFE5DFA6BD3A95EF7" box="[387,453,1504,1526]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (AE: male holotype): AC) dorsal, ventral and lateral and habitus; DF) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="2" pageNumber="153">Figs. 4</figureCitation>
DE, 8F), metatarsomere I width ± length; male genitalia with median lobe arched in lateral view; flagellum length Ĺ length of median lobe and not coiled (
<figureCitation id="E9C92A44FFB7951AFD32FA76D1535D12" box="[748,831,1533,1555]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (AE: male holotype): AC) dorsal, ventral and lateral and habitus; DF) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="2" pageNumber="153">Figs. 4G</figureCitation>
,
<figureCitation id="E9C92A44FFB7951AFC97FA76D10F5D12" box="[841,867,1533,1555]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (AE: female allotype): AC) dorsal, ventral and lateral and habitus; DE) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="2" pageNumber="153">5F</figureCitation>
,
<figureCitation id="E9C92A44FFB7951AFCB2FA76D1E05D12" box="[876,908,1533,1555]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (AF: female allotype): AC) dorsal, ventral and lateral and habitus; DF) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="2" pageNumber="153">6H</figureCitation>
,
<figureCitation id="E9C92A44FFB7951AFC48FA76D1DF5D12" box="[918,947,1533,1555]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (AE: male holotype): AC) dorsal, ventral and lateral and habitus; DE) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="2" pageNumber="153">7G</figureCitation>
,
<figureCitation id="E9C92A44FFB7951AFC62FA76D1B05D12" box="[956,988,1533,1555]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (AD, F: male holotype): AC) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="2" pageNumber="153">8G</figureCitation>
);
<collectingCountry id="09E57651FFB7951AFC30FA76D65B5D12" box="[1006,1079,1533,1555]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
and probably
<collectingCountry id="09E57651FFB7951AFB1CFA76D76E5D12" box="[1218,1282,1533,1555]" name="Belize" pageId="2" pageNumber="153">Belize</collectingCountry>
,
<collectingCountry id="09E57651FFB7951AFAD5FA76D71F5D12" box="[1291,1395,1533,1555]" name="Guatemala" pageId="2" pageNumber="153">Guatemala</collectingCountry>
and
<collectingCountry id="09E57651FFB7951AFF3FF990D3525D30" box="[225,318,1563,1585]" name="Honduras" pageId="2" pageNumber="153">Honduras</collectingCountry>
on
<taxonomicName id="B6F24D42FFB7951AFEBCF997D3B25D30" ID-CoL="3KXP" box="[354,478,1564,1585]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">
<emphasis id="4386EAD3FFB7951AFEBCF997D3B25D30" box="[354,478,1564,1585]" italics="true" pageId="2" pageNumber="153">Ceratozamia</emphasis>
</taxonomicName>
............................................................
<emphasis id="4386EAD3FFB7951AFB6FF990D7F05D30" bold="true" box="[1201,1436,1563,1585]" pageId="2" pageNumber="153">
<taxonomicName id="B6F24D42FFB7951AFB6FF990D74A5D30" authority="Tang &amp; Skelley &amp; Pérez-Farrera, 2018" authorityName="Tang &amp; Skelley &amp; Pérez-Farrera" authorityYear="2018" box="[1201,1318,1563,1585]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="4386EAD3FFB7951AFB6FF990D74A5D30" bold="true" box="[1201,1318,1563,1585]" italics="true" pageId="2" pageNumber="153">Ceratophila</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="58B557A8FFB7951AFAECF997D7F05D30" box="[1330,1436,1564,1585]" pageId="2" pageNumber="153" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="714D36C1FFB7951AFF49F9B2D7F05DC2" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
<keyLead id="CA0395F4FFB7951AFF49F9B2D7F05DC2" pageId="2" pageNumber="153">
- Lateral pronotal carina narrow, same thickness along entire length in lateral view (
<figureCitation id="E9C92A44FFB7951AFC2CF9B3D6505D4F" box="[1010,1084,1592,1614]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Fig. 1C</figureCitation>
); head without supraocular lines and elytra with distinct basal bead (
<figureCitation id="E9C92A44FFB7951AFDCEF9DED01B5D6A" box="[528,631,1621,1643]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">
Fig. 1A;
<taxonomicName id="B6F24D42FFB7951AFDB6F9DDD01B5D6A" authority="Chaves and Genaro" authorityName="Chaves and Genaro" box="[616,631,1622,1643]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="esperanzae">
<emphasis id="4386EAD3FFB7951AFDB6F9DDD01B5D6A" box="[616,631,1622,1643]" italics="true" pageId="2" pageNumber="153">P</emphasis>
</taxonomicName>
</figureCitation>
. esperanzae Chaves and Genaro is the exception); stridulatory files at base of head separated by distance ~ width of scutellar shield (
<figureCitation id="E9C92A44FFB7951AFD1CF9F8D1635D88" box="[706,783,1651,1673]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Fig. 1A</figureCitation>
); tarsomeres IIII narrow, metatarsomere I length always distinctly greater than width; male genitalia with median lobe straight; flagellum length &gt;&gt; length of median lobe and coiled; widespread New World on
<taxonomicName id="B6F24D42FFB7951AFEADF925D3835DC2" ID-CoL="3KXP" box="[371,495,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">
<emphasis id="4386EAD3FFB7951AFEADF925D3835DC2" box="[371,495,1710,1731]" italics="true" pageId="2" pageNumber="153">Ceratozamia</emphasis>
</taxonomicName>
,
<emphasis id="4386EAD3FFB7951AFE25F925D6DF5DC2" box="[507,1203,1710,1731]" italics="true" pageId="2" pageNumber="153">
<taxonomicName id="B6F24D42FFB7951AFE25F925D0555DC2" ID-CoL="62R3L" box="[507,569,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Dioon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">Dioon</taxonomicName>
,
<taxonomicName id="B6F24D42FFB7951AFD9DF925D0D95DC2" ID-CoL="5RPV" box="[579,693,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Microcycas" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">Microcycas</taxonomicName>
,
<taxonomicName id="B6F24D42FFB7951AFD60F925D0975DC2" ID-CoL="63X6Q" box="[702,763,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">Zamia</taxonomicName>
.....................................
</emphasis>
<taxonomicName id="B6F24D42FFB7951AFB61F925D7F05DC2" ID-CoL="6MVR" authority="Reitter" authorityName="Reitter" authorityYear="1875" box="[1215,1436,1710,1732]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="4386EAD3FFB7951AFB61F925D7215DC5" bold="true" box="[1215,1357,1710,1732]" italics="true" pageId="2" pageNumber="153">Pharaxonotha</emphasis>
Reitter
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>