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<document id="C4098F8F35384A7A3EA089C6A3CF373A" ID-DOI="10.11646/zootaxa.3974.3.3" ID-GBIF-Dataset="53679d76-ae41-4651-a3fc-b6823993d5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="231780" ID-ZooBank="408C29FE-AF3B-4D4D-950C-7DFED83B2159" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461219790961" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Anker, Arthur" docDate="2015" docId="F911475EFFE81114EBF8FF53FD3E6460" docLanguage="en" docName="zt03974p360.pdf" docOrigin="Zootaxa 3974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naushonia draconis Anker 2014" docType="treatment" docVersion="16" lastPageNumber="358" masterDocId="05283F26FFF91105EB6FFFC4FFF16112" masterDocTitle="Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key" masterLastPageNumber="360" masterPageNumber="341" pageNumber="358" updateTime="1698603758967" updateUser="plazi">
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<mods:title id="E427B34E243F0C6A3BE850787534AC41">Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key</mods:title>
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<treatment id="F911475EFFE81114EBF8FF53FD3E6460" ID-DOI="http://doi.org/10.5281/zenodo.5073335" ID-GBIF-Taxon="119631466" ID-Zenodo-Dep="5073335" LSID="urn:lsid:plazi:treatment:F911475EFFE81114EBF8FF53FD3E6460" httpUri="http://treatment.plazi.org/id/F911475EFFE81114EBF8FF53FD3E6460" lastPageNumber="358" pageId="17" pageNumber="358">
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<paragraph id="7107F648FFE81114EBF8FF53FDDB61A0" blockId="17.[151,554,151,211]" box="[151,554,151,178]" pageId="17" pageNumber="358">
<heading id="2A4F4124FFE81114EBF8FF53FDDB61A0" bold="true" box="[151,554,151,178]" fontSize="11" level="1" pageId="17" pageNumber="358" reason="1">
<taxonomicName id="B6B88DCBFFE81114EBF8FF53FDDB61A0" ID-CoL="16176e27-ddea-456e-ac3c-e51e8ebff7ac" authority="Anker, 2014" authorityName="Anker" authorityYear="2014" box="[151,554,151,178]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114EBF8FF53FDDB61A0" bold="true" box="[151,554,151,178]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBF8FF53FE7D61A3" bold="true" box="[151,396,151,177]" italics="true" pageId="17" pageNumber="358">Naushonia draconis</emphasis>
<bibRefCitation id="15298BB9FFE81114EAFCFF5CFDDB61A0" author="Anker" box="[403,554,152,178]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker, 2014</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7107F648FFE81114EBF8FF7EFF2861C1" blockId="17.[151,554,151,211]" box="[151,217,186,211]" pageId="17" pageNumber="358">
<figureCitation id="E983EACDFFE81114EBF8FF7EFF2861C1" box="[151,217,186,211]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBF8FEC6FBB9600A" box="[151,1096,258,280]" pageId="17" pageNumber="358" type="reference_group">
<paragraph id="7107F648FFE81114EBF8FEC6FBB9600A" blockId="17.[151,1096,258,280]" box="[151,1096,258,280]" pageId="17" pageNumber="358">
<treatmentCitationGroup id="51A8D166FFE81114EBF8FEC6FBB9600A" box="[151,1096,258,280]" pageId="17" pageNumber="358">
<taxonomicName id="B6B88DCBFFE81114EBF8FEC6FDE6600A" ID-CoL="16176e27-ddea-456e-ac3c-e51e8ebff7ac" authority="Anker 2014: 761" authorityName="Anker" authorityPageNumber="761" authorityYear="2014" box="[151,535,258,280]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114EBF8FEC6FE92600A" box="[151,355,258,280]" italics="true" pageId="17" pageNumber="358">Naushonia draconis</emphasis>
<treatmentCitation id="F019D059FFE81114EA06FEC6FDE6600A" author="Anker" box="[361,535,258,280]" page="761" pageId="17" pageNumber="358" year="2014">
<bibRefCitation id="15298BB9FFE81114EA06FEC6FE17600A" author="Anker" box="[361,486,258,280]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
: 761
</treatmentCitation>
</taxonomicName>
, figs. 15 (
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locality:
<collectingCountry id="09AFB6D8FFE81114E87BFEC6FC60600A" box="[788,913,258,280]" name="Guadeloupe" pageId="17" pageNumber="358">Guadeloupe</collectingCountry>
, French Antilles).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBF8FE81FCBA60B4" pageId="17" pageNumber="358" type="materials_examined">
<paragraph id="7107F648FFE81114EBF8FE81FCBA60B4" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBF8FE81FE8F604C" bold="true" box="[151,382,325,350]" pageId="17" pageNumber="358">Material examined</emphasis>
.
<collectingCountry id="09AFB6D8FFE81114EAFDFE81FE20604C" box="[402,465,325,350]" name="Cuba" pageId="17" pageNumber="358">Cuba</collectingCountry>
:
<specimenCount id="67BE3DC1FFE81114EA8EFE82FDC3604C" box="[481,562,325,350]" pageId="17" pageNumber="358" type="male">1 male</specimenCount>
(pocl
<quantity id="B6405BADFFE81114E9EBFE81FD12604C" box="[644,739,325,350]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="17" pageNumber="358" unit="mm" value="5.1">5.1 mm</quantity>
),
<specimenCount id="67BE3DC1FFE81114E994FE82FC90604C" box="[763,865,325,350]" pageId="17" pageNumber="358" type="female">1 female</specimenCount>
(pocl
<quantity id="B6405BADFFE81114E8DBFE81FBE2604C" box="[948,1043,325,350]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="17" pageNumber="358" unit="mm" value="5.6">5.6 mm</quantity>
),
<collectingCountry id="09AFB6D8FFE81114EF44FE81FBA2604C" box="[1067,1107,325,350]" name="Iceland" pageId="17" pageNumber="358">Isla</collectingCountry>
de la Juventud,
<geoCoordinate id="148C908FFFE81114EE70FE81FA66604C" box="[1311,1431,325,350]" direction="north" orientation="latitude" pageId="17" pageNumber="358" precision="92" value="21.585">21°35.1N</geoCoordinate>
,
<geoCoordinate id="148C908FFFE81114EBF8FEADFEE26093" box="[151,275,361,385]" direction="west" orientation="longitude" pageId="17" pageNumber="358" precision="92" value="-83.17">83°10.2W</geoCoordinate>
,
<quantity id="B6405BADFFE81114EA4EFEADFEAC6093" box="[289,349,361,385]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="358" unit="m" value="11.0">11 m</quantity>
, coral reef, rubble, coll. M. Johnson
<emphasis id="43CC2A5AFFE81114E864FEAEFCB76093" box="[779,838,361,385]" italics="true" pageId="17" pageNumber="358">et al.</emphasis>
,
<date id="0506D088FFE81114E839FEADFC056093" box="[854,1012,361,385]" pageId="17" pageNumber="358" value="2014-06-28">28 June 2014</date>
,
<collectionCode id="17A96E8DFFE81114EF6DFEADFBAE6093" box="[1026,1119,361,385]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="358">MZUSP</collectionCode>
33201; 1 ovigerous female (pocl
<quantity id="B6405BADFFE81114EBB8FE49FEC060B4" box="[215,305,397,422]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="17" pageNumber="358" unit="mm" value="5.2">5.2 mm</quantity>
), same collection data,
<collectionCode id="17A96E8DFFE81114E958FE49FD6060B4" LSID="urn:lsid:biocol.org:col:34992" box="[567,657,397,422]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="358">RMNH</collectionCode>
.
<collectionCode id="17A96E8DFFE81114E9FAFE49FD1360B4" box="[661,738,397,422]" pageId="17" pageNumber="358">CRUS</collectionCode>
.D.57224.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBA8FE75FDE66324" pageId="17" pageNumber="358" type="description">
<paragraph id="7107F648FFE81114EBA8FE75FDDF60DB" blockId="17.[151,1437,325,1394]" box="[199,558,433,458]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FE75FEA060D8" bold="true" box="[199,337,433,458]" pageId="17" pageNumber="358">Description</emphasis>
. See
<bibRefCitation id="15298BB9FFE81114EAE0FE75FDD860DB" author="Anker" box="[399,553,433,458]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker (2014)</bibRefCitation>
.
</paragraph>
<paragraph id="7107F648FFE81114EBA8FE11FDE66324" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FE11FE5C60FC" bold="true" box="[199,429,469,494]" pageId="17" pageNumber="358">Colouration in life</emphasis>
. Body and appendages whitish with pale orange-yellow tinge and some iridescence, especially on chelipeds and pleon (see
<bibRefCitation id="15298BB9FFE81114E921FE3DFD266303" author="Anker" box="[590,727,505,530]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
, fig. 5); eggs in ovigerous females bright orange-red (based on low-resolution field photographs).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBA8FD85FECD636C" pageId="17" pageNumber="358" type="distribution">
<paragraph id="7107F648FFE81114EBA8FD85FECD636C" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FD85FEA86348" bold="true" box="[199,345,577,602]" pageId="17" pageNumber="358">Distribution</emphasis>
. Western Atlantic:
<collectingCountry id="09AFB6D8FFE81114E955FD85FD306348" box="[570,705,577,602]" name="Guadeloupe" pageId="17" pageNumber="358">Guadeloupe</collectingCountry>
(French Antilles) and
<collectingCountry id="09AFB6D8FFE81114E8A6FD85FBF56348" box="[969,1028,577,602]" name="Cuba" pageId="17" pageNumber="358">Cuba</collectingCountry>
(
<collectingCountry id="09AFB6D8FFE81114EF79FD85FBB16348" box="[1046,1088,577,602]" name="Iceland" pageId="17" pageNumber="358">Isla</collectingCountry>
de la Juventud) (
<bibRefCitation id="15298BB9FFE81114EE64FD85FEC1636C" author="Anker" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014; present study</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBA8FD4DFD3E6460" pageId="17" pageNumber="358" type="discussion">
<paragraph id="7107F648FFE81114EBA8FD4DFC726220" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FD4DFEC563B0" bold="true" box="[199,308,649,674]" pageId="17" pageNumber="358">Remarks</emphasis>
.
<taxonomicName id="B6B88DCBFFE81114EA27FD4DFDC063B3" box="[328,561,649,673]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114EA27FD4DFDC063B3" box="[328,561,649,673]" italics="true" pageId="17" pageNumber="358">Naushonia draconis</emphasis>
</taxonomicName>
was previously known from a single specimen, the male
<typeStatus id="AE0348EAFFE81114EF99FD4DFAA963B0" box="[1270,1368,649,674]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
from
<collectingCountry id="09AFB6D8FFE81114EBF8FD69FED063D4" box="[151,289,685,710]" name="Guadeloupe" pageId="17" pageNumber="358">Guadeloupe</collectingCountry>
, French Antilles (
<bibRefCitation id="15298BB9FFE81114EA85FD69FD8663D4" author="Anker" box="[490,631,685,710]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
). The Cuban material agrees almost perfectly with the
<typeStatus id="AE0348EAFFE81114EF8DFD69FAB663D4" box="[1250,1351,685,710]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
, except for some minor details, which all appear to be within an expected intraspecific variation in species of
<taxonomicName id="B6B88DCBFFE81114EE73FD15FA6463FB" box="[1308,1429,721,745]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFE81114EE73FD15FA6463FB" box="[1308,1429,721,745]" italics="true" pageId="17" pageNumber="358">Naushonia</emphasis>
</taxonomicName>
. This variation includes the shape and armature of the mid-dorsal crests on the pleon (
<figureCitation id="E983EACDFFE81114EF34FD31FB6E621C" box="[1115,1183,757,782]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
A, I), ventral armature of the pleomeres (
<figureCitation id="E983EACDFFE81114EA0EFCDDFE526220" box="[353,419,793,818]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
B) and dentition on the chelipeds (
<figureCitation id="E983EACDFFE81114E849FCDDFC986220" box="[806,873,793,818]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
J).
</paragraph>
<paragraph id="7107F648FFE81114EBA8FCF9FECC65AC" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<bibRefCitation id="15298BB9FFE81114EBA8FCF9FEAE6244" author="Anker" box="[199,351,829,854]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker (2014)</bibRefCitation>
dissected and illustrated the third and second maxillipeds of the
<typeStatus id="AE0348EAFFE81114EF5AFCF9FB666244" box="[1077,1175,829,854]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B6B88DCBFFE81114EFD3FCFAFACE6247" box="[1212,1343,829,853]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114EFD3FCFAFACE6247" box="[1212,1343,829,853]" italics="true" pageId="17" pageNumber="358">N. draconis</emphasis>
</taxonomicName>
, but not the rest of the mouthparts in order not to further damage the unique specimen at that time. The new material enables us to illustrate the previously non-figured mandible, maxillule, maxilla and first maxilliped (
<figureCitation id="E983EACDFFE81114EE7CFC41FAA6628C" box="[1299,1367,901,926]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
CG), which are all fairly typical for
<taxonomicName id="B6B88DCBFFE81114EA83FC6DFD9462D3" box="[492,613,937,961]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFE81114EA83FC6DFD9462D3" box="[492,613,937,961]" italics="true" pageId="17" pageNumber="358">Naushonia</emphasis>
</taxonomicName>
. The posterolateral angles of the sixth pleomere are indeed produced into strong sharp spines, i.e. one on each side of the telson; both spines were broken in the
<typeStatus id="AE0348EAFFE81114EF00FC09FB2062F4" box="[1135,1233,973,998]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
(see
<bibRefCitation id="15298BB9FFE81114EE61FC09FA6962F4" author="Anker" box="[1294,1432,973,998]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
: fig. 2E, F). An interesting feature not specified in the original description of
<taxonomicName id="B6B88DCBFFE81114E890FC36FB72651B" box="[1023,1155,1009,1033]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114E890FC36FB72651B" box="[1023,1155,1009,1033]" italics="true" pageId="17" pageNumber="358">N. draconis</emphasis>
</taxonomicName>
is the presence a strong, sharp, posteriorly directed, median tooth on the sternum of the sixth pleomere; this tooth is present in both males and females and can be seen in the ventral view of the
<typeStatus id="AE0348EAFFE81114E870FBFDFC706540" box="[799,897,1081,1106]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="15298BB9FFE81114E8FCFBFDFB716540" author="Anker" box="[915,1152,1081,1106]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014: fig. 1B</bibRefCitation>
). The mesial side of the cheliped coxa is armed with a very stout, anteriorly directed, sharp spine; although not specifically mentioned in the original description, this spine can be seen even in the lateral view of the basal articles of the cheliped (
<bibRefCitation id="15298BB9FFE81114EE3BFB45FEC165AC" author="Anker" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014: fig. 3K</bibRefCitation>
).
</paragraph>
<paragraph id="7107F648FFE81114EBA8FB0DFD3E6460" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
The previously incompletely known gill formula of
<taxonomicName id="B6B88DCBFFE81114E87EFB0EFC6565F3" box="[785,916,1225,1249]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114E87EFB0EFC6565F3" box="[785,916,1225,1249]" italics="true" pageId="17" pageNumber="358">N. draconis</emphasis>
</taxonomicName>
was examined in a male specimen from
<collectingCountry id="09AFB6D8FFE81114EE0EFB0DFA6D65F0" box="[1377,1436,1225,1250]" name="Cuba" pageId="17" pageNumber="358">Cuba</collectingCountry>
(MZUSP 33201) and generally conforms to that of the other five species of the genus reported herein (see Table I). The arthrobranch at the base of the first maxilliped is well developed, with multiple lamellae (
<figureCitation id="E983EACDFFE81114EFDFFAD5FB026438" box="[1200,1267,1297,1322]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
G). The epipod of the fourth pereopod is reduced to a simple lobe; there is apparently no setobranch on the fourth pereopod, corresponding to the epipod on the third pereopod.
</paragraph>
</subSubSection>
</treatment>
</document>