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<document id="5BFA1E0C19AD2A6FB7A099AFD8F4A5D4" ID="10.11646/zootaxa.4170.1.4" ID-DOI="10.11646/zootaxa.4170.1.4" ID-GBIF-Dataset="991162cc-fc47-41dc-b23d-cef3d657efff" ID-ISSN="1175-5326" ID-Zenodo-Dep="254807" ID-ZooBank="F2E2A4C2-0AE0-4E55-916E-D6E1D60060E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1474526355058" checkinUser="plazi" docAuthor="Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai &amp; Panha, Somsak" docDate="2016" docId="F92BAE363418FFFBFF581457E926F87D" docLanguage="en" docName="zootaxa.4170.1.4.pdf" docOrigin="Zootaxa 4170 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Desmoxytes pinnasquali Srisonchai, Enghoff &amp; Panha, sp. n." docType="treatment" docVersion="8" lastPageNumber="111" masterDocId="0512D64E3416FFE9FFCF1730ED1AFFCB" masterDocTitle="Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae)" masterLastPageNumber="113" masterPageNumber="93" originalUpdateDomain="Boston" originalUpdateTime="1545144304724" originalUpdateUser="gglab" pageNumber="107" updateTime="1698386812159" updateUser="plazi">
<mods:mods id="172743A3D4FB92B9DD4E09A3520FF22B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="48A4E11EBEE4E414AE51CD9DC1C3EB83">
<mods:title id="E0702DF2383E447840D2F0B40DD6B3CB">Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="82686165755F777F20CB4E04781D9380" type="personal">
<mods:role id="434CADFFBB98FA0A50F396B7CF947A9D">
<mods:roleTerm id="7A3ECC0FB72BD43B0D6044BDD6A74257">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F7776811202C53ABA2A9E770199C948">Srisonchai, Ruttapon</mods:namePart>
</mods:name>
<mods:name id="CE21B16E72B49A9899B167FE6338D5DE" type="personal">
<mods:role id="049B6E23B2A03B3FFF5C1EBB79418545">
<mods:roleTerm id="ABA2247E692A2C42B4F2A4848D29EF1A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="820A45EDE1601C0431DF0F7089259096">Enghoff, Henrik</mods:namePart>
</mods:name>
<mods:name id="481702692E974E7B1504EC0674AA040B" type="personal">
<mods:role id="AA99B42D5BC89B930AE59F9C46C583E1">
<mods:roleTerm id="FB5A6B16A0AF0EF5EB5FC0E65FFDBA5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9897CE11F041EA6632B2D19C8A169F7">Likhitrakarn, Natdanai</mods:namePart>
</mods:name>
<mods:name id="CA1FB3B4E94CDC5AEED8582C43B18BF9" type="personal">
<mods:role id="0584B0ACFC177B5C460CD70961CC9DB3">
<mods:roleTerm id="6CD75C5561A5036F97316AC202C8F67C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AEAFCB4DEF510F0F1CD9E4191FE92B46">Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource id="2031D17CEB3701E0A7840EF44BD3B692">text</mods:typeOfResource>
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<mods:titleInfo id="CC2402C3935E5FA9648033363807D038">
<mods:title id="428916E12CED140CF7E23FD213B422DB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4E9A177096A8002F137A932D114164DC">
<mods:date id="3AA3962BEC1864C8038A42713E030AF4">2016</mods:date>
<mods:detail id="F6A2CA2118DA1BF29F4E3FF6E3FE9863" type="volume">
<mods:number id="D472027C80463B0F6303B6F513BE3365">4170</mods:number>
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<mods:number id="544BFD010BCCA9FBE184B1331D79CEFF">1</mods:number>
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<mods:start id="0D0B1699042282E9835B29149DF688D5">93</mods:start>
<mods:end id="07CBCF409AB5961A4DA6CE83D5B446E8">113</mods:end>
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<mods:classification id="17B986E0ABBE1B49079567316D50193D">journal article</mods:classification>
<mods:identifier id="F0C1CAFF225595B6BE0514E2F6637447" type="DOI">10.11646/zootaxa.4170.1.4</mods:identifier>
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<mods:identifier id="55FDD3BA12233478227E32F0D51CD051" type="Zenodo-Dep">254807</mods:identifier>
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<treatment id="F92BAE363418FFFBFF581457E926F87D" ID-DOI="http://doi.org/10.5281/zenodo.6059213" ID-GBIF-Taxon="124552648" ID-Zenodo-Dep="6059213" LSID="urn:lsid:plazi:treatment:F92BAE363418FFFBFF581457E926F87D" httpUri="http://treatment.plazi.org/id/F92BAE363418FFFBFF581457E926F87D" lastPageId="18" lastPageNumber="111" pageId="14" pageNumber="107">
<subSubSection id="39984CAB3418FFE7FF581457EC93FC68" pageId="14" pageNumber="107" type="nomenclature">
<paragraph id="713D1F203418FFE7FF581457EE66FC49" blockId="14.[151,892,871,931]" box="[151,892,871,898]" pageId="14" pageNumber="107">
<heading id="2A75A84C3418FFE7FF581457EE66FC49" bold="true" box="[151,892,871,898]" fontSize="11" level="1" pageId="14" pageNumber="107" reason="1">
<emphasis id="43F6C3323418FFE7FF581457EE66FC49" bold="true" box="[151,892,871,898]" pageId="14" pageNumber="107">
<taxonomicName id="B68264A33418FFE7FF581457EE29FC49" ID-CoL="6CNMD" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" box="[151,819,871,898]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="107" phylum="Arthropoda" rank="species" species="pinnasquali" status="sp. nov.">
<emphasis id="43F6C3323418FFE7FF581457ECA6FC4A" bold="true" box="[151,444,871,897]" italics="true" pageId="14" pageNumber="107">Desmoxytes pinnasquali</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="58C57E493418FFE7FCF51458EE66FC49" box="[826,892,872,898]" pageId="14" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="713D1F203418FFE7FF5814BAEC93FC68" blockId="14.[151,892,871,931]" box="[151,393,906,931]" pageId="14" pageNumber="107">
(
<figureCitation id="E9B903A53418FFE7FF5014BAEDF2FC68" box="[159,232,906,931]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[161,1424,565,1778]" captionTargetId="figure@2.[151,1436,553,1790]" captionTargetPageId="2" captionText="FIGURE 1. Live photographs of the new species. A, B: Desmoxytes des sp. n. — A; male holotype, B; female. C, D: Desmoxytes breviverpa sp. n. — male paratype. E, F: Desmoxytes takensis sp. n. — E; male paratype, F; female. G, H: Desmoxytes pinnasquali sp. n. — G: male holotype, H: mating couple." httpUri="https://zenodo.org/record/254808/files/figure.png" pageId="14" pageNumber="107">Figs. 1</figureCitation>
G, 1H, 1113)
</paragraph>
</subSubSection>
<subSubSection id="39984CAB3418FFE7FF5814E2EEF4FB2C" pageId="14" pageNumber="107" type="materials_examined">
<paragraph id="713D1F203418FFE7FF5814E2EF10FBF8" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
<materialsCitation id="C1EA157D3418FFE7FF5814E2EF10FBF8" ID-GBIF-Occurrence="1314597141" collectingDate="2014-08-22" collectionCode="CUMZ" collectorName="S. Panha &amp; C. Sutcharit &amp; U. Bantaowong" country="Thailand" elevation="62" latitude="16.694443" location="Noen Maprang District, near Pra Tham Mans Monastery (Tham Wangdaeng)" longitude="100.67833" pageId="14" pageNumber="107" specimenCount="1" specimenCount-male="1" stateProvince="Phitsanulok" typeStatus="holotype">
<emphasis id="43F6C3323418FFE7FF5814E2EC11FC20" bold="true" box="[151,267,978,1003]" pageId="14" pageNumber="107">
<typeStatus id="AE39A1823418FFE7FF5814E2EC1CFC20" box="[151,262,978,1003]" pageId="14" pageNumber="107" type="holotype">Holotype</typeStatus>
.
</emphasis>
male (
<collectionCode id="179387E53418FFE7FEAC14E3ECADFC20" box="[355,439,979,1003]" country="Thailand" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="14" pageNumber="107">CUMZ</collectionCode>
)
<collectingCountry id="09955FB03418FFE7FE0214E3EF7BFC20" box="[461,609,979,1003]" name="Thailand" pageId="14" pageNumber="107">THAILAND</collectingCountry>
,
<collectingRegion id="B346D1C23418FFE7FDBE14E2EE70FC20" box="[625,874,978,1003]" country="Thailand" name="Phitsanulok" pageId="14" pageNumber="107">Phitsanulok Province</collectingRegion>
,
<collectingMunicipality id="9159855A3418FFE7FCB414E3E994FC20" box="[891,1166,978,1003]" pageId="14" pageNumber="107">Noen Maprang District</collectingMunicipality>
, near
<location id="745D49FB3418FFE7FB1214E3EC15FBC4" LSID="urn:lsid:plazi:treatment:F92BAE363418FFFBFF581457E926F87D:745D49FB3418FFE7FB1214E3EC15FBC4" country="Thailand" latitude="16.694443" longLatPrecision="21" longitude="100.67833" municipality="Noen Maprang District" name="Pra Tham Mans Monastery" pageId="14" pageNumber="107" stateProvince="Phitsanulok Province">Pra Tham Mans Monastery</location>
(Tham Wangdaeng),
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,
<geoCoordinate id="14B679E73418FFE7FD7D14C7EE5EFBC4" box="[690,836,1015,1039]" direction="east" orientation="longitude" pageId="14" pageNumber="107" precision="15" value="100.67833">100°40'42&quot;E</geoCoordinate>
, ca
<elevation id="FAAFF8133418FFE7FCB014C6E91FFBC4" box="[895,1029,1014,1039]" metricMagnitude="1" metricUnit="m" metricValue="6.2" pageId="14" pageNumber="107" unit="m" value="62.0">
<quantity id="B67AB2C53418FFE7FCB014C6EEDAFBC5" box="[895,960,1014,1039]" metricMagnitude="1" metricUnit="m" metricValue="6.2" pageId="14" pageNumber="107" unit="m" value="62.0">62 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="1578C0083418FFE7FBD614C7E9C5FBC4" box="[1049,1247,1015,1039]" pageId="14" pageNumber="107" value="2014-08-22">
<date id="053C39E03418FFE7FBD614C7E9C5FBC4" box="[1049,1247,1015,1039]" pageId="14" pageNumber="107" value="2014-08-22">22 August 2014</date>
</collectingDate>
, leg.
<collectorName id="DC777AF63418FFE7FAE414C7E882FBC4" box="[1323,1432,1014,1039]" pageId="14" pageNumber="107">S. Panha</collectorName>
,
<collectorName id="DC777AF63418FFE7FF58132BEC06FBF8" box="[151,284,1050,1075]" pageId="14" pageNumber="107">C. Sutcharit</collectorName>
and
<collectorName id="DC777AF63418FFE7FE9C132BEF10FBF8" box="[339,522,1051,1075]" pageId="14" pageNumber="107">U. Bantaowong.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="713D1F203418FFE7FF08130EEEF4FB2C" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
<emphasis id="43F6C3323418FFE7FF08130EEC5CFB9C" bold="true" box="[199,326,1086,1111]" pageId="14" pageNumber="107">Paratypes.</emphasis>
<materialsCitation id="C1EA157D3418FFE7FE81130FEFB8FB9C" ID-GBIF-Occurrence="1949262818" box="[334,674,1086,1111]" collectingDate="2014-08-22" collectionCode="CUMZ" collectorName="S. Panha &amp; C. Sutcharit &amp; U. Bantaowong" country="Thailand" elevation="62" latitude="16.694443" location="Noen Maprang District, near Pra Tham Mans Monastery (Tham Wangdaeng)" longitude="100.67833" pageId="14" pageNumber="107" specimenCount="27" specimenCount-female="5" specimenCount-male="22" stateProvince="Phitsanulok" typeStatus="paratype">
<specimenCount id="6784D4A93418FFE7FE81130FECACFB9C" box="[334,438,1086,1111]" pageId="14" pageNumber="107" type="male">22 males</specimenCount>
,
<specimenCount id="6784D4A93418FFE7FE0D130EEF35FB9C" box="[450,559,1086,1111]" pageId="14" pageNumber="107" type="female">5 females</specimenCount>
(
<collectionCode id="179387E53418FFE7FD8F130FEF8DFB9C" box="[576,663,1087,1111]" country="Thailand" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="14" pageNumber="107">CUMZ</collectionCode>
),
</materialsCitation>
<materialsCitation id="C1EA157D3418FFE7FD65130FE9E5FB9C" ID-GBIF-Occurrence="1949262815" box="[682,1279,1086,1111]" collectingDate="2014-08-22" collectionCode="ZMUC" collectorName="S. Panha &amp; C. Sutcharit &amp; U. Bantaowong" country="Thailand" elevation="62" latitude="16.694443" location="Noen Maprang District, near Pra Tham Mans Monastery (Tham Wangdaeng)" longitude="100.67833" pageId="14" pageNumber="107" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Phitsanulok" typeStatus="holotype">
<specimenCount id="6784D4A93418FFE7FD65130FEE1EFB9C" box="[682,772,1086,1111]" pageId="14" pageNumber="107" type="male">2 males</specimenCount>
,
<specimenCount id="6784D4A93418FFE7FCDE130FEE69FB9C" box="[785,883,1086,1111]" pageId="14" pageNumber="107" type="female">1 female</specimenCount>
(
<collectionCode id="179387E53418FFE7FC4C130FEECDFB9C" box="[899,983,1087,1111]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="107">ZMUC</collectionCode>
) same data with
<typeStatus id="AE39A1823418FFE7FB56130EE9E5FB9C" box="[1177,1279,1086,1111]" pageId="14" pageNumber="107" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="C1EA157D3418FFE7FAC4130FE9ABFBB0" ID-GBIF-Occurrence="1314597136" collectingDate="2008-07-23" collectionCode="CUMZ" collectorName="N. Likhitrakarn" country="Thailand" elevation="62" latitude="16.694443" location="Noen Maprang District, near Pra Tham Mans Monastery (Tham Wangdaeng)" longitude="100.67833" pageId="14" pageNumber="107" specimenCount="27" specimenCount-female="17" specimenCount-male="10" stateProvince="Phitsanulok" typeStatus="holotype">
<specimenCount id="6784D4A93418FFE7FAC4130FE86EFB9C" box="[1291,1396,1086,1111]" pageId="14" pageNumber="107" type="male">10 males</specimenCount>
,
<specimenCount id="6784D4A93418FFE7FA4F130FEDF4FBB0" pageId="14" pageNumber="107" type="female">17 females</specimenCount>
(
<collectionCode id="179387E53418FFE7FECF1353EC4EFBB0" box="[256,340,1123,1147]" country="Thailand" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="14" pageNumber="107">CUMZ</collectionCode>
) same locality together with
<typeStatus id="AE39A1823418FFE7FD641352EE0AFBB0" box="[683,784,1122,1147]" pageId="14" pageNumber="107" type="holotype">holotype</typeStatus>
,
<collectingDate id="1578C0083418FFE7FCD11353EEA3FBB0" box="[798,953,1122,1147]" pageId="14" pageNumber="107" value="2008-07-23">
<date id="053C39E03418FFE7FCD11353EEA3FBB0" box="[798,953,1122,1147]" pageId="14" pageNumber="107" value="2008-07-23">23 July 2008</date>
</collectingDate>
, leg.
<collectorName id="DC777AF63418FFE7FC341353E9ABFBB0" box="[1019,1201,1122,1147]" pageId="14" pageNumber="107">N. Likhitrakarn</collectorName>
</materialsCitation>
;
<materialsCitation id="C1EA157D3418FFE7FB701353E9D8FB54" ID-GBIF-Occurrence="1314597128" collectingDate="2009-09-08" collectionCode="CUMZ" collectorName="U. Bantaowong &amp; R. Chanabun" country="Thailand" elevation="62" latitude="16.694443" location="Noen Maprang District, near Pra Tham Mans Monastery (Tham Wangdaeng)" longitude="100.67833" pageId="14" pageNumber="107" specimenCount="2" specimenCount-female="2" stateProvince="Phitsanulok" typeStatus="holotype">
<specimenCount id="6784D4A93418FFE7FB701353E834FBB0" box="[1215,1326,1122,1147]" pageId="14" pageNumber="107" type="female">2 females</specimenCount>
(
<collectionCode id="179387E53418FFE7FA8F1353E88EFBB0" box="[1344,1428,1123,1147]" country="Thailand" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="14" pageNumber="107">CUMZ</collectionCode>
) same locality together with
<typeStatus id="AE39A1823418FFE7FE0213B6EF28FB54" box="[461,562,1158,1183]" pageId="14" pageNumber="107" type="holotype">holotype</typeStatus>
,
<collectingDate id="1578C0083418FFE7FDF213B7EE11FB54" box="[573,779,1158,1183]" pageId="14" pageNumber="107" value="2009-09-08">
<date id="053C39E03418FFE7FDF213B7EE11FB54" box="[573,779,1158,1183]" pageId="14" pageNumber="107" value="2009-09-08">8 September 2009</date>
</collectingDate>
, leg.
<collectorName id="DC777AF63418FFE7FC8913B7EEEFFB54" box="[838,1013,1159,1183]" pageId="14" pageNumber="107">U. Bantaowong</collectorName>
and
<collectorName id="DC777AF63418FFE7FBE313B7E9D8FB54" box="[1068,1218,1158,1183]" pageId="14" pageNumber="107">R. Chanabun</collectorName>
</materialsCitation>
;
<materialsCitation id="C1EA157D3418FFE7FB0313B7EEF0FB2C" ID-GBIF-Occurrence="1314597132" collectingDate="2008-07-23" collectionCode="CUMZ" collectorName="C. Sutcharit &amp; P. Tongkerd" country="Thailand" elevation="92" latitude="16.566668" location="Noen Maprang District, Pa Ma Muang monastery" longitude="100.67722" pageId="14" pageNumber="107" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Phitsanulok" typeStatus="paratype">
<specimenCount id="6784D4A93418FFE7FB0313B7E83FFB54" box="[1228,1317,1158,1183]" pageId="14" pageNumber="107" type="male">3 males</specimenCount>
,
<specimenCount id="6784D4A93418FFE7FAFF13B6E886FB54" box="[1328,1436,1158,1183]" pageId="14" pageNumber="107" type="female">6 females</specimenCount>
(
<collectionCode id="179387E53418FFE7FF51139BEDE8FB08" box="[158,242,1195,1219]" country="Thailand" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="14" pageNumber="107">CUMZ</collectionCode>
)
<collectingCountry id="09955FB03418FFE7FECD139BEC8CFB08" box="[258,406,1195,1219]" name="Thailand" pageId="14" pageNumber="107">THAILAND</collectingCountry>
,
<collectingRegion id="B346D1C23418FFE7FE6E139AEF8EFB08" box="[417,660,1194,1219]" country="Thailand" name="Phitsanulok" pageId="14" pageNumber="107">Phitsanulok Province</collectingRegion>
,
<collectingMunicipality id="9159855A3418FFE7FD50139BEEBDFB08" box="[671,935,1194,1219]" pageId="14" pageNumber="107">Noen Maprang District</collectingMunicipality>
, near
<location id="745D49FB3418FFE7FC24139BE990FB08" LSID="urn:lsid:plazi:treatment:F92BAE363418FFFBFF581457E926F87D:745D49FB3418FFE7FC24139BE990FB08" box="[1003,1162,1195,1219]" country="Thailand" latitude="16.566668" longLatPrecision="21" longitude="100.67722" municipality="Noen Maprang District" name="Pa Ma Muang" pageId="14" pageNumber="107" stateProvince="Phitsanulok Province">Pa Ma Muang</location>
monastery,
<geoCoordinate id="14B679E73418FFE7FADB139AE88DFB08" box="[1300,1431,1194,1219]" direction="north" orientation="latitude" pageId="14" pageNumber="107" precision="15" value="16.566668">16º34'00&quot;N</geoCoordinate>
,
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, ca
<elevation id="FAAFF8133418FFE7FE9B13FFECD3FB2C" box="[340,457,1230,1255]" metricMagnitude="1" metricUnit="m" metricValue="9.2" pageId="14" pageNumber="107" unit="m" value="92.0">
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a.s.l.
</elevation>
,
<collectingDate id="1578C0083418FFE7FE1A13FFEF72FB2C" box="[469,616,1230,1255]" pageId="14" pageNumber="107" value="2008-07-23">
<date id="053C39E03418FFE7FE1A13FFEF72FB2C" box="[469,616,1230,1255]" pageId="14" pageNumber="107" value="2008-07-23">23 July 2008</date>
</collectingDate>
, leg.
<collectorName id="DC777AF63418FFE7FD6B13FFEE30FB2C" box="[676,810,1230,1255]" pageId="14" pageNumber="107">C. Sutcharit</collectorName>
and
<collectorName id="DC777AF63418FFE7FCAF13FFEEF0FB2C" box="[864,1002,1230,1255]" pageId="14" pageNumber="107">P. Tongkerd</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39984CAB3418FFE7FF0813C2ECD5FAE4" pageId="14" pageNumber="107" type="etymology">
<paragraph id="713D1F203418FFE7FF0813C2ECD5FAE4" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
<emphasis id="43F6C3323418FFE7FF0813C2EC54FAC0" bold="true" box="[199,334,1266,1291]" pageId="14" pageNumber="107">Etymology.</emphasis>
The name is a Latin noun in apposition, meaning “shark fin” and referring to the shape of a process on lamina medialis.
</paragraph>
</subSubSection>
<subSubSection id="39984CAB3418FFFBFF08120AEE55F934" lastPageId="18" lastPageNumber="111" pageId="14" pageNumber="107" type="diagnosis">
<paragraph id="713D1F203418FFE7FF08120AEC80FA75" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
<emphasis id="43F6C3323418FFE7FF08120AEC5BFA98" bold="true" box="[199,321,1338,1363]" pageId="14" pageNumber="107">Diagnosis.</emphasis>
Metaterga with 2+2 anterior and 2+2 posterior spines, paraterga wing-like, shocking pink. Similar in these respect to
<taxonomicName id="B68264A33418FFE7FEBA126FECEAFABD" box="[373,496,1374,1398]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="107" phylum="Arthropoda" rank="species" species="planata">
<emphasis id="43F6C3323418FFE7FEBA126FECEAFABD" box="[373,496,1374,1398]" italics="true" pageId="14" pageNumber="107">D. planata</emphasis>
</taxonomicName>
, but differing by pink brown general body color, epiproct with digitiform apical papillae, sternal process between male coxae 4 being subquadrate, lamina medialis (
<emphasis id="43F6C3323418FFE7FB8912B3E978FA50" box="[1094,1122,1411,1435]" italics="true" pageId="14" pageNumber="107">lm</emphasis>
) being highly elevated with a shark finlike process.
</paragraph>
<paragraph id="713D1F203418FFE7FF0812FAEF8DF9CC" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
<emphasis id="43F6C3323418FFE7FF0812FAEC7EFA28" bold="true" box="[199,356,1482,1507]" pageId="14" pageNumber="107">Descriptions.</emphasis>
Length
<quantity id="B67AB2C53418FFE7FE0F12FBEF23FA29" box="[448,569,1483,1507]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="14" pageNumber="107" unit="mm" value="26.5" valueMax="29.0" valueMin="24.0">2429 mm</quantity>
(male),
<quantity id="B67AB2C53418FFE7FD5812FBEE0BFA29" box="[663,785,1483,1507]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="14" pageNumber="107" unit="mm" value="29.0" valueMax="30.0" valueMin="28.0">2830 mm</quantity>
(female); width of midbody prozona and metazona ca 2.0 and 2.0 mm (male), 2.2 and
<quantity id="B67AB2C53418FFE7FE1C12DEEF33F9CD" box="[467,553,1518,1543]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="107" unit="mm" value="2.5">2.5 mm</quantity>
(female).
</paragraph>
<paragraph id="713D1F203418FFE7FF081122ECCCF95C" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
Live coloration vivid pink; paraterga shocking pink; surface below below paraterga and metaterga brown pink; head and antenna dark brown; legs and epiproct pink; sterna brown (
<figureCitation id="E9B903A53418FFE7FC531106EEFFF984" box="[924,997,1590,1615]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[161,1424,565,1778]" captionTargetId="figure@2.[151,1436,553,1790]" captionTargetPageId="2" captionText="FIGURE 1. Live photographs of the new species. A, B: Desmoxytes des sp. n. — A; male holotype, B; female. C, D: Desmoxytes breviverpa sp. n. — male paratype. E, F: Desmoxytes takensis sp. n. — E; male paratype, F; female. G, H: Desmoxytes pinnasquali sp. n. — G: male holotype, H: mating couple." httpUri="https://zenodo.org/record/254808/files/figure.png" pageId="14" pageNumber="107">Figs. 1</figureCitation>
G, 1H). Coloration in alcohol after one year faded to pale brown; paraterga, legs and sterna yellow brown; surface below paraterga, metaterga, head, antenna and epiproct brown.
</paragraph>
<paragraph id="713D1F203418FFE7FF081192E9D2F8EC" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
Width of head &lt;collum &lt;segment 2 &lt;3 = 4 &lt;516, thereafter body gradually tapering towards telson. Head broad ca 2.0 mm (male),
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(female). Clypeolabral region sparsely setose; vertex bare so on; labrum and genae sparsely setose; epicranial suture visible as dark line, quite deep (
<figureCitation id="E9B903A53418FFE7FC0A11DAE93BF8C8" box="[965,1057,1770,1795]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="14" pageNumber="107">Figs. 13</figureCitation>
A, 13D). Antennae very long and slender, reaching back to segments 5 (male) and 4 (female) when expanded dorsally (
<figureCitation id="E9B903A53418FFE7FB9C103EE9BEF8EC" box="[1107,1188,1806,1831]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="14" pageNumber="107">Fig. 13</figureCitation>
D).
</paragraph>
<paragraph id="713D1F203418FFE7FF081002EF52F87C" blockId="14.[151,1437,978,2011]" pageId="14" pageNumber="107">
Collum width ca 3.0 mm (both sexes), surface coarsely microgranulate, with three transverse rows of small setiferous tubercles, 3(4)+3(4) anterior, 1+1 intermediate and 2+2 posterior tubercles; paraterga wing-shaped, elevated at about 30º (male) and 20º (female), directed dorsolaterad, tip pointed, with two distinct setiferous notches on anterior margin (
<figureCitation id="E9B903A53418FFE7FE1C10AEEF39F87C" box="[467,547,1950,1975]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="14" pageNumber="107">Fig. 13</figureCitation>
A).
</paragraph>
<paragraph id="713D1F203418FFE6FF0810F2E977FE43" blockId="14.[151,1437,978,2011]" lastBlockId="15.[151,1437,151,645]" lastPageId="15" lastPageNumber="108" pageId="14" pageNumber="107">
Post-collum segments with surface of prozona finely shagreened; metazona coarsely microgranulate and shining; surface below paraterga finely microgranulate with inconspicuous wrinkles; paraterga and sterna quite smooth (
<figureCitation id="E9B903A53419FFE6FF34178CEC53FF1F" box="[251,329,188,213]" captionStart="FIGURE 8" captionStartId="11.[151,250,1786,1808]" captionTargetBox="[156,1431,624,1763]" captionTargetId="figure@11.[151,1436,619,1769]" captionTargetPageId="11" captionText="FIGURE 8. Desmoxytes takensis sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segment 10 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 10 12 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — ventral view (G) and lateral view (H). I: legs 4 6 — anterior view." httpUri="https://zenodo.org/record/254815/files/figure.png" pageId="15" pageNumber="108">Figs. 8</figureCitation>
AH, 13G). Suture between prozonae and metazonae conspicuous, quite deep and wide, narrow only in segments 25, thereafter much wider (
<figureCitation id="E9B903A53419FFE6FD5617EFEFE9FF33" box="[665,755,223,248]" captionStart="FIGURE 11" captionStartId="15.[151,250,1884,1906]" captionTargetBox="[156,1431,718,1857]" captionTargetId="figure@15.[151,1436,713,1863]" captionTargetPageId="15" captionText="FIGURE 11. Desmoxytes pinnasquali sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segments 10 and 11 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 9 11 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — ventral view (G) and lateral view (H). I: legs 5 7 — anterior view." httpUri="https://zenodo.org/record/254818/files/figure.png" pageId="15" pageNumber="108">Figs. 11</figureCitation>
C, 11E, 13B, 13E, 13G). Metaterga with two transverse rows of setiferous spines; 2+2 anterior and 2+2 posterior spines; spines of anterior rows on metaterga 216 equal in size, lateral spines of posterior rows longer than mesal ones; metaterga 1719 with all spines equal in size. Transverse sulcus on metaterga distinctly expressed on segments 517, quite deep, inconspicuous on segments 4 and 18, missing on segments 2, 3 and 19 (
<figureCitation id="E9B903A53419FFE6FDDC165FEF76FE43" box="[531,620,367,392]" captionStart="FIGURE 11" captionStartId="15.[151,250,1884,1906]" captionTargetBox="[156,1431,718,1857]" captionTargetId="figure@15.[151,1436,713,1863]" captionTargetPageId="15" captionText="FIGURE 11. Desmoxytes pinnasquali sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segments 10 and 11 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 9 11 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — ventral view (G) and lateral view (H). I: legs 5 7 — anterior view." httpUri="https://zenodo.org/record/254818/files/figure.png" pageId="15" pageNumber="108">Figs. 11</figureCitation>
A, 11C, 11F, 13AC). Mid-dorsal line absent.
</paragraph>
<paragraph id="713D1F203419FFE6FF0816A5EEB3FD4F" blockId="15.[151,1437,151,645]" pageId="15" pageNumber="108">
Paraterga strongly developed, especially in male, wing-shaped, directed dorsolaterad at ca 45º (
<figureCitation id="E9B903A53419FFE6FAE816A4E866FE67" box="[1319,1404,404,429]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="15" pageNumber="108">Fig. 13</figureCitation>
F), thereafter curving backward; shoulder present; calluses conspicuous, broad anteriorly and continuing slightly narrowed near tip; lateral margin with two distinct denticles, segments 9, 10, 12,13, 1518 with an additional very small denticle near the tip (
<figureCitation id="E9B903A53419FFE6FE1C16CFEF3CFDD3" box="[467,550,511,536]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="15" pageNumber="108">Fig. 13</figureCitation>
H:
<emphasis id="43F6C3323419FFE6FD841531EF95FDD3" box="[587,655,513,536]" italics="true" pageId="15" pageNumber="108">arrow</emphasis>
); tip of paraterga pointed and sharp, curving dorsolaterad; posterior margin of paraterga curving at base and almost straight near tip (
<figureCitation id="E9B903A53419FFE6FCBC1514EED4FDF7" box="[883,974,548,573]" captionStart="FIGURE 11" captionStartId="15.[151,250,1884,1906]" captionTargetBox="[156,1431,718,1857]" captionTargetId="figure@15.[151,1436,713,1863]" captionTargetPageId="15" captionText="FIGURE 11. Desmoxytes pinnasquali sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segments 10 and 11 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 9 11 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — ventral view (G) and lateral view (H). I: legs 5 7 — anterior view." httpUri="https://zenodo.org/record/254818/files/figure.png" pageId="15" pageNumber="108">Figs. 11</figureCitation>
C, 11E, 13H). Ozopore (
<emphasis id="43F6C3323419FFE6FB2B1514E81BFDF6" bold="true" box="[1252,1281,548,573]" pageId="15" pageNumber="108">op</emphasis>
) visible from above, ovoid (
<figureCitation id="E9B903A53419FFE6FEF11577EC86FDAB" box="[318,412,583,608]" captionStart="FIGURE 11" captionStartId="15.[151,250,1884,1906]" captionTargetBox="[156,1431,718,1857]" captionTargetId="figure@15.[151,1436,713,1863]" captionTargetPageId="15" captionText="FIGURE 11. Desmoxytes pinnasquali sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segments 10 and 11 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 9 11 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — ventral view (G) and lateral view (H). I: legs 5 7 — anterior view." httpUri="https://zenodo.org/record/254818/files/figure.png" pageId="15" pageNumber="108">Figs. 11</figureCitation>
C, 11E, 13H). Pleurosternal carinae forming complete crests on segment 2 (both sexes), small ridge on segment 3 and absent on following segments (
<figureCitation id="E9B903A53419FFE6FC8B155CEE92FD4F" box="[836,904,620,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1396,1418]" captionTargetBox="[156,1431,212,1351]" captionTargetId="figure@3.[151,1436,207,1357]" captionTargetPageId="3" captionText="FIGURE 2. Desmoxytes des sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segments 10 and 11 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 10 12 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — posterior view (G) and lateral view (H). I: legs 5 7 — posterior view." httpUri="https://zenodo.org/record/254809/files/figure.png" pageId="15" pageNumber="108">Fig. 2</figureCitation>
B).
</paragraph>
<caption id="25FD4FA83419FFE6FF58106CEFB6F805" httpUri="https://zenodo.org/record/254818/files/figure.png" pageId="15" pageNumber="108" targetBox="[156,1431,718,1857]" targetPageId="15">
<paragraph id="713D1F203419FFE6FF58106CEFB6F805" blockId="15.[151,1435,1884,1998]" pageId="15" pageNumber="108">
<emphasis id="43F6C3323419FFE6FF58106CEC04F8B9" bold="true" box="[151,286,1884,1906]" pageId="15" pageNumber="108">FIGURE 11.</emphasis>
<taxonomicName id="B68264A33419FFE6FEEB106DEF01F8BA" box="[292,539,1884,1906]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="108" phylum="Arthropoda" rank="species" species="pinnasquali" status="sp. nov.">
<emphasis id="43F6C3323419FFE6FEEB106DEF01F8BA" box="[292,539,1884,1906]" italics="true" pageId="15" pageNumber="108">Desmoxytes pinnasquali</emphasis>
</taxonomicName>
<emphasis id="43F6C3323419FFE6FDED106CEF41F8B9" bold="true" box="[546,603,1884,1906]" pageId="15" pageNumber="108">
<taxonomicNameLabel id="58C57E493419FFE6FDED106CEF41F8B9" box="[546,603,1884,1906]" pageId="15" pageNumber="108" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(male holotype).
<emphasis id="43F6C3323419FFE6FCC0106CEE25F8B9" bold="true" box="[783,831,1884,1906]" pageId="15" pageNumber="108">A, B</emphasis>
: anterior body parts—dorsal view (
<emphasis id="43F6C3323419FFE6FB65106CE9A6F8B9" bold="true" box="[1194,1212,1884,1906]" pageId="15" pageNumber="108">A</emphasis>
) and lateral view (
<emphasis id="43F6C3323419FFE6FAB3106CE897F8B9" bold="true" box="[1404,1421,1884,1906]" pageId="15" pageNumber="108">B</emphasis>
).
<emphasis id="43F6C3323419FFE6FF58104BEDB3F85A" bold="true" box="[151,169,1915,1937]" pageId="15" pageNumber="108">C</emphasis>
: segments 10 and 11 (
<emphasis id="43F6C3323419FFE6FE53104BECADF85A" bold="true" box="[412,439,1915,1937]" pageId="15" pageNumber="108">op</emphasis>
= ozopore)—dorsal view.
<emphasis id="43F6C3323419FFE6FD1E104BEE06F85A" bold="true" box="[721,796,1915,1937]" pageId="15" pageNumber="108">D, F, J</emphasis>
: posterior body parts—lateral view (
<emphasis id="43F6C3323419FFE6FB6D104BE9AEF85A" bold="true" box="[1186,1204,1915,1937]" pageId="15" pageNumber="108">D</emphasis>
), dorsal view (
<emphasis id="43F6C3323419FFE6FA99104BE87FF85A" bold="true" box="[1366,1381,1915,1937]" pageId="15" pageNumber="108">F</emphasis>
) and ventral view (
<emphasis id="43F6C3323419FFE6FEE910A9EC29F864" bold="true" box="[294,307,1945,1967]" pageId="15" pageNumber="108">J</emphasis>
).
<emphasis id="43F6C3323419FFE6FE8610A9EC40F864" bold="true" box="[329,346,1945,1967]" pageId="15" pageNumber="108">E</emphasis>
: segments 911 (
<emphasis id="43F6C3323419FFE6FDDF10A9EF31F864" bold="true" box="[528,555,1945,1967]" pageId="15" pageNumber="108">op</emphasis>
= ozopore)—lateral view.
<emphasis id="43F6C3323419FFE6FC8F10A9EE75F864" bold="true" box="[832,879,1945,1967]" pageId="15" pageNumber="108">G, H</emphasis>
: sternal process between male coxae 4—ventral view (
<emphasis id="43F6C3323419FFE6FF501088EDA9F805" bold="true" box="[159,179,1976,1998]" pageId="15" pageNumber="108">G</emphasis>
) and lateral view (
<emphasis id="43F6C3323419FFE6FEBD1088EC9CF805" bold="true" box="[370,390,1976,1998]" pageId="15" pageNumber="108">H</emphasis>
).
<emphasis id="43F6C3323419FFE6FE541088ECBFF805" bold="true" box="[411,421,1976,1998]" pageId="15" pageNumber="108">I</emphasis>
: legs 57—anterior view.
</paragraph>
</caption>
<caption id="25FD4FA83406FFF9FF581167ECEBF947" httpUri="https://zenodo.org/record/254819/files/figure.png" pageId="16" pageNumber="109" targetBox="[151,1436,193,1601]" targetPageId="16">
<paragraph id="713D1F203406FFF9FF581167ECEBF947" blockId="16.[151,1436,1623,1676]" pageId="16" pageNumber="109">
<emphasis id="43F6C3323406FFF9FF581167EC05F9A6" bold="true" box="[151,287,1623,1645]" pageId="16" pageNumber="109">FIGURE 12.</emphasis>
<taxonomicName id="B68264A33406FFF9FEEB1168EF00F9A6" box="[292,538,1624,1645]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="109" phylum="Arthropoda" rank="species" species="pinnasquali" status="sp. nov.">
<emphasis id="43F6C3323406FFF9FEEB1168EF00F9A6" box="[292,538,1624,1645]" italics="true" pageId="16" pageNumber="109">Desmoxytes pinnasquali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58C57E493406FFF9FDEF1167EF43F9A6" box="[544,601,1623,1645]" pageId="16" pageNumber="109" rank="species">
<emphasis id="43F6C3323406FFF9FDEF1167EF43F9A6" bold="true" box="[544,601,1623,1645]" pageId="16" pageNumber="109">sp. n.</emphasis>
</taxonomicNameLabel>
(male paratype), right gonopod.
<emphasis id="43F6C3323406FFF9FC6C1167EEAFF9A6" bold="true" box="[931,949,1623,1645]" pageId="16" pageNumber="109">A</emphasis>
: lateral view.
<emphasis id="43F6C3323406FFF9FB8C1167E94EF9A6" bold="true" box="[1091,1108,1623,1645]" pageId="16" pageNumber="109">B</emphasis>
: mesal view.
<emphasis id="43F6C3323406FFF9FB131167E9F4F9A6" bold="true" box="[1244,1262,1623,1645]" pageId="16" pageNumber="109">C</emphasis>
: ventral view.
<emphasis id="43F6C3323406FFF9FA4C1167EDBCF947" bold="true" pageId="16" pageNumber="109">D, F</emphasis>
: subdorsal view.
<emphasis id="43F6C3323406FFF9FE981146EC72F947" bold="true" box="[343,360,1654,1676]" pageId="16" pageNumber="109">E</emphasis>
: dorsal view.
</paragraph>
</caption>
<paragraph id="713D1F203406FFF9FF081183EF8CF8DF" blockId="16.[151,1437,1715,1956]" pageId="16" pageNumber="109">
Epiproct conical; tip concave; apical papillae conspicuous, digitiform; with tiny tubercles supporting two pairs of paramedian setae; lateral papillae distinct (
<figureCitation id="E9B903A53406FFF9FD5711E7EFE9F93B" box="[664,755,1751,1776]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="16" pageNumber="109">Figs. 13</figureCitation>
JK). Hypoproct subsemicircular, with prominent setigerous tubercles on convex caudal margin (
<figureCitation id="E9B903A53406FFF9FDE311CBEF64F8DF" box="[556,638,1787,1812]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="16" pageNumber="109">
Fig.
<date id="053C39E03406FFF9FDAF11CCEF64F8DF" box="[608,638,1788,1812]" pageId="16" pageNumber="109" value="2013-01">13</date>
</figureCitation>
I).
</paragraph>
<paragraph id="713D1F203406FFF9FF081010EE76F86F" blockId="16.[151,1437,1715,1956]" pageId="16" pageNumber="109">
Sterna sparsely setose, cross-impression shallow; sternal process between male coxae 4 modified, subquadrate, tip round, sternal pores visible from posterior view (
<figureCitation id="E9B903A53406FFF9FD351073EE4FF897" box="[762,853,1859,1884]" captionStart="FIGURE 11" captionStartId="15.[151,250,1884,1906]" captionTargetBox="[156,1431,718,1857]" captionTargetId="figure@15.[151,1436,713,1863]" captionTargetPageId="15" captionText="FIGURE 11. Desmoxytes pinnasquali sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segments 10 and 11 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 9 11 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — ventral view (G) and lateral view (H). I: legs 5 7 — anterior view." httpUri="https://zenodo.org/record/254818/files/figure.png" pageId="16" pageNumber="109">Figs. 11</figureCitation>
G, 11H, 13M). Legs very long and slender, ca 2.5 (male), 2.0 times (female) as long as midbody height (
<figureCitation id="E9B903A53406FFF9FD331057EE57F84B" box="[764,845,1895,1920]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="16" pageNumber="109">Fig. 13</figureCitation>
P). Male femora 5 and 6 distinctly humped ventrally in middle part, femur 6 a bit bigger than 5 (
<figureCitation id="E9B903A53406FFF9FDB310BBEFCDF86F" box="[636,727,1931,1956]" captionStart="FIGURE 11" captionStartId="15.[151,250,1884,1906]" captionTargetBox="[156,1431,718,1857]" captionTargetId="figure@15.[151,1436,713,1863]" captionTargetPageId="15" captionText="FIGURE 11. Desmoxytes pinnasquali sp. n. (male holotype). A, B: anterior body parts—dorsal view (A) and lateral view (B). C: segments 10 and 11 (op = ozopore) — dorsal view. D, F, J: posterior body parts—lateral view (D), dorsal view (F) and ventral view (J). E: segments 9 11 (op = ozopore) — lateral view. G, H: sternal process between male coxae 4 — ventral view (G) and lateral view (H). I: legs 5 7 — anterior view." httpUri="https://zenodo.org/record/254818/files/figure.png" pageId="16" pageNumber="109">
Figs.
<date id="053C39E03406FFF9FD7410BCEFCDF86F" box="[699,727,1932,1956]" pageId="16" pageNumber="109" value="2011-01">11</date>
</figureCitation>
I, 13N, 13O).
</paragraph>
<caption id="25FD4FA83407FFF8FF5811F2EC08F838" httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="17" pageNumber="110" targetBox="[154,1413,199,1704]" targetPageId="17">
<paragraph id="713D1F203407FFF8FF5811F2EC08F838" blockId="17.[151,1437,1730,2035]" pageId="17" pageNumber="110">
<emphasis id="43F6C3323407FFF8FF5811F2EC3AF913" bold="true" box="[151,288,1730,1752]" pageId="17" pageNumber="110">FIGURE 13.</emphasis>
<taxonomicName id="B68264A33407FFF8FEE911F3EF07F91C" box="[294,541,1730,1752]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="110" phylum="Arthropoda" rank="species" species="pinnasquali" status="sp. nov.">
<emphasis id="43F6C3323407FFF8FEE911F3EF07F91C" box="[294,541,1730,1752]" italics="true" pageId="17" pageNumber="110">Desmoxytes pinnasquali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58C57E493407FFF8FDEB11F2EF44F913" box="[548,606,1730,1752]" pageId="17" pageNumber="110" rank="species">
<emphasis id="43F6C3323407FFF8FDEB11F2EF44F913" bold="true" box="[548,606,1730,1752]" pageId="17" pageNumber="110">sp. n.</emphasis>
</taxonomicNameLabel>
(male holotype).
<emphasis id="43F6C3323407FFF8FCDB11F2EE3CF913" bold="true" box="[788,806,1730,1752]" pageId="17" pageNumber="110">A</emphasis>
: head and segments 13—dorsal view.
<emphasis id="43F6C3323407FFF8FB7311F2E9D7F913" bold="true" box="[1212,1229,1730,1752]" pageId="17" pageNumber="110">B</emphasis>
: segments 810 (
<emphasis id="43F6C3323407FFF8FA4E11F2E881F913" bold="true" box="[1409,1435,1730,1752]" pageId="17" pageNumber="110">op</emphasis>
= ozopore)—dorsal view.
<emphasis id="43F6C3323407FFF8FE6C11D1ECAFF93C" bold="true" box="[419,437,1761,1783]" pageId="17" pageNumber="110">C</emphasis>
: posteriormost segments and telson segments—dorsal view
<emphasis id="43F6C3323407FFF8FBEC11D1E92FF93C" bold="true" box="[1059,1077,1761,1783]" pageId="17" pageNumber="110">D</emphasis>
: head and antenna—anterior view.
<emphasis id="43F6C3323407FFF8FF5811CFEDB2F8DE" bold="true" box="[151,168,1791,1813]" pageId="17" pageNumber="110">E</emphasis>
: segments 810—lateral view.
<emphasis id="43F6C3323407FFF8FE2211CFECE6F8DE" bold="true" box="[493,508,1791,1813]" pageId="17" pageNumber="110">F</emphasis>
: segment 10—posterior view.
<emphasis id="43F6C3323407FFF8FCF611CFEE57F8DE" bold="true" box="[825,845,1791,1813]" pageId="17" pageNumber="110">G</emphasis>
: paraterga and sculpture of segment 10—lateral view.
<emphasis id="43F6C3323407FFF8FA4E11CFE88FF8DE" bold="true" box="[1409,1429,1791,1813]" pageId="17" pageNumber="110">H</emphasis>
: paraterga of segment 10 (
<emphasis id="43F6C3323407FFF8FE51102EECFAF8FF" bold="true" box="[414,480,1822,1844]" pageId="17" pageNumber="110">arrow</emphasis>
= tiny denticle)—lateral view.
<emphasis id="43F6C3323407FFF8FCED102EEE36F8FF" bold="true" box="[802,812,1822,1844]" pageId="17" pageNumber="110">I</emphasis>
: last segment and telson; showing epiproct and hypoproctventral view.
<emphasis id="43F6C3323407FFF8FEE8100CEC4BF899" bold="true" box="[295,337,1852,1874]" pageId="17" pageNumber="110">JL</emphasis>
: epiproct;
<emphasis id="43F6C3323407FFF8FE09100CECC8F899" bold="true" box="[454,466,1852,1874]" pageId="17" pageNumber="110">J</emphasis>
: dorsal view,
<emphasis id="43F6C3323407FFF8FDA4100CEF65F899" bold="true" box="[619,639,1852,1874]" pageId="17" pageNumber="110">K</emphasis>
: lateral view,
<emphasis id="43F6C3323407FFF8FCD6100CEE30F899" bold="true" box="[793,810,1852,1874]" pageId="17" pageNumber="110">L</emphasis>
: ventral view.
<emphasis id="43F6C3323407FFF8FC04100CEEF9F899" bold="true" box="[971,995,1852,1874]" pageId="17" pageNumber="110">M</emphasis>
: sternal process between male coxae 4— posterior view.
<emphasis id="43F6C3323407FFF8FEFB106BEC5CF8BA" bold="true" box="[308,326,1883,1905]" pageId="17" pageNumber="110">N</emphasis>
: male leg 5.
<emphasis id="43F6C3323407FFF8FE04106BECC5F8BA" bold="true" box="[459,479,1883,1905]" pageId="17" pageNumber="110">O</emphasis>
: male leg 6.
<emphasis id="43F6C3323407FFF8FDAB106BEF69F8BA" bold="true" box="[612,627,1883,1905]" pageId="17" pageNumber="110">P</emphasis>
: male right anterior leg of segment 10.
<emphasis id="43F6C3323407FFF8FBC8106BE922F8BA" bold="true" box="[1031,1080,1883,1905]" pageId="17" pageNumber="110">QT</emphasis>
: right gonopod;
<emphasis id="43F6C3323407FFF8FB2D106BE9ECF8BA" bold="true" box="[1250,1270,1883,1905]" pageId="17" pageNumber="110">Q</emphasis>
: mesal view (
<emphasis id="43F6C3323407FFF8FA4A106CE881F8BA" box="[1413,1435,1884,1905]" italics="true" pageId="17" pageNumber="110">cx</emphasis>
= coxa,
<emphasis id="43F6C3323407FFF8FF28104BEDE5F85B" box="[231,255,1915,1936]" italics="true" pageId="17" pageNumber="110">ca</emphasis>
= cannula,
<emphasis id="43F6C3323407FFF8FEBA104AEC89F844" box="[373,403,1914,1935]" italics="true" pageId="17" pageNumber="110">pfe</emphasis>
= prefemorite,
<emphasis id="43F6C3323407FFF8FDFF104AEF58F844" box="[560,578,1914,1935]" italics="true" pageId="17" pageNumber="110">fe</emphasis>
= femorite,
<emphasis id="43F6C3323407FFF8FD70104BEFCCF85B" box="[703,726,1915,1936]" italics="true" pageId="17" pageNumber="110">sg</emphasis>
= seminal groove,
<emphasis id="43F6C3323407FFF8FC57104AEEA6F844" box="[920,956,1914,1935]" italics="true" pageId="17" pageNumber="110">sph</emphasis>
= solenophore,
<emphasis id="43F6C3323407FFF8FB92104BE9ABF85B" bold="true" box="[1117,1201,1915,1936]" italics="true" pageId="17" pageNumber="110">P arrow</emphasis>
= finlike process,
<emphasis id="43F6C3323407FFF8FAA3104BE898F85F" bold="true" box="[1388,1410,1915,1940]" italics="true" pageId="17" pageNumber="110">L1</emphasis>
&amp;
<emphasis id="43F6C3323407FFF8FF5810ACEDB7F87E" bold="true" box="[151,173,1948,1973]" italics="true" pageId="17" pageNumber="110">L2</emphasis>
<emphasis id="43F6C3323407FFF8FF7B10ABEC1AF87A" bold="true" box="[180,256,1947,1969]" pageId="17" pageNumber="110">arrows</emphasis>
= lamellalike lobes),
<emphasis id="43F6C3323407FFF8FE1910ABECF2F87A" bold="true" box="[470,488,1947,1969]" pageId="17" pageNumber="110">R</emphasis>
: lateral view (
<emphasis id="43F6C3323407FFF8FDB310ABEE1AF87A" bold="true" box="[636,768,1947,1969]" pageId="17" pageNumber="110">black arrow</emphasis>
= mesal sulcus,
<emphasis id="43F6C3323407FFF8FC6610ABE936F87A" bold="true" box="[937,1068,1947,1969]" pageId="17" pageNumber="110">white arrow</emphasis>
= lateral sulcus),
<emphasis id="43F6C3323407FFF8FB2D10ABE9EAF87A" bold="true" box="[1250,1264,1947,1969]" pageId="17" pageNumber="110">S</emphasis>
: ventral view (
<emphasis id="43F6C3323407FFF8FA4410ABE886F87B" box="[1419,1436,1947,1968]" italics="true" pageId="17" pageNumber="110">sl</emphasis>
= solenomere,
<emphasis id="43F6C3323407FFF8FEFF108CEC24F81A" box="[304,318,1980,2001]" italics="true" pageId="17" pageNumber="110">ll</emphasis>
= lamina lateralis,
<emphasis id="43F6C3323407FFF8FDC8108CEF3AF81A" box="[519,544,1980,2001]" italics="true" pageId="17" pageNumber="110">lm</emphasis>
= lamina medialis,
<emphasis id="43F6C3323407FFF8FD20108DEE5FF819" bold="true" box="[751,837,1981,2002]" italics="true" pageId="17" pageNumber="110">P arrow</emphasis>
= finlike process,
<emphasis id="43F6C3323407FFF8FBC8108DE907F81D" bold="true" box="[1031,1053,1981,2006]" italics="true" pageId="17" pageNumber="110">L1</emphasis>
&amp;
<emphasis id="43F6C3323407FFF8FB8E108DE94DF81D" bold="true" box="[1089,1111,1981,2006]" italics="true" pageId="17" pageNumber="110">L2</emphasis>
<emphasis id="43F6C3323407FFF8FB90108CE9B1F819" bold="true" box="[1119,1195,1980,2002]" pageId="17" pageNumber="110">arrows</emphasis>
= lamellalike lobes),
<emphasis id="43F6C3323407FFF8FA4A108CE88FF819" bold="true" box="[1413,1429,1980,2002]" pageId="17" pageNumber="110">T</emphasis>
: dorsal view.
</paragraph>
</caption>
<caption id="25FD4FA83404FFFBFF581257EC54FA71" httpUri="https://zenodo.org/record/254821/files/figure.png" pageId="18" pageNumber="111" targetBox="[151,1436,193,1361]" targetPageId="18">
<paragraph id="713D1F203404FFFBFF581257EC54FA71" blockId="18.[151,1436,1383,1466]" pageId="18" pageNumber="111">
<emphasis id="43F6C3323404FFFBFF581257EC3BFAB6" bold="true" box="[151,289,1383,1405]" pageId="18" pageNumber="111">FIGURE 14.</emphasis>
Known distribution of four new species of
<taxonomicName id="B68264A33404FFFBFD251258EE79FAB6" box="[746,867,1384,1405]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="43F6C3323404FFFBFD251258EE79FAB6" box="[746,867,1384,1405]" italics="true" pageId="18" pageNumber="111">Desmoxytes</emphasis>
</taxonomicName>
in northern Thailand. Open triangle =
<emphasis id="43F6C3323404FFFBFB361258E886FAB7" box="[1273,1436,1383,1405]" italics="true" pageId="18" pageNumber="111">
<taxonomicName id="B68264A33404FFFBFB361258E868FAB6" box="[1273,1394,1384,1405]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="111" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
des
</emphasis>
<emphasis id="43F6C3323404FFFBFF5812B6EDCEFA57" bold="true" box="[151,212,1414,1436]" pageId="18" pageNumber="111">sp. n.</emphasis>
: Open star =
<taxonomicName id="B68264A33404FFFBFEA212B7EF47FA50" box="[365,605,1414,1436]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="111" phylum="Arthropoda" rank="species" species="breviverpa" status="sp. nov.">
<emphasis id="43F6C3323404FFFBFEA212B7EF47FA50" box="[365,605,1414,1436]" italics="true" pageId="18" pageNumber="111">Desmoxytes breviverpa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58C57E493404FFFBFDA812B6EFBEFA57" box="[615,676,1414,1436]" pageId="18" pageNumber="111" rank="species">
<emphasis id="43F6C3323404FFFBFDA812B6EFBEFA57" bold="true" box="[615,676,1414,1436]" pageId="18" pageNumber="111">sp. n.</emphasis>
</taxonomicNameLabel>
: Open square =
<taxonomicName id="B68264A33404FFFBFC9412B7E935FA50" box="[859,1071,1414,1436]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="111" phylum="Arthropoda" rank="species" species="takensis" status="sp. nov.">
<emphasis id="43F6C3323404FFFBFC9412B7E935FA50" box="[859,1071,1414,1436]" italics="true" pageId="18" pageNumber="111">Desmoxytes takensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58C57E493404FFFBFBF612B6E96CFA57" box="[1081,1142,1414,1436]" pageId="18" pageNumber="111" rank="species">
<emphasis id="43F6C3323404FFFBFBF612B6E96CFA57" bold="true" box="[1081,1142,1414,1436]" pageId="18" pageNumber="111">sp. n.</emphasis>
</taxonomicNameLabel>
: Open circle =
<taxonomicName id="B68264A33404FFFBFAEC12B7EC15FA72" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="111" phylum="Arthropoda" rank="species" species="pinnasquali" status="sp. nov.">
<emphasis id="43F6C3323404FFFBFAEC12B7EC15FA72" italics="true" pageId="18" pageNumber="111">Desmoxytes pinnasquali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58C57E493404FFFBFEDB1294EC54FA71" box="[276,334,1444,1466]" pageId="18" pageNumber="111" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="713D1F203404FFFBFF0812D7EE55F934" blockId="18.[151,1436,1511,1974]" pageId="18" pageNumber="111">
Gonopods (
<figureCitation id="E9B903A53404FFFBFE8412D7ECB3FA34" box="[331,425,1511,1536]" captionStart="FIGURE 12" captionStartId="16.[151,250,1623,1645]" captionTargetBox="[151,1436,193,1601]" captionTargetId="figure@16.[151,1436,193,1602]" captionTargetPageId="16" captionText="FIGURE 12. Desmoxytes pinnasquali sp. n. (male paratype), right gonopod. A: lateral view. B: mesal view. C: ventral view. D, F: subdorsal view. E: dorsal view." httpUri="https://zenodo.org/record/254819/files/figure.png" pageId="18" pageNumber="111">Figs. 12</figureCitation>
,
<figureCitation id="E9B903A53404FFFBFE7912D7ECCAFA34" box="[438,464,1511,1535]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="18" pageNumber="111">13</figureCitation>
QT) long and slender, suberect, distal parts strongly condensed. Coxa (
<emphasis id="43F6C3323404FFFBFAC812D8E83AFA34" box="[1287,1312,1512,1535]" italics="true" pageId="18" pageNumber="111">cx</emphasis>
) about 1/3 as long as telopodite. Prefemorite (
<emphasis id="43F6C3323404FFFBFDFF113BEF48F9E8" box="[560,594,1547,1571]" italics="true" pageId="18" pageNumber="111">pfe</emphasis>
) about 1/3 as long as telopodite, quite stout. Femorite (
<emphasis id="43F6C3323404FFFBFB2E113BE9EFF9E8" box="[1249,1269,1547,1571]" italics="true" pageId="18" pageNumber="111">fe</emphasis>
) slender, very elongate, with seminal groove running entirely on mesal surface, postfemoral part demarcated by deep mesal and lateral sulcus (
<figureCitation id="E9B903A53404FFFBFE8E1163EC8FF9A7" box="[321,405,1619,1644]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="18" pageNumber="111">Fig. 13</figureCitation>
R:
<emphasis id="43F6C3323404FFFBFE751163EFADF9A0" box="[442,695,1619,1643]" italics="true" pageId="18" pageNumber="111">black &amp; white arrows</emphasis>
). Solenophore strongly developed: lamina lateralis (
<emphasis id="43F6C3323404FFFBFAD51163E833F9A0" box="[1306,1321,1619,1643]" italics="true" pageId="18" pageNumber="111">ll</emphasis>
) swollen, without a lobe: lamina medialis (
<emphasis id="43F6C3323404FFFBFDD81147EF29F944" box="[535,563,1655,1679]" italics="true" pageId="18" pageNumber="111">lm</emphasis>
) broad and highly elevated, with one process and two lobes; process shark finlike, tip obtuse, directed mesad (
<figureCitation id="E9B903A53404FFFBFDE411ABEF92F97F" box="[555,648,1691,1716]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="18" pageNumber="111">Figs. 13</figureCitation>
Q, 13S,
<emphasis id="43F6C3323404FFFBFD2711ACEE5CF978" box="[744,838,1692,1715]" italics="true" pageId="18" pageNumber="111">P arrow</emphasis>
); first lobe lamellalike, terminating in two lamellae, outer one smaller than the inner one (
<figureCitation id="E9B903A53404FFFBFD87118FEFBFF91C" box="[584,677,1727,1752]" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="18" pageNumber="111">Figs. 13</figureCitation>
Q, 13S,
<emphasis id="43F6C3323404FFFBFCCA11F0EE72F91C" box="[773,872,1728,1756]" italics="true" pageId="18" pageNumber="111">L1 arrow</emphasis>
); second lobe lamellalike, thick and broad (
<figureCitation id="E9B903A53404FFFBFAAC118FEDA9F934" captionStart="FIGURE 13" captionStartId="17.[151,250,1730,1752]" captionTargetBox="[154,1413,199,1704]" captionTargetId="figure@17.[150,1436,193,1717]" captionTargetPageId="17" captionText="FIGURE 13. Desmoxytes pinnasquali sp. n. (male holotype). A: head and segments 1 3 — dorsal view. B: segments 8 10 (op = ozopore) — dorsal view. C: posteriormost segments and telson segments—dorsal view D: head and antenna—anterior view. E: segments 8 10 — lateral view. F: segment 10 — posterior view. G: paraterga and sculpture of segment 10 — lateral view. H: paraterga of segment 10 (arrow = tiny denticle) — lateral view. I: last segment and telson; showing epiproct and hypoproctventral view. J L: epiproct; J: dorsal view, K: lateral view, L: ventral view. M: sternal process between male coxae 4 — posterior view. N: male leg 5. O: male leg 6. P: male right anterior leg of segment 10. Q T: right gonopod; Q: mesal view (cx = coxa, ca = cannula, pfe = prefemorite, fe = femorite, sg = seminal groove, sph = solenophore, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), R: lateral view (black arrow = mesal sulcus, white arrow = lateral sulcus), S: ventral view (sl = solenomere, ll = lamina lateralis, lm = lamina medialis, P arrow = finlike process, L 1 &amp; L 2 arrows = lamellalike lobes), T: dorsal view." httpUri="https://zenodo.org/record/254820/files/figure.png" pageId="18" pageNumber="111">Figs. 13</figureCitation>
Q, 13S,
<emphasis id="43F6C3323404FFFBFEC111D7EC6AF935" box="[270,368,1767,1795]" italics="true" pageId="18" pageNumber="111">L2 arrow</emphasis>
). Solenomere flagelliform, curving distad.
</paragraph>
</subSubSection>
<subSubSection id="39984CAB3404FFFBFF08103CEC3DF8A6" pageId="18" pageNumber="111" type="distribution">
<paragraph id="713D1F203404FFFBFF08103CEC3DF8A6" blockId="18.[151,1436,1511,1974]" pageId="18" pageNumber="111">
<emphasis id="43F6C3323404FFFBFF08103CECECF8EE" bold="true" box="[199,502,1804,1829]" pageId="18" pageNumber="111">Distribution and habitat.</emphasis>
This new species occurs in a small limestone area in the west of Thung Salaeng Luang National Park, northern
<collectingCountry id="09955FB03404FFFBFDCA1000EF73F882" box="[517,617,1840,1865]" name="Thailand" pageId="18" pageNumber="111">Thailand</collectingCountry>
and was seen crawling on humid rocks, litter and vegetation under a shading tree.
</paragraph>
</subSubSection>
<subSubSection id="39984CAB3404FFFBFF081048E926F87D" pageId="18" pageNumber="111" type="discussion">
<paragraph id="713D1F203404FFFBFF081048E926F87D" blockId="18.[151,1436,1511,1974]" pageId="18" pageNumber="111">
<emphasis id="43F6C3323404FFFBFF081048EC2AF85A" bold="true" box="[199,304,1912,1937]" pageId="18" pageNumber="111">Remark.</emphasis>
This species shows a bizarre pink body color which is clearly aposematic. It may be closely related to
<taxonomicName id="B68264A33404FFFBFF7B10AEEC36F87E" box="[180,300,1949,1973]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="111" phylum="Arthropoda" rank="species" species="planata">
<emphasis id="43F6C3323404FFFBFF7B10AEEC36F87E" box="[180,300,1949,1973]" italics="true" pageId="18" pageNumber="111">D. planata</emphasis>
</taxonomicName>
with which it shares the same patterns of color and metatergal spines.
</paragraph>
</subSubSection>
</treatment>
</document>