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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.85399" ID-Pensoft-Pub="1313-2970-1110-39" ID-Pensoft-UUID="766F9A94793D5793A9BE1E70F575798B" ID-ZooBank="4681CC6DE5F347C6B1D052DEA78BE7C3" ModsDocID="1313-2970-1110-39" checkinTime="1657006630715" checkinUser="pensoft" docAuthor="Sian Man, Nem, Siriboon, Thanit, Lin, Aung, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="853E2DEF00DF532F88CA302F5F4ECDE5" docLanguage="en" docName="ZooKeys 1110: 39-102" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.85399" docTitle="Oophana laevis" docType="treatment" docVersion="1" id="766F9A94793D5793A9BE1E70F575798B" lastPageNumber="39" masterDocId="766F9A94793D5793A9BE1E70F575798B" masterDocTitle="Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species" masterLastPageNumber="102" masterPageNumber="39" pageNumber="39" updateTime="1657006630715" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sian Man, Nem</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4453-734X</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Siriboon, Thanit</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok, 10110, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Aung</mods:namePart>
<mods:affiliation>Fauna and Flora International, No. 35, 3</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1110</mods:number>
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<mods:extent unit="page">
<mods:start>39</mods:start>
<mods:end>102</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1110.85399</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1110.85399</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1110-39</mods:identifier>
<mods:identifier type="ZooBank">4681CC6DE5F347C6B1D052DEA78BE7C3</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:853E2DEF00DF532F88CA302F5F4ECDE5" httpUri="http://treatment.plazi.org/id/853E2DEF00DF532F88CA302F5F4ECDE5" lastPageNumber="39" pageId="0" pageNumber="39">
<subSubSection pageId="0" pageNumber="39" type="nomenclature">
<paragraph pageId="0" pageNumber="39">
<taxonomicName LSID="853E2DEF-00DF-532F-88CA-302F5F4ECDE5" authority="(Blanford, 1899)" baseAuthorityName="Blanford" baseAuthorityYear="1899" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Oophana laevis" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="laevis">Oophana laevis (Blanford, 1899)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="description">
<paragraph pageId="0" pageNumber="39">
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Oophana mouhoti A possible syntype NHMUK ex. Cuming collection from Camboja and B specimen CUMZ 13005 from Phra (Buddha) Cave, Tanintharyi C, D Oophana obtusus C specimen NHMUK 1888.12.4.788 - 790 from Chauktalon, Moulmein with apertural dentition and D specimen NHMUK 1871.9.23.64 from Moulmein E Oophana laevis, possible syntype NHMUK 1899.7.4.11 from Burma." figureDoi="10.3897/zookeys.1110.85399.figure7" httpUri="https://binary.pensoft.net/fig/710472" pageId="0" pageNumber="39">Fig. 7E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="reference_group">
<paragraph pageId="0" pageNumber="39">
<taxonomicName authorityName="W. T. Blanford" authorityYear="1899" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis laevis" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="laevis">Streptaxis laevis</taxonomicName>
Blanford, 1899: 765, pl. 50, figs 11-12. Type locality: Tenasserim [probably Tanintharyi Region, Myanmar].
<bibRefCitation author="Gude, GK" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="39" pagination="201 - 275" refId="B32" refString="Gude, GK, 1903. A synopsis of the genus Streptaxis and its allies. Proceedings of the Malacological Society of London 5: 201 - 275" title="A synopsis of the genus Streptaxis and its allies." volume="5" year="1903">Gude 1903</bibRefCitation>
: 218.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="39" refId="B10" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
: 13.
</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName authorityName="W. T. Blanford" authorityYear="1899" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis (Odontartemon) laevis" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="laevis" subGenus="Odontartemon">Streptaxis (Odontartemon) laevis</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Journal of the Society for Bibliography of Natural History" pageId="0" pageNumber="39" refId="B37" refString="Kobelt, W, 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1(12b) (2): 1-211 pls42-71 [pp. 1-96 pls 42-59 (1905) ; 97-211, pls 60-71 (1906)]." title="Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (12 b) (2): 1 - 211 pls 42 - 71 [pp. 1 - 96 pls 42 - 59 (1905); 97 - 211, pls 60 - 71 (1906)]." year="1905">Kobelt 1905</bibRefCitation>
: 93, 94, pl. 60, figs 11, 12.
</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Odontartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odontartemon (Oophana) laevis" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="laevis" subGenus="Oophana">Odontartemon (Oophana) laevis</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Jahrbuecher des Nassauischen Vereins fuer Naturkunde" pageId="0" pageNumber="39" pagination="138 - 196" refId="B38" refString="Kobelt, W, 1910. Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 63: 138 - 196" title="Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha." volume="63" year="1910">Kobelt 1910</bibRefCitation>
: 150.
</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName baseAuthorityName="W. T. Blanford" baseAuthorityYear="1899" class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indoartemon laevis" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="laevis">Indoartemon laevis</taxonomicName>
-
<bibRefCitation author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="39" pagination="1 - 326" refId="B70" refString="Richardson, L, 1988. Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia 16: 1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
: 225.
<bibRefCitation author="Siriboon, T" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="39" pagination="161 - 174" refId="B77" refString="Siriboon, T, Sutcharit, C, Naggs, F, Rowson, B, Panha, S, 2014b. Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae). The Raffles Bulletin of Zoology 62: 161 - 174" title="Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)." volume="62" year="2014 b">Siriboon et al. 2014b</bibRefCitation>
: 163.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="materials_examined">
<paragraph pageId="0" pageNumber="39">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">
Possible
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Burma" location="Beddome" specimenCount="1" typeStatus="syntype">
<typeStatus>syntype</typeStatus>
NHMUK 1899.7.4.11 (1 shell; Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Oophana mouhoti A possible syntype NHMUK ex. Cuming collection from Camboja and B specimen CUMZ 13005 from Phra (Buddha) Cave, Tanintharyi C, D Oophana obtusus C specimen NHMUK 1888.12.4.788 - 790 from Chauktalon, Moulmein with apertural dentition and D specimen NHMUK 1871.9.23.64 from Moulmein E Oophana laevis, possible syntype NHMUK 1899.7.4.11 from Burma." figureDoi="10.3897/zookeys.1110.85399.figure7" httpUri="https://binary.pensoft.net/fig/710472" pageId="0" pageNumber="39">7E</figureCitation>
) ex.
<location LSID="urn:lsid:plazi:treatment:853E2DEF00DF532F88CA302F5F4ECDE5:BFEFDFAD1592D067DBADB8B6344CA828" country="Burma" name="Beddome">Beddome</location>
collection from
<collectingCountry name="Myanmar">Burma</collectingCountry>
</materialsCitation>
</emphasis>
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Myanmar" isTailEnumeration="true" location="Myanmar" specimenCount="1" typeStatus="syntype">
[
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="diagnosis">
<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="39">
<taxonomicName baseAuthorityName="Blanford" baseAuthorityYear="1899" class="Gastropoda" family="Streptaxidae" genus="Oophana" kingdom="Animalia" lsidName="Oophana laevis" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="laevis">
<emphasis italics="true" pageId="0" pageNumber="39">Oophana laevis</emphasis>
</taxonomicName>
differs from
<taxonomicName genus="O." lsidName="O. mouhoti" pageId="0" pageNumber="39" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="39">O. mouhoti</emphasis>
</taxonomicName>
and
<taxonomicName genus="O." lsidName="O. obtusus" pageId="0" pageNumber="39" rank="species" species="obtusus">
<emphasis italics="true" pageId="0" pageNumber="39">O. obtusus</emphasis>
</taxonomicName>
by having a rounded penultimate whorl, convex spire, and apertural dentition with parietal and basal lamellae. Although
<taxonomicName genus="O." lsidName="O. mouhoti" pageId="0" pageNumber="39" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="39">O. mouhoti</emphasis>
</taxonomicName>
and
<taxonomicName genus="O." lsidName="O. obtusus" pageId="0" pageNumber="39" rank="species" species="obtusus">
<emphasis italics="true" pageId="0" pageNumber="39">O. obtusus</emphasis>
</taxonomicName>
have a slightly angular penultimate whorl and elevated spire,
<taxonomicName genus="O." lsidName="O. mouhoti" pageId="0" pageNumber="39" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="39">O. mouhoti</emphasis>
</taxonomicName>
usually has only a parietal lamella (sometimes with basal lamella). In contrast,
<taxonomicName genus="O." lsidName="O. obtusus" pageId="0" pageNumber="39" rank="species" species="obtusus">
<emphasis italics="true" pageId="0" pageNumber="39">O. obtusus</emphasis>
</taxonomicName>
has parietal, palatal, and columellar lamellae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="description">
<paragraph pageId="0" pageNumber="39">Description.</paragraph>
<paragraph pageId="0" pageNumber="39">
Shell oblique-heliciform, white, and translucent; whorls
<normalizedToken originalValue="5½">51/2</normalizedToken>
; spire low convex with distinct suture. Shell surface glossy with transverse ridges that diminish below periphery. Embryonic shell ~
<normalizedToken originalValue="2½">21/2</normalizedToken>
whorls with smooth surface; following whorls regularly coiled. Shell periphery rounded; last whorl axially deflected. Aperture subquadrangular with sinulus; peristome discontinuous, thin, expanded and reflected. Apertural dentition with one parietal and one basal lamella, and sometimes columellar lamella is absent. Umbilicus open and shallow (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Oophana mouhoti A possible syntype NHMUK ex. Cuming collection from Camboja and B specimen CUMZ 13005 from Phra (Buddha) Cave, Tanintharyi C, D Oophana obtusus C specimen NHMUK 1888.12.4.788 - 790 from Chauktalon, Moulmein with apertural dentition and D specimen NHMUK 1871.9.23.64 from Moulmein E Oophana laevis, possible syntype NHMUK 1899.7.4.11 from Burma." figureDoi="10.3897/zookeys.1110.85399.figure7" httpUri="https://binary.pensoft.net/fig/710472" pageId="0" pageNumber="39">7E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="distribution">
<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="39">
This species is known only from the type locality, which was mentioned only as
<normalizedToken originalValue="Tenasserim">'Tenasserim'</normalizedToken>
(Blandford 1899). This locality possibly refers to the Tanintharyi Region, southeastern Myanmar.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="remarks">
<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="39">
No new specimens were collected in this study. However, we compared the possible type specimen with the other congeners.
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1899.tb06887.x" author="Blanford, WT" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="39" pagination="764 - 770" refId="B9" refString="Blanford, WT, 1899. On some species of shells of the genera Streptaxis and Ennea from India, Ceylon, and Burma. Proceedings of the Zoological Society of London 67 (3): 764 - 770, DOI: https://doi.org/10.1111/j.1469-7998.1899.tb06887.x" title="On some species of shells of the genera Streptaxis and Ennea from India, Ceylon, and Burma." url="https://doi.org/10.1111/j.1469-7998.1899.tb06887.x" volume="67" year="1899">Blanford (1899)</bibRefCitation>
clearly stated that the type series was collected by Beddome and consisted of three specimens. However, the NHM, London holds a lot of only a single specimen from the Beddome collection, with the word India, probably added at a later date, on the label. Therefore, we consider this lot to be a possible syntype.
</paragraph>
</subSubSection>
</treatment>
</document>