386 lines
50 KiB
XML
386 lines
50 KiB
XML
<document id="41CF72026ED317EA8AC9E01FC8F96430" ID-CLB-Dataset="28054" ID-DOI="10.11646/zootaxa.4508.2.1" ID-GBIF-Dataset="da79b654-bd50-4715-8ed3-80fa7a00d469" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606934" ID-ZooBank="5E2BC894-1919-4F63-8EF5-BAAC91913388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585631772" checkinUser="plazi" docAuthor="Tang, William, Skelley, Paul & Pérez-Farrera, Miguel Angel" docDate="2018" docId="F95B87D7FFA29501FF49FF1CD3BF5C05" docLanguage="en" docName="zootaxa.4508.2.1.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophila (Vovidesa) vazquezi Tang & Skelley & Pérez-Farrera 2018, new species" docType="treatment" docVersion="7" lastPageNumber="176" masterDocId="0562FFAFFFB59518FFDEFF8BD26C5B01" masterDocTitle="Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)" masterLastPageNumber="178" masterPageNumber="151" pageNumber="174" updateTime="1698712984586" updateUser="ExternalLinkService">
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<mods:title id="6ED1D0777F8C90927C366A82747E788D">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<mods:namePart id="341DE69A2AFE8C1204D7C5CD5EE2BDED">Tang, William</mods:namePart>
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<mods:namePart id="1F3A405E727DFC1B6988D3D4589483E9">Skelley, Paul</mods:namePart>
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<mods:name id="30D3B5E5257D40BAA6C49D145BE6E5EA" type="personal">
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<mods:namePart id="955F9728A26477BA6B75CC7007BEE545">Pérez-Farrera, Miguel Angel</mods:namePart>
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<mods:title id="C448A1D0F9BE5FFB7AF12A38D5044258">Zootaxa</mods:title>
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<mods:part id="F9ABB44D2E9C92F4EB09032668B8E2E7">
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<mods:date id="5797E8018286B5F96A1A98A6091AC13C">2018</mods:date>
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<mods:number id="70DDCEFC11F5261A86BE6902D4E27862">2018-10-31</mods:number>
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<mods:number id="6E1F131EDC0C735B307A2B1436DE8373">4508</mods:number>
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<treatment id="F95B87D7FFA29501FF49FF1CD3BF5C05" ID-DOI="http://doi.org/10.5281/zenodo.5958044" ID-GBIF-Taxon="154495722" ID-Zenodo-Dep="5958044" LSID="urn:lsid:plazi:treatment:F95B87D7FFA29501FF49FF1CD3BF5C05" httpUri="http://treatment.plazi.org/id/F95B87D7FFA29501FF49FF1CD3BF5C05" lastPageId="25" lastPageNumber="176" pageId="23" pageNumber="174">
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<subSubSection id="39E8654AFFA2950FFF49FF1CD6785BB1" box="[151,1044,151,176]" pageId="23" pageNumber="174" type="nomenclature">
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<paragraph id="714D36C1FFA2950FFF49FF1CD6785BB1" blockId="23.[151,1044,151,213]" box="[151,1044,151,176]" pageId="23" pageNumber="174">
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<heading id="2A0581ADFFA2950FFF49FF1CD6785BB1" bold="true" box="[151,1044,151,176]" fontSize="10" level="2" pageId="23" pageNumber="174" reason="2">
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<emphasis id="4386EAD3FFA2950FFF49FF1CD6785BB1" bold="true" box="[151,1044,151,176]" pageId="23" pageNumber="174">
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<taxonomicName id="B6F24D42FFA2950FFF49FF1CD1135BB1" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,895,151,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="23" pageNumber="174" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Vovidesa">
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<emphasis id="4386EAD3FFA2950FFF49FF1CD34C5BB1" bold="true" box="[151,288,151,176]" italics="true" pageId="23" pageNumber="174">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFA2950FFEEEFF1CD3FF5BB1" bold="true" box="[304,403,151,176]" italics="true" pageId="23" pageNumber="174">Vovidesa</emphasis>
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)
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<emphasis id="4386EAD3FFA2950FFE7BFF1CD06B5BB1" bold="true" box="[421,519,151,176]" italics="true" pageId="23" pageNumber="174">vazquezi</emphasis>
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Tang, Skelley & Pérez-Farrera
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</taxonomicName>
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,
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<taxonomicNameLabel id="58B557A8FFA2950FFC55FF1CD6785BB1" box="[907,1044,151,176]" pageId="23" pageNumber="174" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFA2950FFF49FF37D3405BD5" box="[151,300,188,213]" pageId="23" pageNumber="174" type="description">
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<paragraph id="714D36C1FFA2950FFF49FF37D3405BD5" blockId="23.[151,1044,151,213]" box="[151,300,188,213]" pageId="23" pageNumber="174">
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(
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<figureCitation id="E9C92A44FFA2950FFF41FF37D2865BD5" box="[159,234,188,213]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8</figureCitation>
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A–H)
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFA2950FFF49FE8FD6FB5A86" pageId="23" pageNumber="174" type="diagnosis">
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<paragraph id="714D36C1FFA2950FFF49FE8FD6FB5A86" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
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<emphasis id="4386EAD3FFA2950FFF49FE8FD3365A1C" bold="true" box="[151,346,260,285]" pageId="23" pageNumber="174">Adult diagnosis.</emphasis>
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Distinguished from other
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<taxonomicName id="B6F24D42FFA2950FFD58FE8FD1E15A1D" authority="(Vovidesa )" baseAuthorityName="Vovidesa" box="[646,909,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="23" pageNumber="155" phylum="Tracheophyta" rank="genus">
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<emphasis id="4386EAD3FFA2950FFD58FE8FD1615A1D" box="[646,781,260,284]" italics="true" pageId="23" pageNumber="174">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFA2950FFCC1FE8FD1EC5A1D" box="[799,896,260,284]" italics="true" pageId="23" pageNumber="174">Vovidesa</emphasis>
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)
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</taxonomicName>
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species by male with subapical emarginations on tibiae, submentum of both sexes with dense punctures and setae, male submental setae projecting laterally, female elytra similarly dulled medially and laterally, male genitalia with median lobe nearly laterally flattened and curved, and with a known distribution in
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<collectingCountry id="09E57651FFA2950FFDBBFEE4D0D15A89" box="[613,701,367,392]" name="Mexico" pageId="23" pageNumber="174">Mexico</collectingCountry>
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, state of
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<collectingRegion id="B336F823FFA2950FFCFFFEFBD1EB5A86" box="[801,903,368,391]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="23" pageNumber="174">Veracruz</collectingRegion>
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, on
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<emphasis id="4386EAD3FFA2950FFC6BFEFBD6FB5A86" box="[949,1175,368,391]" italics="true" pageId="23" pageNumber="174">
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<taxonomicName id="B6F24D42FFA2950FFC6BFEFBD6FF5A86" box="[949,1171,368,391]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="23" pageNumber="174" phylum="Tracheophyta" rank="species" species="tenuis">Ceratozamia tenuis</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFA29501FF19FE1FD28F587E" lastPageId="25" lastPageNumber="176" pageId="23" pageNumber="174" type="description">
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<paragraph id="714D36C1FFA2950FFF19FE1FD1D0593D" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
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<emphasis id="4386EAD3FFA2950FFF19FE1FD38C5AAC" bold="true" box="[199,480,404,429]" pageId="23" pageNumber="174">Adult male description.</emphasis>
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Length
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<quantity id="B60A9B24FFA2950FFDE0FE1FD08F5AAD" box="[574,739,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.51" metricValueMax="4.9" metricValueMin="4.12" pageId="23" pageNumber="174" unit="mm" value="4.51" valueMax="4.9" valueMin="4.12">4.12–4.90 mm</quantity>
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, width
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<quantity id="B60A9B24FFA2950FFCE8FE1FD1B65AAD" box="[822,986,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.84" metricValueMin="1.57" pageId="23" pageNumber="174" unit="mm" value="1.705" valueMax="1.84" valueMin="1.57">1.57–1.84 mm</quantity>
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(n = 10). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
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<figureCitation id="E9C92A44FFA2950FFAD5FE3CD7345AD1" box="[1291,1368,439,464]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8</figureCitation>
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A–F), head and pronotum concolorous and slightly darker than the yellowish-brown elytra; dorsal surface punctate, shining, short procumbent hairs associated with punctation, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
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</paragraph>
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<paragraph id="714D36C1FFA2950FFF19FDCCD3DB58ED" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
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<emphasis id="4386EAD3FFA2950FFF19FDCCD36B5961" bold="true" box="[199,263,583,608]" pageId="23" pageNumber="174">Head</emphasis>
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in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
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<figureCitation id="E9C92A44FFA2950FFEC9FDE7D3CC5985" box="[279,416,620,645]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8A, D</figureCitation>
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), average distance between closest punctures 2× width of puncture; width
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<quantity id="B60A9B24FFA2950FFB2FFDE7D7F45985" box="[1265,1432,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="9.1" metricValueMin="8.1" pageId="23" pageNumber="174" unit="mm" value="0.86" valueMax="0.91" valueMin="0.81">0.81–0.91 mm</quantity>
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; dorsal interocular distance
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<quantity id="B60A9B24FFA2950FFE3DFD04D0FE59A6" box="[483,658,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8999999999999995" metricValueMin="5.2" pageId="23" pageNumber="174" unit="mm" value="0.5549999999999999" valueMax="0.59" valueMin="0.52">0.52–0.59 mm</quantity>
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, head width/dorsal interocular distance ratio 1.45–1.56, ventral interocular distance
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<quantity id="B60A9B24FFA2950FFEA3FD3FD04859CD" box="[381,548,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="23" pageNumber="174" unit="mm" value="0.4" valueMax="0.42" valueMin="0.38">0.38–0.42 mm</quantity>
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, head width/ventral interocular distance ratio 2.04–2.17. Eye with large black facets. Antennal length approximately equal to pronotal width, 1.5× head width; antennomere I (scape) fairly large, slightly elongate, antennomere II subequal to III; IV–VIII small, width equals length; club fairly large, IX and X similar in length, XI slightly longer, globular with acuminate apex. Clypeus truncate anteriorly, moderately punctate, with narrow margin. Mentum with moderate punctation, distance between nearest punctures ~ 1× own width; submentum finely, densely punctate, each puncture with a long seta ~ 2–3× width of eye facet, setae projecting laterally. Gular area smooth, without punctation or setae, border with submentum usually with a shallow depression without punctation, but usually with a shallowly impressed circular or semicircular border along anterior margin (
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<figureCitation id="E9C92A44FFA2950FFE8DFC5FD3C658ED" box="[339,426,980,1005]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Fig. 8E</figureCitation>
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).
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</paragraph>
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<paragraph id="714D36C1FFA2950FFF19FC7CD6665C4D" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
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<emphasis id="4386EAD3FFA2950FFF19FC7CD34C5F11" bold="true" box="[199,288,1015,1040]" pageId="23" pageNumber="174">Thorax</emphasis>
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with pronotum transverse in dorsal view with marginal beads basally and laterally, anterior margin with weak bead, reduced to absent medially; length/width ratio (PL/PW) = 0.76–0.79, rectangular; anterior and posterior angles present, anterior angles projecting; lateral carinae expanding in basal 1/10 to parallel sides, sometimes slightly emarginated, parallel for 50% of length, then gradually converging to anterior angles; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/5 length of posterior margin from posterior corners and touching and just anterior to marginal bead, pores mark base of longitudinal furrows onto disc. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae ~ 1/3 length of eye. Hypomeron laterally apparently lacking punctures, some with minute setae visible; medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.75–1.85, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size, scutellary striole extending ¼ elytral length, with 9–16 punctures; intervals of striae with fine, shallow punctures of similar size to strial punctures. All punctures of elytral bearing a single fine, procumbent seta; seta often only visible in profile, extending posteriorly to, over or beyond next puncture (often abraded), interval puncture setae longer than strial puncture setae. Punctation on meso- and metaventrites moderate to dense, distance between nearest punctures 1–2× width of punctures, puncture depth shallow. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over 3/4 metaventrite length. Legs with procoxa oval; mesocoxa globular; metacoxa transversely elongateoval; trochanters obliquely truncate apically; in males all femora swollen, stout, dorso-ventral width greater than eye length; protibia with shallow medial subapical emargination, meso- and metatibia with a distinct medial subapical emargination; all tibiae obliquely truncate at apex, with complete fringe of fine spinules on anterior and posterior margins; protibia swollen, not flattened in cross section notably larger than other tibiae, triangularly dilated to apex; mesotibia weakly swollen, moderately triangularly dilated to apex; metatibia narrowed, weakly dilating to midlength then slightly narrowing to apex; pro- and mesotarsi greatly enlarged, corresponding tarsomere I width ~ 3× own length, metatarsomere I width ~ own length (
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<figureCitation id="E9C92A44FFA2950FFC84F8BFD1C95C4D" box="[858,933,1844,1869]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8</figureCitation>
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A–C, F).
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</paragraph>
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|
||
<paragraph id="714D36C1FFAD9500FF49F805D7F05CE8" blockId="24.[151,1436,1934,2025]" pageId="24" pageNumber="175">
|
||
<emphasis id="4386EAD3FFAD9500FF49F805D3785CA5" bold="true" box="[151,276,1934,1956]" pageId="24" pageNumber="175">FIGURE 8.</emphasis>
|
||
Photographs of
|
||
<taxonomicName id="B6F24D42FFAD9500FE1FF805D1625CA4" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[449,782,1934,1957]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="24" pageNumber="175" phylum="Tracheophyta" rank="subGenus" subGenus="Uovidesa">
|
||
<emphasis id="4386EAD3FFAD9500FE1FF805D0505CA5" box="[449,572,1934,1956]" italics="true" pageId="24" pageNumber="175">Ceratophila</emphasis>
|
||
(
|
||
<emphasis id="4386EAD3FFAD9500FD92F805D0C85CA5" box="[588,676,1934,1956]" italics="true" pageId="24" pageNumber="175">Uovidesa</emphasis>
|
||
)
|
||
<emphasis id="4386EAD3FFAD9500FD69F81BD1625CA4" box="[695,782,1936,1957]" italics="true" pageId="24" pageNumber="175">vazquezi</emphasis>
|
||
</taxonomicName>
|
||
(A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="714D36C1FFAC9501FF19FF1CD0BB59A9" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FF19FF1CD32F5BB1" bold="true" box="[199,323,151,176]" pageId="25" pageNumber="176">Abdomen.</emphasis>
|
||
Ventrite I with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites strongly alutaceous laterally, less so and glossy medially, bearing dense, moderate, shallow punctation, distance to nearest puncture ~ 1× width of puncture, punctures bearing mostly reclining setae; ventrites I–V with setae length ~ 2–3× width of puncture; I–V each with 2 or more median pairs of longer, semi-erect sensory hairs obscured in dense setation, V with additional 4–5 pairs of long semi-erect hairs located postero-laterally, but not at submargin. Male genitalia with genital capsule fringed with fine setae; tegmen sclerotized, triangular, laterally compressed; anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly (
|
||
<figureCitation id="E9C92A44FFAC9501FEFDFE74D31E5919" box="[291,370,511,536]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="25" pageNumber="176">Figs. 8</figureCitation>
|
||
F–G); apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorso-ventrally compressed, height approximately half its own width, in dorso-ventral view length/width ratio = 2.1–2.8; apically with setae, length of longest setae> width of paramere in dorso-ventral view (
|
||
<figureCitation id="E9C92A44FFAC9501FBECFDCCD6FC5961" box="[1074,1168,583,608]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="25" pageNumber="176">Fig. 8H</figureCitation>
|
||
); median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to a point. Ratio of penile strut length to median lobe length 1.82–1.83 (n = 2) (
|
||
<figureCitation id="E9C92A44FFAC9501FD99FD04D0FE59A9" box="[583,658,655,680]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="25" pageNumber="176">Figs. 8</figureCitation>
|
||
F–G).
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFAC9501FF19FD3FD28F587E" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FF19FD3FD37259CC" bold="true" box="[199,286,692,717]" pageId="25" pageNumber="176">Female</emphasis>
|
||
generally similar to male, except for sexual dimorphism. Female pronotum slightly wider than male, PL/PW = 0.70–0.76; submentum densely punctate, setae short, not projecting; elytral surface dulled medially and laterally; legs unmodified, fore and middle legs not swollen, femora dorso-ventral width about equal to eye length, all tibia gradually dilating to narrowly triangular apex; tibia lacking median subapical emarginations; all legs with tarsomere I width ~ own length; metaventrite flattened medially, not impressed; abdomen less densely punctate and setose.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39E8654AFFAC9501FF19FC07D2835E31" pageId="25" pageNumber="176" type="materials_examined">
|
||
<paragraph id="714D36C1FFAC9501FF19FC07D15258A4" blockId="25.[151,1437,151,1796]" box="[199,830,908,933]" pageId="25" pageNumber="176">
|
||
<materialsCitation id="C19A3C9CFFAC9501FF19FC07D15558A4" ID-GBIF-Occurrence="2028138738" box="[199,825,908,933]" collectorName="El Esquilon" country="Mexico" location="Veracruz" pageId="25" pageNumber="176" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
|
||
<emphasis id="4386EAD3FFAC9501FF19FC07D30958A4" bold="true" box="[199,357,908,933]" pageId="25" pageNumber="176">
|
||
<typeStatus id="AE498863FFAC9501FF19FC07D36E58A4" box="[199,258,908,933]" pageId="25" pageNumber="176">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="09E57651FFAC9501FEB2FC07D3A858A4" box="[364,452,908,933]" name="Mexico" pageId="25" pageNumber="176">Mexico</collectingCountry>
|
||
,
|
||
<collectingRegion id="B336F823FFAC9501FE11FC06D05A58A5" box="[463,566,909,932]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||
, Jilotepec,
|
||
<collectorName id="DC075317FFAC9501FD6DFC07D15558A4" box="[691,825,908,933]" pageId="25" pageNumber="176">El Esquilón</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFAC9501FF19FC24D1A558EC" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FF19FC24D37658C9" bold="true" box="[199,282,943,968]" pageId="25" pageNumber="176">Range.</emphasis>
|
||
As currently understood, this species is known to inhabit
|
||
<taxonomicName id="B6F24D42FFAC9501FC15FC3BD29858EC" authority="(Dyer) D. W. Stev. & Vovides" authorityName="D. W. Stev. & Vovides" baseAuthorityName="Dyer" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FC15FC3BD6C658C6" box="[971,1194,944,967]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
(Dyer) D.W.Stev. & Vovides
|
||
</taxonomicName>
|
||
, which occurs in the vicinity north of Xalapa,
|
||
<collectingRegion id="B336F823FFAC9501FD24FC5ED10D58ED" box="[762,865,981,1004]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry id="09E57651FFAC9501FCB2FC5FD1A958EC" box="[876,965,980,1005]" name="Mexico" pageId="25" pageNumber="176">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFAC9501FF19FC7CD2835E31" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FF19FC7CD04F5F11" bold="true" box="[199,547,1015,1040]" pageId="25" pageNumber="176">
|
||
Material examined.
|
||
<typeStatus id="AE498863FFAC9501FE68FC7CD04F5F11" box="[438,547,1015,1040]" pageId="25" pageNumber="176" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(by designation) male of
|
||
<taxonomicName id="B6F24D42FFAC9501FC9CFC73D1AF5F0E" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[834,963,1016,1039]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="vazquezi">
|
||
<emphasis id="4386EAD3FFAC9501FC9CFC73D1AF5F0E" box="[834,963,1016,1039]" italics="true" pageId="25" pageNumber="176">C. vazquezi</emphasis>
|
||
</taxonomicName>
|
||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||
<collectingCountry id="09E57651FFAC9501FE05FB97D0235F35" box="[475,591,1052,1076]" name="Mexico" pageId="25" pageNumber="176">MEXICO</collectingCountry>
|
||
,
|
||
<collectingRegion id="B336F823FFAC9501FD84FB96D0AD5F35" box="[602,705,1053,1076]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||
, Cerro Coacoatzintla,
|
||
<specimenCount id="67F4FD48FFAC9501FC65FB96D1BC5F35" box="[955,976,1053,1076]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||
cone
|
||
<taxonomicName id="B6F24D42FFAC9501FBCDFB97D6825F35" box="[1043,1262,1052,1076]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FBCDFB97D6825F35" box="[1043,1262,1052,1076]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="054C1001FFAC9501FB22FB97D7275F35" box="[1276,1355,1052,1076]" pageId="25" pageNumber="176" value="1928-02-01">28-II-1</date>
|
||
995, Y. Sanchez Tinoco”; 2) [rectangular; red; printed in black ink]
|
||
<typeStatus id="AE498863FFAC9501FCE6FBCBD1A05F59" box="[824,972,1088,1112]" pageId="25" pageNumber="176" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="67F4FD48FFAC9501FC0DFBCBD1845F56" box="[979,1000,1088,1111]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||
<taxonomicName id="B6F24D42FFAC9501FC31FBCBD3195F7D" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4386EAD3FFAC9501FC31FBCBD6B15F56" box="[1007,1245,1088,1112]" italics="true" pageId="25" pageNumber="176">Ceratophila vazquezi</emphasis>
|
||
<bibRefCitation id="15634B30FFAC9501FB3BFBCBD3195F7D" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." pageId="25" pageNumber="176" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
. Deposited in the FSCA.
|
||
<typeStatus id="AE498863FFAC9501FD40FBEFD16F5F7C" box="[670,771,1124,1149]" pageId="25" pageNumber="176" type="allotype">
|
||
<emphasis id="4386EAD3FFAC9501FD40FBEFD16F5F7C" bold="true" box="[670,771,1124,1149]" pageId="25" pageNumber="176">Allotype</emphasis>
|
||
</typeStatus>
|
||
(FSCA) same data as
|
||
<typeStatus id="AE498863FFAC9501FBD5FBEFD6015F7C" box="[1035,1133,1124,1149]" pageId="25" pageNumber="176" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="67F4FD48FFAC9501FB77FBEFD7795F7C" box="[1193,1301,1124,1149]" pageId="25" pageNumber="176" type="adult">535 adult</specimenCount>
|
||
<typeStatus id="AE498863FFAC9501FAC0FBEFD7F85F7C" box="[1310,1428,1124,1149]" pageId="25" pageNumber="176" type="paratype">
|
||
<emphasis id="4386EAD3FFAC9501FAC0FBEFD7F85F7C" bold="true" box="[1310,1428,1124,1149]" pageId="25" pageNumber="176">paratypes</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<collectingCountry id="09E57651FFAC9501FF49FB0CD3615FA1" box="[151,269,1159,1184]" name="Mexico" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FF49FB0CD3615FA1" bold="true" box="[151,269,1159,1184]" pageId="25" pageNumber="176">MEXICO</emphasis>
|
||
</collectingCountry>
|
||
:
|
||
<collectingRegion id="B336F823FFAC9501FEC5FB0CD3E45FA1" box="[283,392,1159,1184]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FEC5FB0CD3E45FA1" bold="true" box="[283,392,1159,1184]" pageId="25" pageNumber="176">Veracruz</emphasis>
|
||
</collectingRegion>
|
||
, Jilotepec, El Esquilón, ex male cone of
|
||
<taxonomicName id="B6F24D42FFAC9501FC91FB03D63D5F9E" box="[847,1105,1160,1183]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="mexicana">
|
||
<emphasis id="4386EAD3FFAC9501FC91FB03D63D5F9E" box="[847,1105,1160,1183]" italics="true" pageId="25" pageNumber="176">Ceratozamia mexicana</emphasis>
|
||
</taxonomicName>
|
||
[now
|
||
<taxonomicName id="B6F24D42FFAC9501FB46FB03D6905F9E" box="[1176,1276,1160,1183]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FB46FB03D6905F9E" box="[1176,1276,1160,1183]" italics="true" pageId="25" pageNumber="176">C. tenuis</emphasis>
|
||
</taxonomicName>
|
||
],
|
||
<date id="054C1001FFAC9501FACCFB03D7FB5FA1" box="[1298,1431,1160,1184]" pageId="25" pageNumber="176" value="1989-03-10">10-III-1989</date>
|
||
, A. P. Vovides
|
||
<quantity id="B60A9B24FFAC9501FEE8FB27D3EA5FC5" box="[310,390,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="176" unit="m" value="1300.0">1300m</quantity>
|
||
, (97); Cerro Coacoatzintla,
|
||
<specimenCount id="67F4FD48FFAC9501FD1FFB26D0BA5FC5" box="[705,726,1197,1220]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||
cone
|
||
<taxonomicName id="B6F24D42FFAC9501FCC4FB27D1995FC5" box="[794,1013,1196,1220]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FCC4FB27D1995FC5" box="[794,1013,1196,1220]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="054C1001FFAC9501FBDAFB27D6EC5FC4" box="[1028,1152,1196,1221]" pageId="25" pageNumber="176" value="1995-02-28">28-II-1995</date>
|
||
, Y. Sanchez Tinoco (77); W. of Mafafas, [GPS coord. omitted],
|
||
<quantity id="B60A9B24FFAC9501FD99FB5BD0CE5FE9" box="[583,674,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.321" pageId="25" pageNumber="176" unit="m" value="1321.0">1321 m</quantity>
|
||
., ex
|
||
<specimenCount id="67F4FD48FFAC9501FD0BFB5AD0865FE9" box="[725,746,1233,1256]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||
<taxonomicName id="B6F24D42FFAC9501FD2FFB5BD1A15FE6" box="[753,973,1232,1255]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FD2FFB5BD1A15FE6" box="[753,973,1232,1255]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
</taxonomicName>
|
||
late shed cone,
|
||
<date id="054C1001FFAC9501FB5DFB44D7635FE9" box="[1155,1295,1231,1256]" pageId="25" pageNumber="176" value="2018-04-15">15- IV-2018</date>
|
||
, M.A. Perez Farrera & W. Tang (359):
|
||
<typeStatus id="AE498863FFAC9501FE1DFB7ED05C5E0C" box="[451,560,1269,1293]" pageId="25" pageNumber="176" type="paratype">Paratypes</typeStatus>
|
||
deposited at ANIC, BMNH, FSCA, IEXA, INBio, MNHN, NZAC, UNAM, USNM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39E8654AFFAC9501FF19FAB7D0BE5E79" pageId="25" pageNumber="176" type="etymology">
|
||
<paragraph id="714D36C1FFAC9501FF19FAB7D0BE5E79" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FF19FAB7D3225E54" bold="true" box="[199,334,1340,1365]" pageId="25" pageNumber="176">Etymology.</emphasis>
|
||
The species is named in honor of Dr. Mario Vázquez Torres for his work on the taxonomy, ecology and conservation of the cycads of
|
||
<collectingCountry id="09E57651FFAC9501FDABFAD4D0A35E79" box="[629,719,1375,1400]" name="Mexico" pageId="25" pageNumber="176">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39E8654AFFAC9501FF19FA0FD3BF5C05" pageId="25" pageNumber="176" type="discussion">
|
||
<paragraph id="714D36C1FFAC9501FF19FA0FD3BF5C05" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||
<emphasis id="4386EAD3FFAC9501FF19FA0FD3575E9C" bold="true" box="[199,315,1412,1437]" pageId="25" pageNumber="176">Remarks.</emphasis>
|
||
<typeStatus id="AE498863FFAC9501FE9BFA0ED3105E9C" box="[325,380,1413,1437]" pageId="25" pageNumber="176">Type</typeStatus>
|
||
specimens assigned to this species are limited to those collected on
|
||
<taxonomicName id="B6F24D42FFAC9501FB7EFA0FD7125E9D" box="[1184,1406,1412,1436]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FB7EFA0FD7125E9D" box="[1184,1406,1412,1436]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
</taxonomicName>
|
||
, a species recently described by
|
||
<bibRefCitation id="15634B30FFAC9501FE32FA2CD0B25EC1" author="Vovides, A. P. & Stevenson, D. W. & Perez-Farrera, M. A. & Lopez, S. & Avendano, S." box="[492,734,1447,1472]" pageId="25" pageNumber="176" pagination="419 - 429" refId="ref20651" refString="Vovides, A. P., Stevenson, D. W., Perez-Farrera, M. A., Lopez, S. & Avendano, S. (2016) What is Ceratozamia mexicana (Zamiaceae)? Botanical Sciences, 94, 419 - 429. https: // doi. org / 10.17129 / botsci. 449" type="journal article" year="2016">
|
||
Vovides
|
||
<emphasis id="4386EAD3FFAC9501FD90FA22D0E45EC1" box="[590,648,1448,1472]" italics="true" pageId="25" pageNumber="176">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
. Prior to 2016 this host had been erroneously identified as
|
||
<taxonomicName id="B6F24D42FFAC9501FA5CFA23D30B5EE4" authority="Brongn." authorityName="Brongn." class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="mexicana">
|
||
<emphasis id="4386EAD3FFAC9501FA5CFA23D36E5EE5" italics="true" pageId="25" pageNumber="176">C. mexicana</emphasis>
|
||
Brongn.
|
||
</taxonomicName>
|
||
, a species now understood to occur only in the vicinity of El Mirador, further south in
|
||
<collectingRegion id="B336F823FFAC9501FAEFFA46D7F55EE5" box="[1329,1433,1485,1508]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||
. Besides
|
||
<taxonomicName id="B6F24D42FFAC9501FF2BFA7BD3BC5D06" box="[245,464,1520,1543]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FF2BFA7BD3BC5D06" box="[245,464,1520,1543]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
</taxonomicName>
|
||
, several other species of
|
||
<taxonomicName id="B6F24D42FFAC9501FD32FA7BD1115D06" box="[748,893,1520,1543]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4386EAD3FFAC9501FD32FA7BD1115D06" box="[748,893,1520,1543]" italics="true" pageId="25" pageNumber="176">Ceratozamia</emphasis>
|
||
</taxonomicName>
|
||
occur in the rugged terrain of western
|
||
<collectingRegion id="B336F823FFAC9501FAEFFA7AD7F45D09" box="[1329,1432,1521,1544]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||
, including
|
||
<taxonomicName id="B6F24D42FFAC9501FED6F99FD3F65D2D" box="[264,410,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="brevifrons">
|
||
<emphasis id="4386EAD3FFAC9501FED6F99FD3F65D2D" box="[264,410,1556,1580]" italics="true" pageId="25" pageNumber="176">C. brevifrons</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B6F24D42FFAC9501FE77F99FD05D5D2D" box="[425,561,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="delucana">
|
||
<emphasis id="4386EAD3FFAC9501FE77F99FD05D5D2D" box="[425,561,1556,1580]" italics="true" pageId="25" pageNumber="176">C. delucana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B6F24D42FFAC9501FDE1F99FD09D5D2D" box="[575,753,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="haustecorum">
|
||
<emphasis id="4386EAD3FFAC9501FDE1F99FD09D5D2D" box="[575,753,1556,1580]" italics="true" pageId="25" pageNumber="176">C. haustecorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B6F24D42FFAC9501FD21F99FD1E15D2D" box="[767,909,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="mexicana">
|
||
<emphasis id="4386EAD3FFAC9501FD21F99FD1E15D2D" box="[767,909,1556,1580]" italics="true" pageId="25" pageNumber="176">C. mexicana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B6F24D42FFAC9501FC1DF99FD6505D2D" box="[963,1084,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="morettii">
|
||
<emphasis id="4386EAD3FFAC9501FC1DF99FD6505D2D" box="[963,1084,1556,1580]" italics="true" pageId="25" pageNumber="176">C. morettii</emphasis>
|
||
</taxonomicName>
|
||
. The possible occurrence of
|
||
<taxonomicName id="B6F24D42FFAC9501FA5CF99FD3105D4E" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="vazquezi" subGenus="Vovidesa">
|
||
<emphasis id="4386EAD3FFAC9501FA5CF99FD7F95D2A" box="[1410,1429,1556,1579]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||
. (
|
||
<emphasis id="4386EAD3FFAC9501FF7EF9B3D36D5D51" box="[160,257,1592,1616]" italics="true" pageId="25" pageNumber="176">Vovidesa</emphasis>
|
||
)
|
||
<emphasis id="4386EAD3FFAC9501FEC2F9B3D3105D4E" box="[284,380,1592,1615]" italics="true" pageId="25" pageNumber="176">vazquezi</emphasis>
|
||
</taxonomicName>
|
||
in these species is currently being investigated by the authors. In one male cone of
|
||
<taxonomicName id="B6F24D42FFAC9501FF49F9D7D31B5D75" box="[151,375,1628,1652]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FF49F9D7D31B5D75" box="[151,375,1628,1652]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
</taxonomicName>
|
||
sampled at the end of its pollen shed near the town of Mafafas,
|
||
<taxonomicName id="B6F24D42FFAC9501FB4DF9D7D7F05D75" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1171,1436,1628,1652]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="vazquezi" subGenus="Vovidesa">
|
||
<emphasis id="4386EAD3FFAC9501FB4DF9D7D6CA5D72" box="[1171,1190,1628,1651]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||
. (
|
||
<emphasis id="4386EAD3FFAC9501FB1CF9D7D74F5D75" box="[1218,1315,1628,1652]" italics="true" pageId="25" pageNumber="176">Vovidesa</emphasis>
|
||
)
|
||
<emphasis id="4386EAD3FFAC9501FAE2F9D6D7F05D75" box="[1340,1436,1629,1652]" italics="true" pageId="25" pageNumber="176">vazquezi</emphasis>
|
||
</taxonomicName>
|
||
accounted for 86% of
|
||
<specimenCount id="67F4FD48FFAC9501FE72F90BD0705D99" box="[428,540,1663,1688]" pageId="25" pageNumber="176" type="adult">417 adult</specimenCount>
|
||
<taxonomicName id="B6F24D42FFAC9501FDF4F9F4D08A5D99" authorityName="Crowson" authorityYear="1952" box="[554,742,1663,1688]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||
collected, while an undescribed species of
|
||
<taxonomicName id="B6F24D42FFAC9501FB22F90BD7F05D99" authorityName="Reitter" authorityYear="1875" box="[1276,1436,1664,1688]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4386EAD3FFAC9501FB22F90BD7F05D99" box="[1276,1436,1664,1688]" italics="true" pageId="25" pageNumber="176">Pharaxonotha</emphasis>
|
||
</taxonomicName>
|
||
accounted for the remaining 14% of adult beetles in the cone. Adult
|
||
<taxonomicName id="B6F24D42FFAC9501FC7AF92FD6475DBD" box="[932,1067,1700,1724]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||
<emphasis id="4386EAD3FFAC9501FC7AF92FD6475DBD" box="[932,1067,1700,1724]" italics="true" pageId="25" pageNumber="176">Ceratophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="B6F24D42FFAC9501FBEAF92FD6B05DBD" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1076,1244,1700,1724]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="sanchezae" subGenus="Ceratophila">
|
||
(
|
||
<emphasis id="4386EAD3FFAC9501FBE3F92FD63C5DBA" box="[1085,1104,1700,1723]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||
.)
|
||
<emphasis id="4386EAD3FFAC9501FBB7F92FD6B05DBD" box="[1129,1244,1700,1724]" italics="true" pageId="25" pageNumber="176">sanchezae</emphasis>
|
||
</taxonomicName>
|
||
, which are often found with these other two
|
||
<taxonomicName id="B6F24D42FFAC9501FE11F94CD0E75DE1" authorityName="Crowson" authorityYear="1952" box="[463,651,1735,1760]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||
in cones of
|
||
<taxonomicName id="B6F24D42FFAC9501FCC7F943D1995DDE" box="[793,1013,1736,1759]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||
<emphasis id="4386EAD3FFAC9501FCC7F943D1995DDE" box="[793,1013,1736,1759]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||
</taxonomicName>
|
||
, were absent, but see Remarks under
|
||
<taxonomicName id="B6F24D42FFAC9501FF49F967D3735C05" box="[151,287,1772,1796]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||
<emphasis id="4386EAD3FFAC9501FF49F967D3735C05" box="[151,287,1772,1796]" italics="true" pageId="25" pageNumber="176">Ceratophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="B6F24D42FFAC9501FEFBF967D3A35C05" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[293,463,1772,1796]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="sanchezae" subGenus="Ceratophila">
|
||
(
|
||
<emphasis id="4386EAD3FFAC9501FEF0F967D32D5C02" box="[302,321,1772,1795]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||
.)
|
||
<emphasis id="4386EAD3FFAC9501FE86F967D3A35C05" box="[344,463,1772,1796]" italics="true" pageId="25" pageNumber="176">sanchezae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4386EAD3FFAC9501FE11F967D3BF5C05" box="[463,467,1772,1796]" italics="true" pageId="25" pageNumber="176">.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |