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<document id="49249C0055A3E43E82100D1F0728CBA1" ID="10.11646/zootaxa.4147.2.1" ID-DOI="10.11646/zootaxa.4147.2.1" ID-GBIF-Dataset="d7bc7cf8-9d00-462f-8995-f749fb7c09b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="254834" ID-ZooBank="1468C358-6478-46C0-A5DC-4716733244F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470204424906" checkinUser="plazi" docAuthor="Tshernyshev, Sergei E." docDate="2016" docId="F950FC3F5E24FFD2FF59B60C87C3FB0A" docLanguage="en" docName="zootaxa.4147.2.1.pdf" docOrigin="Zootaxa 4147 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Intybia guttata Pascoe 1866" docType="treatment" docVersion="7" lastPageNumber="105" masterDocId="056984475E26FFD6FFCEB2358636FFCF" masterDocTitle="Taxonomic revision of Intybia Pascoe, 1886 species (Coleoptera, Malachiidae) of Thailand and Philippines" masterLastPageNumber="123" masterPageNumber="101" pageNumber="103" updateTime="1698366798311" updateUser="plazi">
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<mods:title id="F62EA0383A4216A5376D133EAD2C496A">Taxonomic revision of Intybia Pascoe, 1886 species (Coleoptera, Malachiidae) of Thailand and Philippines</mods:title>
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<mods:date id="3C08447D18F2D165D4F0C1B6BD4C443A">2016</mods:date>
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<treatment id="F950FC3F5E24FFD2FF59B60C87C3FB0A" ID-DOI="http://doi.org/10.5281/zenodo.6086348" ID-GBIF-Taxon="123514047" ID-Zenodo-Dep="6086348" LSID="urn:lsid:plazi:treatment:F950FC3F5E24FFD2FF59B60C87C3FB0A" httpUri="http://treatment.plazi.org/id/F950FC3F5E24FFD2FF59B60C87C3FB0A" lastPageId="4" lastPageNumber="105" pageId="2" pageNumber="103">
<subSubSection id="39E31EA25E24FFD4FF59B60C873CFBBA" pageId="2" pageNumber="103" type="nomenclature">
<paragraph id="71464D295E24FFD4FF59B60C87D8FB9C" blockId="2.[151,494,1081,1141]" box="[151,494,1081,1107]" pageId="2" pageNumber="103">
<heading id="2A0EFA455E24FFD4FF59B60C87D8FB9C" bold="true" box="[151,494,1081,1107]" fontSize="11" level="1" pageId="2" pageNumber="103" reason="1">
<taxonomicName id="B6F936AA5E24FFD4FF59B60C87D8FB9C" authority="Pascoe, 1866" authorityName="Pascoe" authorityYear="1866" box="[151,494,1081,1107]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="438D913B5E24FFD4FF59B60C87D8FB9C" bold="true" box="[151,494,1081,1107]" pageId="2" pageNumber="103">
<emphasis id="438D913B5E24FFD4FF59B60C877FFB9C" bold="true" box="[151,329,1081,1107]" italics="true" pageId="2" pageNumber="103">Intybia guttata</emphasis>
<bibRefCitation id="156830D85E24FFD4FE9EB60C87D8FB9C" author="Pascoe" box="[336,494,1081,1107]" pageId="2" pageNumber="103" refString="Pascoe, F. P. (1866) Notices of new or little-known genera and species of Coleoptera. Journal of Entomology, 2 (14), 447 - 493, plates 18 - 19." type="journal article" year="1866">Pascoe, 1866</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="71464D295E24FFD4FF59B669873CFBBA" blockId="2.[151,494,1081,1141]" box="[151,266,1116,1141]" pageId="2" pageNumber="103">
(
<figureCitation id="E9C251AC5E24FFD4FF51B6698737FBBA" box="[159,257,1116,1141]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[162,1420,280,1889]" captionTargetId="figure@3.[150,1435,265,1901]" captionTargetPageId="3" captionText="FIGURES 1 5. Intybia guttata Pascoe, 1866, holotype, female, dorsal view (1); lateral view (2); head (3); original labels (4); distribution map (5). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/254835/files/figure.png" pageId="2" pageNumber="103">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39E31EA25E24FFD4FF59B69185E7FB73" box="[151,977,1188,1213]" pageId="2" pageNumber="103" type="materials_examined">
<paragraph id="71464D295E24FFD4FF59B69185E7FB73" blockId="2.[151,1437,1188,2007]" box="[151,977,1188,1213]" pageId="2" pageNumber="103">
<materialsCitation id="C19147745E24FFD4FF59B69185E7FB73" ID-GBIF-Occurrence="1290985015" box="[151,977,1188,1213]" country="Indonesia" location="Bacan Islands" pageId="2" pageNumber="103" specimenCount="1" typeStatus="holotype">
<emphasis id="438D913B5E24FFD4FF59B69187B6FB72" bold="true" box="[151,384,1188,1213]" pageId="2" pageNumber="103">Material examined.</emphasis>
<typeStatus id="AE42F38B5E24FFD4FE49B69187C4FB72" box="[391,498,1188,1213]" pageId="2" pageNumber="103" type="holotype">Holotype</typeStatus>
, female,
<collectingCountry id="09EE0DB95E24FFD4FD99B69184F1FB72" box="[599,711,1188,1213]" name="Indonesia" pageId="2" pageNumber="103">Indonesia</collectingCountry>
,
<location id="74261BF25E24FFD4FD1CB6908559FB72" LSID="urn:lsid:plazi:treatment:F950FC3F5E24FFD2FF59B60C87C3FB0A:74261BF25E24FFD4FD1CB6908559FB72" box="[722,879,1188,1213]" country="Indonesia" name="Bacan Islands" pageId="2" pageNumber="103">Bacan Islands</location>
(
<figureCitation id="E9C251AC5E24FFD4FCB0B69185F5FB73" box="[894,963,1188,1213]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[162,1420,280,1889]" captionTargetId="figure@3.[150,1435,265,1901]" captionTargetPageId="3" captionText="FIGURES 1 5. Intybia guttata Pascoe, 1866, holotype, female, dorsal view (1); lateral view (2); head (3); original labels (4); distribution map (5). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/254835/files/figure.png" pageId="2" pageNumber="103">Fig. 4</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="39E31EA25E24FFD2FF09B6F284ECFD4B" lastPageId="4" lastPageNumber="105" pageId="2" pageNumber="103" type="description">
<paragraph id="71464D295E24FFD4FF09B6F28356FB2F" blockId="2.[151,1437,1188,2007]" box="[199,1376,1223,1248]" pageId="2" pageNumber="103">
<emphasis id="438D913B5E24FFD4FF09B6F28767FB2F" bold="true" box="[199,337,1223,1248]" pageId="2" pageNumber="103">Description</emphasis>
. Female (
<figureCitation id="E9C251AC5E24FFD4FE0FB6F28410FB2F" box="[449,550,1223,1248]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[162,1420,280,1889]" captionTargetId="figure@3.[150,1435,265,1901]" captionTargetPageId="3" captionText="FIGURES 1 5. Intybia guttata Pascoe, 1866, holotype, female, dorsal view (1); lateral view (2); head (3); original labels (4); distribution map (5). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/254835/files/figure.png" pageId="2" pageNumber="103">Figs 1, 2</figureCitation>
). Body slightly elongate, ovoid, expanded posteriorly just behind humeri.
</paragraph>
<paragraph id="71464D295E24FFD4FF09B6D986D0FA12" blockId="2.[151,1437,1188,2007]" pageId="2" pageNumber="103">Body dark brown, except for yellow tarsi and lateral sides of 1st7th antennal segments and white spots in elytral disc. Each elytron with three white spots, one, wide, oval, transverse, in basal half above the middle, not reaching suture or external side but disposed closer to outer side; two spots in apical half below the middle, one, triangular, near suture, second, round near external margin, both spots disposed separate from margins and each other. Underside of body dark brown; legs dark brown with yellow tarsi except for claw segments. Surface evenly covered with short semi-erect white pubescence and sparse erect hairs. Vesicles yellow, thoracic mesepimera brown.</paragraph>
<paragraph id="71464D295E24FFD4FF09B7D2820DF97A" blockId="2.[151,1437,1188,2007]" pageId="2" pageNumber="103">
Head (
<figureCitation id="E9C251AC5E24FFD4FEDAB7D2876FF9CF" box="[276,345,1511,1536]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[162,1420,280,1889]" captionTargetId="figure@3.[150,1435,265,1901]" captionTargetPageId="3" captionText="FIGURES 1 5. Intybia guttata Pascoe, 1866, holotype, female, dorsal view (1); lateral view (2); head (3); original labels (4); distribution map (5). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/254835/files/figure.png" pageId="2" pageNumber="103">Fig. 3</figureCitation>
) slightly wider than pronotum, not elongated, eyes small, round, protruding, but not stretched, frons flat; interocular area with distinct thin longitudinal carina in the middle, genae impressed and oblique; clypeus extremely narrow due to very closely settled antennae, not elongate, transverse, straight; labrum not elongated, transverse; palpi simple with apical segment slightly enlarged, subcylindrical, narrowed posteriorly and cut at the tip, intermediate segments subtriangular, short and transverse; surface of head densely and coarsely punctured, with weak microsculpture, light pubescence dense, short and adpressed.
</paragraph>
<paragraph id="71464D295E24FFD4FF09B4F5870CF840" blockId="2.[151,1437,1188,2007]" pageId="2" pageNumber="103">
Antennae filiform (
<figureCitation id="E9C251AC5E24FFD4FE6FB48A87D0F917" box="[417,486,1727,1752]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[162,1420,280,1889]" captionTargetId="figure@3.[150,1435,265,1901]" captionTargetPageId="3" captionText="FIGURES 1 5. Intybia guttata Pascoe, 1866, holotype, female, dorsal view (1); lateral view (2); head (3); original labels (4); distribution map (5). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/254835/files/figure.png" pageId="2" pageNumber="103">Fig. 2</figureCitation>
),
<quantity id="B601E0CC5E24FFD4FE32B4F58465F918" box="[508,595,1728,1752]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="103" unit="mm" value="1.4">1.4 mm</quantity>
long, reaching the basal quarter of the elytra; segment 1 slightly enlarged, subtriangular, slightly angulate at the outer apical angle, segment 2 small, round-oval, 3rd segment elongated, enlarged and terete; 4th segment short and cylindrical, similar in length to 5th or 6th segment, 711th segments elongated and cylindrical, apical segment somewhat longer than previous, evenly narrowed and pointed at apex; surface evenly covered with short, light pubescence and sparse, semi-erect long light hairs on outer sides of antennomeres.
</paragraph>
<paragraph id="71464D295E24FFD5FF09B5AE8224FF1A" blockId="2.[151,1437,1188,2007]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103">Pronotum not elongated, transversal, somewhat barrel-shaped with not marginate lateral sides, each with small smoothed denticle in middle; slightly narrowed from middle to base, anterior and posterior sides marginate and straight, disc convex anteriorly and narrowly depressed at base; surface dull, densely punctured with distinct microsculpture, surface densely covered with fine, short, white adpressed hairs.</paragraph>
<caption id="25861DA15E25FFD5FF59B5B68410F877" httpUri="https://zenodo.org/record/254835/files/figure.png" pageId="3" pageNumber="104" targetBox="[162,1420,280,1889]" targetPageId="3">
<paragraph id="71464D295E25FFD5FF59B5B68410F877" blockId="3.[151,1436,1923,1976]" pageId="3" pageNumber="104">
<emphasis id="438D913B5E25FFD5FF59B5B6870CF857" bold="true" box="[151,314,1923,1946]" pageId="3" pageNumber="104">FIGURES 15.</emphasis>
<taxonomicName id="B6F936AA5E25FFD5FE8CB5B68451F856" authority="Pascoe, 1866" authorityName="Pascoe" authorityYear="1866" box="[322,615,1923,1945]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="438D913B5E25FFD5FE8CB5B687BCF856" box="[322,394,1923,1945]" italics="true" pageId="3" pageNumber="104">Intybia</emphasis>
<emphasis id="438D913B5E25FFD5FE5EB5B187EEF856" box="[400,472,1924,1945]" italics="true" pageId="3" pageNumber="104">guttata</emphasis>
Pascoe, 1866
</taxonomicName>
, holotype, female, dorsal view (1); lateral view (2); head (3); original labels (4); distribution map (5). Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="71464D295E22FFD2FF09B2A285E7FF1A" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="105">Scutellum small, narrow, rectangular, transversal, with straight apical margin, almost completely covered by pronotum, sparsely punctured, shiny, sparsely pubescent with white hairs.</paragraph>
<paragraph id="71464D295E22FFD2FF09B2EA832FFEAA" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="105">Elytra oval, distinctly widened just behind the humeri, and evenly rounded posteriorly, at base slightly wider than pronotum; humeri distinct, protruding; suture slightly marginate and elevate, more distinctly at basal quarter, which is impressed, and posterior part convex; surface dull, with dense and coarse punctures and distinct microsculpture, evenly covered with white double pubescence—short adpressed and strong erect hairs.</paragraph>
<paragraph id="71464D295E22FFD2FF09B35A87F6FE47" blockId="4.[151,1437,151,1221]" box="[199,448,367,392]" pageId="4" pageNumber="105">Hind wings indistinct.</paragraph>
<paragraph id="71464D295E22FFD2FF09B3A0875FFDD7" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="105">Legs thin; posterior femora not reaching elytral apices; all tibiae thin, straight, femora narrow, slightly compressed, not curved; all tarsi 5-segmented, 1st4th segments short and compressed, claw segment depressed and largest, wide in all legs and equal to 1st3rd segments; claws very small, thin, narrow and sharp, with small membrane at base.</paragraph>
<paragraph id="71464D295E22FFD2FF09B0118527FDAF" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="105">Ventral body surface densely punctured, dull, evenly covered with sparse fine, depressed dark pubescence. Metathorax slightly swollen, simple, lacking appendage.</paragraph>
<paragraph id="71464D295E22FFD2FF09B05984ECFD4B" blockId="4.[151,1437,151,1221]" box="[199,730,620,645]" pageId="4" pageNumber="105">
Length
<quantity id="B601E0CC5E22FFD2FED3B0598743FD4B" box="[285,373,620,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="105" unit="mm" value="2.5">2.5 mm</quantity>
, width (at elytral base)
<quantity id="B601E0CC5E22FFD2FDB3B05984E1FD4B" box="[637,727,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="105" unit="mm" value="0.7">0.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="39E31EA25E22FFD2FF09B0BA837AFD67" box="[199,1356,655,680]" pageId="4" pageNumber="105" type="distribution">
<paragraph id="71464D295E22FFD2FF09B0BA837AFD67" blockId="4.[151,1437,151,1221]" box="[199,1356,655,680]" pageId="4" pageNumber="105">
<emphasis id="438D913B5E22FFD2FF09B0BA8756FD67" bold="true" box="[199,352,655,680]" pageId="4" pageNumber="105">Distribution.</emphasis>
The species remains known only from its
<typeStatus id="AE42F38B5E22FFD2FCF2B0A5855AFD67" box="[828,876,656,680]" pageId="4" pageNumber="105">type</typeStatus>
locality, Bacan Islands,
<collectingCountry id="09EE0DB95E22FFD2FBB0B0BA82DDFD67" box="[1150,1259,655,680]" name="Indonesia" pageId="4" pageNumber="105">Indonesia</collectingCountry>
(
<figureCitation id="E9C251AC5E22FFD2FB34B0BA8308FD67" box="[1274,1342,655,680]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1923,1946]" captionTargetBox="[162,1420,280,1889]" captionTargetId="figure@3.[150,1435,265,1901]" captionTargetPageId="3" captionText="FIGURES 1 5. Intybia guttata Pascoe, 1866, holotype, female, dorsal view (1); lateral view (2); head (3); original labels (4); distribution map (5). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/254835/files/figure.png" pageId="4" pageNumber="105">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39E31EA25E22FFD2FF09B08187C3FB0A" pageId="4" pageNumber="105" type="discussion">
<paragraph id="71464D295E22FFD2FF09B08187C3FB0A" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="105">
<emphasis id="438D913B5E22FFD2FF09B0818779FD02" bold="true" box="[199,335,692,717]" pageId="4" pageNumber="105">Comments.</emphasis>
According to Zoological Nomenclature, characters of
<taxonomicName id="B6F936AA5E22FFD2FC7AB0808246FD03" authority="Pascoe" authorityName="Pascoe" box="[948,1136,693,716]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="438D913B5E22FFD2FC7AB080822DFD03" box="[948,1051,693,716]" italics="true" pageId="4" pageNumber="105">I. guttata</emphasis>
Pascoe
</taxonomicName>
defining the genus
<taxonomicName id="B6F936AA5E22FFD2FA82B08186ECFD20" authority="Pacoe" authorityName="Pacoe" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="438D913B5E22FFD2FA82B08183ADFD02" box="[1356,1435,692,717]" italics="true" pageId="4" pageNumber="105">Intybia</emphasis>
Pacoe
</taxonomicName>
should be precisely given. Female
<typeStatus id="AE42F38B5E22FFD2FDBDB0E284E3FD3F" box="[627,725,727,752]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
of the species complicates this problem because no one male character traditionally used for generic identification. Thus, it is expected that the male of
<taxonomicName id="B6F936AA5E22FFD2FB42B0C8837EFCDB" authority="Pascoe" authorityName="Pascoe" box="[1164,1352,765,788]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="438D913B5E22FFD2FB42B0C882C5FCDB" box="[1164,1267,765,788]" italics="true" pageId="4" pageNumber="105">I. guttata</emphasis>
Pascoe
</taxonomicName>
lacks a tarsal comb in anterior legs, and therefore cannot be considered as
<taxonomicName id="B6F936AA5E22FFD2FC57B12A829DFCF7" authority="Champion" authorityName="Champion" box="[921,1195,799,824]" class="Insecta" family="Melyridae" genus="Dicranolaius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="438D913B5E22FFD2FC57B12A821BFCF7" box="[921,1069,799,824]" italics="true" pageId="4" pageNumber="105">Dicranolaius</emphasis>
Champion
</taxonomicName>
species. Complex of characters, such as not elongate head and simple, not stretched eyes allow one to separate this species from the genus
<emphasis id="438D913B5E22FFD2FF2BB152874BFC4F" box="[229,381,871,896]" italics="true" pageId="4" pageNumber="105">Troglointybia</emphasis>
Tshernyshev. Thus,
<taxonomicName id="B6F936AA5E22FFD2FDB9B15C8509FCB0" authority="Pascoe" authorityName="Pascoe" box="[631,831,872,896]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="438D913B5E22FFD2FDB9B15C84D5FC4F" box="[631,739,873,896]" italics="true" pageId="4" pageNumber="105">I. guttata</emphasis>
Pascoe
</taxonomicName>
belongs to a group of species with the following characters: head elongate in male and/or in female; eyes round, simple, protruding, not stretched; palpae with enlarged apical segment and small triangular intermediate ones; genae impressed and oblique for clear placement of enlarged 1st antennomere; interocular area of head flat, lacking sculpture but possessing longitudinal thin carina in the middle; pronotum and elytra depressed at base and convex at distal parts; tarsi short, 1st4th segment compressed, claw segment depressed, wide, equal in length to 1st3rd segments. This complex of characters is untypical for all representatives of
<taxonomicName id="B6F936AA5E22FFD2FDE2B60A844DFB97" box="[556,635,1087,1112]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="438D913B5E22FFD2FDE2B60A844DFB97" box="[556,635,1087,1112]" italics="true" pageId="4" pageNumber="105">Intybia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE2C405E22FFD2FD4AB67584CCFB97" box="[644,762,1087,1112]" pageId="4" pageNumber="105" sensu="lato">sensu lato</taxonomicNameLabel>
, and good, for example, for
<taxonomicName id="B6F936AA5E22FFD2FB88B67486EEFBB3" authority="Champion, 1921" authorityName="Champion" authorityYear="1921" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="quadristrigatus" status="comb. nov.">
<emphasis id="438D913B5E22FFD2FB88B6748338FB97" box="[1094,1294,1087,1112]" italics="true" pageId="4" pageNumber="105">I. quadristrigatus</emphasis>
(Champion, 1921)
</taxonomicName>
<emphasis id="438D913B5E22FFD2FF2FB6518776FBB2" bold="true" box="[225,320,1124,1149]" pageId="4" pageNumber="105">
<taxonomicNameLabel id="58BE2C405E22FFD2FF2FB6518776FBB2" box="[225,320,1124,1149]" pageId="4" pageNumber="105">comb.n.</taxonomicNameLabel>
</emphasis>
or
<taxonomicName id="B6F936AA5E22FFD2FEA4B6508483FBB3" authority="Champion, 1921" authorityName="Champion" authorityYear="1921" box="[362,693,1124,1149]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="semperi">
<emphasis id="438D913B5E22FFD2FEA4B65087EBFBB3" box="[362,477,1125,1148]" italics="true" pageId="4" pageNumber="105">I. semperi</emphasis>
(Champion, 1921)
</taxonomicName>
discussed below, but reflects the main difference in simple legs, lacking tarsal comb and unsculptured head. More detail results could be gained after investigation of male if it will be found in Indonesian islands.
</paragraph>
</subSubSection>
</treatment>
</document>