147 lines
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147 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.376.6047" ID-GBIF-Dataset="ea01dfa1-5c94-4a89-804b-aa8037d4dacc" ID-PMC="PMC3935421" ID-Pensoft-Pub="1313-2970-376-1" ID-PubMed="24574849" ID-ZBK="2B58574A5FCF40D19A3AFB4D13D33A92" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-376-1" ModsDocOrigin="ZooKeys 376" ModsDocTitle="Review and redescription of species in the Oecetis avara group, with the description of 15 new species (Trichoptera, Leptoceridae)" checkinTime="1451246380649" checkinUser="pensoft" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2014" docId="E10C59FD4524BA419AFD30CFDB7C4551" docLanguage="en" docName="ZooKeys 376: 1-83" docOrigin="ZooKeys 376" docSource="http://dx.doi.org/10.3897/zookeys.376.6047" docTitle="Oecetis tumida Blahnik & Holzenthal, 2014, sp. n." docType="treatment" docUuid="8A57E9CC-856F-4D31-B012-896287F0924C" docUuidSource="ZooBank" docVersion="4" lastPageNumber="59" masterDocId="3A31DE5CFF94FFE90671FF839B6AAD3F" masterDocTitle="Review and redescription of species in the Oecetis avara group, with the description of 15 new species (Trichoptera, Leptoceridae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="57" updateTime="1668157599346" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review and redescription of species in the Oecetis avara group, with the description of 15 new species (Trichoptera, Leptoceridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Blahnik, Roger J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Holzenthal, Ralph W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>376</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.376.6047</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.376.6047</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-376-1</mods:identifier>
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<mods:identifier type="ZBK">2B58574A5FCF40D19A3AFB4D13D33A92</mods:identifier>
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<mods:identifier type="ZooBank">2B58574A5FCF40D19A3AFB4D13D33A92</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050811" LSID="urn:lsid:zoobank.org:act:8A57E9CC-856F-4D31-B012-896287F0924C" httpUri="http://treatment.plazi.org/id/E10C59FD4524BA419AFD30CFDB7C4551" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57">
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<subSubSection pageId="56" pageNumber="57" type="nomenclature">
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName LSID="http://zoobank.org/8A57E9CC-856F-4D31-B012-896287F0924C" class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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<taxonomicNameLabel pageId="56" pageNumber="57">sp. n.</taxonomicNameLabel>
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Figs 24, 48, Map 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="56" pageNumber="57" type="diagnosis">
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<paragraph pageId="56" pageNumber="57">Diagnosis.</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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is more or less intermediate in characters between
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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sp. n., agreeing with the former in the general shape of the inferior appendages, and with the latter in having the ventral apex of the phallobase somewhat convexly bulged preapically. The holotype specimen is from a population that is sympatric with
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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. In general, though, the phallobase is shorter and more bulbous apically than in
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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. In addition to the genitalic differences, the specimens were slightly larger than the co-occurring specimens of
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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. The species is most likely to be confused with
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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, in part due to the similarity in color and size of the 2 species, but especially due to the shape of the inferior appendages, which have an obtusely angled ventral lobe that is very weakly, but angularly projecting. The convex preapical bulge on the phallobase in
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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, and the absence of a distinctly V-shaped apex, in caudal view, should distinguish them.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="adult">
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<paragraph pageId="56" pageNumber="57">Adult.</paragraph>
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<paragraph lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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Forewing length: male (9.0-10 mm) Color pale yellowish to slightly brownish-yellow (similar to
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis mexicana" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="mexicana">
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<pageBreakToken pageId="57" pageNumber="58" start="start">Oecetis</pageBreakToken>
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mexicana
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</taxonomicName>
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); forewing spots distinct, spots at base of discal and thyridial cells large or moderately large, other spots smaller; veins of forewing chord widely spaced, r and r-m veins slightly closer; chord with small spots at juncture of major veins, spots at opposite ends of chord slightly larger; spots at apices of major veins small but distinct, pigmentation largely confined to vein. Setae along veins in apical part of forewing elongate, semi-prostrate, laterally diverging. Fringe of setae along costal margin of forewing moderate in length, somewhat projecting, not conspicuously erect.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="male genitalia">
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<paragraph pageId="57" pageNumber="58">Male genitalia.</paragraph>
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<paragraph lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
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Segment IX very short, with elongate setae along posterolateral margin. Tergum X with narrow, deflexed mesal lobe, lobe relatively elongate, somewhat lobulate (expanded) apically, apex with small sensilla; lobe continuous basoventrally with short paired lateral membranous projections. Preanal appendage moderately elongate, length about 3 times maximum width, simple in structure, apical setae elongate. Inferior appendage with rounded dorsal lobe and angularly projecting ventral lobe; ventral lobe very weakly projecting, forming approximately right to distinctly
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<pageBreakToken pageId="58" pageNumber="59" start="start">obtuse</pageBreakToken>
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angle with dorsal lobe, posterior margin of ventral lobe, as viewed ventrally (Fig. 24F), with distinctly angular bend near base; basomesal projection weakly developed, forming short rounded projection with short, stiff setae; dorsal lobe with stout, mesally-curved setae on dorsal margin and stout, ventrally-curved setae on mesal surface. Phallobase, as viewed laterally, relatively short, distinctly inflated apically; as viewed laterally with weakly developed, convex bulge preapically on ventral margin; as viewed caudally, with apex inflated, broadly U-shaped or V-shaped, not keeled ventrally (Fig. 24C). Phallotremal sclerite prominent, basally forming short tubular collar, ventral margin projecting, apex acute; asymmetrical lateral sclerite apparently absent.
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</paragraph>
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<caption pageId="58" pageNumber="59">
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<paragraph pageId="58" pageNumber="59">
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Figure 24.
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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sp. n., male genitalia: A lateral B phallic apparatus, lateral C phallic apparatus, caudal D phallic apparatus, ventral E inferior appendage, caudal F inferior appendage, ventral G egment IX and tergum X, dorsal.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="holotype">
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<paragraph pageId="58" pageNumber="59">Holotype.</paragraph>
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<paragraph pageId="58" pageNumber="59">
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Male (pinned), COSTA RICA:
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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: Res. Biol. Barbilla,
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Dantas, 15 km (road) S Pacuarito,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.994">9°59.640'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.443">83°26.580'W</geoCoordinate>
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, el 300 m, 27-30.i.1992, Holzenthal,
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<normalizedToken originalValue="Muñoz">Munoz</normalizedToken>
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, Kjer (UMSP) (INBIOCRI000372044).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="paratypes">
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<paragraph pageId="58" pageNumber="59">Paratypes.</paragraph>
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<paragraph pageId="58" pageNumber="59">
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COSTA RICA: Guanacaste: Quebrada Garcia, 10. 6 km ENE Quebrada Grande,
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.862">10.862°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-85.428">85.428°W</geoCoordinate>
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, 470 m, 8.iii.1986, Holzenthal & Fasth, 1 male, 1 female (alcohol) (NMNH);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Los Ahogados, 11.3 km ENE Quebrada Grande,
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.865">10.865°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-85.423">85.423°W</geoCoordinate>
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, el 470 m, 7.iii.1986, Holzenthal & Fasth, 1 male, 2 females (alcohol) (UMSP);
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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: same data as holotype, 1 male (pinned) (UMSP); Puntarenas:
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Singri, ca. 2 km (air) S Finca Helechales,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.057">9°3.420'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.082">83°4.920'W</geoCoordinate>
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, el 720 m, 21.ii.1986, Holzenthal, Morse, Fasth, 1 male (alcohol) (UMSP).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="etymology">
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<paragraph pageId="58" pageNumber="59">Etymology.</paragraph>
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<paragraph pageId="58" pageNumber="59">
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This species is named
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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, after the Latin word tumidus, meaning swollen, and referring to the apex of the phallobase in this species, which is distinctly swollen and usually has a slight, but noticeable, preapical ventral bulge.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |