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<document id="829768DBF3584558CC4FC85F9D5116E3" ID-DOI="10.5281/zenodo.181374" ID-GBIF-Dataset="321f0d1f-9203-4f2e-8594-1dc16a9a1149" ID-ISSN="1175-5326" ID-Zenodo-Dep="181374" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459833167893" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho &amp; Abadie, Esteban Ignácio" docDate="2008" docId="460003706D3AF216FF7344B349CFFEED" docLanguage="en" docName="zt01733p060.pdf" docOrigin="Zootaxa 1733" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Eremobothynus hirsutus Grossi &amp; Abadie, new species" docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="BA397B086D3AF217FFE4470E4A7FFFD4" masterDocTitle="A new species of Eremobothynus Ohaus (Coleoptera: Scarabaeidae: Dynastinae) from Argentina, with notes on Eremobothynus cornutus (Steinheil)" masterLastPageNumber="60" masterPageNumber="57" pageNumber="57" updateTime="1698223136634" updateUser="plazi">
<mods:mods id="7F4093CAF9EFA9D3F82C6CD31E9AC33D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="675E6BA488DE368B858C0955FCBB6CDA">
<mods:title id="202871A91A6D673940CEC3DB31FF64F3">A new species of Eremobothynus Ohaus (Coleoptera: Scarabaeidae: Dynastinae) from Argentina, with notes on Eremobothynus cornutus (Steinheil)</mods:title>
</mods:titleInfo>
<mods:name id="C9B37B7E416F80A24638AAE7C1DBA6AE" type="personal">
<mods:role id="84BB879B1493C0734A93F3D7197236C3">
<mods:roleTerm id="1DEE9E3B075C8F0418753670C5E5CE39">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA674B5536DE0EE66D6F29627F3AC830">Grossi, Paschoal Coelho</mods:namePart>
</mods:name>
<mods:name id="3A127E780BB24067CFA267A0D5E58911" type="personal">
<mods:role id="C634A5A015C7D163C96612DFC83C5B8D">
<mods:roleTerm id="6B5BA84531F79C09360ABB85245B0E3F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F5B004C5C93C978EFE3D2E5E24F72EB">Abadie, Esteban Ignácio</mods:namePart>
</mods:name>
<mods:typeOfResource id="4D288E9B0F49874D3D20109691833E9C">text</mods:typeOfResource>
<mods:relatedItem id="21B6F166AC104197CE24A02C8BC2B028" type="host">
<mods:titleInfo id="5E29129142005CBAFD58A0C1DF57D0D6">
<mods:title id="94FD5CAAEC4D7D3345B77BFCC9131927">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="90F1855826B985629289DE852643C213">
<mods:date id="A5DF2448B64293AC2009A8287DE31122">2008</mods:date>
<mods:detail id="54CF5DF82FDC75AA21EDFE021F917C3E" type="volume">
<mods:number id="8DE6C1DFBA7037344A0FC5815E14982A">1733</mods:number>
</mods:detail>
<mods:extent id="D4051BD5648FAE7A96AE33171A7D62F2" unit="page">
<mods:start id="673960C3496E288784FFC3DAAC22A1C6">57</mods:start>
<mods:end id="644F0EF1CC29A3CAE20DED85B10FB80F">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="792A528563715892F91E0C8479A993B7">journal article</mods:classification>
<mods:identifier id="66B8FEFDCB09A998CC179C0C919608EE" type="DOI">10.5281/zenodo.181374</mods:identifier>
<mods:identifier id="0A99F6C3F3C5F8F957A090585F2A3DE1" type="GBIF-Dataset">321f0d1f-9203-4f2e-8594-1dc16a9a1149</mods:identifier>
<mods:identifier id="13D2EA21B5EE4844A1F2BF9AF1EA953C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D7316F321D3BFC8178A0A23CDFF7DC57" type="Zenodo-Dep">181374</mods:identifier>
</mods:mods>
<treatment id="460003706D3AF216FF7344B349CFFEED" ID-DOI="http://doi.org/10.5281/zenodo.6231146" ID-GBIF-Taxon="119358278" ID-Zenodo-Dep="6231146" LSID="urn:lsid:plazi:treatment:460003706D3AF216FF7344B349CFFEED" httpUri="http://treatment.plazi.org/id/460003706D3AF216FF7344B349CFFEED" lastPageId="1" lastPageNumber="57" pageId="0" pageNumber="57">
<subSubSection id="86B3E1ED6D3AF217FF7344B34946FC03" box="[151,825,957,983]" pageId="0" pageNumber="57" type="nomenclature">
<paragraph id="CE16B2666D3AF217FF7344B34946FC03" blockId="0.[151,825,957,983]" box="[151,825,957,983]" pageId="0" pageNumber="57">
<heading id="955E050A6D3AF217FF7344B34946FC03" bold="true" box="[151,825,957,983]" fontSize="11" level="1" pageId="0" pageNumber="57" reason="1">
<emphasis id="FCDD6E746D3AF217FF7344B34946FC03" bold="true" box="[151,825,957,983]" pageId="0" pageNumber="57">
<taxonomicName id="09A9C9E56D3AF217FF7344B348E3FC03" authority="Grossi &amp; Abadie" authorityName="Grossi &amp; Abadie" box="[151,668,957,983]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="FCDD6E746D3AF217FF7344B34BBEFC03" bold="true" box="[151,449,957,983]" italics="true" pageId="0" pageNumber="57">Eremobothynus hirsutus</emphasis>
Grossi &amp; Abadie
</taxonomicName>
,
<taxonomicNameLabel id="E7EED30F6D3AF217FD4D44B34946FC03" box="[681,825,957,983]" pageId="0" pageNumber="57" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86B3E1ED6D3AF217FF73430C4F1AFB70" pageId="0" pageNumber="57" type="materials_examined">
<paragraph id="CE16B2666D3AF217FF73430C49DCFB8B" blockId="0.[151,1437,1026,2020]" pageId="0" pageNumber="57">
<emphasis id="FCDD6E746D3AF217FF73430C4B41FBCE" bold="true" box="[151,318,1026,1050]" pageId="0" pageNumber="57">
<typeStatus id="11120CC46D3AF217FF73430C4AB0FBCE" box="[151,207,1026,1050]" pageId="0" pageNumber="57">Type</typeStatus>
material.
</emphasis>
<typeStatus id="11120CC46D3AF217FEA1430C4BD3FBCD" box="[325,428,1026,1049]" pageId="0" pageNumber="57" type="holotype">Holotype</typeStatus>
: ɗ, handwritten white label - dorsal “Arg: E. Rios/Pronuncia-/miento”, ventral “11.62./D/149 ɗ” (collected in
<date id="BA1794A66D3AF217FEAB432B487CFBE8" box="[335,515,1061,1084]" pageId="0" pageNumber="57" value="1962-11">November 1962</date>
).
<typeStatus id="11120CC46D3AF217FDFE432B48F9FBE8" box="[538,646,1061,1084]" pageId="0" pageNumber="57" type="paratype">Paratypes</typeStatus>
: (2 ɗɗ) labeled as follows: one with same data as
<typeStatus id="11120CC46D3AF217FB27432B4F5CFBE8" box="[1219,1315,1061,1084]" pageId="0" pageNumber="57" type="holotype">holotype</typeStatus>
, except 64 instead of 62; one with same data as
<typeStatus id="11120CC46D3AF217FDFB43464800FB8B" box="[543,639,1096,1119]" pageId="0" pageNumber="57" type="holotype">holotype</typeStatus>
, except 1966 instead of 62.
</paragraph>
<paragraph id="CE16B2666D3AF217FF2243644F1AFB70" blockId="0.[151,1437,1026,2020]" pageId="0" pageNumber="57">
<typeStatus id="11120CC46D3AF217FF2243644B56FB55" box="[198,297,1130,1153]" pageId="0" pageNumber="57" type="holotype">Holotype</typeStatus>
and one
<typeStatus id="11120CC46D3AF217FE7143644B8EFB55" box="[405,497,1130,1153]" pageId="0" pageNumber="57" type="paratype">paratype</typeStatus>
deposited at Universidade Federal do Paraná, Coleção Padre Jesus Santiago Moure, Curitiba, Paraná,
<collectingCountry id="B6BEF2F66D3AF217FEB543834BE9FB70" box="[337,406,1165,1188]" name="Brazil" pageId="0" pageNumber="57">Brazil</collectingCountry>
; one
<typeStatus id="11120CC46D3AF217FE2A43834855FB70" box="[462,554,1165,1188]" pageId="0" pageNumber="57" type="paratype">paratype</typeStatus>
deposited in the Esteban Abadie private collection, Buenos Aires,
<collectingCountry id="B6BEF2F66D3AF217FB1043834F1EFB70" box="[1268,1377,1165,1188]" name="Argentina" pageId="0" pageNumber="57">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="86B3E1ED6D3AF217FF2243A14EF5F8E3" pageId="0" pageNumber="57" type="description">
<paragraph id="CE16B2666D3AF217FF2243A14887F91B" blockId="0.[151,1437,1026,2020]" pageId="0" pageNumber="57">
<emphasis id="FCDD6E746D3AF217FF2243A14B80FB13" bold="true" box="[198,511,1199,1223]" pageId="0" pageNumber="57">
Description of the
<typeStatus id="11120CC46D3AF217FE7943A14B80FB13" box="[413,511,1199,1223]" pageId="0" pageNumber="57" type="holotype">holotype</typeStatus>
</emphasis>
: Length
<quantity id="09511F836D3AF217FD8643BE48B9FB13" box="[610,710,1200,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="0" pageNumber="57" unit="mm" value="19.4">19.4 mm</quantity>
; greatest pronotal width
<quantity id="09511F836D3AF217FC3043BE4E4AFB13" box="[980,1077,1200,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="0" pageNumber="57" unit="mm" value="10.6">10.6 mm</quantity>
(
<figureCitation id="5692AEE36D3AF217FBA143BE4ED1FB13" box="[1093,1198,1200,1223]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,269,1954,1978]" captionTargetBox="[302,1290,408,1929]" captionTargetId="figure@1.[292,1296,384,1948]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Eremobothynus spp.: 1 - 2: E. hirsutus sp. nov., holotype, dorsal and lateral views; 3: E. cornutus, male dorsal view; 4: female head and pronotum anterolateral view." httpUri="https://zenodo.org/record/181375/files/figure.png" pageId="0" pageNumber="57">Figs. 1–2</figureCitation>
). Color brownish red. Body oval and convex, dorsal surface glabrous, ventrally extremely setose.
<emphasis id="FCDD6E746D3AF217FC2843DC4E71FB3D" box="[972,1038,1234,1257]" italics="true" pageId="0" pageNumber="57">Head:</emphasis>
Short and transverse, nearly 2 times wider than long (
<figureCitation id="5692AEE36D3AF217FEA843FB4BF3FAD8" box="[332,396,1269,1292]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,269,1390,1414]" captionTargetBox="[205,1381,196,1351]" captionTargetId="figure@2.[198,1388,176,1384]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Eremobothynus spp. aedeagus and male head: 5 - 6: E. hirsutus sp. nov. caudal and lateral views; 7 - 8: E. cornutus caudal and lateral views; 9: E. hirsutus sp. nov. head, dorsal view; 10: E. cornutus head, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/181376/files/figure.png" pageId="0" pageNumber="57">Fig. 9</figureCitation>
); frons with sparse, long setae. Frontal horn prominent, wider at posterior edge; surface wrinkled, elongate, curved posteriorly; apex rhomboid. Clypeus short (
<figureCitation id="5692AEE36D3AF217FCA6421649F7FAFB" box="[834,904,1304,1327]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,269,1390,1414]" captionTargetBox="[205,1381,196,1351]" captionTargetId="figure@2.[198,1388,176,1384]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Eremobothynus spp. aedeagus and male head: 5 - 6: E. hirsutus sp. nov. caudal and lateral views; 7 - 8: E. cornutus caudal and lateral views; 9: E. hirsutus sp. nov. head, dorsal view; 10: E. cornutus head, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/181376/files/figure.png" pageId="0" pageNumber="57">Fig. 9</figureCitation>
), quadrate, as wide as horn apical width; apex reflexed. Eye canthus rounded (
<figureCitation id="5692AEE36D3AF217FE0942344852FA85" box="[493,557,1338,1361]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,269,1390,1414]" captionTargetBox="[205,1381,196,1351]" captionTargetId="figure@2.[198,1388,176,1384]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Eremobothynus spp. aedeagus and male head: 5 - 6: E. hirsutus sp. nov. caudal and lateral views; 7 - 8: E. cornutus caudal and lateral views; 9: E. hirsutus sp. nov. head, dorsal view; 10: E. cornutus head, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/181376/files/figure.png" pageId="0" pageNumber="57">Fig. 9</figureCitation>
) with external margin crenulated, apices setose dorsolaterally. Mentum anteriorly projected, triangular; with surface flattened, covered with long setae. Mandibles unarmed, trapezoidal, dorsally convex, with sides subparallel. Antennae with 9 antennomeres, antennal club longer than antennomeres 2-6 combined.
<emphasis id="FCDD6E746D3AF217FAB5428E4AB8FA6D" italics="true" pageId="0" pageNumber="57">Pronotum:</emphasis>
Narrower than elytra; border complete and enlarged anteromedially. Surface completely punctate; punctures deeper, more numerous on anterior portion of disc, coalescing around anterior pronotal declivity. Margin of pronotal declivity sinuous, with concavity below, as wide as width of head. Anterior angles acute, almost projected beneath head.
<emphasis id="FCDD6E746D3AF217FAA042E64FE4FA2B" box="[1348,1435,1512,1535]" italics="true" pageId="0" pageNumber="57">Elytron:</emphasis>
Surface smooth, shiny with sparse, small punctures; each elytron with 2 impressed longitudinal lines. Epipleuron indicated only in basal third.
<emphasis id="FCDD6E746D3AF217FE4141234B9EF990" box="[421,481,1581,1604]" italics="true" pageId="0" pageNumber="57">Legs:</emphasis>
Lateral margin of protibia tridentate, basal tooth smaller; dorsal surface with 2 longitudinal rows of long setae. Meso- and metatibial spurs wide and flattened.
<emphasis id="FCDD6E746D3AF217FC4C415E4E69F9B3" box="[936,1046,1616,1639]" italics="true" pageId="0" pageNumber="57">Abdomen:</emphasis>
Pygidium smooth, slightly convex, completely covered by long yellowish setae except at longitudinal midline. Ventral surface extremely setose with long, yellow setae.
<emphasis id="FCDD6E746D3AF217FECE419B4BE4F978" box="[298,411,1685,1708]" italics="true" pageId="0" pageNumber="57">Aedeagus:</emphasis>
Parameres elongated, symmetrical, ventrally with crenulated expansion (
<figureCitation id="5692AEE36D3AF217FB4B419B4E8EF978" box="[1199,1265,1685,1708]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,269,1390,1414]" captionTargetBox="[205,1381,196,1351]" captionTargetId="figure@2.[198,1388,176,1384]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Eremobothynus spp. aedeagus and male head: 5 - 6: E. hirsutus sp. nov. caudal and lateral views; 7 - 8: E. cornutus caudal and lateral views; 9: E. hirsutus sp. nov. head, dorsal view; 10: E. cornutus head, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/181376/files/figure.png" pageId="0" pageNumber="57">Fig. 5</figureCitation>
). Phallobase on anterolateral basal edge with elevated projection (
<figureCitation id="5692AEE36D3AF217FD4D41B64894F91B" box="[681,747,1720,1743]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,269,1390,1414]" captionTargetBox="[205,1381,196,1351]" captionTargetId="figure@2.[198,1388,176,1384]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Eremobothynus spp. aedeagus and male head: 5 - 6: E. hirsutus sp. nov. caudal and lateral views; 7 - 8: E. cornutus caudal and lateral views; 9: E. hirsutus sp. nov. head, dorsal view; 10: E. cornutus head, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/181376/files/figure.png" pageId="0" pageNumber="57">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="CE16B2666D3AF217FF2241D44EF5F8E3" blockId="0.[151,1437,1026,2020]" pageId="0" pageNumber="57">
<emphasis id="FCDD6E746D3AF217FF2241D44B4AF926" bold="true" box="[198,309,1754,1778]" pageId="0" pageNumber="57">Variation.</emphasis>
Length 15.0–
<quantity id="09511F836D3AF217FE2A41D4484FF925" box="[462,560,1754,1777]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="0" pageNumber="57" unit="mm" value="19.4">19.4 mm</quantity>
; greatest pronotal width 8.0–
<quantity id="09511F836D3AF217FC8D41D449B4F925" box="[873,971,1754,1777]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="0" pageNumber="57" unit="mm" value="10.6">10.6 mm</quantity>
. Structural characters as in
<typeStatus id="11120CC46D3AF217FB1641D44F30F925" box="[1266,1359,1754,1777]" pageId="0" pageNumber="57" type="holotype">holotype</typeStatus>
except for the following.
<emphasis id="FCDD6E746D3AF217FEB941F34BDFF8C0" box="[349,416,1789,1812]" italics="true" pageId="0" pageNumber="57">Head:</emphasis>
Frontal horn smaller. Surface more strongly punctate; eye canthus less rounded, almost quadrate.
<emphasis id="FCDD6E746D3AF217FF28402E4B40F8E3" box="[204,319,1824,1847]" italics="true" pageId="0" pageNumber="57">Pronotum:</emphasis>
Disc with deeper punctures, sometimes becoming wrinkled. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="86B3E1ED6D3AF217FF22404C4900F8A8" pageId="0" pageNumber="57" type="etymology">
<paragraph id="CE16B2666D3AF217FF22404C4900F8A8" blockId="0.[151,1437,1026,2020]" pageId="0" pageNumber="57">
<emphasis id="FCDD6E746D3AF217FF22404C4B38F88E" bold="true" box="[198,327,1858,1882]" pageId="0" pageNumber="57">Etymology.</emphasis>
From the Latin
<taxonomicName id="09A9C9E56D3AF217FE1C404D4831F88E" box="[504,590,1859,1882]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="FCDD6E746D3AF217FE1C404D4831F88E" box="[504,590,1859,1882]" italics="true" pageId="0" pageNumber="57">hirsutus</emphasis>
</taxonomicName>
, meaning “with dense, long and curly hairs;” in reference to the abundance of ventral setae in the species, which does not occur in the other species.
</paragraph>
</subSubSection>
<subSubSection id="86B3E1ED6D3AF217FF2240894E8BF84B" box="[198,1268,1927,1951]" pageId="0" pageNumber="57" type="distribution">
<paragraph id="CE16B2666D3AF217FF2240894E8BF84B" blockId="0.[151,1437,1026,2020]" box="[198,1268,1927,1951]" pageId="0" pageNumber="57">
<emphasis id="FCDD6E746D3AF217FF2240894B26F84B" bold="true" box="[198,345,1927,1951]" pageId="0" pageNumber="57">Distribution.</emphasis>
<taxonomicName id="09A9C9E56D3AF217FEBB4086481DF84B" box="[351,610,1928,1951]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="FCDD6E746D3AF217FEBB4086481DF84B" box="[351,610,1928,1951]" italics="true" pageId="0" pageNumber="57">Eremobothynus hirsutus</emphasis>
</taxonomicName>
is known only from Pronunciamiento, Entre Ríos,
<collectingCountry id="B6BEF2F66D3AF217FB6640864E8FF84B" box="[1154,1264,1928,1951]" name="Argentina" pageId="0" pageNumber="57">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="86B3E1ED6D3AF216FF2240A449CFFEED" lastPageId="1" lastPageNumber="58" pageId="0" pageNumber="57" type="discussion">
<paragraph id="CE16B2666D3AF216FF2240A449CFFEED" blockId="0.[151,1437,1026,2020]" lastBlockId="1.[151,1436,152,313]" lastPageId="1" lastPageNumber="58" pageId="0" pageNumber="57">
<emphasis id="FCDD6E746D3AF217FF2240A44B51F816" bold="true" box="[198,302,1962,1986]" pageId="0" pageNumber="57">Remarks</emphasis>
. The new species shares with
<taxonomicName id="09A9C9E56D3AF217FD8B40A54896F816" box="[623,745,1962,1986]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D3AF217FD8B40A54800F816" box="[623,639,1963,1986]" italics="true" pageId="0" pageNumber="57">E</emphasis>
.
<emphasis id="FCDD6E746D3AF217FD6840A54896F816" box="[652,745,1963,1986]" italics="true" pageId="0" pageNumber="57">cornutus</emphasis>
</taxonomicName>
the presence of a frontal horn in the males. However,
<taxonomicName id="09A9C9E56D3AF217FACD40A54FE3F816" box="[1321,1436,1962,1986]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="FCDD6E746D3AF217FACD40A54F46F816" box="[1321,1337,1963,1986]" italics="true" pageId="0" pageNumber="57">E</emphasis>
.
<emphasis id="FCDD6E746D3AF217FAA240A54FE3F816" box="[1350,1436,1963,1986]" italics="true" pageId="0" pageNumber="57">hirsutus</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName id="09A9C9E56D3AF217FDE040C34800F830" box="[516,639,1997,2020]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D3AF217FDE040C3486BF830" box="[516,532,1997,2020]" italics="true" pageId="0" pageNumber="57">E</emphasis>
.
<emphasis id="FCDD6E746D3AF217FDC640C34800F830" box="[546,639,1997,2020]" italics="true" pageId="0" pageNumber="57">cornutus</emphasis>
</taxonomicName>
by the following characters: (1) more setose body (ventral surface, pygidium, and head); (2) short head and round eye canthus (long head and acutely projected eye canthus in
<taxonomicName id="09A9C9E56D3BF216FB0B47964F14FF7B" box="[1263,1387,152,175]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D3BF216FB0B47964E80FF7B" box="[1263,1279,152,175]" italics="true" pageId="1" pageNumber="58">E</emphasis>
.
<emphasis id="FCDD6E746D3BF216FAEA47964F14FF7B" box="[1294,1387,152,175]" italics="true" pageId="1" pageNumber="58">cornutus</emphasis>
</taxonomicName>
, see
<figureCitation id="5692AEE36D3BF216FF7347B44A98FF05" box="[151,231,186,209]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,269,1390,1414]" captionTargetBox="[205,1381,196,1351]" captionTargetId="figure@2.[198,1388,176,1384]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Eremobothynus spp. aedeagus and male head: 5 - 6: E. hirsutus sp. nov. caudal and lateral views; 7 - 8: E. cornutus caudal and lateral views; 9: E. hirsutus sp. nov. head, dorsal view; 10: E. cornutus head, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/181376/files/figure.png" pageId="1" pageNumber="58">Fig. 10</figureCitation>
); (3) concave anterior pronotal declivity (vertical in
<taxonomicName id="09A9C9E56D3BF216FCC247B449DDFF05" box="[806,930,186,209]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D3BF216FCC247B44949FF05" box="[806,822,186,209]" italics="true" pageId="1" pageNumber="58">E</emphasis>
.
<emphasis id="FCDD6E746D3BF216FCA147B449DDFF05" box="[837,930,186,209]" italics="true" pageId="1" pageNumber="58">cornutus</emphasis>
</taxonomicName>
); (4) flattened mentum (strongly convex in
<taxonomicName id="09A9C9E56D3BF216FA6147B44A8BFF20" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D3BF216FA6147B44FEAFF05" box="[1413,1429,186,209]" italics="true" pageId="1" pageNumber="58">E</emphasis>
.
<emphasis id="FCDD6E746D3BF216FF7347D34A8BFF20" box="[151,244,221,244]" italics="true" pageId="1" pageNumber="58">cornutus</emphasis>
</taxonomicName>
); (5) epipleuron almost absent (entire in
<taxonomicName id="09A9C9E56D3BF216FD5C47D3494AFF20" box="[696,821,221,244]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D3BF216FD5C47D348B7FF20" box="[696,712,221,244]" italics="true" pageId="1" pageNumber="58">E</emphasis>
.
<emphasis id="FCDD6E746D3BF216FD3C47D3494AFF20" box="[728,821,221,244]" italics="true" pageId="1" pageNumber="58">cornutus</emphasis>
</taxonomicName>
). The genitalia are also distinct, as the parameres in the new species are more posteriorly rounded with a more prominent ventral projection, in contrast to the posteriorly elongated parameres and obscure ventral projection of
<taxonomicName id="09A9C9E56D3BF216FD56462C4952FEED" box="[690,813,290,313]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D3BF216FD56462C48BDFEED" box="[690,706,290,313]" italics="true" pageId="1" pageNumber="58">E</emphasis>
.
<emphasis id="FCDD6E746D3BF216FD34462C4952FEED" box="[720,813,290,313]" italics="true" pageId="1" pageNumber="58">cornutus</emphasis>
</taxonomicName>
(
<figureCitation id="5692AEE36D3BF216FCD8462C49DBFEED" box="[828,932,290,313]" captionStart="FIGURES 5 – 10" captionStartId="2.[151,269,1390,1414]" captionTargetBox="[205,1381,196,1351]" captionTargetId="figure@2.[198,1388,176,1384]" captionTargetPageId="2" captionText="FIGURES 5 – 10. Eremobothynus spp. aedeagus and male head: 5 - 6: E. hirsutus sp. nov. caudal and lateral views; 7 - 8: E. cornutus caudal and lateral views; 9: E. hirsutus sp. nov. head, dorsal view; 10: E. cornutus head, dorsal view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/181376/files/figure.png" pageId="1" pageNumber="58">Figs. 7–8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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